316 lines
34 KiB
XML
316 lines
34 KiB
XML
<document id="A431228EC28B7EC55D3141BED209198F" ID-DOI="10.11646/zootaxa.5575.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="14746442" ID-ZooBank="521D404D-EB9D-4110-989E-A8A273C10152" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1737970514015" checkinUser="plazi" docAuthor="Génier, François, Gunter, Nicole L. & Saxton, Natalie A." docDate="2025" docId="038987E8FFFC6E16FF461757FAABFB10" docLanguage="en" docName="zootaxa.5575.1.3.pdf" docOrigin="Zootaxa 5575 (1)" docSource="https://doi.org/10.11646/zootaxa.5575.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tesserodon howdeni Paulian 1985" docType="treatment" docVersion="5" lastPageNumber="100" masterDocId="FFB0FF90FFF76E1BFFD11005FFC3FFB0" masterDocTitle="Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="110" masterPageNumber="87" pageNumber="98" updateTime="1738747596743" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="4AA06840BF7CC064EF09A87720437921">Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="AFC07A2B6F2B2108B27B19BF20C8EE97">Génier, François</mods:namePart>
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<mods:nameIdentifier id="1ECC9C7AC992A25C8AA5C5D52BEFF8AA" type="email">enier@nature.ca</mods:nameIdentifier>
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<mods:namePart id="F1DFFA23BCE755E5D7D036530A9CFF70">Gunter, Nicole L.</mods:namePart>
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<mods:affiliation id="8F7A6C55D21A6DE4314AE8518BED7447">Biodiversity and Geosciences Program, Queensland Museum, South Brisbane, QLD, 4101 Australia.</mods:affiliation>
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<mods:namePart id="ECC191ADE1B713E0EDFBE1C045FBDEC0">Saxton, Natalie A.</mods:namePart>
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<mods:affiliation id="6475BE0C86261ED4CB4FFC509B888BE2">Department of Biology, Case Western Reserve University, 2080 Adelbert Road, Cleveland, OH, 44106, USA. & Corresponding author</mods:affiliation>
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<mods:title id="78B90FECC7908D416761226DBF0023F3">Zootaxa</mods:title>
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<mods:date id="39DC5D2CA7A2B6B98A40A63D379F9168">2025</mods:date>
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<subSubSection id="C33A6575FFFC6E10FF461757FCE7F8DA" box="[151,804,1874,1898]" pageId="11" pageNumber="98" type="nomenclature">
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<paragraph id="8B9F36FEFFFC6E10FF461757FCE7F8DA" blockId="11.[151,804,1874,1898]" box="[151,804,1874,1898]" pageId="11" pageNumber="98">
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<heading id="D0D78192FFFC6E10FF461757FCE7F8DA" bold="true" box="[151,804,1874,1898]" fontSize="10" level="2" pageId="11" pageNumber="98" reason="2">
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<treatmentCitationGroup id="AB3011D0FFFC6E10FF461757FCE7F8DA" box="[151,804,1874,1898]" pageId="11" pageNumber="98">
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<emphasis id="B954EAECFFFC6E10FF461757FCE7F8DA" bold="true" box="[151,804,1874,1898]" pageId="11" pageNumber="98">
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<treatmentCitation id="0A8110EFFFFC6E10FF461757FDFAF8DA" author="Paulian, R." box="[151,569,1874,1898]" page="222" pageId="11" pageNumber="98" year="1985">
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<taxonomicName id="4C204D7DFFFC6E10FF461757FDFAF8DA" ID-CoL="55JGP" authority="Paulian, 1985: 222" authorityName="Paulian" authorityPageNumber="222" authorityYear="1985" box="[151,569,1874,1898]" class="Insecta" family="Scarabaeidae" genus="Tesserodon" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="98" phylum="Arthropoda" rank="species" species="howdeni">
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<emphasis id="B954EAECFFFC6E10FF461757FEAFF8DA" bold="true" box="[151,364,1874,1898]" italics="true" pageId="11" pageNumber="98">Tesserodon howdeni</emphasis>
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<bibRefCitation id="EFB14B0FFFFC6E10FEA31757FDFAF8DA" author="Paulian, R." box="[370,569,1874,1898]" pageId="11" pageNumber="98" pagination="219 - 238" refId="ref16422" refString="Paulian, R. (1985) Les coleopteres Scarabaeidae Canthonides de Nouvelle-Guinee. Annales de la Societe Entomologique de France, Nouvelle Serie, 21 (2), 219 - 238. https: // doi. org / 10.1080 / 21686351.1985.12278409" type="journal article" year="1985">Paulian, 1985: 222</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(original description)
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C33A6575FFFC6E16FF46179DFD5BFED4" lastPageId="13" lastPageNumber="100" pageId="11" pageNumber="98" type="materials_examined">
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<paragraph id="8B9F36FEFFFC6E17FF46179DFB63FF65" blockId="11.[151,1437,1944,2043]" lastBlockId="12.[151,1436,151,213]" lastPageId="12" lastPageNumber="99" pageId="11" pageNumber="98">
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<emphasis id="B954EAECFFFC6E10FF46179DFE09F803" bold="true" box="[151,458,1944,1971]" pageId="11" pageNumber="98">Name-bearing type data.</emphasis>
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<materialsCitation id="3B483CA3FFFC6E17FE07179CFB5AFF64" collectingDate="1974-07-15" collectingDateMax="1974-07-16" collectingDateMin="1974-07-15" collectionCode="CMNC" collectorName="S. Peck" country="Papua N. Guinea" determinerName="R. Paulian" lastPageId="12" lastPageNumber="99" location="Papua N. Guinea" municipality="Musee" pageId="11" pageNumber="98" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus id="549B885CFFFC6E10FE07179CFDFDF803" box="[470,574,1945,1971]" pageId="11" pageNumber="98" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D26FD77FFFC6E10FD9B179CFD9CF803" box="[586,607,1945,1971]" count="1" pageId="11" pageNumber="98" type="male">♂</specimenCount>
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(
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<collectionCode id="ED31AE3BFFFC6E10FDA2179CFD0EF803" box="[627,717,1945,1971]" collectionName="Canada, Ottawa, Canadian Museum of Nature" pageId="11" pageNumber="98">CMNC</collectionCode>
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): ||
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<collectingCountry id="F337766EFFFC6E10FD2D179CFC7BF803" box="[764,952,1945,1971]" pageId="11" pageNumber="98">Papua N.Guinea</collectingCountry>
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| Brown R.
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<quantity id="4CD89B1BFFFC6E10FB97179CFB4BF803" box="[1094,1160,1945,1971]" metricMagnitude="4" metricUnit="m" metricValue="4.828032" pageId="11" pageNumber="98" unit="mi" value="30.0">30mi.</quantity>
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N | Fort Moresby |
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<date id="FF9E103EFFFC6E10FAA6179DFEE3F867" pageId="11" pageNumber="98" value="1974-07-15" valueMax="1974-07-16" valueMin="1974-07-15">
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<collectingDate id="EFDAE9D6FFFC6E10FAA6179DFEE3F867" pageId="11" pageNumber="98" value="1974-07-15" valueMax="1974-07-16" valueMin="1974-07-15">15- 16.VII.1974</collectingDate>
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</date>
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||; ||
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<collectorName id="26D55328FFFC6E10FE8B17B8FE73F867" box="[346,432,1981,2007]" pageId="11" pageNumber="98">S. Peck</collectorName>
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| collector ||; || H. & A. Howden | collection ||; || HOLOTYPE || [red card]; || Canadian Museum of | Musée canadien de la | NATURE | CMNEN 00011854 ||; ||
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<taxonomicName id="4C204D7DFFFC6E10FC1A17E4FB89F84B" authorityName="Hope" authorityYear="1837" box="[971,1098,2017,2043]" class="Insecta" family="Scarabaeidae" genus="Tesserodon" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="98" phylum="Arthropoda" rank="genus">Tesserodon</taxonomicName>
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| howdeni | n. sp. |
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||
<determinerName id="7E43DA3AFFFC6E10FAF617E4FA5FF84B" box="[1319,1436,2017,2043]" pageId="11" pageNumber="98">R. Paulian</determinerName>
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det. || [partly handwritten].
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<emphasis id="B954EAECFFFB6E17FE1C1092FDC6FF01" bold="true" box="[461,517,151,177]" pageId="12" pageNumber="99">Note</emphasis>
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||
. Previously stated as: “Nouvelle-Guinée (Papouasie): Brown River,
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<quantity id="4CD89B1BFFFB6E17FAC91092FABAFF01" box="[1304,1401,151,177]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="12" pageNumber="99" unit="mi" value="20.0">20 miles</quantity>
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||
N. Port Moresby, 14/
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||
<date id="FF9E103EFFFB6E17FEB110BEFE2DFF65" box="[352,494,187,213]" pageId="12" pageNumber="99" value="1974-07-16">16-VII-1974</date>
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||
(S. Peck), piège à excréments, en forêt” by Paulian.
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||
<emphasis id="B954EAECFFFB6E17FBEC10BFFB5AFF64" bold="true" box="[1085,1177,186,212]" pageId="12" pageNumber="99">Studied</emphasis>
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</materialsCitation>
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.
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||
</paragraph>
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||
<caption id="DF5F6676FFFB6E17FF4917A2FE30F853" ID-DOI="http://doi.org/10.5281/zenodo.14746462" ID-Zenodo-Dep="14746462" httpUri="https://zenodo.org/record/14746462/files/figure.png" pageId="12" pageNumber="99" startId="12.[152,255,1959,1983]" targetBox="[155,1431,238,1929]" targetPageId="12" targetType="figure">
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<paragraph id="8B9F36FEFFFB6E17FF4917A2FE30F853" blockId="12.[151,1436,1959,2019]" pageId="12" pageNumber="99">
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<emphasis id="B954EAECFFFB6E17FF4917A2FED4F80F" bold="true" box="[152,279,1959,1983]" pageId="12" pageNumber="99">FIGURE 9.</emphasis>
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Male aedeagus of
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<taxonomicName id="4C204D7DFFFB6E17FE0317A2FD44F80F" box="[466,647,1959,1983]" pageId="12" pageNumber="99">
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<emphasis id="B954EAECFFFB6E17FE0317A2FDF5F80F" box="[466,566,1959,1983]" italics="true" pageId="12" pageNumber="99">Rhytomus</emphasis>
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species
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</taxonomicName>
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||
, left lateral, dorsal, right lateral. A,
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<taxonomicName id="4C204D7DFFFB6E17FC3617ADFBA6F80F" authorityName="Génier & Gunter & Saxton" authorityYear="2025" baseAuthorityName="van Lansberge" baseAuthorityYear="1885" box="[999,1125,1959,1983]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="99" phylum="Arthropoda" rank="species" species="elongatus">
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<emphasis id="B954EAECFFFB6E17FC3617ADFBA6F80F" box="[999,1125,1959,1983]" italics="true" pageId="12" pageNumber="99">R. elongatus</emphasis>
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</taxonomicName>
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; B,
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||
<taxonomicName id="4C204D7DFFFB6E17FB5F17ADFB38F80F" authorityName="Saxton & Genier" authorityYear="2025" box="[1166,1275,1959,1983]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="99" phylum="Arthropoda" rank="species" species="fervidus">
|
||
<emphasis id="B954EAECFFFB6E17FB5F17ADFB38F80F" box="[1166,1275,1959,1983]" italics="true" pageId="12" pageNumber="99">R. fervidus</emphasis>
|
||
</taxonomicName>
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||
; C,
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||
<emphasis id="B954EAECFFFB6E17FAF217ADFA5FF80F" box="[1315,1436,1959,1983]" italics="true" pageId="12" pageNumber="99">
|
||
<taxonomicName id="4C204D7DFFFB6E17FAF217ADFA55F80F" authorityName="Paulian" authorityYear="1985" box="[1315,1430,1959,1983]" class="Insecta" family="Scarabaeidae" genus="Tesserodon" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="99" phylum="Arthropoda" rank="species" species="howdeni">R. howdeni</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
D,
|
||
<emphasis id="B954EAECFFFB6E17FF6717C9FEE3F853" box="[182,288,1995,2019]" italics="true" pageId="12" pageNumber="99">
|
||
<taxonomicName id="4C204D7DFFFB6E17FF6717C9FEDFF853" authorityName="Saxton & Genier" authorityYear="2025" box="[182,284,1995,2019]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="99" phylum="Arthropoda" rank="species" species="larseni">R. larseni</taxonomicName>
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||
.
|
||
</emphasis>
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||
Scale bar = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
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||
<paragraph id="8B9F36FEFFFA6E16FF161093FD5BFED4" blockId="13.[151,1437,150,1185]" pageId="13" pageNumber="100">
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<emphasis id="B954EAECFFFA6E16FF161093FE6BFF00" bold="true" box="[199,424,150,176]" pageId="13" pageNumber="100">Material examined</emphasis>
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(
|
||
<specimenCount id="9D26FD77FFFA6E16FE661093FE34FF01" box="[439,503,150,177]" count="5" pageId="13" pageNumber="100" type="male">5 ♂♂</specimenCount>
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,
|
||
<specimenCount id="9D26FD77FFFA6E16FDD31093FDFEFF00" box="[514,573,150,176]" count="2" pageId="13" pageNumber="100" type="female">2 ♀♀</specimenCount>
|
||
).
|
||
<materialsCitation id="3B483CA3FFFA6E16FD841093FE9AFF48" collectedFrom="forest" collectingDate="1974-07-14" collectingDateMax="1974-07-18" collectingDateMin="1974-07-14" collectingMethod="dung traps" collectionCode="CMNC" collectorName="S. Peck" country="Papua New Guinea" elevation="3" latitude="-9.204166" location="Brown River" longLatPrecision="21" longitude="147.2875" pageId="13" pageNumber="100" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="CENTRAL PROVINCE">
|
||
<emphasis id="B954EAECFFFA6E16FD841093FCACFF00" bold="true" box="[597,879,150,176]" pageId="13" pageNumber="100">
|
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<collectingCountry id="F337766EFFFA6E16FD841093FCACFF00" box="[597,879,150,176]" name="Papua New Guinea" pageId="13" pageNumber="100">PAPUA NEW GUINEA</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<collectingRegion id="49E4F81CFFFA6E16FCAD1093FB52FF00" box="[892,1169,150,176]" country="Papua New Guinea" name="Central" pageId="13" pageNumber="100">CENTRAL PROVINCE</collectingRegion>
|
||
,
|
||
<locationDeviation id="9A2D2FDCFFFA6E16FB4A1093FA5AFF00" box="[1179,1433,150,177]" location="port Moresby" pageId="13" pageNumber="100">
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<quantity id="4CD89B1BFFFA6E16FB4A1093FB22FF00" box="[1179,1249,150,176]" metricMagnitude="4" metricUnit="m" metricValue="2.8968192" pageId="13" pageNumber="100" unit="mi" value="18.0">18 mi.</quantity>
|
||
N port Moresby
|
||
</locationDeviation>
|
||
,
|
||
<location id="8EFF6025FFFA6E16FF4610BEFEE9FF64" LSID="urn:lsid:plazi:treatment:038987E8FFFC6E16FF461757FAABFB10:8EFF6025FFFA6E16FF4610BEFEE9FF64" box="[151,298,186,213]" country="Papua New Guinea" latitude="-9.204166" longLatPrecision="21" longitude="147.2875" name="Brown River" pageId="13" pageNumber="100" stateProvince="CENTRAL PROVINCE">Brown River</location>
|
||
,
|
||
<quantity id="4CD89B1BFFFA6E16FEE410BFFEA2FF65" box="[309,353,186,213]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="100" unit="m" value="3.0">
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<elevation id="000DD1CDFFFA6E16FEE410BFFEA2FF65" box="[309,353,186,213]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="100" unit="m" value="3.0">3 m</elevation>
|
||
</quantity>
|
||
, [
|
||
<geoCoordinate id="EE145039FFFA6E16FEA510BFFDC3FF65" box="[372,512,186,213]" degrees="09" direction="south" minutes="12" orientation="latitude" pageId="13" pageNumber="100" precision="15" seconds="15" value="-9.204166">09°12’15’’S</geoCoordinate>
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,
|
||
<geoCoordinate id="EE145039FFFA6E16FDDD10BFFD6BFF65" box="[524,680,186,213]" degrees="147" direction="east" minutes="17" orientation="longitude" pageId="13" pageNumber="100" precision="15" seconds="15" value="147.2875">147°17’15’’E</geoCoordinate>
|
||
],
|
||
<date id="FF9E103EFFFA6E16FD6A10BFFCA4FF64" box="[699,871,186,212]" pageId="13" pageNumber="100" value="1974-07-14" valueMax="1974-07-18" valueMin="1974-07-14">
|
||
<collectingDate id="EFDAE9D6FFFA6E16FD6A10BFFCA4FF64" box="[699,871,186,212]" pageId="13" pageNumber="100" value="1974-07-14" valueMax="1974-07-18" valueMin="1974-07-14">14–18.vii.1974</collectingDate>
|
||
</date>
|
||
,
|
||
<collectedFrom id="383C7503FFFA6E16FCA310BFFC76FF64" box="[882,949,186,212]" pageId="13" pageNumber="100">forest</collectedFrom>
|
||
,
|
||
<collectingMethod id="52614EE9FFFA6E16FC6E10BFFBF4FF65" box="[959,1079,186,213]" pageId="13" pageNumber="100">dung traps</collectingMethod>
|
||
,
|
||
<collectorName id="26D55328FFFA6E16FB9310BFFB5BFF64" box="[1090,1176,186,212]" pageId="13" pageNumber="100">S. Peck</collectorName>
|
||
, (dung traps 39-42)—
|
||
<specimenCount id="9D26FD77FFFA6E16FA4210BFFF6DFF49" count="1" pageId="13" pageNumber="100" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D26FD77FFFA6E16FF6D10DBFF24FF49" box="[188,231,222,249]" count="1" pageId="13" pageNumber="100" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED31AE3BFFFA6E16FF2910DBFE92FF48" box="[248,337,222,248]" collectionName="Canada, Ottawa, Canadian Museum of Nature" pageId="13" pageNumber="100">CMNC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B483CA3FFFA6E16FEB410DBFD13FEAC" collectingDate="1974-07-16" collectingDateMax="1974-07-18" collectingDateMin="1974-07-16" collectingMethod="dung traps" collectionCode="CMNC" collectorName="S. Peck" country="Papua New Guinea" elevation="3" latitude="-9.191667" location="Brown River" longLatPrecision="21" longitude="147.31667" pageId="13" pageNumber="100" specimenCount="1" specimenCount-female="1" stateProvince="National Capital">
|
||
<locationDeviation id="9A2D2FDCFFFA6E16FEB410DBFDB3FF48" box="[357,624,222,249]" location="Port Moresby" pageId="13" pageNumber="100">
|
||
<quantity id="4CD89B1BFFFA6E16FEB410DBFE6CFF48" box="[357,431,222,248]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="13" pageNumber="100" unit="mi" value="20.0">20 mi.</quantity>
|
||
N Port Moresby
|
||
</locationDeviation>
|
||
,
|
||
<location id="8EFF6025FFFA6E16FDAD10DAFCD2FF48" LSID="urn:lsid:plazi:treatment:038987E8FFFC6E16FF461757FAABFB10:8EFF6025FFFA6E16FDAD10DAFCD2FF48" box="[636,785,222,249]" country="Papua New Guinea" latitude="-9.191667" longLatPrecision="21" longitude="147.31667" name="Brown River" pageId="13" pageNumber="100" stateProvince="National Capital">Brown River</location>
|
||
,
|
||
<quantity id="4CD89B1BFFFA6E16FCCF10DBFC8FFF49" box="[798,844,222,249]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="100" unit="m" value="3.0">
|
||
<elevation id="000DD1CDFFFA6E16FCCF10DBFC8FFF49" box="[798,844,222,249]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="100" unit="m" value="3.0">3 m</elevation>
|
||
</quantity>
|
||
, [
|
||
<geoCoordinate id="EE145039FFFA6E16FCB010DBFC2FFF48" box="[865,1004,222,248]" degrees="09" direction="south" minutes="11" orientation="latitude" pageId="13" pageNumber="100" precision="15" seconds="30" value="-9.191667">09°11’30’’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE145039FFFA6E16FC2B10DBFB55FF48" box="[1018,1174,222,248]" degrees="147" direction="east" minutes="19" orientation="longitude" pageId="13" pageNumber="100" precision="15" seconds="00" value="147.31667">147°19’00’’E</geoCoordinate>
|
||
],
|
||
<date id="FF9E103EFFFA6E16FB7A10DBFA94FF48" box="[1195,1367,222,248]" pageId="13" pageNumber="100" value="1974-07-16" valueMax="1974-07-18" valueMin="1974-07-16">
|
||
<collectingDate id="EFDAE9D6FFFA6E16FB7A10DBFA94FF48" box="[1195,1367,222,248]" pageId="13" pageNumber="100" value="1974-07-16" valueMax="1974-07-18" valueMin="1974-07-16">16–18.vii.1974</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod id="52614EE9FFFA6E16FAB510DBFF13FEAD" pageId="13" pageNumber="100">dung traps</collectingMethod>
|
||
,
|
||
<collectorName id="26D55328FFFA6E16FF0C1107FEF5FEAC" box="[221,310,258,284]" pageId="13" pageNumber="100">S. Peck</collectorName>
|
||
, (dung traps 39-41)—
|
||
<specimenCount id="9D26FD77FFFA6E16FDE61107FD9DFEAD" box="[567,606,258,285]" count="1" pageId="13" pageNumber="100" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED31AE3BFFFA6E16FDBE1107FD0BFEAC" box="[623,712,258,284]" collectionName="Canada, Ottawa, Canadian Museum of Nature" pageId="13" pageNumber="100">CMNC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B483CA3FFFA6E16FD0C1106FDDDFEF0" collectingDate="1974-07-16" collectingDateMax="1974-07-18" collectingDateMin="1974-07-16" collectingMethod="dung trap" collectionCode="MNHN" collectorName="S. Peck" country="Papua New Guinea" elevation="3" latitude="-9.191667" location="Brown River" longLatPrecision="21" longitude="147.31667" pageId="13" pageNumber="100" specimenCount="2" specimenCount-female="1" specimenCount-male="2" stateProvince="Central" typeStatus="paratype">
|
||
same locality,
|
||
<date id="FF9E103EFFFA6E16FC501107FBEEFEAC" box="[897,1069,258,284]" pageId="13" pageNumber="100" value="1974-07-16" valueMax="1974-07-18" valueMin="1974-07-16">
|
||
<collectingDate id="EFDAE9D6FFFA6E16FC501107FBEEFEAC" box="[897,1069,258,284]" pageId="13" pageNumber="100" value="1974-07-16" valueMax="1974-07-18" valueMin="1974-07-16">16–18.vii.1974</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod id="52614EE9FFFA6E16FBEB1107FB6AFEAD" box="[1082,1193,258,285]" pageId="13" pageNumber="100">dung trap</collectingMethod>
|
||
,
|
||
<collectorName id="26D55328FFFA6E16FB671107FACCFEAC" box="[1206,1295,258,284]" pageId="13" pageNumber="100">S. Peck</collectorName>
|
||
, (dung traps 39-41)—
|
||
<specimenCount id="9D26FD77FFFA6E16FF2F1123FEFFFEF1" box="[254,316,294,321]" count="2" pageId="13" pageNumber="100" type="male">2 ♂♂</specimenCount>
|
||
<typeStatus id="549B885CFFFA6E16FE901122FE6FFEF1" box="[321,428,295,321]" pageId="13" pageNumber="100" type="paratype">paratypes</typeStatus>
|
||
(
|
||
<collectionCode id="ED31AE3BFFFA6E16FE6B1123FDD5FEF0" box="[442,534,294,320]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="100">MNHN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B483CA3FFFA6E16FDF91123FE4BFED4" collectingDate="1974-07-15" collectingDateMax="1974-07-16" collectingDateMin="1974-07-15" collectorName="S. Peck" country="Papua New Guinea" latitude="-9.183333" location="Brown River" longLatPrecision="925" longitude="147.34166" pageId="13" pageNumber="100" specimenCount="1" specimenCount-male="1" stateProvince="National Capital" typeStatus="holotype">
|
||
<locationDeviation id="9A2D2FDCFFFA6E16FDF91123FCE4FEF0" box="[552,807,294,321]" location="Port Moresby" pageId="13" pageNumber="100">
|
||
<quantity id="4CD89B1BFFFA6E16FDF91123FDACFEF0" box="[552,623,294,320]" metricMagnitude="4" metricUnit="m" metricValue="4.828032" pageId="13" pageNumber="100" unit="mi" value="30.0">30 mi.</quantity>
|
||
N Port Moresby
|
||
</locationDeviation>
|
||
,
|
||
<location id="8EFF6025FFFA6E16FCE01122FC00FEF0" LSID="urn:lsid:plazi:treatment:038987E8FFFC6E16FF461757FAABFB10:8EFF6025FFFA6E16FCE01122FC00FEF0" box="[817,963,294,321]" country="Papua New Guinea" latitude="-9.183333" longLatPrecision="925" longitude="147.34166" name="Brown River" pageId="13" pageNumber="100" stateProvince="National Capital">Brown River</location>
|
||
, [
|
||
<geoCoordinate id="EE145039FFFA6E16FC051123FBF1FEF0" box="[980,1074,294,320]" degrees="09" direction="south" minutes="11" orientation="latitude" pageId="13" pageNumber="100" precision="925" value="-9.183333">09°11’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE145039FFFA6E16FBEC1123FB19FEF0" box="[1085,1242,294,320]" degrees="147" direction="east" minutes="20" orientation="longitude" pageId="13" pageNumber="100" precision="15" seconds="30" value="147.34166">147°20’30’’E</geoCoordinate>
|
||
],
|
||
<date id="FF9E103EFFFA6E16FB3D1123FA5BFEF1" box="[1260,1432,294,321]" pageId="13" pageNumber="100" value="1974-07-15" valueMax="1974-07-16" valueMin="1974-07-15">
|
||
<collectingDate id="EFDAE9D6FFFA6E16FB3D1123FA5BFEF1" box="[1260,1432,294,321]" pageId="13" pageNumber="100" value="1974-07-15" valueMax="1974-07-16" valueMin="1974-07-15">15–16.vii.1974</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26D55328FFFA6E16FF46114FFF2DFED4" box="[151,238,330,356]" pageId="13" pageNumber="100">S. Peck</collectorName>
|
||
—
|
||
<specimenCount id="9D26FD77FFFA6E16FEDB114FFEDFFED4" box="[266,284,330,356]" count="1" pageId="13" pageNumber="100" type="male">♂</specimenCount>
|
||
holotype
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B483CA3FFFA6E16FE42114FFD57FED4" box="[403,660,330,357]" collectionCode="CMNC" country="Papua New Guinea" pageId="13" pageNumber="100" specimenCount="1" specimenCount-male="1" stateProvince="Central" typeStatus="paratype">
|
||
<specimenCount id="9D26FD77FFFA6E16FE42114FFE7EFED5" box="[403,445,330,357]" count="1" pageId="13" pageNumber="100" type="male">1 ♂</specimenCount>
|
||
<typeStatus id="549B885CFFFA6E16FE15114EFDE7FED5" box="[452,548,331,357]" pageId="13" pageNumber="100" type="paratype">paratype</typeStatus>
|
||
(
|
||
<collectionCode id="ED31AE3BFFFA6E16FDE2114FFD4FFED4" box="[563,652,330,356]" collectionName="Canada, Ottawa, Canadian Museum of Nature" pageId="13" pageNumber="100">CMNC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A6575FFFA6E16FF16116BFD85FC30" pageId="13" pageNumber="100" type="description">
|
||
<paragraph id="8B9F36FEFFFA6E16FF16116BFD85FC30" blockId="13.[151,1437,150,1185]" pageId="13" pageNumber="100">
|
||
<emphasis id="B954EAECFFFA6E16FF16116BFDF0FE38" bold="true" box="[199,563,366,392]" pageId="13" pageNumber="100">Redescription. Measurements.</emphasis>
|
||
Body length
|
||
<quantity id="4CD89B1BFFFA6E16FD1F116BFC9BFE39" box="[718,856,366,393]" metricMagnitude="-3" metricUnit="m" metricValue="3.8499999999999996" metricValueMax="4.1" metricValueMin="3.6" pageId="13" pageNumber="100" unit="mm" value="3.8499999999999996" valueMax="4.1" valueMin="3.6">3.6–4.1 mm</quantity>
|
||
, maximum body width
|
||
<quantity id="4CD89B1BFFFA6E16FBB7116BFB32FE39" box="[1126,1265,366,393]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="13" pageNumber="100" unit="mm" value="2.5" valueMax="2.7" valueMin="2.3">2.3–2.7 mm</quantity>
|
||
.
|
||
<emphasis id="B954EAECFFFA6E16FB2C116BFA83FE38" bold="true" box="[1277,1344,366,392]" pageId="13" pageNumber="100">Body.</emphasis>
|
||
Overall body shape in dorsal view oval, elytral edge evenly rounded from base to apex.
|
||
<emphasis id="B954EAECFFFA6E16FB951197FB48FE1C" bold="true" box="[1092,1163,402,428]" pageId="13" pageNumber="100">Head.</emphasis>
|
||
Clypeal edge lateral to clypeal teeth emarginate (
|
||
<figureCitation id="131B2A7BFFFA6E16FE6F11B3FDD9FE60" box="[446,538,438,464]" captionStart="FIGURE 4" captionStartId="7.[152,255,934,958]" captionTargetBox="[151,1435,182,906]" captionTargetId="figure-17@7.[151,1435,179,909]" captionTargetPageId="7" captionText="FIGURE 4. Diagnostic characters of Rhytomus. A, dorsal head showing emargination and interocular space (R. howdeni); B, dorsal head showing lack of emargination and interocular space (R. fervidus) C, ventral view of male foreleg (R. latidentatus); D, lateral view of elytra (R. vanus); E, dorsal view of male protibia (R. latidentatus)." figureDoi="http://doi.org/10.5281/zenodo.14746450" httpUri="https://zenodo.org/record/14746450/files/figure.png" pageId="13" pageNumber="100">Fig. 4D</figureCitation>
|
||
), head margin at clypeogenal suture of females moderately protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 5.6 times eye width.
|
||
<emphasis id="B954EAECFFFA6E16FBA311DFFB31FE44" bold="true" box="[1138,1266,474,500]" pageId="13" pageNumber="100">Pronotum.</emphasis>
|
||
Pronotal width to length along midline ratio 1.9, lateral pronotal edge sharply defined throughout, pronotal pubescence short and fine, pronotal basal portion along midline lacking fine longitudinal depression.
|
||
<emphasis id="B954EAECFFFA6E16FBFC1227FB42FD8C" bold="true" box="[1069,1153,546,572]" pageId="13" pageNumber="100">Elytra.</emphasis>
|
||
Elytral maximum width to length along suture ratio 1.1, elytral outline moderately convex in lateral view, elytral pubescence short and fine, elytral surface between punctures appearing glossy, at most with ill-defined microsculptures, basal elytral pit slightly wider than stria 10 and shallow, basal surface of pseudepipleuron with sharply defined simple punctures, interstriae 3–5 on basal fifth with scattered unaligned punctures.
|
||
<emphasis id="B954EAECFFFA6E16FC5312B7FC02FD7C" bold="true" box="[898,961,690,716]" pageId="13" pageNumber="100">Legs.</emphasis>
|
||
Profemoral anterior surface flat or might appear furrowed in some individuals, male metatibial apicointernal tooth approximately in line with posterior tibial edge.
|
||
<emphasis id="B954EAECFFFA6E16FF0C12FFFE94FCA4" bold="true" box="[221,343,762,788]" pageId="13" pageNumber="100">Pygidium.</emphasis>
|
||
Pygidial groove transverse, narrowed medially, and terminating in a pit at each end (
|
||
<figureCitation id="131B2A7BFFFA6E16FAE512FFFA4DFCA4" box="[1332,1422,762,788]" captionStart="FIGURE 8" captionStartId="11.[152,255,1592,1616]" captionTargetBox="[154,1432,921,1562]" captionTargetId="figure-331@11.[151,1435,918,1565]" captionTargetPageId="11" captionText="FIGURE 8. Pygidium of Rhytomus species. A, male R. fervidus; B. female R. heurni; C, male R. howdeni; D, male R. larseni; E, male R. latidentatus; F, male R. papuanus; G, female R. setulosus; H, male R. sodalis; I, male R. vanus. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14746460" httpUri="https://zenodo.org/record/14746460/files/figure.png" pageId="13" pageNumber="100">Fig. 8C</figureCitation>
|
||
), pygidial surface along midline with punctures similar in density to rest of discs, pygidial surface along basal groove smooth between punctures, pygidium of female evenly convex.
|
||
<emphasis id="B954EAECFFFA6E16FCBB1347FC27FCEC" bold="true" box="[874,996,834,860]" pageId="13" pageNumber="100">Aedeagus.</emphasis>
|
||
Left paramere apex truncated apically, distinctly angular externally (
|
||
<figureCitation id="131B2A7BFFFA6E16FE301363FDFAFC30" box="[481,569,870,896]" captionStart="FIGURE 9" captionStartId="12.[152,255,1959,1983]" captionTargetBox="[155,1431,238,1929]" captionTargetId="figure-53@12.[151,1435,234,1933]" captionTargetPageId="12" captionText="FIGURE 9. Male aedeagus of Rhytomus species, left lateral, dorsal, right lateral.A, R. elongatus; B, R. fervidus; C, R. howdeni; D, R. larseni. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14746462" httpUri="https://zenodo.org/record/14746462/files/figure.png" pageId="13" pageNumber="100">Fig. 9C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A6575FFFA6E16FF16138FFAABFB10" pageId="13" pageNumber="100" type="discussion">
|
||
<paragraph id="8B9F36FEFFFA6E16FF16138FFAABFB10" blockId="13.[151,1437,150,1185]" pageId="13" pageNumber="100">
|
||
<emphasis id="B954EAECFFFA6E16FF16138FFEF7FC14" bold="true" box="[199,308,906,932]" pageId="13" pageNumber="100">Remarks</emphasis>
|
||
. Examination of the
|
||
<typeStatus id="549B885CFFFA6E16FDF9138FFD41FC14" box="[552,642,906,932]" pageId="13" pageNumber="100" type="allotype">allotype</typeStatus>
|
||
revealed that the specimen labeled by Paulian is actually a male and is here considered a male
|
||
<typeStatus id="549B885CFFFA6E16FE1913AAFDEBFC79" box="[456,552,943,969]" pageId="13" pageNumber="100" type="paratype">paratype</typeStatus>
|
||
in the material examined. Additionally, the date ranges provided by
|
||
<bibRefCitation id="EFB14B0FFFFA6E16FA9913ABFF21FC5D" author="Paulian, R." pageId="13" pageNumber="100" pagination="219 - 238" refId="ref16422" refString="Paulian, R. (1985) Les coleopteres Scarabaeidae Canthonides de Nouvelle-Guinee. Annales de la Societe Entomologique de France, Nouvelle Serie, 21 (2), 219 - 238. https: // doi. org / 10.1080 / 21686351.1985.12278409" type="journal article" year="1985">Paulian (1985)</bibRefCitation>
|
||
in the original description do not match any label data from the specimens studied. Recent examination of the specimens at the MNHN (by FG) confirmed that both are males, indicating that the female specimen described by Paulian is either missing or that Paulian misidentified the gender of the
|
||
<typeStatus id="549B885CFFFA6E16FC38141FFB84FB84" box="[1001,1095,1050,1076]" pageId="13" pageNumber="100" type="allotype">allotype</typeStatus>
|
||
.
|
||
<specimenCount id="9D26FD77FFFA6E16FB83141EFB14FB84" box="[1106,1239,1050,1077]" count="2" pageId="13" pageNumber="100" type="female">Two female</specimenCount>
|
||
specimens in the CMNC are not labeled as
|
||
<typeStatus id="549B885CFFFA6E16FE13143AFDF2FBE9" box="[450,561,1087,1113]" pageId="13" pageNumber="100" type="paratype">paratypes</typeStatus>
|
||
. While one of these could theoretically be the
|
||
<typeStatus id="549B885CFFFA6E16FB94143BFB61FBE8" box="[1093,1186,1086,1112]" pageId="13" pageNumber="100" type="allotype">allotype</typeStatus>
|
||
, it is unclear if Henry Howden ever sent them to Paulian for study, as he often retained some specimens as a precaution against parcels being lost in the mail. Consequently, these specimens cannot be definitively considered part of the type series.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |