222 lines
31 KiB
XML
222 lines
31 KiB
XML
<document id="24D12958014AA5757ABD2B9E60DAFF0B" ID-CLB-Dataset="2961" ID-DOI="10.11646/palaeoentomology.4.6.5" ID-GBIF-Dataset="73a0820d-7503-472f-8c32-3836cf5fe857" ID-ISSN="2624-2834" ID-Zenodo-Dep="5778576" ID-ZooBank="95AA3380-CA95-46E8-B233-E9F0E8D60603" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1639469484134" checkinUser="plazi" docAuthor="Béthoux, Olivier & Anderson, John M." docDate="2021" docId="039DFA6CFFDEFFDD6D65FD87CD59FEF8" docLanguage="en" docName="palaeoentomology.4.6.5.pdf" docOrigin="Palaeoentomology 4 (6)" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Polycytellidae Martynova 1952" docType="treatment" docVersion="5" lastPageNumber="552" masterDocId="FFA48214FFDFFFDF6E4FFF9CC82EFFF8" masterDocTitle="The Polycytellidae viewed as Gondwanan Glosselytrodea" masterLastPageNumber="558" masterPageNumber="550" pageNumber="551" updateTime="1699472406386" updateUser="ExternalLinkService">
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<mods:title id="B3EC915F6BA870EFE3F10B31688B12F6">The Polycytellidae viewed as Gondwanan Glosselytrodea</mods:title>
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<mods:name id="A345C0EB56F28188C9F0E9A20FB033D9" type="personal">
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<mods:namePart id="03AC411C01190F8D92A5D59CD81BED7D">Béthoux, Olivier</mods:namePart>
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<mods:affiliation id="20A22111CE82D68D5B36DC3CBFABDD13">Centre de Recherche sur la Paléontologie - Paris (CR 2 P), Sorbonne Université, MNHN, CNRS, Paris, France</mods:affiliation>
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<mods:namePart id="23A9F2E31DEA7369E42B9D0E2E2447C7">Anderson, John M.</mods:namePart>
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<mods:affiliation id="70A6A0176F44A6C26AFD678BA719BEE1">Evolutionary Studies Institute, Witwatersrand University, Johannesburg, South Africa</mods:affiliation>
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<mods:title id="CF1642D57BB5CF81E946B875482B43BE">Palaeoentomology</mods:title>
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<mods:part id="DA774AA8E495DFF7531A8502C4A15874">
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<mods:date id="3D27C82E2361C60591F26A08ED92D45E">2021</mods:date>
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<mods:number id="9A91305CD1B42688E2ABB4F0C3DE4418">2021-12-14</mods:number>
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<mods:number id="4CD4309FB0D73EA09D39D688B6665191">4</mods:number>
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<mods:identifier id="CA9DA0162E7AC10D8B178DA47BB1DB56" type="DOI">10.11646/palaeoentomology.4.6.5</mods:identifier>
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<treatment id="039DFA6CFFDEFFDD6D65FD87CD59FEF8" ID-DOI="http://doi.org/10.5281/zenodo.5780412" ID-GBIF-Taxon="191571256" ID-Zenodo-Dep="5780412" LSID="urn:lsid:plazi:treatment:039DFA6CFFDEFFDD6D65FD87CD59FEF8" httpUri="http://treatment.plazi.org/id/039DFA6CFFDEFFDD6D65FD87CD59FEF8" lastPageId="2" lastPageNumber="552" pageId="1" pageNumber="551">
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<subSubSection id="C32E18F1FFDEFFDE6D65FD87CD5CFDCE" box="[810,1394,539,566]" pageId="1" pageNumber="551" type="nomenclature">
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<paragraph id="8B8B4B7AFFDEFFDE6D65FD87CD5CFDCE" blockId="1.[810,1394,503,566]" box="[810,1394,539,566]" pageId="1" pageNumber="551">
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<heading id="D0C3FC16FFDEFFDE6D65FD87CD5CFDCE" box="[810,1394,539,566]" fontSize="10" level="2" pageId="1" pageNumber="551" reason="2">
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<emphasis id="B9409768FFDEFFDE6D65FD87CD5CFDCE" bold="true" box="[810,1394,539,566]" pageId="1" pageNumber="551">
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Family
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<taxonomicName id="4C3430F9FFDEFFDE6DCAFD87CCD0FDCD" authority="Martynova, 1952" authorityName="Martynova" authorityYear="1952" box="[901,1278,539,566]" class="Insecta" family="Polycytellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="1" pageNumber="551" phylum="Arthropoda" rank="family">
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Polycytellidae
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<bibRefCitation id="EFA5368BFFDEFFDE6A7DFD80CCD0FDCD" author="Martynova, O. M." box="[1074,1278,539,566]" pageId="1" pageNumber="551" pagination="187 - 196" refId="ref6684" refString="Martynova, O. M. (1952) Otryad Glosselytrodea v permskikh otlozheniyakh Kemerovskoi oblasti. Trudy Paleontologicheskogo instituta, Akademiya Nauk SSSR, 40, 187 - 196." type="journal article" year="1952">Martynova, 1952</bibRefCitation>
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</taxonomicName>
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stat. rest.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E18F1FFDEFFDE6D65FDF8CBACFD14" pageId="1" pageNumber="551" type="type_taxon">
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<paragraph id="8B8B4B7AFFDEFFDE6D65FDF8CBACFD14" blockId="1.[810,1452,612,2092]" pageId="1" pageNumber="551">
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<emphasis id="B9409768FFDEFFDE6D65FDF8CBE9FD86" bold="true" box="[810,967,612,638]" pageId="1" pageNumber="551">Type genus.</emphasis>
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<taxonomicName id="4C3430F9FFDEFFDE6D91FDF8CD0FFD86" ID-CoL="63NBT" authority="Tillyard, 1922" authorityName="Tillyard" authorityYear="1922" box="[990,1313,612,638]" class="Insecta" family="Glosselytridae" genus="Polycytella" kingdom="Animalia" order="Glosselytrodea" pageId="1" pageNumber="551" phylum="Arthropoda" rank="genus">
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<emphasis id="B9409768FFDEFFDE6D91FDF8CC72FD86" box="[990,1116,612,638]" italics="true" pageId="1" pageNumber="551">Polycytella</emphasis>
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<bibRefCitation id="EFA5368BFFDEFFDE6A3DFDF8CD0FFD86" author="Tillyard, R. J." box="[1138,1313,612,638]" pageId="1" pageNumber="551" pagination="447 - 470" refId="ref7128" refString="Tillyard, R. J. (1922) Mesozoic insects of Queensland. No. 9. Orthoptera, and additions to the Protorthoptera, Odonata, Hemiptera and Plannipennia. Proceedings of the Linnean Society of New South Wales, 47 (4), 447 - 470." type="journal article" year="1922">Tillyard, 1922</bibRefCitation>
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</taxonomicName>
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(including
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<taxonomicName id="4C3430F9FFDEFFDE6D65FD15CCEBFD5B" ID-CoL="4L58Y" authority="Tillyard, 1922" authorityName="Tillyard" authorityYear="1922" box="[810,1221,649,675]" class="Insecta" family="Glosselytridae" genus="Polycytella" kingdom="Animalia" order="Glosselytrodea" pageId="1" pageNumber="551" phylum="Arthropoda" rank="species" species="triassica">
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<emphasis id="B9409768FFDEFFDE6D65FD15CC3DFD5A" box="[810,1043,649,675]" italics="true" pageId="1" pageNumber="551">Polycytella triassica</emphasis>
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<bibRefCitation id="EFA5368BFFDEFFDE6A50FD15CCEBFD5B" author="Tillyard, R. J." box="[1055,1221,649,675]" pageId="1" pageNumber="551" pagination="447 - 470" refId="ref7128" refString="Tillyard, R. J. (1922) Mesozoic insects of Queensland. No. 9. Orthoptera, and additions to the Protorthoptera, Odonata, Hemiptera and Plannipennia. Proceedings of the Linnean Society of New South Wales, 47 (4), 447 - 470." type="journal article" year="1922">Tillyard, 1922</bibRefCitation>
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</taxonomicName>
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, Triassic, Australia; and
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<taxonomicName id="4C3430F9FFDEFFDE6D2AFD31CC4BFD3F" authority="Béthoux & Anderson, 2021" authorityName="Béthoux & Anderson" authorityYear="2021" box="[869,1125,685,711]" class="Insecta" family="Glosselytridae" genus="Polycytella" kingdom="Animalia" order="Glosselytrodea" pageId="1" pageNumber="551" phylum="Arthropoda" rank="species" species="rasnitsyni" status="sp. nov.">
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<emphasis id="B9409768FFDEFFDE6D2AFD31CC4BFD3F" box="[869,1125,685,711]" italics="true" pageId="1" pageNumber="551">Polycytella rasnitsyni</emphasis>
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</taxonomicName>
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<emphasis id="B9409768FFDEFFDE6A37FD31CCF0FD3F" bold="true" box="[1144,1246,685,711]" pageId="1" pageNumber="551">
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<taxonomicNameLabel id="A2732A13FFDEFFDE6A37FD31CCF0FD3F" box="[1144,1246,685,711]" pageId="1" pageNumber="551" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, Triassic, South Africa).
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E18F1FFDEFFDE6D10FD69CDA9FC5A" pageId="1" pageNumber="551" type="discussion">
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<paragraph id="8B8B4B7AFFDEFFDE6D10FD69CDA9FC5A" blockId="1.[810,1452,612,2092]" pageId="1" pageNumber="551">
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<emphasis id="B9409768FFDEFFDE6D10FD69CC36FCE8" bold="true" box="[863,1048,757,784]" pageId="1" pageNumber="551">Other genera.</emphasis>
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<taxonomicName id="4C3430F9FFDEFFDE6A7DFD6ACD1CFCE8" authorityName="Martins-Neto & Gallego" authorityYear="2001" box="[1074,1330,758,784]" class="Insecta" family="Glosselytridae" genus="Argentinoglosselytrina" kingdom="Animalia" order="Glosselytrodea" pageId="1" pageNumber="551" phylum="Arthropoda" rank="genus">
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<emphasis id="B9409768FFDEFFDE6A7DFD6ACD1CFCE8" box="[1074,1330,758,784]" italics="true" pageId="1" pageNumber="551">Argentinoglosselytrina</emphasis>
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</taxonomicName>
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Martins- Neto & Gallego, 2001 (including
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<taxonomicName id="4C3430F9FFDEFFDE6AE3FC86CD19FCA1" authority="Martins-Neto and Gallego, 2001" authorityName="Martins-Neto and Gallego" authorityYear="2001" class="Insecta" family="Glosselytridae" genus="Argentinoglosselytrina" kingdom="Animalia" order="Glosselytrodea" pageId="1" pageNumber="551" phylum="Arthropoda" rank="species" species="pulchella">
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<emphasis id="B9409768FFDEFFDE6AE3FC86CBBCFCA0" italics="true" pageId="1" pageNumber="551">Argentinoglosselytrina pulchella</emphasis>
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<bibRefCitation id="EFA5368BFFDEFFDE6DEAFCA2CD19FCA1" author="Martins-Neto, R. G. & Gallego, O. F." box="[933,1335,830,857]" pageId="1" pageNumber="551" pagination="105 - 114" refId="ref6556" refString="Martins-Neto, R. G. & Gallego, O. F. (2001) The Triassic insect fauna from Argentina. IV. Glosselytrodea and complements on Auchenorrhyncha. Acta Geologica Leopoldensia, 24 (52 / 53), 105 - 114." type="journal article" year="2001">Martins-Neto and Gallego, 2001</bibRefCitation>
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</taxonomicName>
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, Triassic, Argentina), and
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<taxonomicName id="4C3430F9FFDEFFDE6DB6FCFFCCB5FC85" authority="Béthoux & Anderson, 2021" authorityName="Béthoux & Anderson" authorityYear="2021" box="[1017,1179,867,893]" class="Insecta" family="Polycytellidae" genus="Moltenojurina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="1" pageNumber="551" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9409768FFDEFFDE6DB6FCFFCCB5FC85" box="[1017,1179,867,893]" italics="true" pageId="1" pageNumber="551">Moltenojurina</emphasis>
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</taxonomicName>
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<emphasis id="B9409768FFDEFFDE6AE0FCFFCD0BFC85" bold="true" box="[1199,1317,867,893]" pageId="1" pageNumber="551">
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<taxonomicNameLabel id="A2732A13FFDEFFDE6AE0FCFFCD0BFC85" box="[1199,1317,867,893]" pageId="1" pageNumber="551" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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(including
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<taxonomicName id="4C3430F9FFDEFFDE6D65FC1BCC3DFC59" authority="Béthoux & Anderson, 2021" authorityName="Béthoux & Anderson" authorityYear="2021" box="[810,1043,903,929]" class="Insecta" family="Polycytellidae" genus="Moltenojurina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="1" pageNumber="551" phylum="Arthropoda" rank="species" species="parva" status="sp. nov.">
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<emphasis id="B9409768FFDEFFDE6D65FC1BCC3DFC59" box="[810,1043,903,929]" italics="true" pageId="1" pageNumber="551">Moltenojurina parva</emphasis>
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</taxonomicName>
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<emphasis id="B9409768FFDEFFDE6A54FC1BCC5AFC59" bold="true" box="[1051,1140,903,929]" pageId="1" pageNumber="551">
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<taxonomicNameLabel id="A2732A13FFDEFFDE6A54FC1BCC5AFC59" box="[1051,1140,903,929]" pageId="1" pageNumber="551" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, Triassic, South Africa).
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E18F1FFDEFFDE6D10FC30CC00FBF7" pageId="1" pageNumber="551" type="diagnosis">
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<paragraph id="8B8B4B7AFFDEFFDE6D10FC30CC00FBF7" blockId="1.[810,1452,612,2092]" pageId="1" pageNumber="551">
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<emphasis id="B9409768FFDEFFDE6D10FC30CC7BFC3D" bold="true" box="[863,1109,939,966]" pageId="1" pageNumber="551">Emended diagnosis.</emphasis>
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Forewing: opposite the end of ScP, area anterior to RA large (accounting for about a quarter of wing width).
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E18F1FFDEFFDD6D10FB84CD59FEF8" lastPageId="2" lastPageNumber="552" pageId="1" pageNumber="551" type="discussion">
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||
<paragraph id="8B8B4B7AFFDEFFDE6D10FB84CBABFB67" blockId="1.[810,1452,612,2092]" pageId="1" pageNumber="551">
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<emphasis id="B9409768FFDEFFDE6D10FB84CBCAFBCA" bold="true" box="[863,996,1048,1074]" pageId="1" pageNumber="551">Discussion.</emphasis>
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Genera herein assigned to the
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<taxonomicName id="4C3430F9FFDEFFDE6D65FBA1CBE4FBAF" authorityName="Martynova" authorityYear="1952" box="[810,970,1085,1111]" class="Insecta" family="Polycytellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="1" pageNumber="551" phylum="Arthropoda" rank="family">Polycytellidae</taxonomicName>
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display two states that have been generally regarded as relevant to the systematics of
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<taxonomicName id="4C3430F9FFDEFFDE6AB3FBFDCD89FB83" authorityName="Martynov" authorityYear="1938" box="[1276,1447,1121,1147]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="1" pageNumber="551" phylum="Arthropoda" rank="order">Glosselytrodea</taxonomicName>
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, namely:
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</paragraph>
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<paragraph id="8B8B4B7AFFDEFFDE6D10FB36CD4AFAF5" blockId="1.[810,1452,612,2092]" pageId="1" pageNumber="551">(1)occurrence of a single mid-vein (or, ‘two mid-veins fused into a single mid-vein, as opposed to ‘occurrence of two, closely adjoined but distinct, mid-veins’); and</paragraph>
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<paragraph id="8B8B4B7AFFDEFFDE6D10FA8BCBE5FA67" blockId="1.[810,1452,612,2092]" pageId="1" pageNumber="551">(2) anterior and posterior ambient veins joining each other at their apices (as opposed to ‘anterior and posterior ambient veins reaching the anterior and posterior margins, respectively’).</paragraph>
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<paragraph id="8B8B4B7AFFDEFFDE6D10FA35CD5AF847" blockId="1.[810,1452,612,2092]" pageId="1" pageNumber="551">
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<bibRefCitation id="EFA5368BFFDEFFDE6D10FA35CC06FA3B" author="Martynova, O. M." box="[863,1064,1448,1475]" pageId="1" pageNumber="551" pagination="187 - 196" refId="ref6684" refString="Martynova, O. M. (1952) Otryad Glosselytrodea v permskikh otlozheniyakh Kemerovskoi oblasti. Trudy Paleontologicheskogo instituta, Akademiya Nauk SSSR, 40, 187 - 196." type="journal article" year="1952">Martynova (1952</bibRefCitation>
|
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; and see
|
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<bibRefCitation id="EFA5368BFFDEFFDE6AD9FA35CD72FA3A" author="Martynova, O. M." box="[1174,1372,1448,1475]" pageId="1" pageNumber="551" pagination="157 - 159" refId="ref6715" refString="Martynova, O. M. (1962) Otryad Glosselytrodea. In: Orlov, Y. A. (Ed.), Osnovy paleontologii. Chlenistonogie, trakheinye i khelitserovye. Akademyia Nauk SSSR, Moscow, Russia, pp. 157 - 159." type="book chapter" year="1962">Martynova, 1962</bibRefCitation>
|
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,
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<bibRefCitation id="EFA5368BFFDEFFDE6B25FA34CD8DFA3A" author="Martynova, O. M." box="[1386,1443,1448,1474]" pageId="1" pageNumber="551" pagination="186 - 189" refId="ref6897" refString="Martynova, O. M. (1991) Order Glosselytrodea. In: Kothekar, V. S. (Ed.), Fundamentals of paleontology. Arthropoda, Tracheata, Chelicerata. Smithsonian Institution Libraries, Washington, D. C., pp. 186 - 189. [English translation of Martynova, 1962]" type="journal article" year="1991">1991</bibRefCitation>
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) regarded these traits as diagnostic of the
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<taxonomicName id="4C3430F9FFDEFFDE6B4BFA51CD89FA1F" authorityName="Martynova" authorityYear="1952" box="[1284,1447,1485,1511]" class="Insecta" family="Polycytellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="1" pageNumber="551" phylum="Arthropoda" rank="family">Polycytellidae</taxonomicName>
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, to which she also assigned
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<taxonomicName id="4C3430F9FFDEFFDE6ADAFA6ECB4CF9C8" authority="Martynova, 1943" authorityName="Martynova" authorityYear="1943" class="Insecta" family="Jurinidae" genus="Mesojurina" kingdom="Animalia" order="Glosselytrodea" pageId="1" pageNumber="551" phylum="Arthropoda" rank="genus">
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<emphasis id="B9409768FFDEFFDE6ADAFA6ECD38F9F3" box="[1173,1302,1522,1547]" italics="true" pageId="1" pageNumber="551">Mesojurina</emphasis>
|
||
<bibRefCitation id="EFA5368BFFDEFFDE6B66FA6ECB4CF9C8" author="Martynova, O. M." pageId="1" pageNumber="551" pagination="312 - 313" refId="ref6646" refString="Martynova, O. M. (1943) Glosselytrodea from the Jurassic shales of the coal bed Sogjuta. Comptes Rendus de l'Academie des Sciences de l'URSS, 39 (7), 312 - 313." type="journal article" year="1943">Martynova, 1943</bibRefCitation>
|
||
</taxonomicName>
|
||
(including
|
||
<taxonomicName id="4C3430F9FFDEFFDE6A4FF98ACB4BF9AC" authority="Martynova, 1943" authorityName="Martynova" authorityYear="1943" class="Insecta" family="Jurinidae" genus="Mesojurina" kingdom="Animalia" order="Glosselytrodea" pageId="1" pageNumber="551" phylum="Arthropoda" rank="species" species="sogjutensis">
|
||
<emphasis id="B9409768FFDEFFDE6A4FF98ACD3DF9D7" box="[1024,1299,1558,1583]" italics="true" pageId="1" pageNumber="551">Mesojurina sogjutensis</emphasis>
|
||
<bibRefCitation id="EFA5368BFFDEFFDE6B66F98ACB4BF9AC" author="Martynova, O. M." pageId="1" pageNumber="551" pagination="312 - 313" refId="ref6646" refString="Martynova, O. M. (1943) Glosselytrodea from the Jurassic shales of the coal bed Sogjuta. Comptes Rendus de l'Academie des Sciences de l'URSS, 39 (7), 312 - 313." type="journal article" year="1943">Martynova, 1943</bibRefCitation>
|
||
</taxonomicName>
|
||
, Jurassic, Kyrgyzstan; and see below). However, Rasnitsyn & Aristov in
|
||
<bibRefCitation id="EFA5368BFFDEFFDE6A73F9C2CD06F980" author="Aristov, D. S. & Bashkuev, A. & Golubev, V. K. & Gorochov, A. V. & Karasev, E. V. & Kopylov, D. S. & Ponomarenko, A. B. & Rasnitsyn, A. P. & Rasnitsyn, D. A. & Sinitshenkova, N. D. & Sukatsheva, I. D. & Vassilenko, D. V." box="[1084,1320,1630,1657]" pageId="1" pageNumber="551" pagination="641 - 832" refId="ref5804" refString="Aristov, D. S., Bashkuev, A., Golubev, V. K., Gorochov, A. V., Karasev, E. V., Kopylov, D. S., Ponomarenko, A. B., Rasnitsyn, A. P., Rasnitsyn, D. A., Sinitshenkova, N. D., Sukatsheva, I. D. & Vassilenko, D. V. (2013) Fossil insects of the Middle and Upper Permian of European Russia. Paleontological Journal, 47 (7), 641 - 832. https: // doi. org / 10.1134 / S 0031030113070010" type="journal article" year="2013">
|
||
Aristov
|
||
<emphasis id="B9409768FFDEFFDE6AD6F9C3CCE3F980" box="[1177,1229,1630,1656]" italics="true" pageId="1" pageNumber="551">et al</emphasis>
|
||
. (2013)
|
||
</bibRefCitation>
|
||
considered that this combination of traits may have been acquired convergently within two distinct families, namely the oligoneurous
|
||
<taxonomicName id="4C3430F9FFDEFFDE6DF0F957CCE3F91D" authority="Zalessky, 1929" authorityName="Zalessky" authorityYear="1929" box="[959,1229,1739,1765]" class="Insecta" family="Jurinidae" kingdom="Animalia" order="Glosselytrodea" pageId="1" pageNumber="551" phylum="Arthropoda" rank="family">
|
||
Jurinidae
|
||
<bibRefCitation id="EFA5368BFFDEFFDE6A66F957CCE3F91D" author="Zalessky, M. D." box="[1065,1229,1739,1765]" pageId="1" pageNumber="551" pagination="1 - 30" refId="ref7290" refString="Zalessky, M. D. (1929) O novykh nasekomykh iz permskikh otlozhenii basseinov rek Kamy, Vyatki i Beloi. Trudy obshchestva estestvoispytatelei pri Kazanskom Universitete, 52 (1), 1 - 30." type="journal article" year="1929">Zalessky,1929</bibRefCitation>
|
||
</taxonomicName>
|
||
and the polyneurous
|
||
<taxonomicName id="4C3430F9FFDEFFDE6D65F973CCBCF8F2" authority="Martynov, 1938" authorityName="Martynov" authorityYear="1938" box="[810,1170,1775,1802]" class="Insecta" family="Glosselytridae" kingdom="Animalia" order="Glosselytrodea" pageId="1" pageNumber="551" phylum="Arthropoda" rank="family">
|
||
Glosselytridae
|
||
<bibRefCitation id="EFA5368BFFDEFFDE6D98F96CCCBCF8F2" author="Martynov, A. V." box="[983,1170,1776,1802]" pageId="1" pageNumber="551" pagination="187 - 206" refId="ref6602" refString="Martynov, A. V. (1938) On a new Permian order of orthopteroid insects, Glosselytrodea. Bulletin de l'Academie des Sciences de l'URSS, Biologie, 1938, 187 - 206. [In Russian with extended English summary]" type="journal article" year="1938">Martynov, 1938</bibRefCitation>
|
||
</taxonomicName>
|
||
. Besides the number of veins, which can be partly related to size, this proposal rests on scenarios of forewing elytrization, achieved either by membrane strengthening or veins strengthening. However, these aspects remain difficult to appreciate based on data currently available on
|
||
<taxonomicName id="4C3430F9FFDEFFDE6A8BF839CD5EF847" authorityName="Martynov" authorityYear="1938" box="[1220,1392,1957,1983]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="1" pageNumber="551" phylum="Arthropoda" rank="order">Glosselytrodea</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8B4B7AFFDEFFDD6D10F856CAA6FE44" blockId="1.[810,1452,612,2092]" lastBlockId="2.[136,778,162,1214]" lastPageId="2" lastPageNumber="552" pageId="1" pageNumber="551">
|
||
Of further interest, it is common that the area between the anterior wing margin and ScP is enlarged in
|
||
<taxonomicName id="4C3430F9FFDEFFDE6D1AF78ECC2FF7D4" authorityName="Martynov" authorityYear="1938" box="[853,1025,2066,2092]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="1" pageNumber="551" phylum="Arthropoda" rank="order">Glosselytrodea</taxonomicName>
|
||
. Consequently, the anterior wing margin has a marked inflexion opposite the end of ScP. However, in genera herein assigned to the
|
||
<taxonomicName id="4C3430F9FFDDFFDD6C2CFF5ACB28FF18" authorityName="Martynova" authorityYear="1952" box="[611,774,198,224]" class="Insecta" family="Polycytellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="2" pageNumber="552" phylum="Arthropoda" rank="family">Polycytellidae</taxonomicName>
|
||
, the area anterior to RA (
|
||
<emphasis id="B9409768FFDDFFDD6FE4FF70C99DFEFD" box="[427,435,236,261]" italics="true" pageId="2" pageNumber="552">i</emphasis>
|
||
.
|
||
<emphasis id="B9409768FFDDFFDD6FF5FF70C9E8FEFD" box="[442,454,236,261]" italics="true" pageId="2" pageNumber="552">e</emphasis>
|
||
., between ScP and RA and then between the anterior wing margin and RA) is also enlarged (opposite the end of ScP, nearly as large as the area between RA and MP+CuA
|
||
<emphasis id="B9409768FFDDFFDD6FACFEC6CA29FE8B" box="[483,519,346,371]" italics="true" pageId="2" pageNumber="552">ant</emphasis>
|
||
), with the consequence that the anterior wing margin regains a more continuous course. This trait can be described as follows:
|
||
</paragraph>
|
||
<paragraph id="8B8B4B7AFFDDFFDD6EF2FE5BCAFEFDD3" blockId="2.[136,778,162,1214]" pageId="2" pageNumber="552">(3) ‘opposite the end of ScP, area anterior to RA large’ (as opposed to ‘RA close to ScP and, distal to the end of this vein, close to the anterior wing margin’).</paragraph>
|
||
<paragraph id="8B8B4B7AFFDDFFDD6EF2FDA9C9F4FBFE" blockId="2.[136,778,162,1214]" pageId="2" pageNumber="552">
|
||
It is present in
|
||
<taxonomicName id="4C3430F9FFDDFFDD6FC3FDA9CA24FDB7" authorityName="Tillyard" authorityYear="1922" box="[396,522,565,591]" class="Insecta" family="Glosselytridae" genus="Polycytella" kingdom="Animalia" order="Glosselytrodea" pageId="2" pageNumber="552" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9409768FFDDFFDD6FC3FDA9CA24FDB7" box="[396,522,565,591]" italics="true" pageId="2" pageNumber="552">Polycytella</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="130F57FFFFDDFFDD6C6BFDA9CA41FDB7" box="[548,623,565,591]" captionStart="FIGURE 1" captionStartId="2.[136,235,1986,2010]" captionTargetBox="[288,1302,1252,1963]" captionTargetId="figure-512@2.[282,1305,1243,1963]" captionTargetPageId="2" captionText="FIGURE 1. Polycytella triassica Tillyard, 1922, holotype specimen (GSQ81a), right forewing (dorsal aspect, but mostly as impression of ventral side). A, Interpretative drawing. B, Photograph (G. Thompson, Queensland Museum, Brisbane, Australia; ‘cf’, claval furrow)." figureDoi="http://doi.org/10.5281/zenodo.5778578" httpUri="https://zenodo.org/record/5778578/files/figure.png" pageId="2" pageNumber="552">Fig. 1</figureCitation>
|
||
),
|
||
<taxonomicName id="4C3430F9FFDDFFDD6CDEFDA9C937FD8B" authorityName="Martins-Neto & Gallego" authorityYear="2001" class="Insecta" family="Glosselytridae" genus="Argentinoglosselytrina" kingdom="Animalia" order="Glosselytrodea" pageId="2" pageNumber="552" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9409768FFDDFFDD6CDEFDA9C937FD8B" italics="true" pageId="2" pageNumber="552">Argentinoglosselytrina</emphasis>
|
||
</taxonomicName>
|
||
and the new material described below (
|
||
<figureCitation id="130F57FFFFDDFFDD6C97FDC5C898FD60" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="3.[136,235,1918,1942]" captionStartId-1="4.[136,235,1039,1063]" captionStartId-2="6.[810,909,1974,1998]" captionTargetBox-0="[189,1398,197,1894]" captionTargetBox-1="[192,1373,208,1007]" captionTargetBox-2="[809,1452,1435,1950]" captionTargetId-0="figure-20@3.[189,1398,189,1894]" captionTargetId-1="figure-435@4.[183,1404,204,1016]" captionTargetId-2="figure-692@6.[809,1452,1430,1950]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="6" captionText-0="FIGURE 2. Polycytella rasnitsyni sp. nov., holotype specimen (PRE/F/10559), forewing pair. A, Interpretative drawing (RFW, right forewing; LFW, left forewing). B–D, Photograph and RTI extracts, part. B, Photograph (light-mirrored; ‘cf’, claval furrow). C, RTI extract, specular enhancement mode. D, RTI extract, normals visualisation mode." captionText-1="FIGURE 3. Polycytella rasnitsyni sp. nov., holotype specimen (PRE/F/10559), interpretative drawings, separated. A, Left forewing, as in Fig. 2A. B, Right forewing, as in Fig. 2A and with reconstructed parts." captionText-2="FIGURE 4. Moltenojurina parva gen. et sp. nov., holotype specimen (PRE/F/20922), right forewing. A, Interpretative drawing. B, Photograph (light-mirrored, flipped horizontally)." figureDoi-0="http://doi.org/10.5281/zenodo.5778582" figureDoi-1="http://doi.org/10.5281/zenodo.5778586" figureDoi-2="http://doi.org/10.5281/zenodo.5778588" httpUri-0="https://zenodo.org/record/5778582/files/figure.png" httpUri-1="https://zenodo.org/record/5778586/files/figure.png" httpUri-2="https://zenodo.org/record/5778588/files/figure.png" pageId="2" pageNumber="552">Figs 2–4</figureCitation>
|
||
). This trait is most likely derived, as it is unique among
|
||
<taxonomicName id="4C3430F9FFDDFFDD6EAEFD3EC9A7FD44" authorityName="Martynov" authorityYear="1938" box="[225,393,674,700]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="2" pageNumber="552" phylum="Arthropoda" rank="order">Glosselytrodea</taxonomicName>
|
||
and, to our knowledge, winged insects as a whole. A seemingly similar condition occurs in Blattodea, in which the area between the anterior wing margin and the stem of R is often very wide (
|
||
<bibRefCitation id="EFA5368BFFDDFFDD6C37FC8CCAD3FCD3" author="Rehn, J. W. H." box="[632,765,784,811]" pageId="2" pageNumber="552" pagination="1 - 134" refId="ref7016" refString="Rehn, J. W. H. (1951) Classification of the Blattaria as indicated by their wings (Orthoptera). Memoirs of the American Entomological Society, 14, 1 - 134." type="journal article" year="1951">Rehn, 1951</bibRefCitation>
|
||
). However, this R stem emits successive anterior branches, the anterior-most ones belonging to RA.As a consequence, the actual area between ScP and RA corresponds to the area between ScP and the first anterior branch of R, and is narrow. The conditions in
|
||
<taxonomicName id="4C3430F9FFDDFFDD6F87FC5BCA46FC19" authorityName="Martynova" authorityYear="1952" box="[456,616,967,993]" class="Insecta" family="Polycytellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="2" pageNumber="552" phylum="Arthropoda" rank="family">Polycytellidae</taxonomicName>
|
||
and Blattodea are therefore not homologous.
|
||
</paragraph>
|
||
<paragraph id="8B8B4B7AFFDDFFDD6EF2FB8CCD59FEF8" blockId="2.[136,778,162,1214]" lastBlockId="2.[810,1452,158,257]" pageId="2" pageNumber="552">
|
||
Considering that
|
||
<taxonomicName id="4C3430F9FFDDFFDD6FF2FB8DCA10FBD2" authorityName="Martynova" authorityYear="1943" box="[445,574,1041,1066]" class="Insecta" family="Jurinidae" genus="Mesojurina" kingdom="Animalia" order="Glosselytrodea" pageId="2" pageNumber="552" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9409768FFDDFFDD6FF2FB8DCA10FBD2" box="[445,574,1041,1066]" italics="true" pageId="2" pageNumber="552">Mesojurina</emphasis>
|
||
</taxonomicName>
|
||
displays the plesiomorphic condition of the character (
|
||
<emphasis id="B9409768FFDDFFDD6C3EFBAACABEFBB7" box="[625,656,1078,1103]" italics="true" pageId="2" pageNumber="552">viz</emphasis>
|
||
., the state ‘RA close to ScP and, distal to the end of this vein, close to the anterior wing margin’), we therefore exclude this genus from the
|
||
<taxonomicName id="4C3430F9FFDDFFDD6F74FB3FC9F0FB45" authorityName="Martynova" authorityYear="1952" box="[315,478,1187,1213]" class="Insecta" family="Polycytellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="2" pageNumber="552" phylum="Arthropoda" rank="family">Polycytellidae</taxonomicName>
|
||
, which, as a consequence, is strictly Gondwanan. Whether the corresponding taxon shall better be considered a subfamily to be nested within
|
||
<taxonomicName id="4C3430F9FFDDFFDD6D65FF7ACBBFFEF8" authorityName="Zalessky" authorityYear="1929" box="[810,913,230,256]" class="Insecta" family="Jurinidae" kingdom="Animalia" order="Glosselytrodea" pageId="2" pageNumber="552" phylum="Arthropoda" rank="family">Jurinidae</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C3430F9FFDDFFDD6DF9FF7ACC76FEF8" authorityName="Martynov" authorityYear="1938" box="[950,1112,230,256]" class="Insecta" family="Glosselytridae" kingdom="Animalia" order="Glosselytrodea" pageId="2" pageNumber="552" phylum="Arthropoda" rank="family">Glosselytridae</taxonomicName>
|
||
remains to be elucidated.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |