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<document id="A8AC93955B5027180A6E029B20207AA2" ID-CLB-Dataset="199799" ID-DOI="10.11646/zootaxa.5214.3.4" ID-GBIF-Dataset="9c8b2ed9-a132-4df0-afa9-b153547b9027" ID-ISSN="1175-5326" ID-Zenodo-Dep="7389267" ID-ZooBank="395A3FB5-DA5E-4145-B9EA-7801C67D6CE3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669970496187" checkinUser="plazi" docAuthor="Huynh, Cuong" docDate="2022" docId="038987F7BC1C087C9495FA4F0CB99AA6" docLanguage="en" docName="zootaxa.5214.3.4.pdf" docOrigin="Zootaxa 5214 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Monographis tamdaoensis Huynh 2022, new species" docType="treatment" docVersion="7" lastPageNumber="409" masterDocId="FFB0FF8FBC12086C9402FF890C769D01" masterDocTitle="Four new species of Monographis Attems, 1907 (Diplopoda, Polyxenida, Polyxenidae) from Vietnam" masterLastPageNumber="420" masterPageNumber="393" pageNumber="407" updateTime="1699412757114" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="326131B5CEA464CEFAEB85FF4842CBF5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="85EC284EC8978C57B822CDACFDBD4CA4">
<mods:title id="5A73510AF027AE9742D7F13E8754CDCD">Four new species of Monographis Attems, 1907 (Diplopoda, Polyxenida, Polyxenidae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="5FC5714FEF9051D30DEDD7EEEDE556FB" type="personal">
<mods:role id="74FCC2DEF3F12345906C797B4BB4E6D8">
<mods:roleTerm id="93122F42D3E63D75B13B22219BF94F0E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="24736904DC2A04CFA73CAB30ADF8FC58">Huynh, Cuong</mods:namePart>
<mods:affiliation id="54CC79C2F44B981B0B135713BCD19901">Centre for Cellular and Molecular Biology (CCMB), Deakin University, 221 Burwood Hwy, Burwood, Melbourne, Australia 3125</mods:affiliation>
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<mods:titleInfo id="39418FCCF64A164EA024DD301B15378B">
<mods:title id="7314D99AF1D4E150FF7C06A92F8D671A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6F9D5AEF2C12236BE967C624BB0B5306">
<mods:date id="7DFB2998AAC8AA977509B88E13059027">2022</mods:date>
<mods:detail id="7687E833150E95ED74EEFA2C4510B9C7" type="pubDate">
<mods:number id="DFF5E0D6516E273B166DB5D68DA0E01E">2022-12-02</mods:number>
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<mods:number id="F84C4C8EA2F31FE455ACFB5211C2636C">5214</mods:number>
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<mods:number id="D657DE3382E3DC000DECA665753B18CB">3</mods:number>
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<mods:start id="F0B819876694C216FA44B9D899350715">393</mods:start>
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<mods:classification id="5F367A8BBDDDE606B56703C21FA3AC49">journal article</mods:classification>
<mods:identifier id="B71F3DCE31523490F56CE33A2569BB4E" type="CLB-Dataset">199799</mods:identifier>
<mods:identifier id="8844F03A3210EA3F433D9188972D7AC5" type="DOI">10.11646/zootaxa.5214.3.4</mods:identifier>
<mods:identifier id="7D0152A781DC0593DCBF87E909A03AEA" type="GBIF-Dataset">9c8b2ed9-a132-4df0-afa9-b153547b9027</mods:identifier>
<mods:identifier id="7323C927D432D8E765535EEEC6AA59B7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6652AD7D5E33ACF110082D1E6B82FD5D" type="Zenodo-Dep">7389267</mods:identifier>
<mods:identifier id="2FAD2EDE9381D9F93B3C2D3E01195DBD" type="ZooBank">395A3FB5-DA5E-4145-B9EA-7801C67D6CE3</mods:identifier>
</mods:mods>
<treatment id="038987F7BC1C087C9495FA4F0CB99AA6" ID-DOI="http://doi.org/10.5281/zenodo.7389482" ID-GBIF-Taxon="205697559" ID-Zenodo-Dep="7389482" LSID="urn:lsid:plazi:treatment:038987F7BC1C087C9495FA4F0CB99AA6" httpUri="http://treatment.plazi.org/id/038987F7BC1C087C9495FA4F0CB99AA6" lastPageId="16" lastPageNumber="409" pageId="14" pageNumber="407">
<subSubSection id="C33A656ABC1C08629495FA4F0E1998E1" box="[151,623,1478,1505]" pageId="14" pageNumber="407" type="nomenclature">
<paragraph id="8B9F36E1BC1C08629495FA4F0E1998E1" blockId="14.[151,623,1478,1540]" box="[151,623,1478,1505]" pageId="14" pageNumber="407">
<heading id="D0D7818DBC1C08629495FA4F0E1998E1" bold="true" box="[151,623,1478,1505]" fontSize="11" level="1" pageId="14" pageNumber="407" reason="1">
<emphasis id="B954EAF3BC1C08629495FA4F0E1998E1" bold="true" box="[151,623,1478,1505]" pageId="14" pageNumber="407">
<taxonomicName id="4C204D62BC1C08629495FA4F0DAE98E0" authority="Huynh, 2022" authorityName="Huynh" authorityYear="2022" box="[151,472,1478,1505]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="tamdaoensis" status="sp. nov.">
<emphasis id="B954EAF3BC1C08629495FA4F0DAE98E0" bold="true" box="[151,472,1478,1505]" italics="true" pageId="14" pageNumber="407">Monographis tamdaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2675788BC1C086295DDFA4F0E1998E1" box="[479,623,1478,1504]" pageId="14" pageNumber="407" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33A656ABC1C08629495FA630D4E9B05" box="[151,312,1514,1540]" pageId="14" pageNumber="407" type="description">
<paragraph id="8B9F36E1BC1C08629495FA630D4E9B05" blockId="14.[151,623,1478,1540]" box="[151,312,1514,1540]" pageId="14" pageNumber="407">
<figureCitation id="131B2A64BC1C08629495FA630D4E9B05" box="[151,312,1514,1540]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStart-2="FIGURE 15" captionStart-3="FIGURE 16" captionStart-4="FIGURE 17" captionStartId-0="15.[152,255,1898,1922]" captionStartId-1="16.[152,255,1358,1382]" captionStartId-2="17.[152,255,1104,1128]" captionStartId-3="17.[152,255,1906,1930]" captionStartId-4="18.[152,255,1080,1104]" captionTargetBox-0="[170,1418,426,1875]" captionTargetBox-1="[151,1417,189,1334]" captionTargetBox-2="[290,1295,188,1070]" captionTargetBox-3="[312,1274,1225,1882]" captionTargetBox-4="[152,1437,182,1056]" captionTargetId-0="figure-136@15.[170,1418,417,1875]" captionTargetId-1="figure-29@16.[151,1436,181,1334]" captionTargetId-2="figure-19@17.[289,1298,181,1080]" captionTargetId-3="figure-91@17.[312,1274,1225,1882]" captionTargetId-4="figure-29@18.[151,1437,181,1056]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="17" captionTargetPageId-4="18" captionText-0="FIGURE 13. Holotype male Monographis tamdaoensis sp. n.. A. Head capsule with eye patterns, posterior vertex (pv) trichome sockets and the trichobothria (a, b and c). B. Enlarged images of Posterior vertex trichome sockets on left and right side; C. Clypeo-labrum with asetose labrum surface (l), lateral labrum (Ll) on each side, anterior setae (sa), posterior setae (Se), backward pointing setae (sb)." captionText-1="FIGURE 14. Holotype male Monographis tamdaoensis sp. n.. A. Section of antennal articles VI–VII and sensory cones (sc), and their sensilla. B. Sensilla on the antennal article VII, with two thick bacilliform sensilla (Ta &amp; Tp), setiform (S) and a conical sensillum (C). C. Sensilla on the article VI, showing thick (T), thin (t) and medium thin bacilliform sensilla (tm)." captionText-2="FIGURE 15. Holotype male Monographis tamdaoensis sp. n.. A. The second leg showing its segments and chaetotaxy, penis (p). B. Telotarsus showing the claw (c) and processes: anterior (a), posterior (p), lamella (la) and anterior setiform process (s) shorter than half length of the claw (dot line). C. The ornamental trichome sockets with trichomes a, b, three c and an indented d." captionText-3="FIGURE 16. SEM images of Monographis tamdaoensis sp. n.. A. Posterior vertex (pv) showing the arrangement of trichome sockets. B. Left eye pattern with eight ommatidia and the trichobothria: a, b and c. C. Gnathochilarium showing the medial palp (Mp) andlateral palp (Lp) with 11 sensilla. D. Clypeo-labrum showing asetose surface (l), backward pointing setae (sb), anterior setae (sa), posterior setae (Se), lateral labrum (Ll) and the medial cleft (mc)." captionText-4="FIGURE 17. SEM images of Monographis tamdaoensis sp. n.. A. Antennal article VI–VIII. B. Arrangement of a setiform (S), conical sensillum (C), thick (T), thin (t) bacilliform sensilla, and the medium length bacilliform sensilla (tm) on article VI. C. Other end of arrangement of sensilla on article VI. D. Telotarsus and its processes: claw (c), anterior setiform process (s), lamella (la), anterior (a) and posterior (p) processes." figureDoi-0="http://doi.org/10.5281/zenodo.7389327" figureDoi-1="http://doi.org/10.5281/zenodo.7389331" figureDoi-2="http://doi.org/10.5281/zenodo.7389333" figureDoi-3="http://doi.org/10.5281/zenodo.7389337" figureDoi-4="http://doi.org/10.5281/zenodo.7389341" httpUri-0="https://zenodo.org/record/7389327/files/figure.png" httpUri-1="https://zenodo.org/record/7389331/files/figure.png" httpUri-2="https://zenodo.org/record/7389333/files/figure.png" httpUri-3="https://zenodo.org/record/7389337/files/figure.png" httpUri-4="https://zenodo.org/record/7389341/files/figure.png" pageId="14" pageNumber="407">Figures 13–17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33A656ABC1C08629495F9BB0FEC9BDD" pageId="14" pageNumber="407" type="materials_examined">
<paragraph id="8B9F36E1BC1C08629495F9BB0FEC9BDD" blockId="14.[151,1437,1586,2009]" pageId="14" pageNumber="407">
<emphasis id="B954EAF3BC1C08629495F9BB0D0F9B4D" bold="true" box="[151,377,1586,1612]" pageId="14" pageNumber="407">Material examined</emphasis>
:
<materialsCitation id="3B483CBCBC1C0862958AF9BB0FEC9BDD" ID-GBIF-Occurrence="3981599307" collectingDate="2017-07-17" country="Vietnam" county="Dr Duc Anh Nguyen" elevation="1017" latitude="21.45861" location="Tam Dao National Park" longLatPrecision="1" longitude="105.649445" municipality="Specimens" pageId="14" pageNumber="407" specimenCount="3" specimenCount-female="1" specimenCount-male="1" specimenCount-subadult="1" stateProvince="Vinh Phuc" typeStatus="holotype">
<typeStatus id="549B8843BC1C0862958AF9BB0D859B4D" box="[392,499,1586,1612]" pageId="14" pageNumber="407" type="holotype">Holotype</typeStatus>
, adult male (IEBR-Myr 989) and
<typeStatus id="549B8843BC1C0862976CF9BA0FA89B4C" box="[878,990,1587,1613]" pageId="14" pageNumber="407" type="paratype">paratypes</typeStatus>
: Adult,
<specimenCount id="9D26FD68BC1C08629037F9BA08D19B4D" box="[1077,1191,1586,1613]" pageId="14" pageNumber="407" type="male">two males</specimenCount>
(IEBR-Myr 990)
<specimenCount id="9D26FD68BC1C08629170F9BA0C989B71" pageId="14" pageNumber="407" type="female">two females</specimenCount>
(IEBR-Myr 991) and,
<specimenCount id="9D26FD68BC1C086295F1F9DE0EF99B71" box="[499,655,1622,1649]" pageId="14" pageNumber="407" type="subadult">two subadults</specimenCount>
with 12 pairs of legs (IEBR-Myr 992) were collected from
<location id="8EFF603ABC1C08629136F9DE0D459B95" LSID="urn:lsid:plazi:treatment:038987F7BC1C087C9495FA4F0CB99AA6:8EFF603ABC1C08629136F9DE0D459B95" country="Vietnam" county="Dr Duc Anh Nguyen" latitude="21.45861" longLatPrecision="1" longitude="105.649445" municipality="Specimens" name="Tam Dao National Park" pageId="14" pageNumber="407" stateProvince="Vinh Phuc">Tam Dao National Park</location>
,
<geoCoordinate id="EE145026BC1C0862953FF9F30DBC9B94" box="[317,458,1658,1685]" degrees="21.458611" direction="north" orientation="latitude" pageId="14" pageNumber="407" precision="1" value="21.45861">21.458611N</geoCoordinate>
,
<geoCoordinate id="EE145026BC1C086295D6F9F30E1B9B94" box="[468,621,1658,1685]" degrees="105.649444" direction="east" orientation="longitude" pageId="14" pageNumber="407" precision="1" value="105.649445">105.649444E</geoCoordinate>
,
<elevation id="000DD1D2BC1C08629675F9F30F4F9B94" box="[631,825,1658,1685]" metricMagnitude="3" metricUnit="m" metricValue="1.017" pageId="14" pageNumber="407" unit="m" value="1017.0">
elevation
<quantity id="4CD89B04BC1C086296E6F9F30F4F9B94" box="[740,825,1658,1685]" metricMagnitude="3" metricUnit="m" metricValue="1.017" pageId="14" pageNumber="407" unit="m" value="1017.0">1017 m</quantity>
</elevation>
,
<collectingRegion id="49E4F803BC1C08629741F9F308559B95" box="[835,1059,1658,1684]" country="Vietnam" name="Vinh Phuc" pageId="14" pageNumber="407">Vinh Phuc Province</collectingRegion>
, northern
<collectingCountry id="F3377671BC1C08629093F9F308829B95" box="[1169,1268,1658,1684]" name="Vietnam" pageId="14" pageNumber="407">Vietnam</collectingCountry>
.
<location id="8EFF603ABC1C086290F9F9F309699B95" LSID="urn:lsid:plazi:treatment:038987F7BC1C087C9495FA4F0CB99AA6:8EFF603ABC1C086290F9F9F309699B95" box="[1275,1311,1658,1684]" country="Vietnam" county="Dr Duc Anh Nguyen" latitude="21.45861" longLatPrecision="1" longitude="105.649445" municipality="Specimens" name="All" pageId="14" pageNumber="407" stateProvince="Vinh Phuc">All</location>
types were deposited in the
<location id="8EFF603ABC1C08629550F9170E449BB9" LSID="urn:lsid:plazi:treatment:038987F7BC1C087C9495FA4F0CB99AA6:8EFF603ABC1C08629550F9170E449BB9" box="[338,562,1694,1720]" country="Vietnam" county="Dr Duc Anh Nguyen" latitude="21.45861" longLatPrecision="1" longitude="105.649445" municipality="Specimens" name="Institute of Ecology" pageId="14" pageNumber="407" stateProvince="Vinh Phuc">Institute of Ecology</location>
and
<location id="8EFF603ABC1C08629668F9170F2B9BB8" LSID="urn:lsid:plazi:treatment:038987F7BC1C087C9495FA4F0CB99AA6:8EFF603ABC1C08629668F9170F2B9BB8" box="[618,861,1694,1721]" country="Vietnam" county="Dr Duc Anh Nguyen" latitude="21.45861" longLatPrecision="1" longitude="105.649445" municipality="Specimens" name="Biological Resources" pageId="14" pageNumber="407" stateProvince="Vinh Phuc">Biological Resources</location>
,
<collectingCountry id="F3377671BC1C0862976AF9170FBD9BB9" box="[872,971,1694,1720]" name="Vietnam" pageId="14" pageNumber="407">Vietnam</collectingCountry>
.
<collectingMunicipality id="6BFBAC9BBC1C086297D4F91708399BB9" box="[982,1103,1694,1720]" pageId="14" pageNumber="407">Specimens</collectingMunicipality>
were collected by author CH in the field trip organised by
<collectingCounty id="62FE4E6DBC1C086295DEF94A0EB19BDC" box="[476,711,1730,1757]" pageId="14" pageNumber="407">Dr Duc Anh Nguyen</collectingCounty>
, on
<date id="FF9E1021BC1C086296F7F94B0FE19BDD" box="[757,919,1730,1756]" pageId="14" pageNumber="407" value="2017-07-17">
<collectingDate id="EFDAE9C9BC1C086296F7F94B0FE19BDD" box="[757,919,1730,1756]" pageId="14" pageNumber="407" value="2017-07-17">
17
<superScript id="7C559BA9BC1C08629713F94B0F689BD1" attach="left" box="[785,798,1730,1744]" fontSize="6" pageId="14" pageNumber="407">th</superScript>
July 2017
</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33A656ABC1C086294C5F96F09F69A6D" pageId="14" pageNumber="407" type="diagnosis">
<paragraph id="8B9F36E1BC1C086294C5F96F09F69A6D" blockId="14.[151,1437,1586,2009]" pageId="14" pageNumber="407">
<emphasis id="B954EAF3BC1C086294C5F96F0D4C9A01" bold="true" box="[199,314,1766,1792]" pageId="14" pageNumber="407">Diagnosis</emphasis>
: Body length
<quantity id="4CD89B04BC1C086295E7F96F0E049A00" box="[485,626,1766,1793]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="3.2" metricValueMin="2.4" pageId="14" pageNumber="407" unit="mm" value="2.8" valueMax="3.2" valueMin="2.4">2.4–3.2 mm</quantity>
. Antennal article VI with 16–18 bacilliform sensilla forming crescent shape, setiform sensillum located in anterior position, conical sensillum present near centre of crescent. Labrum asetose, long slender anterior setae in anterior margin. Telotarsus with posterior lateral process shorter than half-length of claw, anterior lateral and lamella processes present, anterior setiform process shorter than half-length of claw.
</paragraph>
</subSubSection>
<subSubSection id="C33A656ABC1C087C94C5F8FF0CA09A19" lastPageId="16" lastPageNumber="409" pageId="14" pageNumber="407" type="description">
<paragraph id="8B9F36E1BC1C086294C5F8FF0D2C9A90" blockId="14.[151,1437,1586,2009]" box="[199,346,1910,1937]" pageId="14" pageNumber="407">
<emphasis id="B954EAF3BC1C086294C5F8FF0D279A91" bold="true" box="[199,337,1910,1936]" pageId="14" pageNumber="407">Description</emphasis>
:
</paragraph>
<paragraph id="8B9F36E1BC1C086294C5F8130D1C9AD8" blockId="14.[151,1437,1586,2009]" pageId="14" pageNumber="407">
<typeStatus id="549B8843BC1C086294C5F8130D599AB5" box="[199,303,1946,1972]" pageId="14" pageNumber="407" type="holotype">Holotype</typeStatus>
male body
<quantity id="4CD89B04BC1C086295B6F8130E789AB4" box="[436,526,1946,1973]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="14" pageNumber="407" unit="mm" value="2.4">2.4 mm</quantity>
;
<typeStatus id="549B8843BC1C0862961BF8120EFF9AB4" box="[537,649,1947,1973]" pageId="14" pageNumber="407" type="paratype">paratypes</typeStatus>
: males
<quantity id="4CD89B04BC1C086296DFF8130F109AB4" box="[733,870,1946,1973]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="14" pageNumber="407" unit="mm" value="2.3" valueMax="2.4" valueMin="2.2">2.2–2.4 mm</quantity>
, females 3.0–
<quantity id="4CD89B04BC1C08629000F813082D9AB4" box="[1026,1115,1946,1973]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="14" pageNumber="407" unit="mm" value="3.2">3.2 mm</quantity>
; caudal bundle
<quantity id="4CD89B04BC1C08629109F81309159AB4" box="[1291,1379,1946,1973]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="407" unit="mm" value="0.5">0.5 mm</quantity>
, nest trichomes
<quantity id="4CD89B04BC1C0862950FF8370D109AD8" box="[269,358,1982,2009]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="407" unit="mm" value="0.2">0.2 mm</quantity>
.
</paragraph>
<paragraph id="8B9F36E1BC1D086394C5FF1E0E2F9C88" blockId="15.[151,1437,151,393]" pageId="15" pageNumber="408">
<emphasis id="B954EAF3BC1D086394C5FF1E0D719DB0" bold="true" box="[199,263,151,177]" pageId="15" pageNumber="408">Head</emphasis>
:
<materialsCitation id="3B483CBCBC1D08639515FF1E0E239C88" ID-GBIF-Occurrence="3981599312" collectionCode="R" pageId="15" pageNumber="408" specimenCount="4" typeStatus="holotype">
Arrangement of post vertex trichomes as for
<taxonomicName id="4C204D62BC1D08639727FF1E083D9DB1" authorityName="Huynh" authorityYear="2022" box="[805,1099,151,177]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="15" pageNumber="408" phylum="Arthropoda" rank="species" species="cattienensis">
<emphasis id="B954EAF3BC1D08639727FF1E083D9DB1" box="[805,1099,151,177]" italics="true" pageId="15" pageNumber="408">Monographis cattienensis</emphasis>
</taxonomicName>
with the
<typeStatus id="549B8843BC1D086390BFFF1E09699DB0" box="[1213,1311,151,177]" pageId="15" pageNumber="408" type="holotype">holotype</typeStatus>
having 13 sockets (L) and 14 (
<collectionCode id="ED31AE24BC1D08639577FF320DFC9DD4" box="[373,394,187,213]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="408">R</collectionCode>
) in anterior rows, posterior rows with 3 sockets on both sides (
<figureCitation id="131B2A64BC1D08639044FF3209519DD4" box="[1094,1319,187,213]" captionStart="FIGURE 13" captionStartId="15.[152,255,1898,1922]" captionTargetBox="[170,1418,426,1875]" captionTargetId="figure-136@15.[170,1418,417,1875]" captionTargetPageId="15" captionText="FIGURE 13. Holotype male Monographis tamdaoensis sp. n.. A. Head capsule with eye patterns, posterior vertex (pv) trichome sockets and the trichobothria (a, b and c). B. Enlarged images of Posterior vertex trichome sockets on left and right side; C. Clypeo-labrum with asetose labrum surface (l), lateral labrum (Ll) on each side, anterior setae (sa), posterior setae (Se), backward pointing setae (sb)." figureDoi="http://doi.org/10.5281/zenodo.7389327" httpUri="https://zenodo.org/record/7389327/files/figure.png" pageId="15" pageNumber="408">Figures 13A &amp; 13B</figureCitation>
).
<typeStatus id="549B8843BC1D08639138FF3209EA9DD4" box="[1338,1436,187,213]" pageId="15" pageNumber="408" type="paratype">Paratype</typeStatus>
showed anterior rows with 12–14 sockets, posterior rows with 2–3 sockets (
<figureCitation id="131B2A64BC1D086397FAFF5608099DF8" box="[1016,1151,223,249]" captionStart="FIGURE 16" captionStartId="17.[152,255,1906,1930]" captionTargetBox="[312,1274,1225,1882]" captionTargetId="figure-91@17.[312,1274,1225,1882]" captionTargetPageId="17" captionText="FIGURE 16. SEM images of Monographis tamdaoensis sp. n.. A. Posterior vertex (pv) showing the arrangement of trichome sockets. B. Left eye pattern with eight ommatidia and the trichobothria: a, b and c. C. Gnathochilarium showing the medial palp (Mp) andlateral palp (Lp) with 11 sensilla. D. Clypeo-labrum showing asetose surface (l), backward pointing setae (sb), anterior setae (sa), posterior setae (Se), lateral labrum (Ll) and the medial cleft (mc)." figureDoi="http://doi.org/10.5281/zenodo.7389337" httpUri="https://zenodo.org/record/7389337/files/figure.png" pageId="15" pageNumber="408">Figure 16A</figureCitation>
).
<emphasis id="B954EAF3BC1D08639097FF57093D9DF9" bold="true" box="[1173,1355,222,248]" pageId="15" pageNumber="408">Clypeo-labrum</emphasis>
:
<typeStatus id="549B8843BC1D08639159FF560CBC9C1C" pageId="15" pageNumber="408" type="holotype">Holotype</typeStatus>
, labrum posterior margin with 8 setae (Se), these setae longer in length than labrum width (
<typeStatus id="549B8843BC1D086390C1FE8A09539C1C" box="[1219,1317,259,285]" pageId="15" pageNumber="408" type="paratype">paratype</typeStatus>
with 8–10 setae). Labrum surface asetose (l), labrum posterior margin has a row of minute backward-pointing setae (sb). Lateral lamella (Ll) and single broad lamella each side median cleft (mc), 12–14 slender setae (sa) present on anterior margin of labrum (
<figureCitation id="131B2A64BC1D0863956BFEE60D839C88" box="[361,501,367,393]" captionStart="FIGURE 13" captionStartId="15.[152,255,1898,1922]" captionTargetBox="[170,1418,426,1875]" captionTargetId="figure-136@15.[170,1418,417,1875]" captionTargetPageId="15" captionText="FIGURE 13. Holotype male Monographis tamdaoensis sp. n.. A. Head capsule with eye patterns, posterior vertex (pv) trichome sockets and the trichobothria (a, b and c). B. Enlarged images of Posterior vertex trichome sockets on left and right side; C. Clypeo-labrum with asetose labrum surface (l), lateral labrum (Ll) on each side, anterior setae (sa), posterior setae (Se), backward pointing setae (sb)." figureDoi="http://doi.org/10.5281/zenodo.7389327" httpUri="https://zenodo.org/record/7389327/files/figure.png" pageId="15" pageNumber="408">Figures 13C</figureCitation>
&amp;
<figureCitation id="131B2A64BC1D0863961AFEE60E3B9C88" box="[536,589,367,393]" captionStart="FIGURE 16" captionStartId="17.[152,255,1906,1930]" captionTargetBox="[312,1274,1225,1882]" captionTargetId="figure-91@17.[312,1274,1225,1882]" captionTargetPageId="17" captionText="FIGURE 16. SEM images of Monographis tamdaoensis sp. n.. A. Posterior vertex (pv) showing the arrangement of trichome sockets. B. Left eye pattern with eight ommatidia and the trichobothria: a, b and c. C. Gnathochilarium showing the medial palp (Mp) andlateral palp (Lp) with 11 sensilla. D. Clypeo-labrum showing asetose surface (l), backward pointing setae (sb), anterior setae (sa), posterior setae (Se), lateral labrum (Ll) and the medial cleft (mc)." figureDoi="http://doi.org/10.5281/zenodo.7389337" httpUri="https://zenodo.org/record/7389337/files/figure.png" pageId="15" pageNumber="408">16D</figureCitation>
)
</materialsCitation>
.
</paragraph>
<caption id="DF5F6669BC1D0863949AF8E30DE49AEF" ID-DOI="http://doi.org/10.5281/zenodo.7389327" ID-Zenodo-Dep="7389327" httpUri="https://zenodo.org/record/7389327/files/figure.png" pageId="15" pageNumber="408" startId="15.[152,255,1898,1922]" targetBox="[170,1418,426,1875]" targetPageId="15">
<paragraph id="8B9F36E1BC1D0863949AF8E30DE49AEF" blockId="15.[151,1436,1898,2030]" pageId="15" pageNumber="408">
<emphasis id="B954EAF3BC1D0863949AF8E30D559A83" bold="true" box="[152,291,1898,1922]" pageId="15" pageNumber="408">FIGURE 13.</emphasis>
Holotype male
<taxonomicName id="4C204D62BC1D086395BDF8E30EBD9A83" authority="Huynh, 2022" authorityName="Huynh" authorityYear="2022" box="[447,715,1898,1922]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="15" pageNumber="408" phylum="Arthropoda" rank="species" species="tamdaoensis" status="sp. nov.">
<emphasis id="B954EAF3BC1D086395BDF8E30EBD9A83" box="[447,715,1898,1922]" italics="true" pageId="15" pageNumber="408">Monographis tamdaoensis</emphasis>
</taxonomicName>
<emphasis id="B954EAF3BC1D086396CDF8E30F719A83" bold="true" box="[719,775,1898,1922]" pageId="15" pageNumber="408">
<taxonomicNameLabel id="A2675788BC1D086396CDF8E30F719A83" box="[719,775,1898,1922]" pageId="15" pageNumber="408" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
.
<emphasis id="B954EAF3BC1D08639713F8E30F5C9A83" bold="true" box="[785,810,1898,1922]" pageId="15" pageNumber="408">A.</emphasis>
Head capsule with eye patterns, posterior vertex (pv) trichome sockets and the trichobothria (
<emphasis id="B954EAF3BC1D086395D2F8060DAB9AA7" box="[464,477,1935,1958]" italics="true" pageId="15" pageNumber="408">a</emphasis>
,
<emphasis id="B954EAF3BC1D086395E9F8070D8E9AA7" box="[491,504,1934,1958]" italics="true" pageId="15" pageNumber="408">b</emphasis>
and
<emphasis id="B954EAF3BC1D0863962EF8060E419AA7" box="[556,567,1935,1958]" italics="true" pageId="15" pageNumber="408">c</emphasis>
).
<emphasis id="B954EAF3BC1D0863964CF8070E139AA7" bold="true" box="[590,613,1934,1958]" pageId="15" pageNumber="408">B.</emphasis>
Enlarged images of Posterior vertex trichome sockets on left and right side;
<emphasis id="B954EAF3BC1D08639186F80709EA9AA7" bold="true" box="[1412,1436,1934,1958]" pageId="15" pageNumber="408">C.</emphasis>
Clypeo-labrum with asetose labrum surface (l), lateral labrum (Ll) on each side, anterior setae (sa), posterior setae (Se), backward pointing setae (sb).
</paragraph>
</caption>
<caption id="DF5F6669BC02087C949AFAC7096698AE" ID-DOI="http://doi.org/10.5281/zenodo.7389331" ID-Zenodo-Dep="7389331" httpUri="https://zenodo.org/record/7389331/files/figure.png" pageId="16" pageNumber="409" startId="16.[152,255,1358,1382]" targetBox="[151,1417,189,1334]" targetPageId="16">
<paragraph id="8B9F36E1BC02087C949AFAC7096698AE" blockId="16.[151,1436,1358,1455]" pageId="16" pageNumber="409">
<emphasis id="B954EAF3BC02087C949AFAC70D539867" bold="true" box="[152,293,1358,1382]" pageId="16" pageNumber="409">FIGURE 14.</emphasis>
Holotype male
<taxonomicName id="4C204D62BC02087C95CBFAC60EA19866" authority="Huynh, 2022" authorityName="Huynh" authorityYear="2022" box="[457,727,1359,1383]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="16" pageNumber="409" phylum="Arthropoda" rank="species" species="tamdaoensis" status="sp. nov.">
<emphasis id="B954EAF3BC02087C95CBFAC60EA19866" box="[457,727,1359,1383]" italics="true" pageId="16" pageNumber="409">Monographis tamdaoensis</emphasis>
</taxonomicName>
<emphasis id="B954EAF3BC02087C96DCFAC60F6E9866" bold="true" box="[734,792,1359,1383]" pageId="16" pageNumber="409">
<taxonomicNameLabel id="A2675788BC02087C96DCFAC60F6E9866" box="[734,792,1359,1383]" pageId="16" pageNumber="409" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
.
<emphasis id="B954EAF3BC02087C9727FAC70F4B9867" bold="true" box="[805,829,1358,1382]" pageId="16" pageNumber="409">A.</emphasis>
Section of antennal articles VI–VII and sensory cones (sc), and their sensilla.
<emphasis id="B954EAF3BC02087C954EFAFA0D15988A" bold="true" box="[332,355,1395,1419]" pageId="16" pageNumber="409">B.</emphasis>
Sensilla on the antennal article VII, with two thick bacilliform sensilla (Ta &amp; Tp), setiform (S) and a conical sensillum (C).
<emphasis id="B954EAF3BC02087C952FFA1F0D3098AF" bold="true" box="[301,326,1430,1454]" pageId="16" pageNumber="409">C.</emphasis>
Sensilla on the article VI, showing thick (T), thin (t) and medium thin bacilliform sensilla (tm).
</paragraph>
</caption>
<paragraph id="8B9F36E1BC02087C94C5FA540D299B62" blockId="16.[151,1437,1501,1959]" pageId="16" pageNumber="409">
<emphasis id="B954EAF3BC02087C94C5FA540D4C98F6" bold="true" box="[199,314,1501,1527]" pageId="16" pageNumber="409">Antennae</emphasis>
:
<materialsCitation id="3B483CBCBC02087C954AFA540D299B62" ID-GBIF-Occurrence="3981599308" collectionCode="VI, T" pageId="16" pageNumber="409" specimenCount="1" typeStatus="holotype">
Antennal article
<collectionCode id="ED31AE24BC02087C9601FA540E5798F6" box="[515,545,1501,1527]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="16" pageNumber="409">VI</collectionCode>
with 16–18 bacilliform sensilla.
<typeStatus id="549B8843BC02087C9794FA540F8898F6" box="[918,1022,1501,1527]" pageId="16" pageNumber="409" type="holotype">Holotype</typeStatus>
with 16 bacilliform sensilla forming a crescent shape with 3 thick (
<collectionCode id="ED31AE24BC02087C95E9F9880D899B1A" box="[491,511,1537,1563]" name="Tavera, Department of Geology and Geophysics" pageId="16" pageNumber="409">T</collectionCode>
), 9 thin bacilliform sensilla (t) and 4 medium-length thin bacilliform sensilla (tm), setiform sensillum (S) located in anterior position, conical sensillum (C) present near centre of crescent (
<figureCitation id="131B2A64BC02087C9144F9AC0CBE9B62" captionStart="FIGURE 14" captionStartId="16.[152,255,1358,1382]" captionTargetBox="[151,1417,189,1334]" captionTargetId="figure-29@16.[151,1436,181,1334]" captionTargetPageId="16" captionText="FIGURE 14. Holotype male Monographis tamdaoensis sp. n.. A. Section of antennal articles VI–VII and sensory cones (sc), and their sensilla. B. Sensilla on the antennal article VII, with two thick bacilliform sensilla (Ta &amp; Tp), setiform (S) and a conical sensillum (C). C. Sensilla on the article VI, showing thick (T), thin (t) and medium thin bacilliform sensilla (tm)." figureDoi="http://doi.org/10.5281/zenodo.7389331" httpUri="https://zenodo.org/record/7389331/files/figure.png" pageId="16" pageNumber="409">Figures 14C</figureCitation>
,
<figureCitation id="131B2A64BC02087C94D6F9C00D249B62" box="[212,338,1609,1635]" captionStart="FIGURE 17" captionStartId="18.[152,255,1080,1104]" captionTargetBox="[152,1437,182,1056]" captionTargetId="figure-29@18.[151,1437,181,1056]" captionTargetPageId="18" captionText="FIGURE 17. SEM images of Monographis tamdaoensis sp. n.. A. Antennal article VI–VIII. B. Arrangement of a setiform (S), conical sensillum (C), thick (T), thin (t) bacilliform sensilla, and the medium length bacilliform sensilla (tm) on article VI. C. Other end of arrangement of sensilla on article VI. D. Telotarsus and its processes: claw (c), anterior setiform process (s), lamella (la), anterior (a) and posterior (p) processes." figureDoi="http://doi.org/10.5281/zenodo.7389341" httpUri="https://zenodo.org/record/7389341/files/figure.png" pageId="16" pageNumber="409">17B &amp;17C</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8B9F36E1BC02087C94C5F9E409EC9BAA" blockId="16.[151,1437,1501,1959]" pageId="16" pageNumber="409">
<emphasis id="B954EAF3BC02087C94C5F9E40D349B86" bold="true" box="[199,322,1645,1671]" pageId="16" pageNumber="409">Telotarsus</emphasis>
: Claw slender, posterior lateral process (p) shorter than half-length of claw, anterior lateral (a) and lamella processes (la) present, anterior setiform process (s) less than half-length of claw (c). (
<figureCitation id="131B2A64BC02087C90A8F91809409BAD" box="[1194,1334,1681,1708]" captionStart="FIGURE 15" captionStartId="17.[152,255,1104,1128]" captionTargetBox="[290,1295,188,1070]" captionTargetId="figure-19@17.[289,1298,181,1080]" captionTargetPageId="17" captionText="FIGURE 15. Holotype male Monographis tamdaoensis sp. n.. A. The second leg showing its segments and chaetotaxy, penis (p). B. Telotarsus showing the claw (c) and processes: anterior (a), posterior (p), lamella (la) and anterior setiform process (s) shorter than half length of the claw (dot line). C. The ornamental trichome sockets with trichomes a, b, three c and an indented d." figureDoi="http://doi.org/10.5281/zenodo.7389333" httpUri="https://zenodo.org/record/7389333/files/figure.png" pageId="16" pageNumber="409">Figures 15B</figureCitation>
&amp;
<figureCitation id="131B2A64BC02087C9158F91809F89BAA" box="[1370,1422,1681,1707]" captionStart="FIGURE 17" captionStartId="18.[152,255,1080,1104]" captionTargetBox="[152,1437,182,1056]" captionTargetId="figure-29@18.[151,1437,181,1056]" captionTargetPageId="18" captionText="FIGURE 17. SEM images of Monographis tamdaoensis sp. n.. A. Antennal article VI–VIII. B. Arrangement of a setiform (S), conical sensillum (C), thick (T), thin (t) bacilliform sensilla, and the medium length bacilliform sensilla (tm) on article VI. C. Other end of arrangement of sensilla on article VI. D. Telotarsus and its processes: claw (c), anterior setiform process (s), lamella (la), anterior (a) and posterior (p) processes." figureDoi="http://doi.org/10.5281/zenodo.7389341" httpUri="https://zenodo.org/record/7389341/files/figure.png" pageId="16" pageNumber="409">17D</figureCitation>
).
</paragraph>
<paragraph id="8B9F36E1BC02087C94C5F93C0CA09A19" blockId="16.[151,1437,1501,1959]" pageId="16" pageNumber="409">
<emphasis id="B954EAF3BC02087C94C5F93C0D629BCE" bold="true" box="[199,276,1717,1743]" pageId="16" pageNumber="409">Telson</emphasis>
: Arrangement of dorsal ornamental trichome sockets similar to
<taxonomicName id="4C204D62BC02087C97ECF93F08EE9BCE" authorityName="Huynh" authorityYear="2022" box="[1006,1176,1718,1743]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="16" pageNumber="409" phylum="Arthropoda" rank="species" species="cattienensis">
<emphasis id="B954EAF3BC02087C97ECF93F08EE9BCE" box="[1006,1176,1718,1743]" italics="true" pageId="16" pageNumber="409">M. cattienensis</emphasis>
</taxonomicName>
with 17 sockets of trichomes
<emphasis id="B954EAF3BC02087C94F6F9500D749BF2" bold="true" box="[244,258,1753,1779]" italics="true" pageId="16" pageNumber="409">a</emphasis>
left side and 18 sockets on right side of the telson in the
<typeStatus id="549B8843BC02087C9782F9500F949BF2" box="[896,994,1753,1779]" pageId="16" pageNumber="409" type="holotype">holotype</typeStatus>
(
<typeStatus id="549B8843BC02087C97F3F95008259BF2" box="[1009,1107,1753,1779]" pageId="16" pageNumber="409" type="paratype">paratype</typeStatus>
with 16–18 sockets). (
<figureCitation id="131B2A64BC02087C9153F9500CBF9A19" captionStart="FIGURE 15" captionStartId="17.[152,255,1104,1128]" captionTargetBox="[290,1295,188,1070]" captionTargetId="figure-19@17.[289,1298,181,1080]" captionTargetPageId="17" captionText="FIGURE 15. Holotype male Monographis tamdaoensis sp. n.. A. The second leg showing its segments and chaetotaxy, penis (p). B. Telotarsus showing the claw (c) and processes: anterior (a), posterior (p), lamella (la) and anterior setiform process (s) shorter than half length of the claw (dot line). C. The ornamental trichome sockets with trichomes a, b, three c and an indented d." figureDoi="http://doi.org/10.5281/zenodo.7389333" httpUri="https://zenodo.org/record/7389333/files/figure.png" pageId="16" pageNumber="409">Figure 15C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33A656ABC02087C94C5F8A80EB79A5E" pageId="16" pageNumber="409" type="discussion">
<paragraph id="8B9F36E1BC02087C94C5F8A80EB79A5E" blockId="16.[151,1437,1501,1959]" pageId="16" pageNumber="409">
<emphasis id="B954EAF3BC02087C94C5F8A80D429A3A" bold="true" box="[199,308,1825,1851]" pageId="16" pageNumber="409">Remarks</emphasis>
:
<taxonomicName id="4C204D62BC02087C9541F8A80E189A3A" authority="Huynh, 2022" authorityName="Huynh" authorityYear="2022" box="[323,622,1825,1851]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="16" pageNumber="409" phylum="Arthropoda" rank="species" species="tamdaoensis" status="sp. nov.">
<emphasis id="B954EAF3BC02087C9541F8A80E189A3A" box="[323,622,1825,1851]" italics="true" pageId="16" pageNumber="409">Monographis tamdaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2675788BC02087C9677F8A80EC39A3A" box="[629,693,1825,1851]" pageId="16" pageNumber="409" rank="species">
<emphasis id="B954EAF3BC02087C9677F8A80ED89A3A" bold="true" box="[629,686,1825,1851]" pageId="16" pageNumber="409">sp. n</emphasis>
.
</taxonomicNameLabel>
is distinguished from congeners in its labrum structures and anterior setiform process equal to half-length of claw.
</paragraph>
</subSubSection>
<subSubSection id="C33A656ABC02087C94C5F8E00CB99AA6" pageId="16" pageNumber="409" type="etymology">
<paragraph id="8B9F36E1BC02087C94C5F8E00CB99AA6" blockId="16.[151,1437,1501,1959]" pageId="16" pageNumber="409">
<emphasis id="B954EAF3BC02087C94C5F8E00D3E9A82" bold="true" box="[199,328,1897,1923]" pageId="16" pageNumber="409">Etymology</emphasis>
:
<taxonomicName id="4C204D62BC02087C955BF8E00EF39A82" authorityName="Huynh" authorityYear="2022" box="[345,645,1897,1923]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="16" pageNumber="409" phylum="Arthropoda" rank="species" species="tamdaoensis">
<emphasis id="B954EAF3BC02087C955BF8E00EF39A82" box="[345,645,1897,1923]" italics="true" pageId="16" pageNumber="409">Monographis tamdaoensis</emphasis>
</taxonomicName>
is named after Tam Dao National Park in
<collectingRegion id="49E4F803BC02087C9073F8E009219A82" box="[1137,1367,1897,1923]" country="Vietnam" name="Vinh Phuc" pageId="16" pageNumber="409">Vinh Phuc Province</collectingRegion>
,
<collectingCountry id="F3377671BC02087C9166F8E00CBD9AA6" name="Vietnam" pageId="16" pageNumber="409">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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