411 lines
40 KiB
XML
411 lines
40 KiB
XML
<document ID-DOI="10.1016/j.mycres.2006.05.012" ID-GBIF-Dataset="60fc6c5d-bde4-4e23-b84c-519bc8fefb0e" ID-GBIF-Taxon="159154736" ID-Zenodo-Dep="3366470" approvalRequired="57" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="56" checkinTime="1565675472132" checkinUser="plazi" docAuthor="Frøslev, Tobias G., Jeppesen, Thomas S. & Laessøe, Thomas" docDate="2006" docId="0393543E7A5B4A73FCD3FC8DDE05FC93" docLanguage="en" docName="mycologicalResearch.110.1046-1058.pdf" docOrigin="Mycological Research 110" docStyle="DocumentStyle{}" docTitle="Cortinarius langeorum Frøslev & Jeppesen & Laessøe 2006, sp. nov." docType="treatment" docVersion="8" lastPageId="7" lastPageNumber="1053" masterDocId="FFAA2C467A5D4A74FFF8FFC0DA23FFE3" masterDocTitle="Seven new calochroid and fulvoid species of Cortinarius" masterLastPageNumber="1058" masterPageNumber="1046" pageId="6" pageNumber="1052" updateTime="1643584967406" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Seven new calochroid and fulvoid species of Cortinarius</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Frøslev, Tobias G.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jeppesen, Thomas S.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Laessøe, Thomas</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Mycological Research</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2006</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2006-12-31</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>110</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1046</mods:start>
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<mods:end>1058</mods:end>
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</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1016/j.mycres.2006.05.012</mods:identifier>
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<mods:identifier type="GBIF-Dataset">60fc6c5d-bde4-4e23-b84c-519bc8fefb0e</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3366470</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159154736" LSID="urn:lsid:plazi:treatment:0393543E7A5B4A73FCD3FC8DDE05FC93" httpUri="http://treatment.plazi.org/id/0393543E7A5B4A73FCD3FC8DDE05FC93" lastPageId="7" lastPageNumber="1053" pageId="6" pageNumber="1052">
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<subSubSection pageId="6" pageNumber="1052" type="nomenclature">
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||
<paragraph blockId="6.[811,1449,845,896]" box="[811,1399,845,865]" pageId="6" pageNumber="1052">
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<taxonomicName authority="Froslev & T. S. Jeppesen" authorityName="Frøslev & Jeppesen & Laessøe" authorityYear="2006" box="[811,1309,845,865]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="6" pageNumber="1052" phylum="Basidiomycota" rank="species" species="langeorum" status="sp. nov.">
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<emphasis bold="true" box="[811,1050,845,865]" pageId="6" pageNumber="1052">Cortinarius langeorum</emphasis>
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Frøslev & T. S. Jeppesen
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</taxonomicName>
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,
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<taxonomicNameLabel box="[1317,1399,845,865]" pageId="6" pageNumber="1052" rank="species">
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<emphasis bold="true" box="[1317,1399,845,865]" pageId="6" pageNumber="1052">sp. nov.</emphasis>
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</taxonomicNameLabel>
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</paragraph>
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<paragraph blockId="6.[811,1449,845,896]" box="[811,1080,876,896]" pageId="6" pageNumber="1052">
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MycoBank no.:
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<accessionNumber box="[970,1080,876,896]" httpUri="http://www.mycobank.org/MB/500730" pageId="6" pageNumber="1052">
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<collectionCode box="[970,1003,876,896]" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="6" pageNumber="1052">MB</collectionCode>
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500730
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</accessionNumber>
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</paragraph>
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</subSubSection>
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<subSubSection box="[1264,1449,876,896]" pageId="6" pageNumber="1052" type="description">
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<paragraph blockId="6.[811,1449,845,896]" box="[1264,1449,876,896]" pageId="6" pageNumber="1052">
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(
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<figureCitation box="[1269,1348,876,896]" captionStart="Fig 1" captionStartId="2.[126,158,1858,1878]" captionTargetBox="[191,1384,210,1826]" captionTargetId="figure@2.[191,1385,210,1826]" captionTargetPageId="2" captionText="Fig 1 – Fruit bodies of the new species of Cortinarius. (A) C. albertii, TF2000-058 (holotype). (B) C. chailluzii, TSJ2000-085 (holotype). (C) C. cisticola, TSJ2002-081 (holotype). (D) C. langeorum, TF2000-012 (holotype). (E) C. sancti-felicis, TSJ2002-072 (holotype). (F) C. selandicus, TF2004-104 (holotype). (G) C. vesterholtii, TT1997-009 (holotype). Photographs A, D and G by T.G.F., B, C, E and F by T.S.J." httpUri="https://zenodo.org/record/3366472/files/figure.png" pageId="6" pageNumber="1052">Figs 1D</figureCitation>
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,
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<figureCitation box="[1357,1385,876,896]" captionStart="Fig 2" captionStartId="3.[138,170,1896,1916]" captionTargetBox="[245,1355,178,1864]" captionTargetId="figure@3.[245,1355,178,1864]" captionTargetPageId="3" captionText="Fig 2 – Basidiospores. (A) Cortinarius albertii, TF2000-058 (holotype). (B) C. chailluzii, TSJ2000-085 (holotype). (C) C. cisticola, TSJ2002-081 (holotype). (D) C. langeorum, TF2000-012 (holotype). (E) C. sancti-felicis, TSJ2002-072 (holotype) (F) C. selandicus, TF2004-104 (holotype). (G) C. vesterholtii, TT1997-009 (holotype). (H) C. catharinae, TF2004-105. Scale bar = 10 Mm in each case." httpUri="https://zenodo.org/record/3366474/files/figure.png" pageId="6" pageNumber="1052">2D</figureCitation>
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&
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<figureCitation box="[1415,1443,876,896]" captionStart="Fig 4" captionStartId="4.[811,843,874,894]" captionTargetBox="[814,1448,178,842]" captionTargetId="figure@4.[814,1448,178,842]" captionTargetPageId="4" captionText="Fig 4 – Fruit body habit of the new Cortinarius species. C. albertii, TF2000-058 (holotype). (B) C. chailluzii, TSJ2000- 085 (holotype). (C) C. cisticola, TSJ2002-081 (holotype). (D) C. langeorum, TSJ2003-057 (middle) and TSJ1999-089. (E) C. sancti-felicis, TSJ2002-072 (holotype). (F) C. vesterholtii, TSJ1999-039 (right) and TF2004-101. Drawn to scale. Scale bar = 1 cm in each case." httpUri="https://zenodo.org/record/3366476/files/figure.png" pageId="6" pageNumber="1052">4D</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection box="[843,1335,921,941]" pageId="6" pageNumber="1052" type="etymology">
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<paragraph blockId="6.[843,1335,921,941]" box="[843,1335,921,941]" pageId="6" pageNumber="1052">Etym.: langeorum (Latin), after the family Lange.</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="1052" type="description">
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<paragraph blockId="6.[811,1451,965,1394]" pageId="6" pageNumber="1052">
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Pileo
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<quantity box="[861,971,966,984]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="6" pageNumber="1052" unit="mm" value="75.0" valueMax="100.0" valueMin="50.0">50–100 mm</quantity>
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lato, hemisphaerico, dein plano-convexo, glutinoso, primo luteo, dein a centro ochraceo-luteo vel brunneo, interdum centro e velo pallide virido-flavescenti maculato, in laesionibus safrano-ochrascenti, KOH ope brunneo, dein roseobrunneo. Velo universale virido-flavo, KOH ope obscuro roseobrunneo. Lamellis emarginatis, in juventute pallide eburneis, nec luteis. Stipite pallido, albido, non luteo, bulboso, bulbo distincte marginato. Margine bulbi e velo virido-luteo, KOH ope obscuro roseo-brunneo. Facie externa mycelioque lutea, KOH ope nullo. Velo partiale albido vel virido-flavo. Carne albida, interdum in apice stipitis pallide ochraceo-brunnea, sapore miti, odore subnullo, KOH ope nullo. Sporis limoniformibus, grosse verrucosis, 11-12.5 X
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<quantity box="[904,988,1285,1303]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="8.0" metricValueMin="6.5" pageId="6" pageNumber="1052" unit="mm" value="7.25" valueMax="8.0" valueMin="6.5">6.5-8 µm</quantity>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="1052" type="materials_examined">
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<paragraph blockId="6.[811,1451,965,1394]" pageId="6" pageNumber="1052">
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Typus:
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<materialsCitation ID-GBIF-Occurrence="2331580475" collectedFrom="under Fagus sylvatica" collectingDate="2000-09-15" collectionCode="C" collectorName="T. G. FrOslev" country="Denmark" location="Arhusskovene" municipality="Ostjylland" pageId="6" pageNumber="1052" specimenCode="TF2000- 012" specimenCount="1" typeStatus="holotype">
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<collectingCountry box="[913,1011,1313,1333]" name="Denmark" pageId="6" pageNumber="1052">
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<emphasis bold="true" box="[913,1011,1313,1333]" pageId="6" pageNumber="1052">Denmark</emphasis>
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</collectingCountry>
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:
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<collectingMunicipality box="[1023,1121,1313,1333]" pageId="6" pageNumber="1052">Østjylland</collectingMunicipality>
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:
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<location LSID="urn:lsid:plazi:treatment:0393543E7A5B4A73FCD3FC8DDE05FC93:8EE5B3F37A5B4A72FB94FADDDF27FAD6" box="[1132,1284,1309,1333]" country="Denmark" municipality="Ostjylland" name="Arhusskovene" pageId="6" pageNumber="1052">Århusskovene</location>
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,
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<location LSID="urn:lsid:plazi:treatment:0393543E7A5B4A73FCD3FC8DDE05FC93:8EE5B3F37A5B4A72FAF4FAE1D979FAB0" country="Denmark" municipality="Ostjylland" name="Jens Baeks Vedkast" pageId="6" pageNumber="1052">Jens Baeks Vedkast</location>
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,
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<collectedFrom box="[867,1084,1343,1363]" pageId="6" pageNumber="1052">under Fagus sylvatica</collectedFrom>
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,
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<collectingDate box="[1097,1227,1343,1363]" pageId="6" pageNumber="1052" value="2000-09-15">
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<date box="[1097,1227,1343,1363]" pageId="6" pageNumber="1052" value="2000-09-15">15 Sep. 2000</date>
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</collectingDate>
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,
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<collectorName box="[1236,1357,1343,1363]" pageId="6" pageNumber="1052">T. G. FrØslev</collectorName>
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,
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<specimenCode pageId="6" pageNumber="1052">TF2000- 012</specimenCode>
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(
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<collectionCode box="[858,874,1374,1394]" pageId="6" pageNumber="1052">C</collectionCode>
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–
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<typeStatus box="[897,1002,1374,1394]" pageId="6" pageNumber="1052">holotypus</typeStatus>
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)
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="7" lastPageNumber="1053" pageId="6" pageNumber="1052" type="description">
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<paragraph blockId="6.[811,1450,1431,2001]" lastBlockId="7.[138,777,178,2001]" lastPageId="7" lastPageNumber="1053" pageId="6" pageNumber="1052">
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Pileus 60–120(–140) mm, hemispherical then plano-convex, glutinous, glabrous, with age slightly innately fibrillose, at first yellow (2A4–2A5–3A4–3A5–4A6), soon becoming ochre–yellow (4B6–4B7) then brownish (4C7–5D7), margin remains yellow (3A4–3A5), often with veil remnants seen as pale greenish yellow to whitish patches, or sometimes as small brownish scales, where damaged often more or less saffron-coloured to red brown (7E7). Lamellae emarginate, greyish white to cream when young. Stipe 50–120
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<date box="[1170,1249,1675,1695]" pageId="6" pageNumber="1052" value="1925-10-10">
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X
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<quantity box="[1191,1297,1675,1695]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="6" pageNumber="1052" unit="mm" value="17.5" valueMax="25.0" valueMin="10.0">10–25 mm</quantity>
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</date>
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with a broad marginate bulb (–
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<quantity box="[994,1072,1706,1726]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="1052" unit="mm" value="45.0">45 mm</quantity>
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), whitish. Bulbipellis and mycelium characteristically pale cream yellow to bright greenish yellow (1A3–1A4), bulb margin often bright greenish yellow from veil remnants. Cortina whitish to greenish yellow, fairly abundant. Veil pale greenish yellow to white. Context whitish, sometimes with a very pale greyish brown tinge in stipe apex, a thin greenish yellow line is seen in the lower bulb cortex when the bulbipellis is strongly coloured. Taste mild, indistinct, smell somewhat yeast-like (like many phlegmacioid species, e.g.
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<taxonomicName authorityName="sensu Brandrud et al." box="[1001,1122,1981,2001]" class="Polycystina" family="Stephaniidae" genus="Cortina" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="6" pageNumber="1052" phylum="Radiozoa" rank="species" species="saporatus">C. saporatus</taxonomicName>
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), sometimes with a faint curry component. Spores citriform, with distinct net-like ornamentation, 11–12.5(–13) X
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<quantity box="[351,441,209,229]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="8.0" metricValueMin="6.5" pageId="7" pageNumber="1053" unit="mm" value="7.25" valueMax="8.0" valueMin="6.5">6.5–8 µm</quantity>
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, average 12.0 ± 0.5 X 7.1 ±
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<quantity box="[704,772,209,229]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="1053" unit="mm" value="0.4">0.4 µm</quantity>
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; Q ¼ 1.69 ± 0.07.
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</paragraph>
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<paragraph blockId="7.[138,777,178,2001]" pageId="7" pageNumber="1053">Chemical reactions: KOH on pileus at first brown (7E8), becoming deep red brown (8E8–9E8) after 2–3 min, on universal veil on bulb margin deeper red brown (9E8–10E8), in context and on bulbipellis negative.</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="1053" type="biology_ecology">
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<paragraph blockId="7.[138,777,178,2001]" pageId="7" pageNumber="1053">
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Habitat: Known from temperate forests dominated by Fagus sylvatica on calcareous or mineral rich soil in
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<collectingCountry box="[631,728,423,443]" name="Denmark" pageId="7" pageNumber="1053">Denmark</collectingCountry>
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and
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<collectingCountry box="[138,224,453,473]" name="Sweden" pageId="7" pageNumber="1053">Sweden</collectingCountry>
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, almost always found in bare soil in erosion zones or on soils with a very thin litter layer.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="1053" type="discussion">
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<paragraph blockId="7.[138,777,178,2001]" lastBlockId="7.[824,1463,178,881]" pageId="7" pageNumber="1053">
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Observations/Discussion:
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<taxonomicName authorityName="Frøslev & Jeppesen & Laessøe" authorityYear="2006" box="[428,655,514,534]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="langeorum">Cortinarius langeorum</taxonomicName>
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is distinguished from all other fulvoid (and calochroid) species by the lack of yellow colour on lamellae, context and stipe, a rather strong alkaline reaction on the pileus in combination with the characteristic yellow veil (seen as remnants on the bulb margin) and a pale greenish yellow bulbipellis. It is most similar to the less pigment rich fulvoid species
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<emphasis bold="true" box="[715,736,701,717]" italics="true" pageId="7" pageNumber="1053">̅</emphasis>
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i.e.
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<taxonomicName authorityName="Brandrud" authorityYear="1998" box="[138,255,728,748]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="humolens">C. humolens</taxonomicName>
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(syn.
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<taxonomicName authority="sensu Moser" authorityName="sensu Moser" box="[314,577,728,748]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="claroflavus">C. claroflavus sensu Moser</taxonomicName>
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) and
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<taxonomicName authorityName="in Flora" box="[632,770,728,748]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="sulphurinus">C. sulphurinus</taxonomicName>
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. The most similar species,
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<taxonomicName authorityName="Brandrud" authorityYear="1998" box="[422,542,759,779]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="humolens">C. humolens</taxonomicName>
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, differs in several respects.
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<taxonomicName authorityName="Brandrud" authorityYear="1998" box="[215,333,789,809]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="humolens">C. humolens</taxonomicName>
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has an earth-like smell, whereas
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<taxonomicName authorityName="Frøslev & Jeppesen & Laessøe" authorityYear="2006" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="langeorum">C. langeorum</taxonomicName>
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has a neutral Phlegmacium- like smell, sometimes with a faint curry-like component.
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<taxonomicName authorityName="Brandrud" authorityYear="1998" box="[449,566,850,870]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="humolens">C. humolens</taxonomicName>
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has a weak alkaline reaction (negative to pale brownish) on the pileus, whereas
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<taxonomicName authorityName="Frøslev & Jeppesen & Laessøe" authorityYear="2006" box="[138,271,912,932]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="langeorum">C. langeorum</taxonomicName>
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reacts strongly brown to red–brown. Furthermore,
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<taxonomicName authorityName="Brandrud" authorityYear="1998" box="[210,333,942,962]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="humolens">C. humolens</taxonomicName>
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has yellowish colours on both lamellae and usually also on the stipe, where
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<taxonomicName authorityName="Frøslev & Jeppesen & Laessøe" authorityYear="2006" box="[524,651,973,993]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="langeorum">C. langeorum</taxonomicName>
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has colourless lamellae and stipe contrasting the yellow bulbipellis and veil remnants on the bulb margin.
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<taxonomicName authorityName="in Flora" box="[522,665,1034,1054]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="sulphurinus">C. sulphurinus</taxonomicName>
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differs by the parsley-like smell, greenish yellow colours on the lamellae, stipe and context, and has a strong red alkaline reaction on the mycelial strands.
|
||
<taxonomicName authorityName="Rob. Henry" authorityYear="1939" box="[403,534,1125,1145]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="flavovirens">C. flavovirens</taxonomicName>
|
||
also has weak colours on the lamellae and in the context, but has a strong farinose smell and much smaller spores. Seen from above the pileus of
|
||
<taxonomicName authorityName="Frøslev & Jeppesen & Laessøe" authorityYear="2006" box="[170,304,1217,1237]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="langeorum">C. langeorum</taxonomicName>
|
||
oxidises in a way that makes it resemble more pigment-rich species (e.g.
|
||
<taxonomicName authorityName="Rob. Henry" authorityYear="1952" box="[470,626,1248,1268]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="alcalinophilus">C. alcalinophilus</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Rob. Henry" authorityYear="1987" box="[638,763,1248,1268]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="olearioides">C. olearioides</taxonomicName>
|
||
), which all, however, have yellow context and lamellae. Due to the lack of colours in the context and lamellae, it seems possible that the species could have been treated in the section Multiformes or Calochroi – but no names published in these sections fit the species to our knowledge. Aberrant specimens of
|
||
<taxonomicName authorityName="E. Campo & Vizzini" authorityYear="2011" baseAuthorityName="Chevassut & Rob. Henry" box="[138,319,1431,1451]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="caesiocortinatus">C. caesiocortinatus</taxonomicName>
|
||
may show a certain resemblance, but are easily identified by the (sub-)globose spores. The illustration of
|
||
<taxonomicName authority="in Flora" authorityName="in Flora" box="[165,388,1492,1512]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="sulphurinus">C. sulphurinus in Flora</taxonomicName>
|
||
<taxonomicName authority="Danica" authorityName="Danica" box="[394,571,1492,1512]" class="Anthozoa" family="Agariciidae" genus="Agaricina" kingdom="Animalia" order="Scleractinia" pageId="7" pageNumber="1053" phylum="Cnidaria" rank="genus">Agaricina Danica</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Lange JE" box="[583,697,1492,1512]" journalOrPublisher="Vol. 3 Kobenhavn" pageId="7" pageNumber="1053" refId="ref14063" refString="Lange JE, 1938. Flora Agaricina Danica, Vol. 3 Kobenhavn." title="Flora Agaricina Danica" type="book" year="1938">Lange 1938</bibRefCitation>
|
||
) shows an expanded fruit body with brownish yellow colours on a pileus with velar patches. The original description of
|
||
<taxonomicName authorityName="in Flora" box="[138,279,1584,1604]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="sulphurinus">C. sulphurinus</taxonomicName>
|
||
presents a more pigment rich species than the one collected and illustrated by Lange. Presently
|
||
<taxonomicName authority="sensu Lange" authorityName="sensu Lange. Contrarily" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="sulphurinus">C. sulphurinus sensu Lange</taxonomicName>
|
||
is widely perceived as a synonym of
|
||
<taxonomicName authority="sensu Brandrud et al." authorityName="sensu Brandrud et al." class="Polycystina" family="Stephaniidae" genus="Cortina" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="7" pageNumber="1053" phylum="Radiozoa" rank="species" species="saporatus">C. saporatus sensu Brandrud et al.</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Brandrud TE & Lindstrom H & Marklund H & Melot J & Muskos S" box="[364,564,1675,1695]" journalOrPublisher="Cortinarius HB, Matfors" pageId="7" pageNumber="1053" refId="ref13736" refString="Brandrud TE, Lindstrom H, Marklund H, Melot J, Muskos S, 1989 - 1998. Cortinarius Flora Photographica I-IV. Cortinarius HB, Matfors." title="Cortinarius Flora Photographica I-IV" type="book" year="1989">
|
||
Brandrud et al.
|
||
<date box="[517,621,1675,1695]" pageId="7" pageNumber="1053" value="1989" valueMax="1998">
|
||
1989–
|
||
<bibRefCitation author="Brandrud TE" box="[574,621,1675,1695]" journalOrPublisher="Journal des J. E. C" pageId="7" pageNumber="1053" pagination="4 - 9" part="0" refId="ref13705" refString="Brandrud TE, 1998. Cortinarius subgen. Phlegmacium sect. Fulvi ‾ chemotaxonomy versus morphological taxonomy. Journal des J. E. C 0: 4 - 9." title="Cortinarius subgen. Phlegmacium sect. Fulvi ‾ chemotaxonomy versus morphological taxonomy" type="book" year="1998">1998</bibRefCitation>
|
||
</date>
|
||
</bibRefCitation>
|
||
) syn.
|
||
<taxonomicName authorityName="Rob.Henry ex P.D Orton" authorityYear="1939" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="subturbinatus">C. subturbinatus</taxonomicName>
|
||
. This interpretation, however, has to be rejected as the drawing of Lange clearly shows an agaric with bright yellow colours on the bulb. In the many years after Lange drew his plate
|
||
<taxonomicName authorityName="sensu Brandrud et al." box="[196,315,1797,1817]" class="Polycystina" family="Stephaniidae" genus="Cortina" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="7" pageNumber="1053" phylum="Radiozoa" rank="species" species="saporatus">C. saporatus</taxonomicName>
|
||
has never been found on or near the locality of
|
||
<taxonomicName authority="sensu Lange. Contrarily" authorityName="sensu Lange. Contrarily" box="[164,555,1828,1848]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="sulphurinus">C. sulphurinus sensu Lange. Contrarily</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Frøslev & Jeppesen & Laessøe" authorityYear="2006" box="[564,691,1828,1848]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="langeorum">C. langeorum</taxonomicName>
|
||
has one of its localities there. Therefore, as both the macroscopic and microscopic features fits well with
|
||
<taxonomicName authorityName="Frøslev & Jeppesen & Laessøe" authorityYear="2006" box="[516,646,1889,1909]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="langeorum">C. langeorum</taxonomicName>
|
||
, we dare to postulate that
|
||
<taxonomicName authority="sensu Lange" authorityName="sensu Lange. Contrarily" box="[286,556,1920,1940]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="sulphurinus">C. sulphurinus sensu Lange</taxonomicName>
|
||
has found its final interpretation. It has only been found in localities hosting several other rare species of phlegmacioid
|
||
<taxonomicName authorityName="S.F.Gray" authorityYear="1821" baseAuthorityName="Person" box="[607,714,1981,2001]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="genus">Cortinarius</taxonomicName>
|
||
, and seems to be among the top ten rarest phlegmacioid species in
|
||
<collectingCountry box="[849,949,209,229]" name="Denmark" pageId="7" pageNumber="1053">Denmark</collectingCountry>
|
||
. Although the species is well known to us and easily recognisable, we have never encountered the species in southern parts of Europe or with other hosts than Fagus sylvatica, nor in the literature. We have chosen to name our species after the family Lange, which has given us three generations of prominent figures in mycology, and especially as an appreciation of Jakob E. Lange’s monumental work on agarics. The species was included in the study of
|
||
<bibRefCitation author="Froslev TG & Matheny PB & Hibbett D" box="[1227,1430,423,443]" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="7" pageNumber="1053" pagination="602 - 618" part="37" refId="ref13830" refString="Froslev TG, Matheny PB, Hibbett D, 2005. Lower level relationships in the mushroom genus Cortinarius (Basidiomycota, Agaricales): A comparison of RPB 1, RPB 2, and ITS phylogenies. Molecular Phylogenetics and Evolution 37: 602 - 618." title="Lower level relationships in the mushroom genus Cortinarius (Basidiomycota, Agaricales): A comparison of RPB 1, RPB 2, and ITS phylogenies" type="journal article" year="2005">Frøslev et al. (2005)</bibRefCitation>
|
||
as C. neolangei ined. (
|
||
<collectionCode box="[1014,1042,453,473]" country="Thailand" name="Department of Mineral Resources" pageId="7" pageNumber="1053">TF</collectionCode>
|
||
2000-012, TSJ2003-078 and TSJ2003-057). An ITS sequence from the
|
||
<typeStatus box="[1094,1185,484,504]" pageId="7" pageNumber="1053">holotype</typeStatus>
|
||
of
|
||
<taxonomicName authorityName="Frøslev & Jeppesen & Laessøe" authorityYear="2006" box="[1216,1342,484,504]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="1053" phylum="Basidiomycota" rank="species" species="langeorum">C. langeorum</taxonomicName>
|
||
is available at GenBank (accession
|
||
<accessionNumber box="[1063,1166,514,534]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/DQ083798" pageId="7" pageNumber="1053">
|
||
<collectionCode box="[1063,1093,514,534]" pageId="7" pageNumber="1053">DQ</collectionCode>
|
||
083798
|
||
</accessionNumber>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="7" pageNumber="1053" type="materials_examined">
|
||
<paragraph blockId="7.[824,1463,178,881]" pageId="7" pageNumber="1053">
|
||
Additional specimens examined:
|
||
<materialsCitation ID-GBIF-Occurrence="2459009034" collectedFrom="Fagus sylvatica on calcareous soil" collectingDate="2003-10-04" collectionCode="TSJ" collectorName="T. S. J." country="Denmark" location="Allindelille Fredskov" municipality="Allindelille Fredskov" pageId="7" pageNumber="1053" specimenCode="TSJ2003-093" specimenCount="1" stateProvince="Sjaelland">
|
||
<collectingCountry box="[1152,1245,544,562]" name="Denmark" pageId="7" pageNumber="1053">
|
||
<emphasis bold="true" box="[1152,1245,544,562]" pageId="7" pageNumber="1053">Denmark</emphasis>
|
||
</collectingCountry>
|
||
:
|
||
<collectingRegion box="[1262,1338,543,562]" country="Denmark" name="Sjaelland" pageId="7" pageNumber="1053">Sjaelland</collectingRegion>
|
||
:
|
||
<collectingMunicipality pageId="7" pageNumber="1053">Allindelille Fredskov</collectingMunicipality>
|
||
,
|
||
<collectedFrom box="[927,1267,569,588]" pageId="7" pageNumber="1053">Fagus sylvatica on calcareous soil</collectedFrom>
|
||
,
|
||
<date box="[1280,1397,570,588]" pageId="7" pageNumber="1053" value="2003-10-04">
|
||
<collectingDate box="[1280,1397,570,588]" pageId="7" pageNumber="1053" value="2003-10-04">4 Oct. 2003</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[1409,1457,569,588]" pageId="7" pageNumber="1053">T.S.J.</collectorName>
|
||
,
|
||
<specimenCode box="[824,938,596,615]" pageId="7" pageNumber="1053">TSJ2003-093</specimenCode>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="2459009041" box="[948,1395,596,615]" collectingDate="2004-09-20" collectionCode="TSJ" collectorName="T. G. F. & T. S. J." country="Denmark" pageId="7" pageNumber="1053" specimenCode="TSJ2004-011" specimenCount="1">
|
||
loc.cit.,
|
||
<date box="[1014,1133,596,615]" pageId="7" pageNumber="1053" value="2004-09-20">
|
||
<collectingDate box="[1014,1133,596,615]" pageId="7" pageNumber="1053" value="2004-09-20">20 Sep. 2004</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[1140,1196,596,615]" pageId="7" pageNumber="1053">T.G.F.</collectorName>
|
||
&
|
||
<collectorName box="[1219,1267,596,615]" pageId="7" pageNumber="1053">T.S.J.</collectorName>
|
||
,
|
||
<specimenCode box="[1275,1389,596,615]" pageId="7" pageNumber="1053">TSJ2004-011</specimenCode>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="2459009032" collectingDate="2004-10-08" collectionCode="TSJ" collectorName="T. S. J. & T. G. F." pageId="7" pageNumber="1053" specimenCode="TSJ2004-065" specimenCount="1">
|
||
loc.cit.,
|
||
<date box="[824,934,623,641]" pageId="7" pageNumber="1053" value="2004-10-08">
|
||
<collectingDate box="[824,934,623,641]" pageId="7" pageNumber="1053" value="2004-10-08">8 Oct. 2004</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[944,991,623,642]" pageId="7" pageNumber="1053">T.S.J.</collectorName>
|
||
&
|
||
<collectorName box="[1019,1074,623,642]" pageId="7" pageNumber="1053">T.G.F.</collectorName>
|
||
,
|
||
<specimenCode box="[1085,1200,623,642]" pageId="7" pageNumber="1053">TSJ2004-065</specimenCode>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="2459009036" collectedFrom="F. sylvatica on mineral rich to calcareous soil" collectingDate="2003-09-19" collectionCode="TSJ" collectorName="T. S. J." location="Lellinge Skovhusvaenge at Koge" pageId="7" pageNumber="1053" specimenCode="TSJ2003-023" specimenCount="1">
|
||
loc. cit.,
|
||
<location LSID="urn:lsid:plazi:treatment:0393543E7A5B4A73FCD3FC8DDE05FC93:8EE5B3F37A5A4A73FAFFFDAFD9EFFD7F" name="Lellinge Skovhusvaenge at Koge" pageId="7" pageNumber="1053">Lellinge Skovhusvaenge at Køge</location>
|
||
,
|
||
<collectedFrom box="[982,1429,649,668]" pageId="7" pageNumber="1053">F. sylvatica on mineral rich to calcareous soil</collectedFrom>
|
||
,
|
||
<date pageId="7" pageNumber="1053" value="2003-09-19">
|
||
<collectingDate pageId="7" pageNumber="1053" value="2003-09-19">19 Sep. 2003</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[926,974,676,695]" pageId="7" pageNumber="1053">T.S.J.</collectorName>
|
||
,
|
||
<specimenCode box="[984,1099,676,695]" pageId="7" pageNumber="1053">TSJ2003-023</specimenCode>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="2459009031" collectingDate="2003-09-25" collectionCode="TSJ" collectorName="T. S. J." pageId="7" pageNumber="1053" specimenCode="TSJ2003- 057, TSJ2003-058" specimenCount="1">
|
||
loc. cit.,
|
||
<date box="[1184,1314,676,694]" pageId="7" pageNumber="1053" value="2003-09-25">
|
||
<collectingDate box="[1184,1314,676,694]" pageId="7" pageNumber="1053" value="2003-09-25">25 Sept. 2003</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[1322,1370,676,695]" pageId="7" pageNumber="1053">T.S.J.</collectorName>
|
||
,
|
||
<specimenCode pageId="7" pageNumber="1053">TSJ2003- 057</specimenCode>
|
||
,
|
||
<specimenCode box="[868,982,702,721]" pageId="7" pageNumber="1053">TSJ2003-058</specimenCode>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="2459009039" box="[993,1359,702,721]" collectingDate="2003-10-05" collectionCode="TSJ" collectorName="T. S. J" pageId="7" pageNumber="1053" specimenCode="TSJ2003-098" specimenCount="1">
|
||
loc. cit.,
|
||
<date box="[1066,1173,703,721]" pageId="7" pageNumber="1053" value="2003-10-05">
|
||
<collectingDate box="[1066,1173,703,721]" pageId="7" pageNumber="1053" value="2003-10-05">5 Oct. 2003</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[1181,1225,702,721]" pageId="7" pageNumber="1053">T.S.J</collectorName>
|
||
.,
|
||
<specimenCode box="[1239,1353,702,721]" pageId="7" pageNumber="1053">TSJ2003-098</specimenCode>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="2459009033" collectedFrom="F. sylvatica on mineral rich soil" collectingDate="1997-10-08" collectionCode="TT" collectorName="T. G. F. & T. S. J." country="Denmark" location="Arhusskovene" municipality="Ostjylland" pageId="7" pageNumber="1053" specimenCode="TT1997-040" specimenCount="1">
|
||
<collectingMunicipality box="[1364,1456,702,721]" pageId="7" pageNumber="1053">Østjylland</collectingMunicipality>
|
||
:
|
||
<location LSID="urn:lsid:plazi:treatment:0393543E7A5B4A73FCD3FC8DDE05FC93:8EE5B3F37A5A4A73FCC0FD14D9E4FD09" box="[824,967,724,746]" municipality="Ostjylland" name="Arhusskovene" pageId="7" pageNumber="1053">Århusskovene</location>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:0393543E7A5B4A73FCD3FC8DDE05FC93:8EE5B3F37A5A4A73FC36FD19DEA6FD08" box="[974,1157,729,747]" municipality="Ostjylland" name="Jens Baeks Vedkast" pageId="7" pageNumber="1053">Jens Baeks Vedkast</location>
|
||
,
|
||
<collectedFrom box="[1163,1458,729,748]" pageId="7" pageNumber="1053">F. sylvatica on mineral rich soil</collectedFrom>
|
||
,
|
||
<date box="[824,932,756,774]" pageId="7" pageNumber="1053" value="1997-10-08">
|
||
<collectingDate box="[824,932,756,774]" pageId="7" pageNumber="1053" value="1997-10-08">8 Oct. 1997</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[940,995,755,774]" pageId="7" pageNumber="1053">T.G.F.</collectorName>
|
||
&
|
||
<collectorName box="[1021,1069,755,774]" pageId="7" pageNumber="1053">T.S.J.</collectorName>
|
||
,
|
||
<specimenCode box="[1079,1188,755,774]" pageId="7" pageNumber="1053">TT1997-040</specimenCode>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="2459009035" collectingDate="1999-10-16" collectionCode="TSJ" collectorName="T. S. J. & T. G. F." pageId="7" pageNumber="1053" specimenCode="TSJ1999-089" specimenCount="1">
|
||
loc.cit.,
|
||
<date box="[1266,1386,756,774]" pageId="7" pageNumber="1053" value="1999-10-16">
|
||
<collectingDate box="[1266,1386,756,774]" pageId="7" pageNumber="1053" value="1999-10-16">16 Oct. 1999</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[1394,1442,755,774]" pageId="7" pageNumber="1053">T.S.J.</collectorName>
|
||
&
|
||
<collectorName box="[824,879,782,801]" pageId="7" pageNumber="1053">T.G.F.</collectorName>
|
||
,
|
||
<specimenCode box="[890,1004,782,801]" pageId="7" pageNumber="1053">TSJ1999-089</specimenCode>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="2459009042" box="[1017,1207,782,801]" pageId="7" pageNumber="1053">
|
||
loc. cit.,
|
||
<specimenCode box="[1093,1170,782,801]" pageId="7" pageNumber="1053">
|
||
<collectionCode box="[1093,1128,782,801]" collectionName="TSJ" pageId="7" pageNumber="1053">TSJ</collectionCode>
|
||
1999
|
||
</specimenCode>
|
||
-090
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2459009037" collectingDate="2004-10-03" collectionCode="TSJ" collectorName="T. S. J." pageId="7" pageNumber="1053" specimenCode="TSJ2004-058" specimenCount="1">
|
||
loc.cit.,
|
||
<date box="[1289,1400,782,801]" pageId="7" pageNumber="1053" value="2004-10-03">
|
||
<collectingDate box="[1289,1400,782,801]" pageId="7" pageNumber="1053" value="2004-10-03">3 Oct. 2004</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[1409,1457,782,801]" pageId="7" pageNumber="1053">T.S.J.</collectorName>
|
||
,
|
||
<specimenCode box="[824,938,808,827]" pageId="7" pageNumber="1053">TSJ2004-058</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
<emphasis bold="true" box="[951,1056,809,827]" pageId="7" pageNumber="1053">
|
||
<emphasis bold="true" box="[951,971,812,827]" italics="true" pageId="7" pageNumber="1053">̅</emphasis>
|
||
<materialsCitation ID-GBIF-Occurrence="2459009040" collectedFrom="F. sylvatica on mineral rich soil" collectingDate="2003-09-30" collectionCode="TSJ" collectorName="B. Hagg & T. S. J." country="Sweden" location="Vittskovle" pageId="7" pageNumber="1053" specimenCode="TSJ2003- 078" specimenCount="1" stateProvince="Skane">
|
||
<collectingCountry box="[978,1056,809,827]" name="Sweden" pageId="7" pageNumber="1053">Sweden</collectingCountry>
|
||
:
|
||
<collectingRegion box="[1069,1122,808,827]" country="Sweden" name="Skane" pageId="7" pageNumber="1053">Skåne</collectingRegion>
|
||
:
|
||
<location LSID="urn:lsid:plazi:treatment:0393543E7A5B4A73FCD3FC8DDE05FC93:8EE5B3F37A5A4A73FB97FCEADEF7FCDF" box="[1135,1236,810,828]" country="Sweden" name="Vittskovle" pageId="7" pageNumber="1053" stateProvince="Skane">Vittskövle</location>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:0393543E7A5B4A73FCD3FC8DDE05FC93:8EE5B3F37A5A4A73FB26FCEADF7EFCDF" box="[1246,1373,810,828]" country="Sweden" name="Herremollan" pageId="7" pageNumber="1053" stateProvince="Skane">Herremöllan</location>
|
||
,
|
||
<collectedFrom pageId="7" pageNumber="1053">F. sylvatica on mineral rich soil</collectedFrom>
|
||
,
|
||
<date box="[1077,1204,836,854]" pageId="7" pageNumber="1053" value="2003-09-30">
|
||
<collectingDate box="[1077,1204,836,854]" pageId="7" pageNumber="1053" value="2003-09-30">30 Sep. 2003</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[1215,1368,835,854]" pageId="7" pageNumber="1053">B. Hägg & T.S.J.</collectorName>
|
||
,
|
||
<specimenCode pageId="7" pageNumber="1053">TSJ2003- 078</specimenCode>
|
||
</materialsCitation>
|
||
</emphasis>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2459009044" box="[868,1062,862,881]" collectionCode="TSJ" pageId="7" pageNumber="1053" specimenCode="TSJ2003-079" specimenCount="1">
|
||
loc. cit.,
|
||
<specimenCode box="[942,1057,862,881]" pageId="7" pageNumber="1053">TSJ2003-079</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |