951 lines
151 KiB
XML
951 lines
151 KiB
XML
<document id="881B660CBE94801A6E5952A525BE22C6" ID-CLB-Dataset="20794" ID-DOI="10.11646/zootaxa.5094.2.4" ID-GBIF-Dataset="4a011d42-96e0-4206-966e-2819094fbbd5" ID-ISSN="1175-5326" ID-Zenodo-Dep="5965221" ID-ZooBank="9655C635-FF92-4A87-AE47-2610CED08B7D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1643872797008" checkinUser="plazi" docAuthor="Anker, Arthur & Ďuriš, Zdeněk" docDate="2022" docId="03A387EF2669FFFAE39BFC4A79BAE9D7" docLanguage="en" docName="zootaxa.5094.2.4.pdf" docOrigin="Zootaxa 5094 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Athanas philippei Anker & Ďuriš 2022, n. sp." docType="treatment" docVersion="7" lastPageNumber="298" masterDocId="FF9AFF972668FFF0E30CFFE47840EA1E" masterDocTitle="Athanas philippei n. sp., a new alpheid shrimp associated with stomatopods in New Caledonia (Decapoda: Caridea)" masterLastPageNumber="300" masterPageNumber="288" pageNumber="289" updateTime="1699306174581" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="02311492F62BB649C62FC635794154E4">Athanas philippei n. sp., a new alpheid shrimp associated with stomatopods in New Caledonia (Decapoda: Caridea)</mods:title>
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<mods:name id="E53C5DEEC1410DBDBC63F3BBBCA8640F" type="personal">
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<mods:namePart id="C6E7267206102578DAEED1348431BA70">Anker, Arthur</mods:namePart>
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<mods:affiliation id="CB76DA9B2B78341EF8278C28157B430D">Universidade Federal de Goiás, Campus Samambaia, Instituto de Ciências Biológicas-ICB- 5. Av. Esperança, s / n., 74690 - 900, Goiânia, Goiás, Brazil.</mods:affiliation>
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<mods:namePart id="3BAFC4EC3E58724AAEB9905FDE90F689">Ďuriš, Zdeněk</mods:namePart>
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<mods:affiliation id="391C4416F9A3EA0884CCEF21CAA36CC1">Department of Biology and Ecology and Institute of Environmental Technologies, Faculty of Science, University of Ostrava, Chittussiho 10, 710 00 Ostrava, Czech Republic.</mods:affiliation>
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<mods:title id="A96923043F5CB48D6802495FCEEE3151">Zootaxa</mods:title>
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<mods:part id="92F48CCCE023F440DF89AF28DFD38B26">
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<mods:date id="F4F4929E4699D601518B32CB286BB87D">2022</mods:date>
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<mods:number id="4EAD739BC0FE1441673E37CB887AB215">2022-02-03</mods:number>
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<mods:number id="CE72246B9EF202DF226C2DCD62EBF16F">5094</mods:number>
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<treatment id="03A387EF2669FFFAE39BFC4A79BAE9D7" ID-DOI="http://doi.org/10.5281/zenodo.6301624" ID-GBIF-Taxon="193276984" ID-Zenodo-Dep="6301624" LSID="urn:lsid:plazi:treatment:03A387EF2669FFFAE39BFC4A79BAE9D7" httpUri="http://treatment.plazi.org/id/03A387EF2669FFFAE39BFC4A79BAE9D7" lastPageId="10" lastPageNumber="298" pageId="1" pageNumber="289">
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<subSubSection id="C31065722669FFF1E39BFC4A79F7E9D7" box="[151,439,942,969]" pageId="1" pageNumber="289" type="nomenclature">
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<heading id="D0FD81952669FFF1E39BFC4A79F7E9D7" bold="true" box="[151,439,942,969]" fontSize="11" level="1" pageId="1" pageNumber="289" reason="1">
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<emphasis id="B97EEAEB2669FFF1E39BFC4A79F7E9D7" bold="true" box="[151,439,942,969]" pageId="1" pageNumber="289">
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<taxonomicName id="4C0A4D7A2669FFF1E39BFC4A792EE9D7" authority="Anker & Ďuriš, 2022" authorityName="Anker & Ďuriš" authorityYear="2022" box="[151,366,942,969]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="289" phylum="Arthropoda" rank="species" species="philippei" status="sp. nov.">
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<emphasis id="B97EEAEB2669FFF1E39BFC4A792EE9D7" bold="true" box="[151,366,942,969]" italics="true" pageId="1" pageNumber="289">Athanas philippei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24D57902669FFF1E279FC4B79F7E9D7" box="[373,439,943,969]" pageId="1" pageNumber="289" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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<subSubSection id="C31065722669FFF1E39BFC377923E9F3" box="[151,355,978,1005]" pageId="1" pageNumber="289" type="description">
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<paragraph id="8BB536F92669FFF1E39BFC377923E9F3" blockId="1.[151,439,942,1005]" box="[151,355,978,1005]" pageId="1" pageNumber="289">
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(
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<figureCitation id="13312A7C2669FFF1E393FC37794BE9F3" box="[159,267,978,1005]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1030,1054]" captionStartId-1="3.[151,250,1713,1737]" captionStartId-2="4.[151,250,1910,1934]" captionStartId-3="5.[151,250,1361,1385]" captionStartId-4="6.[151,250,1684,1708]" captionTargetBox-0="[158,1429,387,996]" captionTargetBox-1="[176,1411,211,1650]" captionTargetBox-2="[172,1412,532,1871]" captionTargetBox-3="[165,1423,188,1329]" captionTargetBox-4="[185,1402,181,1660]" captionTargetId-0="figure-86@2.[151,1436,378,1004]" captionTargetId-1="figure-31@3.[169,1418,181,1689]" captionTargetId-2="figure-169@4.[164,1423,525,1886]" captionTargetId-3="figure-31@5.[154,1434,181,1338]" captionTargetId-4="figure-17@6.[185,1402,181,1660]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURE 1. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124), habitus, lateral view." captionText-1="FIGURE 2. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) rostrum and left eye and orbital teeth, lateral view; (B) anterior region, lateral view; (C) same, dorsal view; (D) telson and uropods, dorsal view; (E) telson, posterior half, dorsal view; (F) third maxilliped, lateral view; (G) left uropod, detail of distolateral region, dorsal view. All scales as indicated, except for G drawn from a microscope photograph without scale." captionText-2="FIGURE 3. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) right (minor) cheliped, fully stretched, lateral view; (B) same, proximal articles (mainly ischium, merus and carpus), mesial view; (C) same, folded, lateral view; (D) left (major) cheliped, distal articles (mainly carpus and chela), lateral view; (E) same, distal articles (mainly carpus and chela), dorsolateral view; (F) same, ischium, mesial view." captionText-3="FIGURE 4. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) second pereiopod, lateral view; (B) third pereiopod, lateral view; (C) same, distal portion of propodus and dactylus; (D) fourth pereiopod, lateral view; (E) fifth pereiopod, lateral view; (F) same, distal portion of propodus and dactylus, lateral view; (G) same, distal portion of propodus and dactylus, mesial view." captionText-4="FIGURE 5. Athanas philippei n. sp., paratype, female (cl 5.2 mm) from Pouébo, New Caledonia (MNHN-IU-2018-1017): (A) shrimp alive, dorsal view; (B) same, lateral view; (C) same, frontal view. Photographs by A. Anker." figureDoi-0="http://doi.org/10.5281/zenodo.5965225" figureDoi-1="http://doi.org/10.5281/zenodo.5965227" figureDoi-2="http://doi.org/10.5281/zenodo.5965231" figureDoi-3="http://doi.org/10.5281/zenodo.5965237" figureDoi-4="http://doi.org/10.5281/zenodo.5965241" httpUri-0="https://zenodo.org/record/5965225/files/figure.png" httpUri-1="https://zenodo.org/record/5965227/files/figure.png" httpUri-2="https://zenodo.org/record/5965231/files/figure.png" httpUri-3="https://zenodo.org/record/5965237/files/figure.png" httpUri-4="https://zenodo.org/record/5965241/files/figure.png" pageId="1" pageNumber="289">Figs. 1–5</figureCitation>
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,
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<figureCitation id="13312A7C2669FFF1E21BFC37791AE9F3" box="[279,346,979,1005]" captionStart="FIGURE 6" captionStartId="8.[151,250,1865,1889]" captionTargetBox="[197,1390,415,1842]" captionTargetId="figure-134@8.[197,1390,415,1842]" captionTargetPageId="8" captionText="FIGURE 6. Athanas philippei n. sp. [A, B]: (A) holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018- 1124): shrimp alive, lateral view; (B) paratype, female (cl 7.2 mm) from Saint-Mathieu, New Caledonia (MNHN-IU-2019- 2986), shrimp alive, anterolateral view. Lysiosquillina maculata (Fabricius, 1793) (Lysiosquillidae) [C]: host (MNHN-IU-2018- 1087) of the female paratype of A. philippei n. sp. (MNHN-IU-2018-1017) from Pouébo, New Caledonia. Photographs by Z. Ďuriš." figureDoi="http://doi.org/10.5281/zenodo.5965243" httpUri="https://zenodo.org/record/5965243/files/figure.png" pageId="1" pageNumber="289">6A, B</figureCitation>
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)
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</paragraph>
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<subSubSection id="C31065722669FFF1E39BFBFF7930EFDF" pageId="1" pageNumber="289" type="materials_examined">
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<emphasis id="B97EEAEB2669FFF1E39BFBFF7906EE2A" bold="true" box="[151,326,1050,1077]" pageId="1" pageNumber="289">Type material.</emphasis>
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<typeStatus id="54B1885B2669FFF1E243FBFF79F6EE2B" box="[335,438,1051,1077]" pageId="1" pageNumber="289" type="holotype">Holotype</typeStatus>
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</emphasis>
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: female (cl
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<quantity id="4CF29B1C2669FFF1E14CFBFF7ADBEE2B" box="[576,667,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="1" pageNumber="289" unit="mm" value="6.9">6.9 mm</quantity>
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), MNHN-IU-2018-1124,
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, north-eastern coast,
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<location id="8ED560222669FFF1E66CFBFF7DD8EE2B" LSID="urn:lsid:plazi:treatment:03A387EF2669FFFAE39BFC4A79BAE9D7:8ED560222669FFF1E66CFBFF7DD8EE2B" box="[1376,1432,1051,1077]" country="New Caledonia" latitude="-20.256666" longLatPrecision="126" longitude="164.37833" name="Tiari" pageId="1" pageNumber="289">Tiari</location>
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, sta.
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<specimenCode id="DBAC9E822669FFF1E3CAFBDB7960EE47" box="[198,288,1087,1113]" collectionCode="KM" country="Russia" name="Kotel'nich Museum" pageId="1" pageNumber="289">KM608</specimenCode>
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,
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,
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<geoCoordinate id="EE3E503E2669FFF1E2A1FBDB7A70EE47" box="[429,560,1087,1113]" degrees="164" direction="east" minutes="22.7" orientation="longitude" pageId="1" pageNumber="289" precision="92" value="164.37833">164°22.7’E</geoCoordinate>
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, coastal platform composed of sand and fragmented coral rubble interspaced with shallow sandy pools along mangrove coast, depth less than
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<quantity id="4CF29B1C2669FFF1E062FB867BF0EE63" box="[878,944,1122,1149]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="289" unit="m" value="0.5">
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<elevation id="0027D1CA2669FFF1E062FB867BF0EE63" box="[878,944,1122,1149]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="289" unit="m" value="0.5">0.5 m</elevation>
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</quantity>
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, suction pump, in burrow, leg.
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<collectorName id="26FF532F2669FFF1E607FB877DDCEE61" box="[1291,1436,1123,1151]" pageId="1" pageNumber="289">A. Šobáňová</collectorName>
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&
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<collectorName id="26FF532F2669FFF1E3B8FB637954EEBD" box="[180,276,1159,1187]" pageId="1" pageNumber="289">Z. Ďuriš</collectorName>
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,
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<date id="FFB410392669FFF1E213FB6379E1EEBF" box="[287,417,1159,1185]" pageId="1" pageNumber="289" value="2018-09-23">
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<collectingDate id="EFF0E9D12669FFF1E213FB6379E1EEBF" box="[287,417,1159,1185]" pageId="1" pageNumber="289" value="2018-09-23">23.09.2018</collectingDate>
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</date>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BB536F92669FFF1E3CBFB4F7930EFDF" blockId="1.[151,1437,1050,2013]" pageId="1" pageNumber="289">
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<typeStatus id="54B1885B2669FFF1E3CBFB4F7979EEDA" box="[199,313,1195,1220]" pageId="1" pageNumber="289" type="paratype">Paratypes</typeStatus>
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</emphasis>
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:
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<specimenCount id="9D0CFD702669FFF1E245FB4F79ECEEDB" box="[329,428,1195,1221]" pageId="1" pageNumber="289" type="female">1 female</specimenCount>
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(cl 6.0 mm), MNHN-IU-2018-608, same collection details as for
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<typeStatus id="54B1885B2669FFF1E7ADFB4F7D47EEDB" box="[1185,1287,1195,1221]" pageId="1" pageNumber="289" type="holotype">holotype</typeStatus>
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;
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<specimenCount id="9D0CFD702669FFF1E61FFB4F7D36EEDB" box="[1299,1398,1195,1221]" pageId="1" pageNumber="289" type="female">1 female</specimenCount>
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(cl
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<quantity id="4CF29B1C2669FFF1E39BFB2B78B2EEF7" box="[151,242,1231,1257]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="1" pageNumber="289" unit="mm" value="7.8">7.8 mm</quantity>
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; freshly moulted), MNHN-IU-2018-1123, same collection details as for
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<typeStatus id="54B1885B2669FFF1E723FB2B7CD5EEF7" box="[1071,1173,1231,1257]" pageId="1" pageNumber="289" type="holotype">holotype</typeStatus>
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;
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<materialsCitation id="3B623CA42669FFF1E7AEFB2B7DC4EF2F" ID-GBIF-Occurrence="3462295302" collectingDate="2019-01-11" collectionCode="KM" collectorName="Z. Duris" country="New Caledonia" elevation="5" latitude="-20.38" location="Saint-Mathieu" longLatPrecision="872" longitude="164.58333" pageId="1" pageNumber="289" specimenCode="KM606" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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<specimenCount id="9D0CFD702669FFF1E7AEFB2B7D45EEF7" box="[1186,1285,1231,1257]" pageId="1" pageNumber="289" type="female">1 female</specimenCount>
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(cl
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<quantity id="4CF29B1C2669FFF1E638FB2B7DD0EEF7" box="[1332,1424,1231,1257]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="1" pageNumber="289" unit="mm" value="7.2">7.2 mm</quantity>
|
||
), MNHN-IU-2019-2986,
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<collectingCountry id="F31D76692669FFF1E2ADFB177A0EEF13" box="[417,590,1267,1293]" name="New Caledonia" pageId="1" pageNumber="289">New Caledonia</collectingCountry>
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, north-eastern coast,
|
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<location id="8ED560222669FFF1E03EFB177B95EF13" LSID="urn:lsid:plazi:treatment:03A387EF2669FFFAE39BFC4A79BAE9D7:8ED560222669FFF1E03EFB177B95EF13" box="[818,981,1267,1293]" country="New Caledonia" latitude="-20.38" longLatPrecision="872" longitude="164.58333" name="Saint-Mathieu" pageId="1" pageNumber="289">Saint-Mathieu</location>
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, sta.
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<specimenCode id="DBAC9E822669FFF1E70BFB177C21EF13" box="[1031,1121,1267,1293]" collectionCode="KM" country="Russia" name="Kotel'nich Museum" pageId="1" pageNumber="289">KM606</specimenCode>
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,
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|
||
,
|
||
<geoCoordinate id="EE3E503E2669FFF1E7E8FB167D13EF13" box="[1252,1363,1266,1293]" degrees="164" direction="east" minutes="35" orientation="longitude" pageId="1" pageNumber="289" precision="925" value="164.58333">164°35’E</geoCoordinate>
|
||
, sandy coastal platform along mangrove coast, depth less than
|
||
<quantity id="4CF29B1C2669FFF1E00CFAF27B01EF2F" box="[768,833,1302,1329]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="289" unit="m" value="0.5">
|
||
<elevation id="0027D1CA2669FFF1E00CFAF27B01EF2F" box="[768,833,1302,1329]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="289" unit="m" value="0.5">0.5 m</elevation>
|
||
</quantity>
|
||
, suction pump, in burrow, leg.
|
||
<collectorName id="26FF532F2669FFF1E795FAF37CB8EF2D" box="[1177,1272,1303,1331]" pageId="1" pageNumber="289">Z. Ďuriš</collectorName>
|
||
,
|
||
<date id="FFB410392669FFF1E60FFAF37DC4EF2F" box="[1283,1412,1303,1329]" pageId="1" pageNumber="289" value="2019-01-11">
|
||
<collectingDate id="EFF0E9D12669FFF1E60FFAF37DC4EF2F" box="[1283,1412,1303,1329]" pageId="1" pageNumber="289" value="2019-01-11">01.11.2019</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B623CA42669FFF1E682FAF3792BEFDF" ID-GBIF-Occurrence="3462295303" collectingDate="2018-09-19" collectionCode="KM" collectorName="A. Anker" country="New Caledonia" elevation="5" latitude="-20.293333" location="Pouebo" longLatPrecision="126" longitude="164.485" pageId="1" pageNumber="289" specimenCode="KM607" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<specimenCount id="9D0CFD702669FFF1E682FAF378A3EF4B" pageId="1" pageNumber="289" type="female">1 female</specimenCount>
|
||
(cl
|
||
<quantity id="4CF29B1C2669FFF1E21CFADE792BEF4B" box="[272,363,1338,1365]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="1" pageNumber="289" unit="mm" value="5.2">5.2 mm</quantity>
|
||
), MNHN-IU-2018-1017,
|
||
<collectingCountry id="F31D76692669FFF1E181FADF7B7DEF4B" box="[653,829,1339,1365]" name="New Caledonia" pageId="1" pageNumber="289">New Caledonia</collectingCountry>
|
||
, north-eastern coast,
|
||
<location id="8ED560222669FFF1E722FADF7CC5EF4B" LSID="urn:lsid:plazi:treatment:03A387EF2669FFFAE39BFC4A79BAE9D7:8ED560222669FFF1E722FADF7CC5EF4B" box="[1070,1157,1339,1365]" country="New Caledonia" latitude="-20.293333" longLatPrecision="126" longitude="164.485" name="Pouebo" pageId="1" pageNumber="289">Pouébo</location>
|
||
, sta.
|
||
<specimenCode id="DBAC9E822669FFF1E7B2FADF7D58EF4B" box="[1214,1304,1339,1365]" collectionCode="KM" country="Russia" name="Kotel'nich Museum" pageId="1" pageNumber="289">KM607</specimenCode>
|
||
,
|
||
<geoCoordinate id="EE3E503E2669FFF1E628FADF7DD7EF4B" box="[1316,1431,1339,1365]" degrees="20" direction="south" minutes="17.6" orientation="latitude" pageId="1" pageNumber="289" precision="92" value="-20.293333">20°17.6’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3E503E2669FFF1E39BFABB795AEF67" box="[151,282,1375,1401]" degrees="164" direction="east" minutes="29.1" orientation="longitude" pageId="1" pageNumber="289" precision="92" value="164.485">164°29.1’E</geoCoordinate>
|
||
, shallow subtidal sand flat with nearby patches of seagrass and fragmented coral rubble, depth less than
|
||
<quantity id="4CF29B1C2669FFF1E39BFA66789AEF83" box="[151,218,1410,1437]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="289" unit="m" value="0.5">
|
||
<elevation id="0027D1CA2669FFF1E39BFA66789AEF83" box="[151,218,1410,1437]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="289" unit="m" value="0.5">0.5 m</elevation>
|
||
</quantity>
|
||
, suction pump, in burrow of
|
||
<taxonomicName id="4C0A4D7A2669FFF1E12BFA677B75EF83" baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[551,821,1411,1437]" class="Malacostraca" family="Lysiosquillidae" genus="Lysiosquillina" kingdom="Animalia" order="Stomatopoda" pageId="1" pageNumber="289" phylum="Arthropoda" rank="species" species="maculata">
|
||
<emphasis id="B97EEAEB2669FFF1E12BFA677B75EF83" box="[551,821,1411,1437]" italics="true" pageId="1" pageNumber="289">Lysiosquillina maculata</emphasis>
|
||
</taxonomicName>
|
||
[host deposited under MNHN-IU-2018-1087], leg.
|
||
<collectorName id="26FF532F2669FFF1E68DFA67789FEFDF" pageId="1" pageNumber="289">A. Anker</collectorName>
|
||
,
|
||
<date id="FFB410392669FFF1E3E6FA43792BEFDF" box="[234,363,1447,1473]" pageId="1" pageNumber="289" value="2018-09-19">
|
||
<collectingDate id="EFF0E9D12669FFF1E3E6FA43792BEFDF" box="[234,363,1447,1473]" pageId="1" pageNumber="289" value="2018-09-19">19.09.2018</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31065722669FFF7E3CBFA2E7961EBB5" lastPageId="7" lastPageNumber="295" pageId="1" pageNumber="289" type="description">
|
||
<paragraph id="8BB536F92669FFF1E3CBFA2E7994EC87" blockId="1.[151,1437,1050,2013]" pageId="1" pageNumber="289">
|
||
<emphasis id="B97EEAEB2669FFF1E3CBFA2E79A7EFFB" bold="true" box="[199,487,1482,1509]" pageId="1" pageNumber="289">
|
||
Description of
|
||
<typeStatus id="54B1885B2669FFF1E275FA2F79A2EFFB" box="[377,482,1483,1509]" pageId="1" pageNumber="289" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Carapace (
|
||
<figureCitation id="13312A7C2669FFF1E16BFA2F7AF5EFFB" box="[615,693,1483,1509]" captionStart="FIGURE 1" captionStartId="2.[151,250,1030,1054]" captionTargetBox="[158,1429,387,996]" captionTargetId="figure-86@2.[151,1436,378,1004]" captionTargetPageId="2" captionText="FIGURE 1. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124), habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5965225" httpUri="https://zenodo.org/record/5965225/files/figure.png" pageId="1" pageNumber="289">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="13312A7C2669FFF1E1C8FA2F7B48EFFB" box="[708,776,1483,1509]" captionStart="FIGURE 2" captionStartId="3.[151,250,1713,1737]" captionTargetBox="[176,1411,211,1650]" captionTargetId="figure-31@3.[169,1418,181,1689]" captionTargetPageId="3" captionText="FIGURE 2. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) rostrum and left eye and orbital teeth, lateral view; (B) anterior region, lateral view; (C) same, dorsal view; (D) telson and uropods, dorsal view; (E) telson, posterior half, dorsal view; (F) third maxilliped, lateral view; (G) left uropod, detail of distolateral region, dorsal view. All scales as indicated, except for G drawn from a microscope photograph without scale." figureDoi="http://doi.org/10.5281/zenodo.5965227" httpUri="https://zenodo.org/record/5965227/files/figure.png" pageId="1" pageNumber="289">2B, C</figureCitation>
|
||
) glabrous, with low median post-rostral tubercle and deep cardiac notch. Rostrum (
|
||
<figureCitation id="13312A7C2669FFF1E2BEFA0B7A6EEC17" box="[434,558,1519,1545]" captionStart="FIGURE 2" captionStartId="3.[151,250,1713,1737]" captionTargetBox="[176,1411,211,1650]" captionTargetId="figure-31@3.[169,1418,181,1689]" captionTargetPageId="3" captionText="FIGURE 2. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) rostrum and left eye and orbital teeth, lateral view; (B) anterior region, lateral view; (C) same, dorsal view; (D) telson and uropods, dorsal view; (E) telson, posterior half, dorsal view; (F) third maxilliped, lateral view; (G) left uropod, detail of distolateral region, dorsal view. All scales as indicated, except for G drawn from a microscope photograph without scale." figureDoi="http://doi.org/10.5281/zenodo.5965227" httpUri="https://zenodo.org/record/5965227/files/figure.png" pageId="1" pageNumber="289">Fig. 2B, C</figureCitation>
|
||
) reaching to mid-length of second article of antennular peduncle, somewhat broadened at base, tapering distally; rostral carina distinct, flattening posteriorly, with 5 minute blunt teeth on anterior half. Extra-corneal teeth (
|
||
<figureCitation id="13312A7C2669FFF1E11AF9D37ACFEC4F" box="[534,655,1591,1617]" captionStart="FIGURE 2" captionStartId="3.[151,250,1713,1737]" captionTargetBox="[176,1411,211,1650]" captionTargetId="figure-31@3.[169,1418,181,1689]" captionTargetPageId="3" captionText="FIGURE 2. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) rostrum and left eye and orbital teeth, lateral view; (B) anterior region, lateral view; (C) same, dorsal view; (D) telson and uropods, dorsal view; (E) telson, posterior half, dorsal view; (F) third maxilliped, lateral view; (G) left uropod, detail of distolateral region, dorsal view. All scales as indicated, except for G drawn from a microscope photograph without scale." figureDoi="http://doi.org/10.5281/zenodo.5965227" httpUri="https://zenodo.org/record/5965227/files/figure.png" pageId="1" pageNumber="289">Fig. 2A–C</figureCitation>
|
||
) small, acute, laterally concealing basal-most portion of eye corneas. Infra-corneal teeth (
|
||
<figureCitation id="13312A7C2669FFF1E274F9BF79B1EC6B" box="[376,497,1627,1653]" captionStart="FIGURE 2" captionStartId="3.[151,250,1713,1737]" captionTargetBox="[176,1411,211,1650]" captionTargetId="figure-31@3.[169,1418,181,1689]" captionTargetPageId="3" captionText="FIGURE 2. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) rostrum and left eye and orbital teeth, lateral view; (B) anterior region, lateral view; (C) same, dorsal view; (D) telson and uropods, dorsal view; (E) telson, posterior half, dorsal view; (F) third maxilliped, lateral view; (G) left uropod, detail of distolateral region, dorsal view. All scales as indicated, except for G drawn from a microscope photograph without scale." figureDoi="http://doi.org/10.5281/zenodo.5965227" httpUri="https://zenodo.org/record/5965227/files/figure.png" pageId="1" pageNumber="289">Fig. 2A–C</figureCitation>
|
||
) poorly developed as small bumps. Pterygostomial angle (
|
||
<figureCitation id="13312A7C2669FFF1E784F9BF7C9FEC6B" box="[1160,1247,1627,1653]" captionStart="FIGURE 2" captionStartId="3.[151,250,1713,1737]" captionTargetBox="[176,1411,211,1650]" captionTargetId="figure-31@3.[169,1418,181,1689]" captionTargetPageId="3" captionText="FIGURE 2. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) rostrum and left eye and orbital teeth, lateral view; (B) anterior region, lateral view; (C) same, dorsal view; (D) telson and uropods, dorsal view; (E) telson, posterior half, dorsal view; (F) third maxilliped, lateral view; (G) left uropod, detail of distolateral region, dorsal view. All scales as indicated, except for G drawn from a microscope photograph without scale." figureDoi="http://doi.org/10.5281/zenodo.5965227" httpUri="https://zenodo.org/record/5965227/files/figure.png" pageId="1" pageNumber="289">Fig. 2B</figureCitation>
|
||
) rounded, not or slightly produced anteriorly.
|
||
</paragraph>
|
||
<paragraph id="8BB536F92669FFF1E3CBF9477A26ED53" blockId="1.[151,1437,1050,2013]" pageId="1" pageNumber="289">
|
||
Pleon (
|
||
<figureCitation id="13312A7C2669FFF1E218F9477913ECA3" box="[276,339,1699,1725]" captionStart="FIGURE 1" captionStartId="2.[151,250,1030,1054]" captionTargetBox="[158,1429,387,996]" captionTargetId="figure-86@2.[151,1436,378,1004]" captionTargetPageId="2" captionText="FIGURE 1. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124), habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5965225" httpUri="https://zenodo.org/record/5965225/files/figure.png" pageId="1" pageNumber="289">Fig. 1</figureCitation>
|
||
) with first to fifth pleura rounded distoventrally; sixth pleonite with small subacute distal projection near telson base and subtriangular articulated plate. Telson (
|
||
<figureCitation id="13312A7C2669FFF1E048F9237BFEECFF" box="[836,958,1735,1761]" captionStart="FIGURE 2" captionStartId="3.[151,250,1713,1737]" captionTargetBox="[176,1411,211,1650]" captionTargetId="figure-31@3.[169,1418,181,1689]" captionTargetPageId="3" captionText="FIGURE 2. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) rostrum and left eye and orbital teeth, lateral view; (B) anterior region, lateral view; (C) same, dorsal view; (D) telson and uropods, dorsal view; (E) telson, posterior half, dorsal view; (F) third maxilliped, lateral view; (G) left uropod, detail of distolateral region, dorsal view. All scales as indicated, except for G drawn from a microscope photograph without scale." figureDoi="http://doi.org/10.5281/zenodo.5965227" httpUri="https://zenodo.org/record/5965227/files/figure.png" pageId="1" pageNumber="289">Fig. 2D, E</figureCitation>
|
||
) subrectangular, slightly tapering distally, nearly twice as long as proximal width, with 2 pairs of stout cuspidate setae situated approximately at 0.3–0.4 (anterior) and 0.6–0.7 (posterior) of telson length; posterior margin slightly convex, with 2 pairs of spiniform setae, mesial more than twice as long as lateral.
|
||
</paragraph>
|
||
<paragraph id="8BB536F92669FFF1E3CBF8B37CB3ED8B" blockId="1.[151,1437,1050,2013]" pageId="1" pageNumber="289">
|
||
Eyes (
|
||
<figureCitation id="13312A7C2669FFF1E207F8B379C1ED6F" box="[267,385,1879,1905]" captionStart="FIGURE 2" captionStartId="3.[151,250,1713,1737]" captionTargetBox="[176,1411,211,1650]" captionTargetId="figure-31@3.[169,1418,181,1689]" captionTargetPageId="3" captionText="FIGURE 2. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) rostrum and left eye and orbital teeth, lateral view; (B) anterior region, lateral view; (C) same, dorsal view; (D) telson and uropods, dorsal view; (E) telson, posterior half, dorsal view; (F) third maxilliped, lateral view; (G) left uropod, detail of distolateral region, dorsal view. All scales as indicated, except for G drawn from a microscope photograph without scale." figureDoi="http://doi.org/10.5281/zenodo.5965227" httpUri="https://zenodo.org/record/5965227/files/figure.png" pageId="1" pageNumber="289">Fig. 2A–C</figureCitation>
|
||
) large, largely exposed, with well-developed and normally pigmented corneas, latter occupying most of distal and distolateral area of eyestalk; anteromesial margin of eyestalks rounded, unarmed.
|
||
</paragraph>
|
||
<paragraph id="8BB536F92669FFF2E3CBF87B7A20EACB" blockId="1.[151,1437,1050,2013]" lastBlockId="2.[151,1436,151,357]" lastPageId="2" lastPageNumber="290" pageId="1" pageNumber="289">
|
||
Antennule (
|
||
<figureCitation id="13312A7C2669FFF1E240F87B7986EDA7" box="[332,454,1951,1977]" captionStart="FIGURE 2" captionStartId="3.[151,250,1713,1737]" captionTargetBox="[176,1411,211,1650]" captionTargetId="figure-31@3.[169,1418,181,1689]" captionTargetPageId="3" captionText="FIGURE 2. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) rostrum and left eye and orbital teeth, lateral view; (B) anterior region, lateral view; (C) same, dorsal view; (D) telson and uropods, dorsal view; (E) telson, posterior half, dorsal view; (F) third maxilliped, lateral view; (G) left uropod, detail of distolateral region, dorsal view. All scales as indicated, except for G drawn from a microscope photograph without scale." figureDoi="http://doi.org/10.5281/zenodo.5965227" httpUri="https://zenodo.org/record/5965227/files/figure.png" pageId="1" pageNumber="289">Fig. 2B, C</figureCitation>
|
||
) moderately stout; stylocerite relatively slender, with acute tip, reaching mid-length of second article; second and third article subequal in length; second article about 1.3 times as long as wide; lateral flagellum with fused portion composed of about 6 units, accessory ramus subdivided into 3 more or less distinct units, with several groups of aesthetascs.
|
||
</paragraph>
|
||
<paragraph id="8BB536F9266AFFF2E3CBFF3B7A4DEB7B" blockId="2.[151,1436,151,357]" pageId="2" pageNumber="290">
|
||
Antenna (
|
||
<figureCitation id="13312A7C266AFFF2E230FF3B7980EAE7" box="[316,448,223,249]" captionStart="FIGURE 2" captionStartId="3.[151,250,1713,1737]" captionTargetBox="[176,1411,211,1650]" captionTargetId="figure-31@3.[169,1418,181,1689]" captionTargetPageId="3" captionText="FIGURE 2. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) rostrum and left eye and orbital teeth, lateral view; (B) anterior region, lateral view; (C) same, dorsal view; (D) telson and uropods, dorsal view; (E) telson, posterior half, dorsal view; (F) third maxilliped, lateral view; (G) left uropod, detail of distolateral region, dorsal view. All scales as indicated, except for G drawn from a microscope photograph without scale." figureDoi="http://doi.org/10.5281/zenodo.5965227" httpUri="https://zenodo.org/record/5965227/files/figure.png" pageId="2" pageNumber="290">Fig. 2B, C</figureCitation>
|
||
) with basicerite robust, armed with stout subacute distoventral tooth; scaphocerite broad, overreaching end of antennular peduncle; lateral margin straight; blade broadly convex anteriorly, slightly overreaching small distolateral tooth; carpocerite stout, overreaching antennular peduncle, but falling short of distolateral tooth of scaphocerite.
|
||
</paragraph>
|
||
<caption id="DF756671266AFFF2E39BFBE27954EE5C" ID-DOI="http://doi.org/10.5281/zenodo.5965225" ID-Zenodo-Dep="5965225" httpUri="https://zenodo.org/record/5965225/files/figure.png" pageId="2" pageNumber="290" startId="2.[151,250,1030,1054]" targetBox="[158,1429,387,996]" targetPageId="2">
|
||
<paragraph id="8BB536F9266AFFF2E39BFBE27954EE5C" blockId="2.[151,1436,1030,1090]" pageId="2" pageNumber="290">
|
||
<emphasis id="B97EEAEB266AFFF2E39BFBE2794CEE00" bold="true" box="[151,268,1030,1054]" pageId="2" pageNumber="290">FIGURE 1</emphasis>
|
||
.
|
||
<taxonomicName id="4C0A4D7A266AFFF2E21BFBE27989EE00" authority="Anker & Ďuriš, 2022" authorityName="Anker & Ďuriš" authorityYear="2022" box="[279,457,1030,1054]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="290" phylum="Arthropoda" rank="species" species="philippei" status="sp. nov.">
|
||
<emphasis id="B97EEAEB266AFFF2E21BFBE27989EE00" box="[279,457,1030,1054]" italics="true" pageId="2" pageNumber="290">Athanas philippei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAEB266AFFF2E2C2FBE27A46EE00" bold="true" box="[462,518,1030,1054]" pageId="2" pageNumber="290">
|
||
<taxonomicNameLabel id="A24D5790266AFFF2E2C2FBE27A46EE00" box="[462,518,1030,1054]" pageId="2" pageNumber="290" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124), habitus, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB536F9266AFFF2E3CBFB947C7AEEE8" blockId="2.[151,1437,1136,2027]" pageId="2" pageNumber="290">
|
||
Mouthparts not dissected, typical for genus in external observation.Third maxilliped (
|
||
<figureCitation id="13312A7C266AFFF2E778FB947C85EE95" box="[1140,1221,1136,1163]" captionStart="FIGURE 2" captionStartId="3.[151,250,1713,1737]" captionTargetBox="[176,1411,211,1650]" captionTargetId="figure-31@3.[169,1418,181,1689]" captionTargetPageId="3" captionText="FIGURE 2. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) rostrum and left eye and orbital teeth, lateral view; (B) anterior region, lateral view; (C) same, dorsal view; (D) telson and uropods, dorsal view; (E) telson, posterior half, dorsal view; (F) third maxilliped, lateral view; (G) left uropod, detail of distolateral region, dorsal view. All scales as indicated, except for G drawn from a microscope photograph without scale." figureDoi="http://doi.org/10.5281/zenodo.5965227" httpUri="https://zenodo.org/record/5965227/files/figure.png" pageId="2" pageNumber="290">Fig. 2F</figureCitation>
|
||
) slender, pediform; coxa with distally subacute lateral plate; antepenultimate article about 5 times as long as greatest width; penultimate article slightly widening distally, about 4 times as long as distal width; ultimate article slender, tapering distally, with unarmed tip; exopod well developed, slightly overreaching antepenultimate article.
|
||
</paragraph>
|
||
<paragraph id="8BB536F9266AFFF2E3CBFAE47BF4ECD4" blockId="2.[151,1437,1136,2027]" pageId="2" pageNumber="290">
|
||
First pereiopods (= cheliped) subequal in size/length and somewhat asymmetrical in shape (
|
||
<figureCitation id="13312A7C266AFFF2E7E3FAE47D75EF05" box="[1263,1333,1280,1307]" captionStart="FIGURE 3" captionStartId="4.[151,250,1910,1934]" captionTargetBox="[172,1412,532,1871]" captionTargetId="figure-169@4.[164,1423,525,1886]" captionTargetPageId="4" captionText="FIGURE 3. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) right (minor) cheliped, fully stretched, lateral view; (B) same, proximal articles (mainly ischium, merus and carpus), mesial view; (C) same, folded, lateral view; (D) left (major) cheliped, distal articles (mainly carpus and chela), lateral view; (E) same, distal articles (mainly carpus and chela), dorsolateral view; (F) same, ischium, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5965231" httpUri="https://zenodo.org/record/5965231/files/figure.png" pageId="2" pageNumber="290">Fig. 3</figureCitation>
|
||
), carried folded ventrally when at rest (
|
||
<figureCitation id="13312A7C266AFFF2E2F7FAC07A03EF21" box="[507,579,1316,1343]" captionStart="FIGURE 5" captionStartId="6.[151,250,1684,1708]" captionTargetBox="[185,1402,181,1660]" captionTargetId="figure-17@6.[185,1402,181,1660]" captionTargetPageId="6" captionText="FIGURE 5. Athanas philippei n. sp., paratype, female (cl 5.2 mm) from Pouébo, New Caledonia (MNHN-IU-2018-1017): (A) shrimp alive, dorsal view; (B) same, lateral view; (C) same, frontal view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.5965241" httpUri="https://zenodo.org/record/5965241/files/figure.png" pageId="2" pageNumber="290">Fig. 5</figureCitation>
|
||
,
|
||
<figureCitation id="13312A7C266AFFF2E15FFAC07ADCEF21" box="[595,668,1316,1343]" captionStart="FIGURE 6" captionStartId="8.[151,250,1865,1889]" captionTargetBox="[197,1390,415,1842]" captionTargetId="figure-134@8.[197,1390,415,1842]" captionTargetPageId="8" captionText="FIGURE 6. Athanas philippei n. sp. [A, B]: (A) holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018- 1124): shrimp alive, lateral view; (B) paratype, female (cl 7.2 mm) from Saint-Mathieu, New Caledonia (MNHN-IU-2019- 2986), shrimp alive, anterolateral view. Lysiosquillina maculata (Fabricius, 1793) (Lysiosquillidae) [C]: host (MNHN-IU-2018- 1087) of the female paratype of A. philippei n. sp. (MNHN-IU-2018-1017) from Pouébo, New Caledonia. Photographs by Z. Ďuriš." figureDoi="http://doi.org/10.5281/zenodo.5965243" httpUri="https://zenodo.org/record/5965243/files/figure.png" pageId="2" pageNumber="290">6A, B</figureCitation>
|
||
). Right (minor) cheliped (
|
||
<figureCitation id="13312A7C266AFFF2E0DDFAC07C0EEF21" box="[977,1102,1316,1343]" captionStart="FIGURE 3" captionStartId="4.[151,250,1910,1934]" captionTargetBox="[172,1412,532,1871]" captionTargetId="figure-169@4.[164,1423,525,1886]" captionTargetPageId="4" captionText="FIGURE 3. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) right (minor) cheliped, fully stretched, lateral view; (B) same, proximal articles (mainly ischium, merus and carpus), mesial view; (C) same, folded, lateral view; (D) left (major) cheliped, distal articles (mainly carpus and chela), lateral view; (E) same, distal articles (mainly carpus and chela), dorsolateral view; (F) same, ischium, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5965231" httpUri="https://zenodo.org/record/5965231/files/figure.png" pageId="2" pageNumber="290">Fig. 3A–C</figureCitation>
|
||
) moderately robust. Ischium somewhat flattened ventrally, with about 12 slender spiniform setae, most organised in 2 rows, 1 row along dorsal margin and 1 more irregular row on mesial surface, in addition to 1 spiniform seta situated on proximo-mesial surface; ventrolateral margin distinctly rugose. Merus relatively slender, almost twice as long as ischium, almost 8 times as long as wide at mid-length, not swollen, shallowly excavated ventrally; ventromesial and ventrolateral margins each with row of 3 poorly marked, low, blunt teeth subdistally. Carpus relatively short, cup-shaped, constricted proximally. Palm subcylindrical, slightly more than twice as long as maximal width, with row of 6 small tubercles on ventrolateral surface and shallow sinus near base of pollex. Fingers somewhat twisted away from main chela axis, subequal, about 0.6 length of palm; tips crossing when chela closed; cutting edge of pollex with 2 teeth, proximal tooth larger and more rounded, distal tooth smaller and more triangular, distal portion smooth; cutting edge of dactylus with 2 teeth, proximal tooth small and rounded, more distal tooth broader, situated at mid-length of cutting edge; all 4 teeth fitting into broad hiatuses on opposed margins.
|
||
</paragraph>
|
||
<paragraph id="8BB536F9266AFFF2E3CBF9307B9DEDF4" blockId="2.[151,1437,1136,2027]" pageId="2" pageNumber="290">
|
||
Left (major) cheliped (
|
||
<figureCitation id="13312A7C266AFFF2E2CEF9307A77ECF1" box="[450,567,1748,1775]" captionStart="FIGURE 3" captionStartId="4.[151,250,1910,1934]" captionTargetBox="[172,1412,532,1871]" captionTargetId="figure-169@4.[164,1423,525,1886]" captionTargetPageId="4" captionText="FIGURE 3. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) right (minor) cheliped, fully stretched, lateral view; (B) same, proximal articles (mainly ischium, merus and carpus), mesial view; (C) same, folded, lateral view; (D) left (major) cheliped, distal articles (mainly carpus and chela), lateral view; (E) same, distal articles (mainly carpus and chela), dorsolateral view; (F) same, ischium, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5965231" httpUri="https://zenodo.org/record/5965231/files/figure.png" pageId="2" pageNumber="290">Fig. 3D–F</figureCitation>
|
||
) slightly more robust than right cheliped. Ischium, merus and carpus generally similar to those of right cheliped; chela slightly more robust and with somewhat different palm-finger ratio. Ischium with 11 spiniform setae, most arranged in 2 rows as on right cheliped, in addition to 2 spiniform setae on proximomesial surface. Merus with ventromesial margin bearing row of 3 low blunt teeth subdistally; ventrolateral margin unarmed subdistally, but with somewhat irregular margin at about distal third. Palm noticeably more swollen than that of right cheliped, with 3 tubercles disposed in row on ventrolateral surface and distinct sinus near base of pollex. Fingers proportionally somewhat longer compared to those of minor chela, about 0.7 length of palm, with dentition generally similar to that of right cheliped, but with more pronounced teeth.
|
||
</paragraph>
|
||
<caption id="DF756671266BFFF3E39BF9557D1FED28" ID-DOI="http://doi.org/10.5281/zenodo.5965227" ID-Zenodo-Dep="5965227" httpUri="https://zenodo.org/record/5965227/files/figure.png" pageId="3" pageNumber="291" startId="3.[151,250,1713,1737]" targetBox="[176,1411,211,1650]" targetPageId="3">
|
||
<paragraph id="8BB536F9266BFFF3E39BF9557D1FED28" blockId="3.[151,1437,1713,1846]" pageId="3" pageNumber="291">
|
||
<emphasis id="B97EEAEB266BFFF3E39BF9557951ECD7" bold="true" box="[151,273,1713,1737]" pageId="3" pageNumber="291">FIGURE 2</emphasis>
|
||
.
|
||
<taxonomicName id="4C0A4D7A266BFFF3E22EF956799AECD4" authority="Anker & Ďuriš, 2022" authorityName="Anker & Ďuriš" authorityYear="2022" box="[290,474,1714,1738]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="291" phylum="Arthropoda" rank="species" species="philippei" status="sp. nov.">
|
||
<emphasis id="B97EEAEB266BFFF3E22EF956799AECD4" box="[290,474,1714,1738]" italics="true" pageId="3" pageNumber="291">Athanas philippei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAEB266BFFF3E2E8F9567A62ECD4" bold="true" box="[484,546,1714,1738]" pageId="3" pageNumber="291">
|
||
<taxonomicNameLabel id="A24D5790266BFFF3E2E8F9567A62ECD4" box="[484,546,1714,1738]" pageId="3" pageNumber="291" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) rostrum and left eye and orbital teeth, lateral view; (B) anterior region, lateral view; (C) same, dorsal view; (D) telson and uropods, dorsal view; (E) telson, posterior half, dorsal view; (F) third maxilliped, lateral view; (G) left uropod, detail of distolateral region, dorsal view. All scales as indicated, except for G drawn from a microscope photograph without scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB536F9266BFFF4E3CBF8807CA4EBCF" blockId="3.[151,1437,1892,2026]" lastBlockId="4.[151,1436,150,465]" lastPageId="4" lastPageNumber="292" pageId="3" pageNumber="291">
|
||
Second pereiopod (
|
||
<figureCitation id="13312A7C266BFFF3E2AEF88079BAED60" box="[418,506,1892,1918]" captionStart="FIGURE 4" captionStartId="5.[151,250,1361,1385]" captionTargetBox="[165,1423,188,1329]" captionTargetId="figure-31@5.[154,1434,181,1338]" captionTargetPageId="5" captionText="FIGURE 4. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) second pereiopod, lateral view; (B) third pereiopod, lateral view; (C) same, distal portion of propodus and dactylus; (D) fourth pereiopod, lateral view; (E) fifth pereiopod, lateral view; (F) same, distal portion of propodus and dactylus, lateral view; (G) same, distal portion of propodus and dactylus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5965237" httpUri="https://zenodo.org/record/5965237/files/figure.png" pageId="3" pageNumber="291">Fig. 4A</figureCitation>
|
||
) slender; ischium about 5 times as long as wide; merus about 1.3 times as long as ischium; carpus slender, with five subdivisions, proximal slightly longer than sum of 4 others; chela simple, with fingers subequal to palm. Third pereiopod (
|
||
<figureCitation id="13312A7C266BFFF3E189F8487ABFEDD8" box="[645,767,1964,1990]" captionStart="FIGURE 4" captionStartId="5.[151,250,1361,1385]" captionTargetBox="[165,1423,188,1329]" captionTargetId="figure-31@5.[154,1434,181,1338]" captionTargetPageId="5" captionText="FIGURE 4. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) second pereiopod, lateral view; (B) third pereiopod, lateral view; (C) same, distal portion of propodus and dactylus; (D) fourth pereiopod, lateral view; (E) fifth pereiopod, lateral view; (F) same, distal portion of propodus and dactylus, lateral view; (G) same, distal portion of propodus and dactylus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5965237" httpUri="https://zenodo.org/record/5965237/files/figure.png" pageId="3" pageNumber="291">Fig. 4B, C</figureCitation>
|
||
) relatively slender; ischium about 5 times as long as wide, with 2 cuspidate setae on ventrolateral surface and 1 slender spiniform seta on distodorsal margin; merus somewhat slender, about twice as long as ischium and about 6.5 times as long as wide; carpus noticeably more slender than merus, about half as long as merus, with 1 slender spiniform seta on distoventral margin; propodus slender, unarmed ventrally, except for single small spiniform seta distally, near propodo-dactylar articulation; dactylus simple, slender, gently curved distally, almost half-length of propodus. Fourth pereiopod (
|
||
<figureCitation id="13312A7C266CFFF4E0CDFEE77C58EB03" box="[961,1048,259,285]" captionStart="FIGURE 4" captionStartId="5.[151,250,1361,1385]" captionTargetBox="[165,1423,188,1329]" captionTargetId="figure-31@5.[154,1434,181,1338]" captionTargetPageId="5" captionText="FIGURE 4. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) second pereiopod, lateral view; (B) third pereiopod, lateral view; (C) same, distal portion of propodus and dactylus; (D) fourth pereiopod, lateral view; (E) fifth pereiopod, lateral view; (F) same, distal portion of propodus and dactylus, lateral view; (G) same, distal portion of propodus and dactylus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5965237" httpUri="https://zenodo.org/record/5965237/files/figure.png" pageId="4" pageNumber="292">Fig. 4D</figureCitation>
|
||
) generally similar to third; ischium bearing 2 cuspidate setae on ventrolateral surface. Fifth pereiopod (
|
||
<figureCitation id="13312A7C266CFFF4E099FEC37C4AEB5F" box="[917,1034,295,321]" captionStart="FIGURE 4" captionStartId="5.[151,250,1361,1385]" captionTargetBox="[165,1423,188,1329]" captionTargetId="figure-31@5.[154,1434,181,1338]" captionTargetPageId="5" captionText="FIGURE 4. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) second pereiopod, lateral view; (B) third pereiopod, lateral view; (C) same, distal portion of propodus and dactylus; (D) fourth pereiopod, lateral view; (E) fifth pereiopod, lateral view; (F) same, distal portion of propodus and dactylus, lateral view; (G) same, distal portion of propodus and dactylus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5965237" httpUri="https://zenodo.org/record/5965237/files/figure.png" pageId="4" pageNumber="292">Fig. 4E, F</figureCitation>
|
||
) shortest and slenderest of walking legs; ischium with 1 cuspidate seta on ventrolateral surface and 1 slender spiniform seta on distodorsal margin; merus slightly more than twice as long as ischium; carpus about 0.6 length of merus; propodus slightly longer than merus, bearing 4 rows of serrulate grooming setae on distolateral surface and 1 pair of short spiniform setae distally, near propodo-dactylar articulation; dactylus slender, simple, curved, about 0.4 length of propodus.
|
||
</paragraph>
|
||
<caption id="DF756671266CFFF4E39BF8927C76EDE4" ID-DOI="http://doi.org/10.5281/zenodo.5965231" ID-Zenodo-Dep="5965231" httpUri="https://zenodo.org/record/5965231/files/figure.png" pageId="4" pageNumber="292" startId="4.[151,250,1910,1934]" targetBox="[172,1412,532,1871]" targetPageId="4">
|
||
<paragraph id="8BB536F9266CFFF4E39BF8927C76EDE4" blockId="4.[151,1436,1910,2042]" pageId="4" pageNumber="292">
|
||
<emphasis id="B97EEAEB266CFFF4E39BF892794EED90" bold="true" box="[151,270,1910,1934]" pageId="4" pageNumber="292">FIGURE 3</emphasis>
|
||
.
|
||
<taxonomicName id="4C0A4D7A266CFFF4E210F8927991ED90" authority="Anker & Ďuriš, 2022" authorityName="Anker & Ďuriš" authorityYear="2022" box="[284,465,1910,1934]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="292" phylum="Arthropoda" rank="species" species="philippei" status="sp. nov.">
|
||
<emphasis id="B97EEAEB266CFFF4E210F8927991ED90" box="[284,465,1910,1934]" italics="true" pageId="4" pageNumber="292">Athanas philippei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAEB266CFFF4E2D5F8927A54ED90" bold="true" box="[473,532,1910,1934]" pageId="4" pageNumber="292">
|
||
<taxonomicNameLabel id="A24D5790266CFFF4E2D5F8927A54ED90" box="[473,532,1910,1934]" pageId="4" pageNumber="292" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) right (minor) cheliped, fully stretched, lateral view; (B) same, proximal articles (mainly ischium, merus and carpus), mesial view; (C) same, folded, lateral view; (D) left (major) cheliped, distal articles (mainly carpus and chela), lateral view; (E) same, distal articles (mainly carpus and chela), dorsolateral view; (F) same, ischium, mesial view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF756671266DFFF5E39BFAB57AB0EFC8" ID-DOI="http://doi.org/10.5281/zenodo.5965237" ID-Zenodo-Dep="5965237" httpUri="https://zenodo.org/record/5965237/files/figure.png" pageId="5" pageNumber="293" startId="5.[151,250,1361,1385]" targetBox="[165,1423,188,1329]" targetPageId="5">
|
||
<paragraph id="8BB536F9266DFFF5E39BFAB57AB0EFC8" blockId="5.[151,1437,1361,1494]" pageId="5" pageNumber="293">
|
||
<emphasis id="B97EEAEB266DFFF5E39BFAB5794EEF77" bold="true" box="[151,270,1361,1385]" pageId="5" pageNumber="293">FIGURE 4</emphasis>
|
||
.
|
||
<taxonomicName id="4C0A4D7A266DFFF5E210FAB67991EF74" authority="Anker & Ďuriš, 2022" authorityName="Anker & Ďuriš" authorityYear="2022" box="[284,465,1362,1386]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="293" phylum="Arthropoda" rank="species" species="philippei" status="sp. nov.">
|
||
<emphasis id="B97EEAEB266DFFF5E210FAB67991EF74" box="[284,465,1362,1386]" italics="true" pageId="5" pageNumber="293">Athanas philippei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAEB266DFFF5E2D5FAB67A54EF74" bold="true" box="[473,532,1362,1386]" pageId="5" pageNumber="293">
|
||
<taxonomicNameLabel id="A24D5790266DFFF5E2D5FAB67A54EF74" box="[473,532,1362,1386]" pageId="5" pageNumber="293" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) second pereiopod, lateral view; (B) third pereiopod, lateral view; (C) same, distal portion of propodus and dactylus; (D) fourth pereiopod, lateral view; (E) fifth pereiopod, lateral view; (F) same, distal portion of propodus and dactylus, lateral view; (G) same, distal portion of propodus and dactylus, mesial view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB536F9266DFFF5E3CBF9E07B56EC78" blockId="5.[151,1437,1540,1963]" pageId="5" pageNumber="293">
|
||
Uropod (
|
||
<figureCitation id="13312A7C266DFFF5E23DF9E079F5EC00" box="[305,437,1540,1566]" captionStart="FIGURE 2" captionStartId="3.[151,250,1713,1737]" captionTargetBox="[176,1411,211,1650]" captionTargetId="figure-31@3.[169,1418,181,1689]" captionTargetPageId="3" captionText="FIGURE 2. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) rostrum and left eye and orbital teeth, lateral view; (B) anterior region, lateral view; (C) same, dorsal view; (D) telson and uropods, dorsal view; (E) telson, posterior half, dorsal view; (F) third maxilliped, lateral view; (G) left uropod, detail of distolateral region, dorsal view. All scales as indicated, except for G drawn from a microscope photograph without scale." figureDoi="http://doi.org/10.5281/zenodo.5965227" httpUri="https://zenodo.org/record/5965227/files/figure.png" pageId="5" pageNumber="293">Fig. 2D, G</figureCitation>
|
||
) with lateral lobe of protopod subacute; exopod with distolateral tooth and adjacent spiniform seta both rather small; lateral portion of diaeresis straight, with small, articulated, spiniform seta (referred to as spinule below) mesial to distolateral spiniform seta.
|
||
</paragraph>
|
||
<paragraph id="8BB536F9266DFFF5E3CBF9947A96ECB0" blockId="5.[151,1437,1540,1963]" pageId="5" pageNumber="293">Gill-exopod formula: 5 pleurobranchs (P1–5); 0 podobranchs; 0 arthrobranchs; 4 strap-like epipods (Mxp3, P1–3); 4 setobranchs (P1–4); 3 exopods (Mxp1–3).</paragraph>
|
||
<paragraph id="8BB536F9266DFFF5E3CBF95C7A04EDB4" blockId="5.[151,1437,1540,1963]" pageId="5" pageNumber="293">
|
||
<emphasis id="B97EEAEB266DFFF5E3CBF95C79C2ECCC" bold="true" box="[199,386,1720,1746]" pageId="5" pageNumber="293">Colour pattern</emphasis>
|
||
. Body generally semi-transparent; carapace and pleon flanks with more or less evenly distributed, irregular clusters of bright red chromatophores, some reminiscent of paired chromosomes of mammals, their density and colour intensity increasing towards posterior end of pleon; mid-dorsal area with broad, opaque, white or pale yellow band extending from tip of rostrum to telson, widest on carapace, tapering towards posterior pleonites; basal parts of antennae and antennules whitish with patches of red chromatophores; chelipeds hyaline white; pereiopods colourless, semi-transparent; telson entirely opaque white; uropod with transparent endopod and opaque white exopod (
|
||
<figureCitation id="13312A7C266DFFF5E29AF87479A5EDB5" box="[406,485,1936,1963]" captionStart="FIGURE 5" captionStartId="6.[151,250,1684,1708]" captionTargetBox="[185,1402,181,1660]" captionTargetId="figure-17@6.[185,1402,181,1660]" captionTargetPageId="6" captionText="FIGURE 5. Athanas philippei n. sp., paratype, female (cl 5.2 mm) from Pouébo, New Caledonia (MNHN-IU-2018-1017): (A) shrimp alive, dorsal view; (B) same, lateral view; (C) same, frontal view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.5965241" httpUri="https://zenodo.org/record/5965241/files/figure.png" pageId="5" pageNumber="293">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation id="13312A7C266DFFF5E2FDF8747A76EDB4" box="[497,566,1936,1962]" captionStart="FIGURE 6" captionStartId="8.[151,250,1865,1889]" captionTargetBox="[197,1390,415,1842]" captionTargetId="figure-134@8.[197,1390,415,1842]" captionTargetPageId="8" captionText="FIGURE 6. Athanas philippei n. sp. [A, B]: (A) holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018- 1124): shrimp alive, lateral view; (B) paratype, female (cl 7.2 mm) from Saint-Mathieu, New Caledonia (MNHN-IU-2019- 2986), shrimp alive, anterolateral view. Lysiosquillina maculata (Fabricius, 1793) (Lysiosquillidae) [C]: host (MNHN-IU-2018- 1087) of the female paratype of A. philippei n. sp. (MNHN-IU-2018-1017) from Pouébo, New Caledonia. Photographs by Z. Ďuriš." figureDoi="http://doi.org/10.5281/zenodo.5965243" httpUri="https://zenodo.org/record/5965243/files/figure.png" pageId="5" pageNumber="293">6A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF756671266EFFF6E39BF9707CC7ECCE" ID-DOI="http://doi.org/10.5281/zenodo.5965241" ID-Zenodo-Dep="5965241" httpUri="https://zenodo.org/record/5965241/files/figure.png" pageId="6" pageNumber="294" startId="6.[151,250,1684,1708]" targetBox="[185,1402,181,1660]" targetPageId="6">
|
||
<paragraph id="8BB536F9266EFFF6E39BF9707CC7ECCE" blockId="6.[151,1436,1684,1744]" pageId="6" pageNumber="294">
|
||
<emphasis id="B97EEAEB266EFFF6E39BF970794DECB2" bold="true" box="[151,269,1684,1708]" pageId="6" pageNumber="294">FIGURE 5</emphasis>
|
||
.
|
||
<taxonomicName id="4C0A4D7A266EFFF6E215F970798CECB2" authority="Anker & Ďuriš, 2022" authorityName="Anker & Ďuriš" authorityYear="2022" box="[281,460,1684,1708]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="294" phylum="Arthropoda" rank="species" species="philippei" status="sp. nov.">
|
||
<emphasis id="B97EEAEB266EFFF6E215F970798CECB2" box="[281,460,1684,1708]" italics="true" pageId="6" pageNumber="294">Athanas philippei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAEB266EFFF6E2DEF9707A4BECB2" bold="true" box="[466,523,1684,1708]" pageId="6" pageNumber="294">
|
||
<taxonomicNameLabel id="A24D5790266EFFF6E2DEF9707A4BECB2" box="[466,523,1684,1708]" pageId="6" pageNumber="294" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype, female (cl 5.2 mm) from Pouébo, New Caledonia (MNHN-IU-2018-1017): (A) shrimp alive, dorsal view; (B) same, lateral view; (C) same, frontal view. Photographs by A. Anker.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB536F9266EFFF7E3CBF91A7961EBB5" blockId="6.[151,1437,1790,2033]" lastBlockId="7.[151,1437,151,2023]" lastPageId="7" lastPageNumber="295" pageId="6" pageNumber="294">
|
||
<emphasis id="B97EEAEB266EFFF6E3CBF91A7978ED06" bold="true" box="[199,312,1790,1816]" pageId="6" pageNumber="294">Variation</emphasis>
|
||
. The cl size range of the type series is
|
||
<quantity id="4CF29B1C266EFFF6E009F91A7BCEED07" box="[773,910,1790,1817]" metricMagnitude="-3" metricUnit="m" metricValue="6.449999999999999" metricValueMax="7.8" metricValueMin="5.1" pageId="6" pageNumber="294" unit="mm" value="6.449999999999999" valueMax="7.8" valueMin="5.1">5.1–7.8 mm</quantity>
|
||
(
|
||
<typeStatus id="54B1885B266EFFF6E0ACF91A7C48ED06" box="[928,1032,1790,1816]" pageId="6" pageNumber="294" type="holotype">holotype</typeStatus>
|
||
: cl
|
||
<quantity id="4CF29B1C266EFFF6E73EF91A7CCFED07" box="[1074,1167,1790,1817]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="6" pageNumber="294" unit="mm" value="6.9">6.9 mm</quantity>
|
||
). The rostrum typically reaches at least to the distal half of the second article of the antennular peduncle, as in the
|
||
<typeStatus id="54B1885B266EFFF6E79DF8C67CB6ED22" box="[1169,1270,1826,1852]" pageId="6" pageNumber="294" type="holotype">holotype</typeStatus>
|
||
, but falls short of it in some
|
||
<typeStatus id="54B1885B266EFFF6E225F8A379C9ED7F" box="[297,393,1863,1889]" pageId="6" pageNumber="294" type="paratype">paratype</typeStatus>
|
||
specimens. The number of teeth on the rostral carina varies from 5 (
|
||
<typeStatus id="54B1885B266EFFF6E78CF8A27CA5ED7F" box="[1152,1253,1862,1889]" pageId="6" pageNumber="294" type="holotype">holotype</typeStatus>
|
||
) to 10 (
|
||
<typeStatus id="54B1885B266EFFF6E636F8A27DDCED7E" box="[1338,1436,1862,1888]" pageId="6" pageNumber="294" type="paratype">paratype</typeStatus>
|
||
MNHN-IU-2018-1123); in the latter case they extend almost to the base of the rostral carina (
|
||
<figureCitation id="13312A7C266EFFF6E7BDF88E7D41ED9A" box="[1201,1281,1898,1924]" captionStart="FIGURE 2" captionStartId="3.[151,250,1713,1737]" captionTargetBox="[176,1411,211,1650]" captionTargetId="figure-31@3.[169,1418,181,1689]" captionTargetPageId="3" captionText="FIGURE 2. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) rostrum and left eye and orbital teeth, lateral view; (B) anterior region, lateral view; (C) same, dorsal view; (D) telson and uropods, dorsal view; (E) telson, posterior half, dorsal view; (F) third maxilliped, lateral view; (G) left uropod, detail of distolateral region, dorsal view. All scales as indicated, except for G drawn from a microscope photograph without scale." figureDoi="http://doi.org/10.5281/zenodo.5965227" httpUri="https://zenodo.org/record/5965227/files/figure.png" pageId="6" pageNumber="294">Fig. 2I</figureCitation>
|
||
). The number of ventral tubercles on the chela palm varies from 1 (untypical condition, on both chelae of the
|
||
<typeStatus id="54B1885B266EFFF6E7D9F86B7D75EDB7" box="[1237,1333,1935,1961]" pageId="6" pageNumber="294" type="paratype">paratype</typeStatus>
|
||
MNHN- IU-2019-2986) to 3 (minor) and 6 (major), as in the
|
||
<typeStatus id="54B1885B266EFFF6E1FDF8567B13EDD2" box="[753,851,1970,1996]" pageId="6" pageNumber="294" type="holotype">holotype</typeStatus>
|
||
and
|
||
<specimenCount id="9D0CFD70266EFFF6E081F8577C6AEDD3" box="[909,1066,1971,1997]" pageId="6" pageNumber="294" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
|
||
(MNHN-IU-2018-608, MNHN- IU-2018-1087). The number of teeth on the ventrolateral margin of the cheliped merus varies from 3 (
|
||
<typeStatus id="54B1885B266EFFF6E61EF8327D37EDEF" box="[1298,1399,2006,2033]" pageId="6" pageNumber="294" type="holotype">holotype</typeStatus>
|
||
) to 6 (
|
||
<typeStatus id="54B1885B266FFFF7E3B8FF737956EAAF" box="[180,278,151,177]" pageId="7" pageNumber="295" type="paratype">paratype</typeStatus>
|
||
MNHN-IU-2018-608). The finger/palm length ratio of the minor chela is also variable to some degree, with the fingers being subequal to the palm in one of the
|
||
<typeStatus id="54B1885B266FFFF7E004FF5F7B33EACB" box="[776,883,187,213]" pageId="7" pageNumber="295" type="paratype">paratypes</typeStatus>
|
||
(MNHN-IU-2018-608). The number of cuspidate setae on the ischia of the third to fifth pereiopods (P3–5) is relatively consistent: two on P3 and P4, and one on P5 (except on left P
|
||
<quantity id="4CF29B1C266FFFF7E25AFEE779C2EB03" box="[342,386,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="7" pageNumber="295" unit="in" value="3.0">3 in</quantity>
|
||
the
|
||
<typeStatus id="54B1885B266FFFF7E2B9FEE77A55EB03" box="[437,533,259,285]" pageId="7" pageNumber="295" type="paratype">paratype</typeStatus>
|
||
MNHN-IU-2019-1123, with a single spiniform seta). The small spinule on the uropodal exopod, which is present on both sides in the
|
||
<typeStatus id="54B1885B266FFFF7E001FEC27B32EB5E" box="[781,882,294,320]" pageId="7" pageNumber="295" type="holotype">holotype</typeStatus>
|
||
, is present only on the right side in
|
||
<specimenCount id="9D0CFD70266FFFF7E607FEC37DDCEB5F" box="[1291,1436,295,321]" pageId="7" pageNumber="295" type="generic" typeStatus="paratype">one paratype</specimenCount>
|
||
(MNHN-IU-2018-608), being replaced on the left side by a short, feebly plumose seta. This seta, instead of a spinule, is also present on both uropods in at least two other
|
||
<typeStatus id="54B1885B266FFFF7E052FE8A7B89EB96" box="[862,969,366,392]" pageId="7" pageNumber="295" type="paratype">paratypes</typeStatus>
|
||
(MNHN-IU-2018-1123 and MNHN-IU- 2019-2986).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3106572266FFFF7E3CBFE507B71E80B" pageId="7" pageNumber="295" type="etymology">
|
||
<paragraph id="8BB536F9266FFFF7E3CBFE507B71E80B" blockId="7.[151,1437,151,2023]" pageId="7" pageNumber="295">
|
||
<emphasis id="B97EEAEB266FFFF7E3CBFE507908EBD0" bold="true" box="[199,328,436,462]" pageId="7" pageNumber="295">Etymology</emphasis>
|
||
. The new species is named after our colleague, Prof. Philippe Bouchet (MNHN), a prominent malacologist and expert of marine biodiversity, and the main organiser of several “Our Planet Reviewed” expeditions, to which the authors were generously invited to participate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3106572266FFFF7E3CBFDFA7AF2E842" pageId="7" pageNumber="295" type="distribution">
|
||
<paragraph id="8BB536F9266FFFF7E3CBFDFA7AF2E842" blockId="7.[151,1437,151,2023]" pageId="7" pageNumber="295">
|
||
<emphasis id="B97EEAEB266FFFF7E3CBFDFA7919E826" bold="true" box="[199,345,542,568]" pageId="7" pageNumber="295">Distribution</emphasis>
|
||
. Presently known only from three localities on the north-eastern coast of
|
||
<collectingCountry id="F31D7669266FFFF7E7BEFDFB7D25E826" box="[1202,1381,542,569]" name="New Caledonia" pageId="7" pageNumber="295">New Caledonia</collectingCountry>
|
||
, viz. Tiari (
|
||
<typeStatus id="54B1885B266FFFF7E3D7FDA6794DE842" box="[219,269,578,604]" pageId="7" pageNumber="295">type</typeStatus>
|
||
locality), Saint-Mathieu and Pouébo.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3106572266FFFF7E3CBFD817C90E8D9" pageId="7" pageNumber="295" type="biology_ecology">
|
||
<paragraph id="8BB536F9266FFFF7E3CBFD817C90E8D9" blockId="7.[151,1437,151,2023]" pageId="7" pageNumber="295">
|
||
<emphasis id="B97EEAEB266FFFF7E3CBFD817966E861" bold="true" box="[199,294,613,639]" pageId="7" pageNumber="295">Ecology</emphasis>
|
||
. Intertidal and shallow subtidal (
|
||
<quantity id="4CF29B1C266FFFF7E19EFD817A93E89E" box="[658,723,613,640]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="295" unit="m" value="0.5">0.5 m</quantity>
|
||
at low tide) sand flats adjacent to seagrass beds and coral rubble (
|
||
<figureCitation id="13312A7C266FFFF7E3ACFD6D78A5E8BD" box="[160,229,649,675]" captionStart="FIGURE 7" captionStartId="9.[151,250,1615,1639]" captionTargetBox="[151,1435,181,1591]" captionTargetId="figure-31@9.[151,1435,181,1591]" captionTargetPageId="9" captionText="FIGURE 7. One of the collection localities of Athanas philippei n. sp., near Pouébo, north-eastern coast of New Caledonia: (A) view from the beach, with black arrow approximately indicating the area of the collection of the first specimen (female paratype, MNHN-IU-2018-1017), (B) view of the habitat during sampling. Photographs by A. Anker (A) and Z. Ďuriš (B)." figureDoi="http://doi.org/10.5281/zenodo.5965247" httpUri="https://zenodo.org/record/5965247/files/figure.png" pageId="7" pageNumber="295">Fig. 7</figureCitation>
|
||
); associated with burrows of the stomatopod
|
||
<taxonomicName id="4C0A4D7A266FFFF7E1EBFD6D7BB5E8BD" baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[743,1013,649,675]" class="Malacostraca" family="Lysiosquillidae" genus="Lysiosquillina" kingdom="Animalia" order="Stomatopoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="maculata">
|
||
<emphasis id="B97EEAEB266FFFF7E1EBFD6D7BB5E8BD" box="[743,1013,649,675]" italics="true" pageId="7" pageNumber="295">Lysiosquillina maculata</emphasis>
|
||
</taxonomicName>
|
||
; the host of the
|
||
<typeStatus id="54B1885B266FFFF7E7A2FD6D7D4EE8BD" box="[1198,1294,649,675]" pageId="7" pageNumber="295" type="paratype">paratype</typeStatus>
|
||
MNHN-IU- 2018-1017 (deposited under MNHN-IU-2018-1087) had a total length of about
|
||
<quantity id="4CF29B1C266FFFF7E719FD487C1AE8D9" box="[1045,1114,684,711]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="295" unit="cm" value="15.0">15 cm</quantity>
|
||
(
|
||
<figureCitation id="13312A7C266FFFF7E766FD487C82E8D9" box="[1130,1218,684,711]" captionStart="FIGURE 6" captionStartId="8.[151,250,1865,1889]" captionTargetBox="[197,1390,415,1842]" captionTargetId="figure-134@8.[197,1390,415,1842]" captionTargetPageId="8" captionText="FIGURE 6. Athanas philippei n. sp. [A, B]: (A) holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018- 1124): shrimp alive, lateral view; (B) paratype, female (cl 7.2 mm) from Saint-Mathieu, New Caledonia (MNHN-IU-2019- 2986), shrimp alive, anterolateral view. Lysiosquillina maculata (Fabricius, 1793) (Lysiosquillidae) [C]: host (MNHN-IU-2018- 1087) of the female paratype of A. philippei n. sp. (MNHN-IU-2018-1017) from Pouébo, New Caledonia. Photographs by Z. Ďuriš." figureDoi="http://doi.org/10.5281/zenodo.5965243" httpUri="https://zenodo.org/record/5965243/files/figure.png" pageId="7" pageNumber="295">Fig. 6C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3106572266FFFFAE3CBFD3479BAE9D7" lastPageId="10" lastPageNumber="298" pageId="7" pageNumber="295" type="discussion">
|
||
<paragraph id="8BB536F9266FFFF7E3CBFD347BEEE92F" blockId="7.[151,1437,151,2023]" pageId="7" pageNumber="295">
|
||
<emphasis id="B97EEAEB266FFFF7E3CBFD347974E8F4" bold="true" box="[199,308,720,746]" pageId="7" pageNumber="295">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C0A4D7A266FFFF7E244FD347A54E8F4" authority="Anker & Ďuriš, 2022" authorityName="Anker & Ďuriš" authorityYear="2022" box="[328,532,720,746]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="philippei" status="sp. nov.">
|
||
<emphasis id="B97EEAEB266FFFF7E244FD347A54E8F4" box="[328,532,720,746]" italics="true" pageId="7" pageNumber="295">Athanas philippei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAEB266FFFF7E12CFD347A24E8F4" bold="true" box="[544,612,720,746]" pageId="7" pageNumber="295">
|
||
<taxonomicNameLabel id="A24D5790266FFFF7E12CFD347A24E8F4" box="[544,612,720,746]" pageId="7" pageNumber="295" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is morphologically closest to two other Indo-West Pacific species of
|
||
<taxonomicName id="4C0A4D7A266FFFF7E39BFD1778B3E913" authorityName="Leach" authorityYear="1814" box="[151,243,755,781]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97EEAEB266FFFF7E39BFD1778B3E913" box="[151,243,755,781]" italics="true" pageId="7" pageNumber="295">Athanas</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0A4D7A266FFFF7E20DFD10793AE913" authorityName="Anker & Komai" authorityYear="2010" box="[257,378,755,782]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="ahyongi">
|
||
<emphasis id="B97EEAEB266FFFF7E20DFD10793AE913" box="[257,378,755,782]" italics="true" pageId="7" pageNumber="295">A. ahyongi</emphasis>
|
||
</taxonomicName>
|
||
(suspected to be associated with stomatopods) and
|
||
<taxonomicName id="4C0A4D7A266FFFF7E0B3FD107C19E913" authorityName="Duris & Anker" authorityYear="2014" box="[959,1113,755,782]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="manticolus">
|
||
<emphasis id="B97EEAEB266FFFF7E0B3FD107C19E913" box="[959,1113,755,782]" italics="true" pageId="7" pageNumber="295">A. manticolus</emphasis>
|
||
</taxonomicName>
|
||
(associated with
|
||
<taxonomicName id="4C0A4D7A266FFFF7E611FD17795DE92F" baseAuthorityName="Fabricius" baseAuthorityYear="1798" class="Malacostraca" family="Nannosquillidae" genus="Bigelowina" kingdom="Animalia" order="Stomatopoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="phalangium">
|
||
<emphasis id="B97EEAEB266FFFF7E611FD17795DE92F" italics="true" pageId="7" pageNumber="295">Bigelowina phalangium</emphasis>
|
||
</taxonomicName>
|
||
, see below) (
|
||
<bibRefCitation id="EF9B4B08266FFFF7E2BEFCF37AEDE92F" author="Anker, A. & Komai, T." box="[434,685,791,817]" pageId="7" pageNumber="295" pagination="45 - 54" refId="ref6296" refString="Anker, A. & Komai, T. (2010) Description of a new species of Athanas Leach, 1814 (Crustacea: Decapoda: Alpheidae) from Madagascar and Japan. Zootaxa, 2680 (1), 45 - 54. https: // doi. org / 10.11646 / zootaxa. 2680.1.4" type="journal article" year="2010">Anker & Komai 2010</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9B4B08266FFFF7E1B4FCFC7BE2E92F" author="Duris, Z. & Anker, A." box="[696,930,791,819]" pageId="7" pageNumber="295" pagination="550 - 558" refId="ref7010" refString="Duris, Z. & Anker, A. (2014) Athanas manticolus sp. nov., a new stomatopod-associated alpheid shrimp from Vietnam (Crustacea, Decapoda). Zootaxa, 3784 (5), 550 - 558. https: // doi. org / 10.11646 / zootaxa. 3784.5.4" type="journal article" year="2014">Ďuriš & Anker 2014</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB536F9266FFFF7E3CBFCDE7882EF76" blockId="7.[151,1437,151,2023]" pageId="7" pageNumber="295">
|
||
The new species can be separated from
|
||
<taxonomicName id="4C0A4D7A266FFFF7E183FCDE7B4AE94A" authorityName="Anker & Komai" authorityYear="2010" box="[655,778,826,852]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="ahyongi">
|
||
<emphasis id="B97EEAEB266FFFF7E183FCDE7B4AE94A" box="[655,778,826,852]" italics="true" pageId="7" pageNumber="295">A. ahyongi</emphasis>
|
||
</taxonomicName>
|
||
by the presence of a post-rostral tubercle on the carapace (
|
||
<emphasis id="B97EEAEB266FFFF7E3ADFCBA78FFE969" box="[161,191,862,887]" italics="true" pageId="7" pageNumber="295">vs.</emphasis>
|
||
absent in
|
||
<taxonomicName id="4C0A4D7A266FFFF7E23DFCBA79EAE969" authorityName="Anker & Komai" authorityYear="2010" box="[305,426,861,888]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="ahyongi">
|
||
<emphasis id="B97EEAEB266FFFF7E23DFCBA79EAE969" box="[305,426,861,888]" italics="true" pageId="7" pageNumber="295">A. ahyongi</emphasis>
|
||
</taxonomicName>
|
||
); the rostrum with a series of minute, spaced dorsal teeth (
|
||
<emphasis id="B97EEAEB266FFFF7E731FCBA7C14E969" box="[1085,1108,862,887]" italics="true" pageId="7" pageNumber="295">vs</emphasis>
|
||
. unarmed in
|
||
<taxonomicName id="4C0A4D7A266FFFF7E7E4FCBA7D22E969" authorityName="Anker & Komai" authorityYear="2010" box="[1256,1378,861,888]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="ahyongi">
|
||
<emphasis id="B97EEAEB266FFFF7E7E4FCBA7D22E969" box="[1256,1378,861,888]" italics="true" pageId="7" pageNumber="295">A. ahyongi</emphasis>
|
||
</taxonomicName>
|
||
); the rostrum reaching mid-length of the second article of the antennular peduncle (
|
||
<emphasis id="B97EEAEB266FFFF7E706FC667C61E985" box="[1034,1057,898,923]" italics="true" pageId="7" pageNumber="295">vs</emphasis>
|
||
. falling short of the distal margin of the first article in
|
||
<taxonomicName id="4C0A4D7A266FFFF7E28AFC417A41E9A0" authorityName="Anker & Komai" authorityYear="2010" box="[390,513,932,959]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="ahyongi">
|
||
<emphasis id="B97EEAEB266FFFF7E28AFC417A41E9A0" box="[390,513,932,959]" italics="true" pageId="7" pageNumber="295">A. ahyongi</emphasis>
|
||
</taxonomicName>
|
||
); the ultimate article of the third maxilliped very slender (
|
||
<emphasis id="B97EEAEB266FFFF7E7AEFC417CF9E9A0" box="[1186,1209,933,958]" italics="true" pageId="7" pageNumber="295">vs</emphasis>
|
||
. much stouter in
|
||
<taxonomicName id="4C0A4D7A266FFFF7E688FC4178B1E9FC" authorityName="Anker & Komai" authorityYear="2010" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="ahyongi">
|
||
<emphasis id="B97EEAEB266FFFF7E688FC4178B1E9FC" italics="true" pageId="7" pageNumber="295">A. ahyongi</emphasis>
|
||
</taxonomicName>
|
||
); the cheliped merus with few (typically three) low blunt teeth in subdistal position on both ventrolateral and ventromesial margins (
|
||
<emphasis id="B97EEAEB266FFFF7E295FC0879F0EE1B" box="[409,432,1004,1029]" italics="true" pageId="7" pageNumber="295">vs</emphasis>
|
||
. armed with much stronger teeth along almost entire ventrolateral margin in
|
||
<taxonomicName id="4C0A4D7A266FFFF7E61EFC087DCBEE1B" authorityName="Anker & Komai" authorityYear="2010" box="[1298,1419,1003,1030]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="ahyongi">
|
||
<emphasis id="B97EEAEB266FFFF7E61EFC087DCBEE1B" box="[1298,1419,1003,1030]" italics="true" pageId="7" pageNumber="295">A. ahyongi</emphasis>
|
||
</taxonomicName>
|
||
); the ventral surface of the cheliped palm with one to six small but distinct tubercles (
|
||
<emphasis id="B97EEAEB266FFFF7E74AFBF47C1DEE37" box="[1094,1117,1040,1065]" italics="true" pageId="7" pageNumber="295">vs</emphasis>
|
||
. smooth in
|
||
<taxonomicName id="4C0A4D7A266FFFF7E7EEFBEB7D1BEE37" authorityName="Anker & Komai" authorityYear="2010" box="[1250,1371,1039,1065]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="ahyongi">
|
||
<emphasis id="B97EEAEB266FFFF7E7EEFBEB7D1BEE37" box="[1250,1371,1039,1065]" italics="true" pageId="7" pageNumber="295">A. ahyongi</emphasis>
|
||
</taxonomicName>
|
||
); and the third pereiopod propodus unarmed ventrally, except for a single distoventral spiniform seta (vs. with at least two additional spiniform setae on the ventral margin of the propodus in
|
||
<taxonomicName id="4C0A4D7A266FFFF7E0B7FBB27C74EE6E" authorityName="Anker & Komai" authorityYear="2010" box="[955,1076,1110,1136]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="ahyongi">
|
||
<emphasis id="B97EEAEB266FFFF7E0B7FBB27C74EE6E" box="[955,1076,1110,1136]" italics="true" pageId="7" pageNumber="295">A. ahyongi</emphasis>
|
||
</taxonomicName>
|
||
) (cf.
|
||
<figureCitation id="13312A7C266FFFF7E77DFBB27C9DEE6E" box="[1137,1245,1110,1136]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="3.[151,250,1713,1737]" captionStartId-1="4.[151,250,1910,1934]" captionStartId-2="5.[151,250,1361,1385]" captionTargetBox-0="[176,1411,211,1650]" captionTargetBox-1="[172,1412,532,1871]" captionTargetBox-2="[165,1423,188,1329]" captionTargetId-0="figure-31@3.[169,1418,181,1689]" captionTargetId-1="figure-169@4.[164,1423,525,1886]" captionTargetId-2="figure-31@5.[154,1434,181,1338]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 2. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) rostrum and left eye and orbital teeth, lateral view; (B) anterior region, lateral view; (C) same, dorsal view; (D) telson and uropods, dorsal view; (E) telson, posterior half, dorsal view; (F) third maxilliped, lateral view; (G) left uropod, detail of distolateral region, dorsal view. All scales as indicated, except for G drawn from a microscope photograph without scale." captionText-1="FIGURE 3. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) right (minor) cheliped, fully stretched, lateral view; (B) same, proximal articles (mainly ischium, merus and carpus), mesial view; (C) same, folded, lateral view; (D) left (major) cheliped, distal articles (mainly carpus and chela), lateral view; (E) same, distal articles (mainly carpus and chela), dorsolateral view; (F) same, ischium, mesial view." captionText-2="FIGURE 4. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) second pereiopod, lateral view; (B) third pereiopod, lateral view; (C) same, distal portion of propodus and dactylus; (D) fourth pereiopod, lateral view; (E) fifth pereiopod, lateral view; (F) same, distal portion of propodus and dactylus, lateral view; (G) same, distal portion of propodus and dactylus, mesial view." figureDoi-0="http://doi.org/10.5281/zenodo.5965227" figureDoi-1="http://doi.org/10.5281/zenodo.5965231" figureDoi-2="http://doi.org/10.5281/zenodo.5965237" httpUri-0="https://zenodo.org/record/5965227/files/figure.png" httpUri-1="https://zenodo.org/record/5965231/files/figure.png" httpUri-2="https://zenodo.org/record/5965237/files/figure.png" pageId="7" pageNumber="295">Figs. 2–4</figureCitation>
|
||
;
|
||
<bibRefCitation id="EF9B4B08266FFFF7E7EAFBB27894EE8D" author="Anker, A. & Komai, T." pageId="7" pageNumber="295" pagination="45 - 54" refId="ref6296" refString="Anker, A. & Komai, T. (2010) Description of a new species of Athanas Leach, 1814 (Crustacea: Decapoda: Alpheidae) from Madagascar and Japan. Zootaxa, 2680 (1), 45 - 54. https: // doi. org / 10.11646 / zootaxa. 2680.1.4" type="journal article" year="2010">Anker & Komai 2010</bibRefCitation>
|
||
). The colour patterns of
|
||
<taxonomicName id="4C0A4D7A266FFFF7E2EAFB9E7A2AEE8D" authority="Anker & Ďuriš, 2022" authorityName="Anker & Ďuriš" authorityYear="2022" box="[486,618,1145,1172]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="philippei" status="sp. nov.">
|
||
<emphasis id="B97EEAEB266FFFF7E2EAFB9E7A2AEE8D" box="[486,618,1145,1172]" italics="true" pageId="7" pageNumber="295">A. philippei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAEB266FFFF7E17DFB9D7AF0EE8D" bold="true" box="[625,688,1145,1171]" pageId="7" pageNumber="295">
|
||
<taxonomicNameLabel id="A24D5790266FFFF7E17DFB9D7AF0EE8D" box="[625,688,1145,1171]" pageId="7" pageNumber="295" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13312A7C266FFFF7E1CCFB9D7B4FEE8A" box="[704,783,1145,1172]" captionStart="FIGURE 5" captionStartId="6.[151,250,1684,1708]" captionTargetBox="[185,1402,181,1660]" captionTargetId="figure-17@6.[185,1402,181,1660]" captionTargetPageId="6" captionText="FIGURE 5. Athanas philippei n. sp., paratype, female (cl 5.2 mm) from Pouébo, New Caledonia (MNHN-IU-2018-1017): (A) shrimp alive, dorsal view; (B) same, lateral view; (C) same, frontal view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.5965241" httpUri="https://zenodo.org/record/5965241/files/figure.png" pageId="7" pageNumber="295">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation id="13312A7C266FFFF7E017FB9D7B1FEE8A" box="[795,863,1145,1172]" captionStart="FIGURE 6" captionStartId="8.[151,250,1865,1889]" captionTargetBox="[197,1390,415,1842]" captionTargetId="figure-134@8.[197,1390,415,1842]" captionTargetPageId="8" captionText="FIGURE 6. Athanas philippei n. sp. [A, B]: (A) holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018- 1124): shrimp alive, lateral view; (B) paratype, female (cl 7.2 mm) from Saint-Mathieu, New Caledonia (MNHN-IU-2019- 2986), shrimp alive, anterolateral view. Lysiosquillina maculata (Fabricius, 1793) (Lysiosquillidae) [C]: host (MNHN-IU-2018- 1087) of the female paratype of A. philippei n. sp. (MNHN-IU-2018-1017) from Pouébo, New Caledonia. Photographs by Z. Ďuriš." figureDoi="http://doi.org/10.5281/zenodo.5965243" httpUri="https://zenodo.org/record/5965243/files/figure.png" pageId="7" pageNumber="295">6A, B</figureCitation>
|
||
) and
|
||
<taxonomicName id="4C0A4D7A266FFFF7E093FB9E7C58EE8D" authorityName="Anker & Komai" authorityYear="2010" box="[927,1048,1145,1172]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="ahyongi">
|
||
<emphasis id="B97EEAEB266FFFF7E093FB9E7C58EE8D" box="[927,1048,1145,1172]" italics="true" pageId="7" pageNumber="295">A. ahyongi</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<bibRefCitation id="EF9B4B08266FFFF7E740FB9D7D05EE8D" author="Anker, A. & Komai, T." box="[1100,1349,1145,1171]" pageId="7" pageNumber="295" pagination="45 - 54" refId="ref6296" refString="Anker, A. & Komai, T. (2010) Description of a new species of Athanas Leach, 1814 (Crustacea: Decapoda: Alpheidae) from Madagascar and Japan. Zootaxa, 2680 (1), 45 - 54. https: // doi. org / 10.11646 / zootaxa. 2680.1.4" type="journal article" year="2010">Anker & Komai 2010</bibRefCitation>
|
||
: fig. 6), although generally similar, are different in several important aspects. In the new species, the red chromatophores are present on most of the carapace and pleon flanks, whereas in
|
||
<taxonomicName id="4C0A4D7A266FFFF7E088FB257BBEEEC4" authorityName="Anker & Komai" authorityYear="2010" box="[900,1022,1216,1243]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="ahyongi">
|
||
<emphasis id="B97EEAEB266FFFF7E088FB257BBEEEC4" box="[900,1022,1216,1243]" italics="true" pageId="7" pageNumber="295">A. ahyongi</emphasis>
|
||
</taxonomicName>
|
||
, they form a broad red lateral band, extending the full body length of the shrimp, with two colourless bands or areas, one between the red band and the white or pale-yellow mid-dorsal band, and another below the red band. Furthermore, in
|
||
<taxonomicName id="4C0A4D7A266FFFF7E7ADFAEC7D65EF3F" authority="Anker & Ďuriš, 2022" authorityName="Anker & Ďuriš" authorityYear="2022" box="[1185,1317,1287,1314]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="philippei" status="sp. nov.">
|
||
<emphasis id="B97EEAEB266FFFF7E7ADFAEC7D65EF3F" box="[1185,1317,1287,1314]" italics="true" pageId="7" pageNumber="295">A. philippei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAEB266FFFF7E620FAE37D2BEF3F" bold="true" box="[1324,1387,1287,1313]" pageId="7" pageNumber="295">
|
||
<taxonomicNameLabel id="A24D5790266FFFF7E620FAE37D2BEF3F" box="[1324,1387,1287,1313]" pageId="7" pageNumber="295" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, the uropodal exopod and endopod are opaque white and colourless, respectively, whereas in
|
||
<taxonomicName id="4C0A4D7A266FFFF7E78DFACF7CBAEF5B" authorityName="Anker & Komai" authorityYear="2010" box="[1153,1274,1323,1349]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="ahyongi">
|
||
<emphasis id="B97EEAEB266FFFF7E78DFACF7CBAEF5B" box="[1153,1274,1323,1349]" italics="true" pageId="7" pageNumber="295">A. ahyongi</emphasis>
|
||
</taxonomicName>
|
||
, they are both red.
|
||
</paragraph>
|
||
<paragraph id="8BB536F9266FFFF8E3CBFA957B22EB99" blockId="7.[151,1437,151,2023]" lastBlockId="8.[151,1437,150,391]" lastPageId="8" lastPageNumber="296" pageId="7" pageNumber="295">
|
||
<taxonomicName id="4C0A4D7A266FFFF7E3CBFA9579D1EF95" authority="Anker & Ďuriš, 2022" authorityName="Anker & Ďuriš" authorityYear="2022" box="[199,401,1393,1419]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="philippei" status="sp. nov.">
|
||
<emphasis id="B97EEAEB266FFFF7E3CBFA9579D1EF95" box="[199,401,1393,1419]" italics="true" pageId="7" pageNumber="295">Athanas philippei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAEB266FFFF7E296FA96799BEF92" bold="true" box="[410,475,1394,1420]" pageId="7" pageNumber="295">
|
||
<taxonomicNameLabel id="A24D5790266FFFF7E296FA96799BEF92" box="[410,475,1394,1420]" pageId="7" pageNumber="295" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C0A4D7A266FFFF7E11BFA967AF3EF95" authorityName="Duris & Anker" authorityYear="2014" box="[535,691,1393,1420]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="manticolus">
|
||
<emphasis id="B97EEAEB266FFFF7E11BFA967AF3EF95" box="[535,691,1393,1420]" italics="true" pageId="7" pageNumber="295">A. manticolus</emphasis>
|
||
</taxonomicName>
|
||
share the presence of a minutely toothed rostrum, a feature also reported for the more distantly related
|
||
<taxonomicName id="4C0A4D7A266FFFF7E141FA717C65EFB1" authority="Anker, Jeng & Chan, 2001" authorityName="Anker, Jeng & Chan" authorityYear="2001" box="[589,1061,1429,1456]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="dentirostris">
|
||
<emphasis id="B97EEAEB266FFFF7E141FA717AADEFB1" box="[589,749,1429,1455]" italics="true" pageId="7" pageNumber="295">A. dentirostris</emphasis>
|
||
<bibRefCitation id="EF9B4B08266FFFF7E1F8FA717C65EFB1" author="Anker, A. & Jeng, M. - S. & Chan, T. - Y." box="[756,1061,1429,1456]" pageId="7" pageNumber="295" pagination="1049 - 1061" refId="ref6228" refString="Anker, A., Jeng, M. - S. & Chan, T. - Y. (2001) Two unusual species of Alpheidae (Decapoda: Caridea) associated with upogebiid mudshrimps in the mudflats of Taiwan and Vietnam. Journal of Crustacean Biology, 21, 1049 - 1061. https: // doi. org / 10.1163 / 20021975 - 99990197" type="journal article" year="2001">Anker, Jeng & Chan, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
, but generally highly unusual for alpheid shrimps (
|
||
<bibRefCitation id="EF9B4B08266FFFF7E251FA5C7A6FEFCD" author="Anker, A. & Jeng, M. - S. & Chan, T. - Y." box="[349,559,1464,1491]" pageId="7" pageNumber="295" pagination="1049 - 1061" refId="ref6228" refString="Anker, A., Jeng, M. - S. & Chan, T. - Y. (2001) Two unusual species of Alpheidae (Decapoda: Caridea) associated with upogebiid mudshrimps in the mudflats of Taiwan and Vietnam. Journal of Crustacean Biology, 21, 1049 - 1061. https: // doi. org / 10.1163 / 20021975 - 99990197" type="journal article" year="2001">
|
||
Anker
|
||
<emphasis id="B97EEAEB266FFFF7E2A3FA5D79AAEFCC" box="[431,490,1464,1490]" italics="true" pageId="7" pageNumber="295">et al.</emphasis>
|
||
2001
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9B4B08266FFFF7E131FA5E7B70EFCD" author="Duris, Z. & Anker, A." box="[573,816,1464,1493]" pageId="7" pageNumber="295" pagination="550 - 558" refId="ref7010" refString="Duris, Z. & Anker, A. (2014) Athanas manticolus sp. nov., a new stomatopod-associated alpheid shrimp from Vietnam (Crustacea, Decapoda). Zootaxa, 3784 (5), 550 - 558. https: // doi. org / 10.11646 / zootaxa. 3784.5.4" type="journal article" year="2014">Ďuriš & Anker 2014</bibRefCitation>
|
||
). However, the new species can be separated from
|
||
<taxonomicName id="4C0A4D7A266FFFF7E688FA5D7952EFE8" authorityName="Duris & Anker" authorityYear="2014" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="manticolus">
|
||
<emphasis id="B97EEAEB266FFFF7E688FA5D7952EFE8" italics="true" pageId="7" pageNumber="295">A. manticolus</emphasis>
|
||
</taxonomicName>
|
||
by the noticeably longer rostrum, reaching to the middle of the second article of the antennular peduncle (
|
||
<emphasis id="B97EEAEB266FFFF7E3ADF9E478F8EC07" box="[161,184,1536,1561]" italics="true" pageId="7" pageNumber="295">vs</emphasis>
|
||
. reaching only to the distal margin of the first article in
|
||
<taxonomicName id="4C0A4D7A266FFFF7E020F9E47B86EC07" authorityName="Duris & Anker" authorityYear="2014" box="[812,966,1535,1562]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="manticolus">
|
||
<emphasis id="B97EEAEB266FFFF7E020F9E47B86EC07" box="[812,966,1535,1562]" italics="true" pageId="7" pageNumber="295">A. manticolus</emphasis>
|
||
</taxonomicName>
|
||
); the extra-corneal teeth well-marked and acute (
|
||
<emphasis id="B97EEAEB266FFFF7E3EEF9C078B9EC23" box="[226,249,1572,1597]" italics="true" pageId="7" pageNumber="295">vs</emphasis>
|
||
. reduced to barely noticeable convexities in
|
||
<taxonomicName id="4C0A4D7A266FFFF7E1E0F9C77BC6EC23" authorityName="Duris & Anker" authorityYear="2014" box="[748,902,1571,1597]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="manticolus">
|
||
<emphasis id="B97EEAEB266FFFF7E1E0F9C77BC6EC23" box="[748,902,1571,1597]" italics="true" pageId="7" pageNumber="295">A. manticolus</emphasis>
|
||
</taxonomicName>
|
||
); the cheliped ischium armed with several teeth on the ventrolateral margin (
|
||
<emphasis id="B97EEAEB266FFFF7E2D3F9A379B6EC7E" box="[479,502,1607,1632]" italics="true" pageId="7" pageNumber="295">vs</emphasis>
|
||
. with one rather large triangular tooth in
|
||
<taxonomicName id="4C0A4D7A266FFFF7E0D8F9A37C30EC7E" authorityName="Duris & Anker" authorityYear="2014" box="[980,1136,1606,1633]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="manticolus">
|
||
<emphasis id="B97EEAEB266FFFF7E0D8F9A37C30EC7E" box="[980,1136,1606,1633]" italics="true" pageId="7" pageNumber="295">A. manticolus</emphasis>
|
||
</taxonomicName>
|
||
); the cheliped merus with some low subdistal teeth on ventrolateral and ventromesial margins (
|
||
<emphasis id="B97EEAEB266FFFF7E0ACF98F7BF7EC9A" box="[928,951,1643,1668]" italics="true" pageId="7" pageNumber="295">vs</emphasis>
|
||
. with small teeth along entire ventrolateral margin in
|
||
<taxonomicName id="4C0A4D7A266FFFF7E21CF96A79EDECB9" authorityName="Duris & Anker" authorityYear="2014" box="[272,429,1677,1704]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="manticolus">
|
||
<emphasis id="B97EEAEB266FFFF7E21CF96A79EDECB9" box="[272,429,1677,1704]" italics="true" pageId="7" pageNumber="295">A. manticolus</emphasis>
|
||
</taxonomicName>
|
||
); the cheliped carpus being much shorter, almost cup-shaped (
|
||
<emphasis id="B97EEAEB266FFFF7E789F96A7CDCECB9" box="[1157,1180,1678,1703]" italics="true" pageId="7" pageNumber="295">vs</emphasis>
|
||
. elongate, more vaseshaped in
|
||
<taxonomicName id="4C0A4D7A266FFFF7E201F95579EAECD5" authorityName="Duris & Anker" authorityYear="2014" box="[269,426,1713,1739]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="manticolus">
|
||
<emphasis id="B97EEAEB266FFFF7E201F95579EAECD5" box="[269,426,1713,1739]" italics="true" pageId="7" pageNumber="295">A. manticolus</emphasis>
|
||
</taxonomicName>
|
||
); the ventral surface of the cheliped palm with one to six relatively small tubercles (
|
||
<emphasis id="B97EEAEB266FFFF7E672F9567DD5ECD5" box="[1406,1429,1714,1739]" italics="true" pageId="7" pageNumber="295">vs</emphasis>
|
||
. with one fairly large tubercle in
|
||
<taxonomicName id="4C0A4D7A266FFFF7E101F9317AE9ECF0" authorityName="Duris & Anker" authorityYear="2014" box="[525,681,1748,1775]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="manticolus">
|
||
<emphasis id="B97EEAEB266FFFF7E101F9317AE9ECF0" box="[525,681,1748,1775]" italics="true" pageId="7" pageNumber="295">A. manticolus</emphasis>
|
||
</taxonomicName>
|
||
); the cheliped fingers armed with very stout teeth (
|
||
<emphasis id="B97EEAEB266FFFF7E7F7F9317D5AECF0" box="[1275,1306,1749,1774]" italics="true" pageId="7" pageNumber="295">vs.</emphasis>
|
||
with much weaker teeth in
|
||
<taxonomicName id="4C0A4D7A266FFFF7E246F91C79A5ED0C" authorityName="Duris & Anker" authorityYear="2014" box="[330,485,1784,1810]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="manticolus">
|
||
<emphasis id="B97EEAEB266FFFF7E246F91C79A5ED0C" box="[330,485,1784,1810]" italics="true" pageId="7" pageNumber="295">A. manticolus</emphasis>
|
||
</taxonomicName>
|
||
); the noticeably slenderer third pereiopod, especially in the length-width ratio of the ischium and merus (for instance, merus about 6.5 times as long as wide in
|
||
<taxonomicName id="4C0A4D7A266FFFF7E0C6F8F87C0CED2B" authority="Anker & Ďuriš, 2022" authorityName="Anker & Ďuriš" authorityYear="2022" box="[970,1100,1819,1846]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="philippei" status="sp. nov.">
|
||
<emphasis id="B97EEAEB266FFFF7E0C6F8F87C0CED2B" box="[970,1100,1819,1846]" italics="true" pageId="7" pageNumber="295">A. philippei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAEB266FFFF7E75DF8FF7CCFED2B" bold="true" box="[1105,1167,1819,1845]" pageId="7" pageNumber="295">
|
||
<taxonomicNameLabel id="A24D5790266FFFF7E75DF8FF7CCFED2B" box="[1105,1167,1819,1845]" pageId="7" pageNumber="295" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
<emphasis id="B97EEAEB266FFFF7E799F8F87CECED2B" box="[1173,1196,1820,1845]" italics="true" pageId="7" pageNumber="295">vs</emphasis>
|
||
. about 4.5 times in
|
||
<taxonomicName id="4C0A4D7A266FFFF7E688F8F87952ED47" authorityName="Duris & Anker" authorityYear="2014" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="manticolus">
|
||
<emphasis id="B97EEAEB266FFFF7E688F8F87952ED47" italics="true" pageId="7" pageNumber="295">A. manticolus</emphasis>
|
||
</taxonomicName>
|
||
); and the ischia of the fourth and fifth pereiopods armed with two and one cuspidate seta(e), respectively (
|
||
<emphasis id="B97EEAEB266FFFF7E3ADF88778F8ED62" box="[161,184,1891,1916]" italics="true" pageId="7" pageNumber="295">vs</emphasis>
|
||
. one and two, respectively, in
|
||
<taxonomicName id="4C0A4D7A266FFFF7E118F8867AF0ED62" authorityName="Duris & Anker" authorityYear="2014" box="[532,688,1890,1916]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="manticolus">
|
||
<emphasis id="B97EEAEB266FFFF7E118F8867AF0ED62" box="[532,688,1890,1916]" italics="true" pageId="7" pageNumber="295">A. manticolus</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<bibRefCitation id="EF9B4B08266FFFF7E1C5F8807BF9ED62" author="Duris, Z. & Anker, A." box="[713,953,1890,1919]" pageId="7" pageNumber="295" pagination="550 - 558" refId="ref7010" refString="Duris, Z. & Anker, A. (2014) Athanas manticolus sp. nov., a new stomatopod-associated alpheid shrimp from Vietnam (Crustacea, Decapoda). Zootaxa, 3784 (5), 550 - 558. https: // doi. org / 10.11646 / zootaxa. 3784.5.4" type="journal article" year="2014">Ďuriš & Anker 2014</bibRefCitation>
|
||
). The colour patterns of
|
||
<taxonomicName id="4C0A4D7A266FFFF7E7DCF8867D14ED62" authority="Anker & Ďuriš, 2022" authorityName="Anker & Ďuriš" authorityYear="2022" box="[1232,1364,1890,1916]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="philippei" status="sp. nov.">
|
||
<emphasis id="B97EEAEB266FFFF7E7DCF8867D14ED62" box="[1232,1364,1890,1916]" italics="true" pageId="7" pageNumber="295">A. philippei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAEB266FFFF7E650F8867DDCED62" bold="true" box="[1372,1436,1890,1916]" pageId="7" pageNumber="295">
|
||
<taxonomicNameLabel id="A24D5790266FFFF7E650F8867DDCED62" box="[1372,1436,1890,1916]" pageId="7" pageNumber="295" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C0A4D7A266FFFF7E3CBF8627922ED81" authorityName="Duris & Anker" authorityYear="2014" box="[199,354,1925,1952]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="manticolus">
|
||
<emphasis id="B97EEAEB266FFFF7E3CBF8627922ED81" box="[199,354,1925,1952]" italics="true" pageId="7" pageNumber="295">A. manticolus</emphasis>
|
||
</taxonomicName>
|
||
are similar, especially in the red chromatophores clustering in short, randomly oriented lines on the sides of the body, the presence of a broad white to pale yellow mid-dorsal band and the mostly white tail fan. However, the broader chromosome-like clusters of
|
||
<taxonomicName id="4C0A4D7A266FFFF7E1D5F8297B1CEDF8" authority="Anker & Ďuriš, 2022" authorityName="Anker & Ďuriš" authorityYear="2022" box="[729,860,1996,2023]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="philippei" status="sp. nov.">
|
||
<emphasis id="B97EEAEB266FFFF7E1D5F8297B1CEDF8" box="[729,860,1996,2023]" italics="true" pageId="7" pageNumber="295">A. philippei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAEB266FFFF7E06FF8297BE2EDF9" bold="true" box="[867,930,1997,2023]" pageId="7" pageNumber="295">
|
||
<taxonomicNameLabel id="A24D5790266FFFF7E06FF8297BE2EDF9" box="[867,930,1997,2023]" pageId="7" pageNumber="295" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are quite different from the narrower streaks of
|
||
<taxonomicName id="4C0A4D7A2660FFF8E3BAFF737911EAAF" authorityName="Duris & Anker" authorityYear="2014" box="[182,337,151,177]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="296" phylum="Arthropoda" rank="species" species="manticolus">
|
||
<emphasis id="B97EEAEB2660FFF8E3BAFF737911EAAF" box="[182,337,151,177]" italics="true" pageId="8" pageNumber="296">A. manticolus</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<figureCitation id="13312A7C2660FFF8E28BFF737996EAAF" box="[391,470,150,177]" captionStart="FIGURE 5" captionStartId="6.[151,250,1684,1708]" captionTargetBox="[185,1402,181,1660]" captionTargetId="figure-17@6.[185,1402,181,1660]" captionTargetPageId="6" captionText="FIGURE 5. Athanas philippei n. sp., paratype, female (cl 5.2 mm) from Pouébo, New Caledonia (MNHN-IU-2018-1017): (A) shrimp alive, dorsal view; (B) same, lateral view; (C) same, frontal view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.5965241" httpUri="https://zenodo.org/record/5965241/files/figure.png" pageId="8" pageNumber="296">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation id="13312A7C2660FFF8E2EFFF737A69EAAF" box="[483,553,151,177]" captionStart="FIGURE 6" captionStartId="8.[151,250,1865,1889]" captionTargetBox="[197,1390,415,1842]" captionTargetId="figure-134@8.[197,1390,415,1842]" captionTargetPageId="8" captionText="FIGURE 6. Athanas philippei n. sp. [A, B]: (A) holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018- 1124): shrimp alive, lateral view; (B) paratype, female (cl 7.2 mm) from Saint-Mathieu, New Caledonia (MNHN-IU-2019- 2986), shrimp alive, anterolateral view. Lysiosquillina maculata (Fabricius, 1793) (Lysiosquillidae) [C]: host (MNHN-IU-2018- 1087) of the female paratype of A. philippei n. sp. (MNHN-IU-2018-1017) from Pouébo, New Caledonia. Photographs by Z. Ďuriš." figureDoi="http://doi.org/10.5281/zenodo.5965243" httpUri="https://zenodo.org/record/5965243/files/figure.png" pageId="8" pageNumber="296">6A, B</figureCitation>
|
||
;
|
||
<bibRefCitation id="EF9B4B082660FFF8E13AFF7C7B68EAAF" author="Duris, Z. & Anker, A." box="[566,808,151,179]" pageId="8" pageNumber="296" pagination="550 - 558" refId="ref7010" refString="Duris, Z. & Anker, A. (2014) Athanas manticolus sp. nov., a new stomatopod-associated alpheid shrimp from Vietnam (Crustacea, Decapoda). Zootaxa, 3784 (5), 550 - 558. https: // doi. org / 10.11646 / zootaxa. 3784.5.4" type="journal article" year="2014">Ďuriš & Anker, 2014</bibRefCitation>
|
||
: fig. 6A–C). In addition, the hosts of
|
||
<taxonomicName id="4C0A4D7A2660FFF8E7DDFF737D15EAAF" authority="Anker & Ďuriš, 2022" authorityName="Anker & Ďuriš" authorityYear="2022" box="[1233,1365,151,177]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="296" phylum="Arthropoda" rank="species" species="philippei" status="sp. nov.">
|
||
<emphasis id="B97EEAEB2660FFF8E7DDFF737D15EAAF" box="[1233,1365,151,177]" italics="true" pageId="8" pageNumber="296">A. philippei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAEB2660FFF8E650FF737DDCEAAF" bold="true" box="[1372,1436,151,177]" pageId="8" pageNumber="296">
|
||
<taxonomicNameLabel id="A24D57902660FFF8E650FF737DDCEAAF" box="[1372,1436,151,177]" pageId="8" pageNumber="296" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C0A4D7A2660FFF8E3C5FF5F7926EACA" authorityName="Duris & Anker" authorityYear="2014" box="[201,358,186,213]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="296" phylum="Arthropoda" rank="species" species="manticolus">
|
||
<emphasis id="B97EEAEB2660FFF8E3C5FF5F7926EACA" box="[201,358,186,213]" italics="true" pageId="8" pageNumber="296">A. manticolus</emphasis>
|
||
</taxonomicName>
|
||
are two quite different mantis shrimps.
|
||
<taxonomicName id="4C0A4D7A2660FFF8E035FF5E7D53EACB" baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[825,1299,186,213]" class="Malacostraca" family="Lysiosquillidae" genus="Lysiosquillina" kingdom="Animalia" order="Stomatopoda" pageId="8" pageNumber="296" phylum="Arthropoda" rank="species" species="maculata">
|
||
<emphasis id="B97EEAEB2660FFF8E035FF5E7C08EACA" box="[825,1096,186,212]" italics="true" pageId="8" pageNumber="296">Lysiosquillina maculata</emphasis>
|
||
(Lysiosquillidae)
|
||
</taxonomicName>
|
||
, the host of
|
||
<taxonomicName id="4C0A4D7A2660FFF8E39BFF3A795CEAE6" authority="Anker & Ďuriš, 2022" authorityName="Anker & Ďuriš" authorityYear="2022" box="[151,284,222,248]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="296" phylum="Arthropoda" rank="species" species="philippei" status="sp. nov.">
|
||
<emphasis id="B97EEAEB2660FFF8E39BFF3A795CEAE6" box="[151,284,222,248]" italics="true" pageId="8" pageNumber="296">A. philippei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAEB2660FFF8E228FF3A7925EAE6" bold="true" box="[292,357,222,248]" pageId="8" pageNumber="296">
|
||
<taxonomicNameLabel id="A24D57902660FFF8E228FF3A7925EAE6" box="[292,357,222,248]" pageId="8" pageNumber="296" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13312A7C2660FFF8E27AFF3A7990EAE6" box="[374,464,221,248]" captionStart="FIGURE 6" captionStartId="8.[151,250,1865,1889]" captionTargetBox="[197,1390,415,1842]" captionTargetId="figure-134@8.[197,1390,415,1842]" captionTargetPageId="8" captionText="FIGURE 6. Athanas philippei n. sp. [A, B]: (A) holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018- 1124): shrimp alive, lateral view; (B) paratype, female (cl 7.2 mm) from Saint-Mathieu, New Caledonia (MNHN-IU-2019- 2986), shrimp alive, anterolateral view. Lysiosquillina maculata (Fabricius, 1793) (Lysiosquillidae) [C]: host (MNHN-IU-2018- 1087) of the female paratype of A. philippei n. sp. (MNHN-IU-2018-1017) from Pouébo, New Caledonia. Photographs by Z. Ďuriš." figureDoi="http://doi.org/10.5281/zenodo.5965243" httpUri="https://zenodo.org/record/5965243/files/figure.png" pageId="8" pageNumber="296">Fig. 6C</figureCitation>
|
||
), is a very large and powerful stomatopod popularly known as “tiger mantis shrimp” (maximal total length:
|
||
<quantity id="4CF29B1C2660FFF8E294FEE579A1EB02" box="[408,481,257,284]" metricMagnitude="-1" metricUnit="m" metricValue="3.8" pageId="8" pageNumber="296" unit="cm" value="38.0">38 cm</quantity>
|
||
), which excavates large, spacious, U-shaped burrows (
|
||
<bibRefCitation id="EF9B4B082660FFF8E74BFEE57CB0EB05" author="Manning, R. V." box="[1095,1264,257,283]" pageId="8" pageNumber="296" refId="ref7271" refString="Manning, R. V. (1998) Stomatopods. In: Carpenter, K. E. & Niem, V. H. (Eds.), FAO species identification guide for fishery purposes. The living marine resources of the western central Pacific. Vol. 2. Cephalopods, crustaceans, holothurians and sharks. FAO, Rome, pp. 827 - 849." type="book" year="1998">Manning 1998</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9B4B082660FFF8E7F7FEE57DD7EB05" author="Ahyong, S. T." box="[1275,1431,257,283]" pageId="8" pageNumber="296" pagination="1 - 326" refId="ref5804" refString="Ahyong, S. T. (2001) Revision of the Australian stomatopod Crustacea. Records of the Australian Museum, Supplement 26, 1 - 326. https: // doi. org / 10.3853 / j. 0812 - 7387.26.2001.1333" type="journal article" year="2001">Ahyong 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9B4B082660FFF8E39BFEC07939EB21" author="Ahyong, S. T. & Caldwell, R. L. & Erdmann, M. V." box="[151,377,292,319]" pageId="8" pageNumber="296" pagination="109 - 114" refId="ref5847" refString="Ahyong, S. T., Caldwell, R. L. & Erdmann, M. V. (2017) Collecting and processing stomatopods. Journal of Crustacean Biology, 37, 109 - 114. https: // doi. org / 10.1093 / jcbiol / ruw 005" type="journal article" year="2017">
|
||
Ahyong
|
||
<emphasis id="B97EEAEB2660FFF8E3F7FEC17975EB20" box="[251,309,292,318]" italics="true" pageId="8" pageNumber="296">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
; A. Anker, pers. obs.). In contrast,
|
||
<taxonomicName id="4C0A4D7A2660FFF8E002FEC07CB3EB21" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[782,1267,292,319]" class="Malacostraca" family="Nannosquillidae" genus="Bigelowina" kingdom="Animalia" order="Stomatopoda" pageId="8" pageNumber="296" phylum="Arthropoda" rank="species" species="phalangium">
|
||
<emphasis id="B97EEAEB2660FFF8E002FEC07C5DEB20" box="[782,1053,292,318]" italics="true" pageId="8" pageNumber="296">Bigelowina phalangium</emphasis>
|
||
(Nannosquillidae)
|
||
</taxonomicName>
|
||
, the host of
|
||
<taxonomicName id="4C0A4D7A2660FFF8E688FEC17952EB7C" authorityName="Duris & Anker" authorityYear="2014" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="296" phylum="Arthropoda" rank="species" species="manticolus">
|
||
<emphasis id="B97EEAEB2660FFF8E688FEC17952EB7C" italics="true" pageId="8" pageNumber="296">A. manticolus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF9B4B082660FFF8E22CFEAE7A4AEB7D" author="Duris, Z. & Anker, A." box="[288,522,328,357]" pageId="8" pageNumber="296" pagination="550 - 558" refId="ref7010" refString="Duris, Z. & Anker, A. (2014) Athanas manticolus sp. nov., a new stomatopod-associated alpheid shrimp from Vietnam (Crustacea, Decapoda). Zootaxa, 3784 (5), 550 - 558. https: // doi. org / 10.11646 / zootaxa. 3784.5.4" type="journal article" year="2014">Ďuriš & Anker 2014</bibRefCitation>
|
||
: fig. 6D), is a significantly smaller stomatopod (maximal total length: ca.
|
||
<quantity id="4CF29B1C2660FFF8E64CFEAD7DD0EB7D" box="[1344,1424,329,355]" metricMagnitude="-2" metricUnit="m" metricValue="9.4" pageId="8" pageNumber="296" unit="cm" value="9.4">9.4 cm</quantity>
|
||
), which lives in much smaller U-shaped burrows (
|
||
<bibRefCitation id="EF9B4B082660FFF8E1BBFE887B16EB99" author="Ahyong, S. T." box="[695,854,364,391]" pageId="8" pageNumber="296" pagination="1 - 326" refId="ref5804" refString="Ahyong, S. T. (2001) Revision of the Australian stomatopod Crustacea. Records of the Australian Museum, Supplement 26, 1 - 326. https: // doi. org / 10.3853 / j. 0812 - 7387.26.2001.1333" type="journal article" year="2001">Ahyong 2001</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF7566712660FFF8E39BF8AD7897EDED" ID-DOI="http://doi.org/10.5281/zenodo.5965243" ID-Zenodo-Dep="5965243" httpUri="https://zenodo.org/record/5965243/files/figure.png" pageId="8" pageNumber="296" startId="8.[151,250,1865,1889]" targetBox="[197,1390,415,1842]" targetPageId="8">
|
||
<paragraph id="8BB536F92660FFF8E39BF8AD7897EDED" blockId="8.[151,1437,1865,2035]" pageId="8" pageNumber="296">
|
||
<emphasis id="B97EEAEB2660FFF8E39BF8AD794EED7F" bold="true" box="[151,270,1865,1889]" pageId="8" pageNumber="296">FIGURE 6</emphasis>
|
||
.
|
||
<taxonomicName id="4C0A4D7A2660FFF8E217F8AD798FED7F" authority="Anker & Ďuriš, 2022" authorityName="Anker & Ďuriš" authorityYear="2022" box="[283,463,1865,1889]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="296" phylum="Arthropoda" rank="species" species="philippei" status="sp. nov.">
|
||
<emphasis id="B97EEAEB2660FFF8E217F8AD798FED7F" box="[283,463,1865,1889]" italics="true" pageId="8" pageNumber="296">Athanas philippei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAEB2660FFF8E2DBF8AD7A51ED7F" bold="true" box="[471,529,1865,1889]" pageId="8" pageNumber="296">
|
||
<taxonomicNameLabel id="A24D57902660FFF8E2DBF8AD7A51ED7F" box="[471,529,1865,1889]" pageId="8" pageNumber="296" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
[A, B]: (A) holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018- 1124): shrimp alive, lateral view; (B) paratype, female (cl 7.2 mm) from Saint-Mathieu, New Caledonia (MNHN-IU-2019- 2986), shrimp alive, anterolateral view.
|
||
<taxonomicName id="4C0A4D7A2660FFF8E12AF8757C3DEDB7" authority="(Fabricius, 1793)" baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[550,1149,1937,1961]" class="Malacostraca" family="Lysiosquillidae" genus="Lysiosquillina" kingdom="Animalia" order="Stomatopoda" pageId="8" pageNumber="296" phylum="Arthropoda" rank="species" species="maculata">
|
||
<emphasis id="B97EEAEB2660FFF8E12AF8757B58EDB7" box="[550,792,1937,1961]" italics="true" pageId="8" pageNumber="296">Lysiosquillina maculata</emphasis>
|
||
(Fabricius, 1793) (Lysiosquillidae)
|
||
</taxonomicName>
|
||
[C]: host (MNHN-IU-2018- 1087) of the female paratype of
|
||
<taxonomicName id="4C0A4D7A2660FFF8E2EFF8517A1AEDD3" authority="Anker & Ďuriš, 2022" authorityName="Anker & Ďuriš" authorityYear="2022" box="[483,602,1973,1997]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="296" phylum="Arthropoda" rank="species" species="philippei" status="sp. nov.">
|
||
<emphasis id="B97EEAEB2660FFF8E2EFF8517A1AEDD3" box="[483,602,1973,1997]" italics="true" pageId="8" pageNumber="296">A. philippei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAEB2660FFF8E16EF8517ADCEDD3" bold="true" box="[610,668,1973,1997]" pageId="8" pageNumber="296">
|
||
<taxonomicNameLabel id="A24D57902660FFF8E16EF8517ADCEDD3" box="[610,668,1973,1997]" pageId="8" pageNumber="296" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(MNHN-IU-2018-1017) from Pouébo, New Caledonia. Photographs by Z. Ďuriš.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7566712661FFF9E39BF9AB7D4FECB1" ID-DOI="http://doi.org/10.5281/zenodo.5965247" ID-Zenodo-Dep="5965247" httpUri="https://zenodo.org/record/5965247/files/figure.png" pageId="9" pageNumber="297" startId="9.[151,250,1615,1639]" targetBox="[151,1435,181,1591]" targetPageId="9">
|
||
<paragraph id="8BB536F92661FFF9E39BF9AB7D4FECB1" blockId="9.[151,1437,1615,1713]" pageId="9" pageNumber="297">
|
||
<emphasis id="B97EEAEB2661FFF9E39BF9AB794CEC79" bold="true" box="[151,268,1615,1639]" pageId="9" pageNumber="297">FIGURE 7</emphasis>
|
||
. One of the collection localities of
|
||
<taxonomicName id="4C0A4D7A2661FFF9E160F9AB7B5EEC79" authority="Anker & Ďuriš, 2022" authorityName="Anker & Ďuriš" authorityYear="2022" box="[620,798,1615,1639]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="297" phylum="Arthropoda" rank="species" species="philippei" status="sp. nov.">
|
||
<emphasis id="B97EEAEB2661FFF9E160F9AB7B5EEC79" box="[620,798,1615,1639]" italics="true" pageId="9" pageNumber="297">Athanas philippei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAEB2661FFF9E028F9AB7B1CEC79" bold="true" box="[804,860,1615,1639]" pageId="9" pageNumber="297">
|
||
<taxonomicNameLabel id="A24D57902661FFF9E028F9AB7B1CEC79" box="[804,860,1615,1639]" pageId="9" pageNumber="297" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, near Pouébo, north-eastern coast of New Caledonia: (A) view from the beach, with black arrow approximately indicating the area of the collection of the first specimen (female paratype, MNHN-IU-2018-1017), (B) view of the habitat during sampling. Photographs by A. Anker (A) and Z. Ďuriš (B).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB536F92661FFFAE3CBF93A7D28EACB" blockId="9.[151,1437,1757,2036]" lastBlockId="10.[151,1437,151,969]" lastPageId="10" lastPageNumber="298" pageId="9" pageNumber="297">
|
||
Two other species of
|
||
<taxonomicName id="4C0A4D7A2661FFF9E2B4F9397A54ECE9" authorityName="Leach" authorityYear="1814" box="[440,532,1757,1783]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="297" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97EEAEB2661FFF9E2B4F9397A54ECE9" box="[440,532,1757,1783]" italics="true" pageId="9" pageNumber="297">Athanas</emphasis>
|
||
</taxonomicName>
|
||
, viz.
|
||
<taxonomicName id="4C0A4D7A2661FFF9E15FF93A7C46ECE9" authority="Anker & Ahyong, 2007" authorityName="Anker & Ahyong" authorityYear="2007" box="[595,1030,1757,1784]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="297" phylum="Arthropoda" rank="species" species="sydneyensis">
|
||
<emphasis id="B97EEAEB2661FFF9E15FF93A7AB6ECE9" box="[595,758,1757,1784]" italics="true" pageId="9" pageNumber="297">A. sydneyensis</emphasis>
|
||
<bibRefCitation id="EF9B4B082661FFF9E1F0F9397C46ECE9" author="Anker, A. & Ahyong, S. T." box="[764,1030,1757,1783]" pageId="9" pageNumber="297" pagination="17 - 30" refId="ref6098" refString="Anker, A. & Ahyong, S. T. (2007) Description of two species in the alpheid shrimp genus Athanas Leach, 1814, with remarks on A. amazone Holthuis, 1951 (Decapoda, Caridea). Zootaxa, 1563 (1), 17 - 30. https: // doi. org / 10.11646 / zootaxa. 1563.1.2" type="journal article" year="2007">Anker & Ahyong, 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
from south-eastern
|
||
<collectingCountry id="F31D76692661FFF9E7EBF9397D0EECE9" box="[1255,1358,1757,1783]" name="Australia" pageId="9" pageNumber="297">Australia</collectingCountry>
|
||
and
|
||
<taxonomicName id="4C0A4D7A2661FFF9E688F93A7A52ED05" authority="Anker & Ahyong, 2007" authorityName="Anker & Ahyong" authorityYear="2007" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="297" phylum="Arthropoda" rank="species" species="ivoiriensis">
|
||
<emphasis id="B97EEAEB2661FFF9E688F93A794CED05" italics="true" pageId="9" pageNumber="297">A. ivoiriensis</emphasis>
|
||
<bibRefCitation id="EF9B4B082661FFF9E203F8E57A52ED05" author="Anker, A. & Ahyong, S. T." box="[271,530,1793,1819]" pageId="9" pageNumber="297" pagination="17 - 30" refId="ref6098" refString="Anker, A. & Ahyong, S. T. (2007) Description of two species in the alpheid shrimp genus Athanas Leach, 1814, with remarks on A. amazone Holthuis, 1951 (Decapoda, Caridea). Zootaxa, 1563 (1), 17 - 30. https: // doi. org / 10.11646 / zootaxa. 1563.1.2" type="journal article" year="2007">Anker & Ahyong, 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
from western Africa, share several morphological features with
|
||
<taxonomicName id="4C0A4D7A2661FFF9E7DEF8E67D13ED05" authority="Anker & Ďuriš, 2022" authorityName="Anker & Ďuriš" authorityYear="2022" box="[1234,1363,1793,1820]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="297" phylum="Arthropoda" rank="species" species="philippei" status="sp. nov.">
|
||
<emphasis id="B97EEAEB2661FFF9E7DEF8E67D13ED05" box="[1234,1363,1793,1820]" italics="true" pageId="9" pageNumber="297">A. philippei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAEB2661FFF9E654F8E57DD5ED05" bold="true" box="[1368,1429,1793,1819]" pageId="9" pageNumber="297">
|
||
<taxonomicNameLabel id="A24D57902661FFF9E654F8E57DD5ED05" box="[1368,1429,1793,1819]" pageId="9" pageNumber="297" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, such as the general configuration of the frontal margin, the degree of symmetry and general shape of the chelipeds, and the proportions and armature of the third to fifth pereiopods (
|
||
<bibRefCitation id="EF9B4B082661FFF9E078F8AD7C3DED7D" author="Anker, A. & Ahyong, S. T." box="[884,1149,1865,1891]" pageId="9" pageNumber="297" pagination="17 - 30" refId="ref6098" refString="Anker, A. & Ahyong, S. T. (2007) Description of two species in the alpheid shrimp genus Athanas Leach, 1814, with remarks on A. amazone Holthuis, 1951 (Decapoda, Caridea). Zootaxa, 1563 (1), 17 - 30. https: // doi. org / 10.11646 / zootaxa. 1563.1.2" type="journal article" year="2007">Anker & Ahyong 2007</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C0A4D7A2661FFF9E783F8AD7D16ED7D" authority="Anker & Ďuriš, 2022" authorityName="Anker & Ďuriš" authorityYear="2022" box="[1167,1366,1865,1891]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="297" phylum="Arthropoda" rank="species" species="philippei" status="sp. nov.">
|
||
<emphasis id="B97EEAEB2661FFF9E783F8AD7D16ED7D" box="[1167,1366,1865,1891]" italics="true" pageId="9" pageNumber="297">Athanas philippei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAEB2661FFF9E651F8AD7DDCED7D" bold="true" box="[1373,1436,1865,1891]" pageId="9" pageNumber="297">
|
||
<taxonomicNameLabel id="A24D57902661FFF9E651F8AD7DDCED7D" box="[1373,1436,1865,1891]" pageId="9" pageNumber="297" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
may be easily distinguished from both of them by the dorsally minutely toothed rostrum; the presence of a distinct post-rostral tubercle; and the reduced armature of the cheliped merus (which is armed with more or less strong teeth on the ventrolateral margin in the other two species).
|
||
<taxonomicName id="4C0A4D7A2661FFF9E1E7F8517BF1EDD1" authority="Anker & Ďuriš, 2022" authorityName="Anker & Ďuriš" authorityYear="2022" box="[747,945,1973,1999]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="297" phylum="Arthropoda" rank="species" species="philippei" status="sp. nov.">
|
||
<emphasis id="B97EEAEB2661FFF9E1E7F8517BF1EDD1" box="[747,945,1973,1999]" italics="true" pageId="9" pageNumber="297">Athanas philippei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAEB2661FFF9E0BBF8517BB6EDD1" bold="true" box="[951,1014,1973,1999]" pageId="9" pageNumber="297">
|
||
<taxonomicNameLabel id="A24D57902661FFF9E0BBF8517BB6EDD1" box="[951,1014,1973,1999]" pageId="9" pageNumber="297" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be also distinguished specifically from
|
||
<taxonomicName id="4C0A4D7A2661FFF9E3D8F83E7936EDED" authorityName="Anker & Ahyong" authorityYear="2007" box="[212,374,2009,2036]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="297" phylum="Arthropoda" rank="species" species="sydneyensis">
|
||
<emphasis id="B97EEAEB2661FFF9E3D8F83E7936EDED" box="[212,374,2009,2036]" italics="true" pageId="9" pageNumber="297">A. sydneyensis</emphasis>
|
||
</taxonomicName>
|
||
by the presence of tubercles on the ventromesial surface of the palm (
|
||
<emphasis id="B97EEAEB2661FFF9E771F83E7CD4EDED" box="[1149,1172,2010,2035]" italics="true" pageId="9" pageNumber="297">vs</emphasis>
|
||
. without tubercles in
|
||
<taxonomicName id="4C0A4D7A2661FFFAE688F83E795BEAAF" authorityName="Anker & Ahyong" authorityYear="2007" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="10" lastPageNumber="298" order="Decapoda" pageId="9" pageNumber="297" phylum="Arthropoda" rank="species" species="sydneyensis">
|
||
<emphasis id="B97EEAEB2661FFFAE688F83E795BEAAF" italics="true" lastPageId="10" lastPageNumber="298" pageId="9" pageNumber="297">A. sydneyensis</emphasis>
|
||
</taxonomicName>
|
||
); and from
|
||
<taxonomicName id="4C0A4D7A2662FFFAE293FF737A72EAAE" authorityName="Anker & Ahyong" authorityYear="2007" box="[415,562,151,177]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="298" phylum="Arthropoda" rank="species" species="ivoiriensis">
|
||
<emphasis id="B97EEAEB2662FFFAE293FF737A72EAAE" box="[415,562,151,177]" italics="true" pageId="10" pageNumber="298">A. ivoiriensis</emphasis>
|
||
</taxonomicName>
|
||
by the ventrally unarmed carpus (
|
||
<emphasis id="B97EEAEB2662FFFAE0A3FF737B86EAAE" box="[943,966,151,176]" italics="true" pageId="10" pageNumber="298">vs</emphasis>
|
||
. with proximal teeth in
|
||
<taxonomicName id="4C0A4D7A2662FFFAE7DDFF737D24EAAE" authorityName="Anker & Ahyong" authorityYear="2007" box="[1233,1380,151,177]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="298" phylum="Arthropoda" rank="species" species="ivoiriensis">
|
||
<emphasis id="B97EEAEB2662FFFAE7DDFF737D24EAAE" box="[1233,1380,151,177]" italics="true" pageId="10" pageNumber="298">A. ivoiriensis</emphasis>
|
||
</taxonomicName>
|
||
) and finger cutting edges with a strong dentition (
|
||
<emphasis id="B97EEAEB2662FFFAE186FF5F7AE1EACA" box="[650,673,187,212]" italics="true" pageId="10" pageNumber="298">vs</emphasis>
|
||
. almost unarmed in
|
||
<taxonomicName id="4C0A4D7A2662FFFAE08AFF5F7C5AEACA" authorityName="Anker & Ahyong" authorityYear="2007" box="[902,1050,187,213]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="298" phylum="Arthropoda" rank="species" species="ivoiriensis">
|
||
<emphasis id="B97EEAEB2662FFFAE08AFF5F7C5AEACA" box="[902,1050,187,213]" italics="true" pageId="10" pageNumber="298">A. ivoiriensis</emphasis>
|
||
</taxonomicName>
|
||
) (cf.
|
||
<bibRefCitation id="EF9B4B082662FFFAE75AFF5F7D1CEACB" author="Anker, A. & Ahyong, S. T." box="[1110,1372,187,213]" pageId="10" pageNumber="298" pagination="17 - 30" refId="ref6098" refString="Anker, A. & Ahyong, S. T. (2007) Description of two species in the alpheid shrimp genus Athanas Leach, 1814, with remarks on A. amazone Holthuis, 1951 (Decapoda, Caridea). Zootaxa, 1563 (1), 17 - 30. https: // doi. org / 10.11646 / zootaxa. 1563.1.2" type="journal article" year="2007">Anker & Ahyong 2007</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB536F92662FFFAE3CBFF3B795FEBB0" blockId="10.[151,1437,151,969]" pageId="10" pageNumber="298">
|
||
The presence of a small spinule mesial to the distolateral spiniform seta of the uropod in
|
||
<specimenCount id="9D0CFD702662FFFAE7BBFF3B7D1EEAE7" box="[1207,1374,223,249]" pageId="10" pageNumber="298" type="generic">two specimens</specimenCount>
|
||
of
|
||
<taxonomicName id="4C0A4D7A2662FFFAE688FF3B78BBEB03" authority="Anker & Ďuriš, 2022" authorityName="Anker & Ďuriš" authorityYear="2022" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="298" phylum="Arthropoda" rank="species" species="philippei" status="sp. nov.">
|
||
<emphasis id="B97EEAEB2662FFFAE688FF3B78BBEB03" italics="true" pageId="10" pageNumber="298">A. philippei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAEB2662FFFAE20DFEE7797FEB03" bold="true" box="[257,319,259,285]" pageId="10" pageNumber="298">
|
||
<taxonomicNameLabel id="A24D57902662FFFAE20DFEE7797FEB03" box="[257,319,259,285]" pageId="10" pageNumber="298" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13312A7C2662FFFAE241FEE77985EB03" box="[333,453,259,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1713,1737]" captionTargetBox="[176,1411,211,1650]" captionTargetId="figure-31@3.[169,1418,181,1689]" captionTargetPageId="3" captionText="FIGURE 2. Athanas philippei n. sp., holotype, female (cl 6.9 mm) from Tiari, New Caledonia (MNHN-IU-2018-1124): (A) rostrum and left eye and orbital teeth, lateral view; (B) anterior region, lateral view; (C) same, dorsal view; (D) telson and uropods, dorsal view; (E) telson, posterior half, dorsal view; (F) third maxilliped, lateral view; (G) left uropod, detail of distolateral region, dorsal view. All scales as indicated, except for G drawn from a microscope photograph without scale." figureDoi="http://doi.org/10.5281/zenodo.5965227" httpUri="https://zenodo.org/record/5965227/files/figure.png" pageId="10" pageNumber="298">Fig. 2D, H</figureCitation>
|
||
) is an interesting and possibly novel morphological feature within
|
||
<taxonomicName id="4C0A4D7A2662FFFAE7A7FEE77D47EB03" authorityName="Leach" authorityYear="1814" box="[1195,1287,259,285]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="298" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97EEAEB2662FFFAE7A7FEE77D47EB03" box="[1195,1287,259,285]" italics="true" pageId="10" pageNumber="298">Athanas</emphasis>
|
||
</taxonomicName>
|
||
. This spinule is absent or at least has not been illustrated in
|
||
<taxonomicName id="4C0A4D7A2662FFFAE19CFEC37B49EB5F" authorityName="Anker & Komai" authorityYear="2010" box="[656,777,295,321]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="298" phylum="Arthropoda" rank="species" species="ahyongi">
|
||
<emphasis id="B97EEAEB2662FFFAE19CFEC37B49EB5F" box="[656,777,295,321]" italics="true" pageId="10" pageNumber="298">A. ahyongi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0A4D7A2662FFFAE01AFEC37BEFEB5F" authorityName="Duris & Anker" authorityYear="2014" box="[790,943,295,321]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="298" phylum="Arthropoda" rank="species" species="manticolus">
|
||
<emphasis id="B97EEAEB2662FFFAE01AFEC37BEFEB5F" box="[790,943,295,321]" italics="true" pageId="10" pageNumber="298">A. manticolus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0A4D7A2662FFFAE0B0FEC37C1EEB5F" authorityName="Anker & Ahyong" authorityYear="2007" box="[956,1118,295,321]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="298" phylum="Arthropoda" rank="species" species="sydneyensis">
|
||
<emphasis id="B97EEAEB2662FFFAE0B0FEC37C1EEB5F" box="[956,1118,295,321]" italics="true" pageId="10" pageNumber="298">A. sydneyensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0A4D7A2662FFFAE79FFEC37D66EB5E" authorityName="Anker & Ahyong" authorityYear="2007" box="[1171,1318,295,321]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="298" phylum="Arthropoda" rank="species" species="ivoiriensis">
|
||
<emphasis id="B97EEAEB2662FFFAE79FFEC37D66EB5E" box="[1171,1318,295,321]" italics="true" pageId="10" pageNumber="298">A. ivoiriensis</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<bibRefCitation id="EF9B4B082662FFFAE65AFEC37910EB7B" author="Anker, A. & Ahyong, S. T." pageId="10" pageNumber="298" pagination="17 - 30" refId="ref6098" refString="Anker, A. & Ahyong, S. T. (2007) Description of two species in the alpheid shrimp genus Athanas Leach, 1814, with remarks on A. amazone Holthuis, 1951 (Decapoda, Caridea). Zootaxa, 1563 (1), 17 - 30. https: // doi. org / 10.11646 / zootaxa. 1563.1.2" type="journal article" year="2007">Anker & Ahyong 2007</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9B4B082662FFFAE256FEAF7A15EB7B" author="Anker, A. & Komai, T." box="[346,597,331,357]" pageId="10" pageNumber="298" pagination="45 - 54" refId="ref6296" refString="Anker, A. & Komai, T. (2010) Description of a new species of Athanas Leach, 1814 (Crustacea: Decapoda: Alpheidae) from Madagascar and Japan. Zootaxa, 2680 (1), 45 - 54. https: // doi. org / 10.11646 / zootaxa. 2680.1.4" type="journal article" year="2010">Anker & Komai 2010</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9B4B082662FFFAE16CFEA87B0BEB7B" author="Duris, Z. & Anker, A." box="[608,843,331,359]" pageId="10" pageNumber="298" pagination="550 - 558" refId="ref7010" refString="Duris, Z. & Anker, A. (2014) Athanas manticolus sp. nov., a new stomatopod-associated alpheid shrimp from Vietnam (Crustacea, Decapoda). Zootaxa, 3784 (5), 550 - 558. https: // doi. org / 10.11646 / zootaxa. 3784.5.4" type="journal article" year="2014">Ďuriš & Anker 2014</bibRefCitation>
|
||
), representing an additional distinguishing feature of the new species, albeit a somewhat inconsistent one, since it is absent on one or both uropods in some
|
||
<typeStatus id="54B1885B2662FFFAE63DFE8B7DDCEB97" box="[1329,1436,367,393]" pageId="10" pageNumber="298" type="paratype">paratypes</typeStatus>
|
||
(see above).
|
||
</paragraph>
|
||
<paragraph id="8BB536F92662FFFAE3CBFE537A5FE87F" blockId="10.[151,1437,151,969]" pageId="10" pageNumber="298">
|
||
None of the remaining species of
|
||
<taxonomicName id="4C0A4D7A2662FFFAE15AFE537AF2EBCF" authorityName="Leach" authorityYear="1814" box="[598,690,439,465]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="298" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97EEAEB2662FFFAE15AFE537AF2EBCF" box="[598,690,439,465]" italics="true" pageId="10" pageNumber="298">Athanas</emphasis>
|
||
</taxonomicName>
|
||
appears to have close affinities to the present new species (e.g.
|
||
<bibRefCitation id="EF9B4B082662FFFAE39BFE3F7979EBEB" author="Coutiere, H." box="[151,313,475,501]" pageId="10" pageNumber="298" pagination="72 - 90" refId="ref6884" refString="Coutiere, H. (1903) Note sur quelques Alpheidae des Maldives et Laquedives. Bulletin de la Societe Philomathique de Paris, 9 (5), 72 - 90." type="journal article" year="1903">Coutière 1903</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9B4B082662FFFAE249FE3F79A2EBEB" author="De Man, J. G." box="[325,482,475,501]" pageId="10" pageNumber="298" pagination="287 - 319" refId="ref6969" refString="De Man, J. G. (1910) Diagnoses of new species of macrurous decapod Crustacea from the " Siboga-Expedition ", V. Tijdschrift der Nederlandsche Dierkundige Vereeniging, 2 (11), 287 - 319." type="journal article" year="1910">De Man 1910</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9B4B082662FFFAE2E2FE3F7AD0EBEB" author="Holthuis, L. B." box="[494,656,474,501]" pageId="10" pageNumber="298" pagination="7 - 187" refId="ref7168" refString="Holthuis, L. B. (1951) The caridean Crustacea of tropical West Africa. Atlantide Report Scientific Results of the Danish Expedition to the Coasts of Tropical West Africa 1945 - 1946, 2, 7 - 187." type="journal article" year="1951">Holthuis 1951</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9B4B082662FFFAE190FE3F7B63EBEB" author="Chace, F. A. Jr." box="[668,803,474,501]" pageId="10" pageNumber="298" pagination="1 - 22" refId="ref6779" refString="Chace, F. A. Jr. (1955) Notes on shrimps from the Marshall lslands. Proceedings of the United States National Museum, 105, 1 - 22. https: // doi. org / 10.5479 / si. 00963801.105 - 3349.1" type="journal article" year="1955">Chace 1955</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9B4B082662FFFAE023FE3F7C76EBEB" author="Banner, A. H. & Banner, D. M." box="[815,1078,475,501]" pageId="10" pageNumber="298" pagination="129 - 155" refId="ref6463" refString="Banner, A. H. & Banner, D. M. (1960) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean. Part V. The Indo-Pacific members of the genus Athanas. Pacific Science, 14, 129 - 155." type="journal article" year="1960">Banner & Banner 1960</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9B4B082662FFFAE74DFE3F7C3CEBEB" author="Banner, A. H. & Banner, D. M." box="[1089,1148,475,501]" pageId="10" pageNumber="298" pagination="1 - 168" refId="ref6512" refString="Banner, A. H. & Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">1966</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9B4B082662FFFAE78BFE3F7C82EBEB" author="Banner, D. M. & Banner, A. H." box="[1159,1218,475,501]" pageId="10" pageNumber="298" pagination="291 - 382" refId="ref6587" refString="Banner, D. M. & Banner, A. H. (1973) The alpheid shrimp of Australia. Part I: The lower genera. Records of the Australian Museum, 28, 291 - 382. https: // doi. org / 10.3853 / j. 0067 - 1975.28.1973.407" type="journal article" year="1973">1973</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9B4B082662FFFAE7C1FE3F7D48EBEB" author="Banner, D. M. & Banner, A. H." box="[1229,1288,475,501]" pageId="10" pageNumber="298" pagination="215 - 257" refId="ref6642" refString="Banner, D. M. & Banner, A. H. (1978) Annotated checklist of alpheid and ogyridid shrimp from the Philippine archipelago and the South China Sea. Micronesica, 14, 215 - 257." type="journal article" year="1978">1978</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9B4B082662FFFAE61FFE3F7D0EEBEB" author="Banner, A. H. & Banner, D. M." box="[1299,1358,475,501]" pageId="10" pageNumber="298" pagination="1 - 164" refId="ref6546" refString="Banner, A. H. & Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">1983</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9B4B082662FFFAE655FE3F7892E807" author="Bruce, A." pageId="10" pageNumber="298" refId="ref6711" refString="Bruce, A. J (1990) Additions to the marine shrimp fauna of Hong Kong. In: Morton, B. (Ed.), Proceedings of the Second International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China. Vol. 2. Hong Kong University Press, Hong Kong, pp. 611 - 648." type="book" year="1990">Bruce 1990</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9B4B082662FFFAE3D7FE1B7922E807" author="Anker, A." box="[219,354,511,537]" pageId="10" pageNumber="298" pagination="283 - 314" refId="ref5984" refString="Anker, A. (2003) Alpheid shrimps from the mangroves and mudflats of Singapore. Part I. Genera Salmoneus, Athanas and Potamalpheops, with the description of two new species (Crustacea: Decapoda: Caridea). Raffles Bulletin of Zoology, 51, 283 - 314." type="journal article" year="2003">Anker 2003</bibRefCitation>
|
||
; Anker
|
||
<emphasis id="B97EEAEB2662FFFAE2BAFE1B79ADE807" box="[438,493,511,537]" italics="true" pageId="10" pageNumber="298">et al.</emphasis>
|
||
2010;
|
||
<bibRefCitation id="EF9B4B082662FFFAE134FE1B7AFEE807" author="Marin, I. N." box="[568,702,511,537]" pageId="10" pageNumber="298" pagination="50 - 62" refId="ref7342" refString="Marin, I. N. (2017) Athanas alpheusophilus sp. nov. (Decapoda: Alpheidae) - a new Alpheus - associated shrimp from the Russian coast of the Sea of Japan. Zootaxa, 4324 (1), 50 - 62. https: // doi. org / 10.11646 / zootaxa. 4324.1.3" type="journal article" year="2017">Marin 2017</bibRefCitation>
|
||
), including
|
||
<taxonomicName id="4C0A4D7A2662FFFAE033FE1B7B9DE807" authorityName="Anker, Jeng & Chan" authorityYear="2001" box="[831,989,511,537]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="298" phylum="Arthropoda" rank="species" species="dentirostris">
|
||
<emphasis id="B97EEAEB2662FFFAE033FE1B7B9DE807" box="[831,989,511,537]" italics="true" pageId="10" pageNumber="298">A. dentirostris</emphasis>
|
||
</taxonomicName>
|
||
, the only other species of the genus with a toothed rostrum, and
|
||
<taxonomicName id="4C0A4D7A2662FFFAE2A9FDC77A18E823" authorityName="Hayashi" authorityYear="2002" box="[421,600,547,573]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="298" phylum="Arthropoda" rank="species" species="squillophilus">
|
||
<emphasis id="B97EEAEB2662FFFAE2A9FDC77A18E823" box="[421,600,547,573]" italics="true" pageId="10" pageNumber="298">A. squillophilus</emphasis>
|
||
</taxonomicName>
|
||
, a morphologically very distinctive species associated with stomatopods (
|
||
<bibRefCitation id="EF9B4B082662FFFAE393FDA37927E87F" author="Anker, A. & Jeng, M. - S. & Chan, T. - Y." box="[159,359,583,609]" pageId="10" pageNumber="298" pagination="1049 - 1061" refId="ref6228" refString="Anker, A., Jeng, M. - S. & Chan, T. - Y. (2001) Two unusual species of Alpheidae (Decapoda: Caridea) associated with upogebiid mudshrimps in the mudflats of Taiwan and Vietnam. Journal of Crustacean Biology, 21, 1049 - 1061. https: // doi. org / 10.1163 / 20021975 - 99990197" type="journal article" year="2001">
|
||
Anker
|
||
<emphasis id="B97EEAEB2662FFFAE3E1FDA37966E87F" box="[237,294,583,609]" italics="true" pageId="10" pageNumber="298">et al.</emphasis>
|
||
2001
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9B4B082662FFFAE27FFDA37A53E87F" author="Hayashi, K." box="[371,531,583,609]" pageId="10" pageNumber="298" pagination="395 - 403" refId="ref7120" refString="Hayashi, K. (2002) A new species of the genus Athanas (Decapoda, Caridea, Alpheidae) living in the burrows of a mantis shrimp. Crustaceana, 75, 395 - 403. https: // doi. org / 10.1163 / 156854002760095462" type="journal article" year="2002">Hayashi 2002</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB536F92662FFFAE3CBFD8F79BAE9D7" blockId="10.[151,1437,151,969]" pageId="10" pageNumber="298">
|
||
From an ecological point of view,
|
||
<taxonomicName id="4C0A4D7A2662FFFAE169FD8F7AADE89B" authority="Anker & Ďuriš, 2022" authorityName="Anker & Ďuriš" authorityYear="2022" box="[613,749,619,645]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="298" phylum="Arthropoda" rank="species" species="philippei" status="sp. nov.">
|
||
<emphasis id="B97EEAEB2662FFFAE169FD8F7AADE89B" box="[613,749,619,645]" italics="true" pageId="10" pageNumber="298">A. philippei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAEB2662FFFAE1F4FD8F7B7BE89B" bold="true" box="[760,827,619,645]" pageId="10" pageNumber="298">
|
||
<taxonomicNameLabel id="A24D57902662FFFAE1F4FD8F7B7BE89B" box="[760,827,619,645]" pageId="10" pageNumber="298" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is the fourth species of the genus
|
||
<taxonomicName id="4C0A4D7A2662FFFAE7D3FD8F7D7BE89B" authorityName="Leach" authorityYear="1814" box="[1247,1339,619,645]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="298" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97EEAEB2662FFFAE7D3FD8F7D7BE89B" box="[1247,1339,619,645]" italics="true" pageId="10" pageNumber="298">Athanas</emphasis>
|
||
</taxonomicName>
|
||
directly encountered in a mantis shrimp burrow, suggesting a direct symbiotic association, the others being
|
||
<taxonomicName id="4C0A4D7A2662FFFAE61CFD6B7DD6E8B6" authorityName="Holthuis" authorityYear="1951" box="[1296,1430,655,681]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="298" phylum="Arthropoda" rank="species" species="amazone">
|
||
<emphasis id="B97EEAEB2662FFFAE61CFD6B7DD6E8B6" box="[1296,1430,655,681]" italics="true" pageId="10" pageNumber="298">A. amazone</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0A4D7A2662FFFAE39BFD577909E8D3" authorityName="Hayashi" authorityYear="2002" box="[151,329,691,717]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="298" phylum="Arthropoda" rank="species" species="squillophilus">
|
||
<emphasis id="B97EEAEB2662FFFAE39BFD577909E8D3" box="[151,329,691,717]" italics="true" pageId="10" pageNumber="298">A. squillophilus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0A4D7A2662FFFAE28DFD577A5DE8D3" authorityName="Duris & Anker" authorityYear="2014" box="[385,541,691,717]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="298" phylum="Arthropoda" rank="species" species="manticolus">
|
||
<emphasis id="B97EEAEB2662FFFAE28DFD577A5DE8D3" box="[385,541,691,717]" italics="true" pageId="10" pageNumber="298">A. manticolus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF9B4B082662FFFAE120FD577B0EE8D3" author="Froglia, C. & Atkinson, R. J. A." box="[556,846,691,717]" pageId="10" pageNumber="298" pagination="529 - 532" refId="ref7065" refString="Froglia, C. & Atkinson, R. J. A. (1998) Association between Athanas amazone (Decapoda: Alpheidae) and Squilla mantis (Stomatopoda: Squillidae). Journal of Crustacean Biology, 18, 529 - 532. https: // doi. org / 10.2307 / 1549417" type="journal article" year="1998">Froglia & Atkinson 1998</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9B4B082662FFFAE057FD577BBBE8D3" author="Hayashi, K." box="[859,1019,691,717]" pageId="10" pageNumber="298" pagination="395 - 403" refId="ref7120" refString="Hayashi, K. (2002) A new species of the genus Athanas (Decapoda, Caridea, Alpheidae) living in the burrows of a mantis shrimp. Crustaceana, 75, 395 - 403. https: // doi. org / 10.1163 / 156854002760095462" type="journal article" year="2002">Hayashi 2002</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9B4B082662FFFAE70BFD507CB5E8D3" author="Duris, Z. & Anker, A." box="[1031,1269,691,719]" pageId="10" pageNumber="298" pagination="550 - 558" refId="ref7010" refString="Duris, Z. & Anker, A. (2014) Athanas manticolus sp. nov., a new stomatopod-associated alpheid shrimp from Vietnam (Crustacea, Decapoda). Zootaxa, 3784 (5), 550 - 558. https: // doi. org / 10.11646 / zootaxa. 3784.5.4" type="journal article" year="2014">Ďuriš & Anker 2014</bibRefCitation>
|
||
). Based on the presently available evidence,
|
||
<taxonomicName id="4C0A4D7A2662FFFAE2EAFD337A20E8EF" authorityName="Anker & Komai" authorityYear="2010" box="[486,608,727,753]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="298" phylum="Arthropoda" rank="species" species="ahyongi">
|
||
<emphasis id="B97EEAEB2662FFFAE2EAFD337A20E8EF" box="[486,608,727,753]" italics="true" pageId="10" pageNumber="298">A. ahyongi</emphasis>
|
||
</taxonomicName>
|
||
also appears to be an inquiline shrimp, dwelling in stomatopod burrows, although no specimens have been collected so far together with a host (
|
||
<bibRefCitation id="EF9B4B082662FFFAE0CCFD1F7CFFE90B" author="Anker, A. & Komai, T." box="[960,1215,763,789]" pageId="10" pageNumber="298" pagination="45 - 54" refId="ref6296" refString="Anker, A. & Komai, T. (2010) Description of a new species of Athanas Leach, 1814 (Crustacea: Decapoda: Alpheidae) from Madagascar and Japan. Zootaxa, 2680 (1), 45 - 54. https: // doi. org / 10.11646 / zootaxa. 2680.1.4" type="journal article" year="2010">Anker & Komai 2010</bibRefCitation>
|
||
). The hypothesis of the likely association between
|
||
<taxonomicName id="4C0A4D7A2662FFFAE2FCFCFB7A29E927" authorityName="Anker & Komai" authorityYear="2010" box="[496,617,799,825]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="298" phylum="Arthropoda" rank="species" species="ahyongi">
|
||
<emphasis id="B97EEAEB2662FFFAE2FCFCFB7A29E927" box="[496,617,799,825]" italics="true" pageId="10" pageNumber="298">A. ahyongi</emphasis>
|
||
</taxonomicName>
|
||
and stomatopods is based on the collection data of the Nosy-Bé material (subtidal sand flat with rich infaunal community, including large stomatopods, AA, pers. obs.), the species’ general morphology, as well as its characteristic colour pattern, which is almost identical to the colour pattern of several shrimps, presumably
|
||
<taxonomicName id="4C0A4D7A2662FFFAE282FC6F7A55E9BB" box="[398,533,907,933]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="298" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B97EEAEB2662FFFAE282FC6F79AAE9BB" box="[398,490,907,933]" italics="true" pageId="10" pageNumber="298">Athanas</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
, photographed near entrances of stomatopod burrows (e.g.
|
||
<bibRefCitation id="EF9B4B082662FFFAE7CBFC6F794BE9D7" author="Kuiter, R. H. & Debelius, H." pageId="10" pageNumber="298" refId="ref7210" refString="Kuiter, R. H. & Debelius, H. (2009) World Atlas of Marine Fauna. IKAN, Frankfurt, 725 pp." type="book" year="2009">Kuiter & Debelius 2009: 159</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9B4B082662FFFAE21BFC4B79ADE9D7" author="Ryanskiy, A. S." box="[279,493,943,969]" pageId="10" pageNumber="298" refId="ref7540" refString="Ryanskiy, A. S. (2020) Coral Reef Crustaceans from Red Sea to Papua: Reef ID Books. Produced by the author, Moscow, 149 pp." type="book" year="2020">Ryanskiy 2020: 29</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |