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<mods:title id="497766FC9A4438E0AFA8E4EC18D3FEC8">A new species and subgenus of Elseya (Testudines: Pleurodira: Chelidae) from New Guinea</mods:title>
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<heading id="D0D781F0A811FFE0FF01FF4EFC0AFF6F" bold="true" box="[151,929,151,178]" fontSize="11" level="1" pageId="9" pageNumber="68" reason="1">
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<taxonomicName id="4C204D1FA811FFE0FF01FF4EFCAAFF6F" authority="Vogt 1911: 410" authorityName="Vogt" authorityPageNumber="410" authorityYear="1911" box="[151,769,151,178]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="68" phylum="Chordata" rank="species" species="schultzei" subGenus="Hanwarachelys">
<emphasis id="B954EA8EA811FFE0FF01FF4EFF4CFF6C" bold="true" box="[151,231,151,177]" italics="true" pageId="9" pageNumber="68">Elseya</emphasis>
(
<emphasis id="B954EA8EA811FFE0FF6EFF4EFE1EFF6C" bold="true" box="[248,437,151,177]" italics="true" pageId="9" pageNumber="68">Hanwarachelys</emphasis>
)
<emphasis id="B954EA8EA811FFE0FE53FF4EFD84FF6C" bold="true" box="[453,559,151,177]" italics="true" pageId="9" pageNumber="68">schultzei</emphasis>
(
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<bibRefCitation id="EFB14B6DA811FFE0FDA8FF41FD52FF6F" author="Vogt" box="[574,761,152,178]" pageId="9" pageNumber="68" refString="Vogt, T. (1911) Emydura schultzei, sp. nov. Reptilien und Amphibien aus Neu Guinea. Sber. ges. naturf. Freunde, Berlin, 9, 410 - 412." type="journal article" year="1911">Vogt 1911: 410</bibRefCitation>
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)
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(
<figureCitation id="131B2A19A811FFE0FC86FF41FC32FF6F" box="[784,921,152,178]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,250,1981,2003]" captionStartId-1="9.[151,250,1979,2001]" captionTargetBox-0="[170,1414,1206,1942]" captionTargetBox-1="[167,1408,1195,1937]" captionTargetId-0="figure@8.[151,1436,1184,1959]" captionTargetId-1="figure@9.[151,1435,1179,1957]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Dorsal views of the holotypes of A. Elseya (H) schultzei (ZMB 22182) and B. Elseya (H) novaeguineae (MTD D 8222)." captionText-1="FIGURE 7. Ventral views of the holotypes of A. Elseya (H) schultzei (ZMB 22182) and B. Elseya (H) novaeguineae (MTD D 8222)." httpUri-0="https://zenodo.org/record/289268/files/figure.png" httpUri-1="https://zenodo.org/record/289269/files/figure.png" pageId="9" pageNumber="68">Figure 67</figureCitation>
)
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Northern New
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Stream Turtle
</paragraph>
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1911
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<taxonomicName id="4C204D1FA811FFE0FEB8FF2FFE20FED1" box="[302,395,246,268]" class="Reptilia" family="Chelidae" genus="Emydura" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="68" phylum="Chordata" rank="genus">Emydura</taxonomicName>
schultzei—
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<treatmentCitation id="0A81108DA811FFE0FD96FF2EFD3FFED1" author="Vogt" box="[512,660,246,268]" page="410" pageId="9" pageNumber="68" year="1911">
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:410
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<treatmentCitationGroup id="AB3011B2A811FFE0FF74FECCFBA3FE5A" pageId="9" pageNumber="68">
2000
<emphasis id="B954EA8EA811FFE0FEB8FECCFE4CFEF6" box="[302,487,277,299]" italics="true" pageId="9" pageNumber="68">
<taxonomicName id="4C204D1FA811FFE0FEB8FECCFED9FEF6" box="[302,370,277,299]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="68" phylum="Chordata" rank="genus">Elseya</taxonomicName>
schultzei—
</emphasis>
Samedi and
<treatmentCitation id="0A81108DA811FFE0FDF2FECCFCB4FEF6" author="Iskandar" box="[612,799,277,299]" page="111" pageId="9" pageNumber="68" year="2000">
<bibRefCitation id="EFB14B6DA811FFE0FDF2FECCFD52FEF6" author="Iskandar" box="[612,761,277,299]" pageId="9" pageNumber="68" refString="Iskandar, D. T. (2000) Turtles and Crocodiles of Insular Southeast Asia and New Guinea. Institute of Technology, Bandung, 191 pp." type="book" year="2000">Iskandar 2000</bibRefCitation>
:111
</treatmentCitation>
2010
<taxonomicName id="4C204D1FA811FFE0FEB8FEEDFD86FE97" box="[302,557,308,330]" class="Reptilia" family="Chelidae" genus="Myuchelys" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="68" phylum="Chordata" rank="species" species="novaeguineae">
<emphasis id="B954EA8EA811FFE0FEB8FEEDFD86FE97" box="[302,557,308,330]" italics="true" pageId="9" pageNumber="68">Myuchelys novaeguineae</emphasis>
</taxonomicName>
<treatmentCitation id="0A81108DA811FFE0FDD1FEEDFC2EFE97" author="Georges" box="[583,901,308,330]" page="29" pageId="9" pageNumber="68" year="2010">
<bibRefCitation id="EFB14B6DA811FFE0FDD1FEEDFCC3FE97" author="Georges" box="[583,872,308,330]" pageId="9" pageNumber="68" refString="Georges, A. &amp; Thomson, S. (2010) Diversity of Australasian freshwater turtles, with an annotated synonymy and keys to species. Zootaxa, 2496, 1 - 37." type="journal article" year="2010">Georges and Thomson 2010</bibRefCitation>
:29
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<taxonomicName id="4C204D1FA811FFE0FEB8FE8BFDAFFEB5" box="[302,516,338,360]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="68" phylum="Chordata" rank="species" species="novaeguineae">
<emphasis id="B954EA8EA811FFE0FEB8FE8BFDAFFEB5" box="[302,516,338,360]" italics="true" pageId="9" pageNumber="68">Elseya novaeguineae</emphasis>
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<treatmentCitation id="0A81108DA811FFE0FD88FE8BFCADFEB5" author="Rhodin" box="[542,774,338,360]" page="131" pageId="9" pageNumber="68" year="2010">
<bibRefCitation id="EFB14B6DA811FFE0FD88FE8BFD75FEB5" author="Rhodin" box="[542,734,338,360]" pageId="9" pageNumber="68" refString="Rhodin, A. G. J., van Dijk, P. P., Iverson, J. B. &amp; Shaffer, H. B. (2010) Turtles of the World, 2010 Update: Annotated Checklist of Taxonomy, Synonymy, Distribution, and Conservation Status Cheloinian Research Monographs, 5, 85 - 161. http: // dx. doi. org / 10.3854 / crm. 5.000. checklist. v 3.2010" type="journal article" year="2010">Rhodin et al. 2010</bibRefCitation>
:131
</treatmentCitation>
2015
<taxonomicName id="4C204D1FA811FFE0FEB8FEA8FD2CFE5A" box="[302,647,369,391]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="68" phylum="Chordata" rank="species" species="schultzei" subGenus="Hanwarachelys">
<emphasis id="B954EA8EA811FFE0FEB8FEA8FED9FE5A" box="[302,370,369,391]" italics="true" pageId="9" pageNumber="68">Elseya</emphasis>
(
<emphasis id="B954EA8EA811FFE0FE16FEA8FDB5FE5A" box="[384,542,369,391]" italics="true" pageId="9" pageNumber="68">Hanwarachelys</emphasis>
)
<emphasis id="B954EA8EA811FFE0FDB8FEA8FD2CFE5A" box="[558,647,369,391]" italics="true" pageId="9" pageNumber="68">schultzei</emphasis>
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resurrected, Thomson et al. this study
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data
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: Holotype—ZMB 22182 collected near Sae village, Seko coast, near Skosai, ca
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W. mouth of Tami River,
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,
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(2˚37'S,140˚54'E), restricted this study.
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<paragraph id="8B9F369CA811FFE0FF51FE2CFDDFFDA5" blockId="9.[151,1437,429,1129]" pageId="9" pageNumber="68">
<emphasis id="B954EA8EA811FFE0FF51FE2CFE91FDD3" bold="true" box="[199,314,501,526]" pageId="9" pageNumber="68">Diagnosis</emphasis>
—Eyes typically green in the sclera with a bright gold iris which is distinct. Head shield entire, extending down the parietal arches to broadly contact the tympanum. Head shield narrower than in
<taxonomicName id="4C204D1FA811FFE0FAC7FDC0FE98FD89" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="68" phylum="Chordata" rank="species" species="novaeguineae">
<emphasis id="B954EA8EA811FFE0FAC7FDC0FE98FD89" italics="true" pageId="9" pageNumber="68">Elseya novaeguineae</emphasis>
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but wider than in
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<emphasis id="B954EA8EA811FFE0FD8BFDE4FD85FD89" box="[541,558,573,596]" italics="true" pageId="9" pageNumber="68">E</emphasis>
.
<emphasis id="B954EA8EA811FFE0FDD4FDE5FD3FFD89" box="[578,660,572,596]" italics="true" pageId="9" pageNumber="68">rhodini</emphasis>
</taxonomicName>
. No contact between the vomer and the pterygoids of the skull, distinguishing the species from
<taxonomicName id="4C204D1FA811FFE0FE6DFDB8FDDBFDA5" box="[507,624,608,632]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="68" phylum="Chordata" rank="species" species="rhodini">
<emphasis id="B954EA8EA811FFE0FE6DFDB8FDA7FDA5" box="[507,524,609,632]" italics="true" pageId="9" pageNumber="68">E</emphasis>
.
<emphasis id="B954EA8EA811FFE0FD8CFDB9FDDBFDA5" box="[538,624,608,632]" italics="true" pageId="9" pageNumber="68">rhodini</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
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<emphasis id="B954EA8EA811FFE0FF51FD5AFEF2FD41" bold="true" box="[199,345,643,668]" pageId="9" pageNumber="68">Distribution</emphasis>
—Streams draining into the rivers of New
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north of the central ranges—i.e. not including the Vogelkopf and Bomberai peninsulas (Birds Head)—and east to at least Madang.
</paragraph>
<paragraph id="8B9F369CA811FFE0FF51FD13FBC0FBB5" blockId="9.[151,1437,429,1129]" pageId="9" pageNumber="68">
<emphasis id="B954EA8EA811FFE0FF51FD13FE0CFD3E" bold="true" box="[199,423,714,739]" pageId="9" pageNumber="68">Collection History</emphasis>
<emphasis id="B954EA8EA811FFE0FE31FD12FE6BFD3F" box="[423,448,715,738]" italics="true" pageId="9" pageNumber="68"></emphasis>
This taxon was described by
<bibRefCitation id="EFB14B6DA811FFE0FCB4FD12FC06FD3F" author="Vogt" box="[802,941,714,739]" pageId="9" pageNumber="68" refString="Vogt, T. (1911) Emydura schultzei, sp. nov. Reptilien und Amphibien aus Neu Guinea. Sber. ges. naturf. Freunde, Berlin, 9, 410 - 412." type="journal article" year="1911">Vogt (1911)</bibRefCitation>
based on a single specimen collected by Leonhard Schultze in an unnamed river west of the Tami River in northeastern
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,
<collectingCountry id="F337760CA811FFE0FB1BFD34FB51FCDB" box="[1165,1274,749,774]" name="Indonesia" pageId="9" pageNumber="68">Indonesia</collectingCountry>
(
<typeStatus id="549B883EA811FFE0FA9AFD37FA95FCDB" box="[1292,1342,750,774]" pageId="9" pageNumber="68">type</typeStatus>
locality “Fluss westlich der Tamimündung” [river west of the mouth of the Tami]). Although it is impossible to give a precise locality,
<bibRefCitation id="EFB14B6DA811FFE0FED8FCEDFDABFC90" author="Schultze" box="[334,512,820,845]" pageId="9" pageNumber="68" refString="Schultze, L. (1914) Forschungen im Innern der Insel Neu Guinea (Bericht des Fuhrers uber die wissenschaftlichen Ergebnisse der deutschen Grenzexpedition in das westliche Kaiser-Wilhelmsland 1910). Mitteilungen aus den Deutschen Schutzgebiet, 11, 1 - 100." type="journal article" year="1914">Schultze (1914)</bibRefCitation>
gave an account of his travels and a detailed map from which one can deduce that the
<typeStatus id="549B883EA811FFE0FF54FC81FF59FCAD" box="[194,242,856,880]" pageId="9" pageNumber="68">type</typeStatus>
specimen was probably collected near Sae village, Seko coast, near Skosai, ca
<quantity id="4CD89B79A811FFE0FB10FC8EFB6AFCAD" box="[1158,1217,855,880]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="68" unit="km" value="5.0">5 km</quantity>
W. mouth of Tami River,
<collectingCountry id="F337760CA811FFE0FF70FCA5FE86FC4E" box="[230,301,892,915]" name="Papua New Guinea" pageId="9" pageNumber="68">Papua</collectingCountry>
,
<collectingCountry id="F337760CA811FFE0FEAAFCA2FE02FC49" box="[316,425,891,916]" name="Indonesia" pageId="9" pageNumber="68">Indonesia</collectingCountry>
(2˚37'S,140˚54'E); and the
<typeStatus id="549B883EA811FFE0FD7BFCA5FCB6FC49" box="[749,797,892,916]" pageId="9" pageNumber="68">type</typeStatus>
locality is hereby so restricted. The
<typeStatus id="549B883EA811FFE0FB59FCA2FA9AFC49" box="[1231,1329,891,916]" pageId="9" pageNumber="68" type="holotype">holotype</typeStatus>
is in the Berlin collection (ZMB 22182, figured by
<bibRefCitation id="EFB14B6DA811FFE0FDE3FC47FCA9FC6A" author="Goode" box="[629,770,926,951]" pageId="9" pageNumber="68" refString="Goode, J. (1967) Freshwater Tortoises of Australia and New Guinea (in the Family Chelidae). Lansdowne Press, Melbourne, 154 pp." type="book" year="1967">Goode 1967</bibRefCitation>
). When described,
<taxonomicName id="4C204D1FA811FFE0FC4EFC46FB03FC6A" box="[984,1192,927,951]" class="Reptilia" family="Chelidae" genus="Emydura" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="68" phylum="Chordata" rank="species" species="schultzei">
<emphasis id="B954EA8EA811FFE0FC4EFC46FB03FC6A" box="[984,1192,927,951]" italics="true" pageId="9" pageNumber="68">Emydura schultzei</emphasis>
</taxonomicName>
was compared to and distinguished from a specimen of “
<taxonomicName id="4C204D1FA811FFE0FD8AFC1BFCA4FC07" box="[540,783,962,986]" class="Reptilia" family="Chelidae" genus="Emydura" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="68" phylum="Chordata" rank="species" species="macquariae">
<emphasis id="B954EA8EA811FFE0FD8AFC1BFCA4FC07" box="[540,783,962,986]" italics="true" pageId="9" pageNumber="68">Emydura macquariae</emphasis>
</taxonomicName>
” (ZMB 22210, figured by
<bibRefCitation id="EFB14B6DA811FFE0FBADFC1BFB63FC06" author="Goode" box="[1083,1224,962,987]" pageId="9" pageNumber="68" refString="Goode, J. (1967) Freshwater Tortoises of Australia and New Guinea (in the Family Chelidae). Lansdowne Press, Melbourne, 154 pp." type="book" year="1967">Goode 1967</bibRefCitation>
:67) from the upper Sepik River basin south of the headwaters of the Tami River, near the Indonesia-Papua New
<collectingCountry id="F337760CA811FFE0FB56FC3CFABAFC23" box="[1216,1297,997,1022]" name="Guinea-Bissau" pageId="9" pageNumber="68">Guinea</collectingCountry>
border. The species was recognized as distinct until synonymized with
<taxonomicName id="4C204D1FA811FFE0FCA9FBD3FC56FBFC" box="[831,1021,1034,1057]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="68" phylum="Chordata" rank="species" species="novaeguineae">
<emphasis id="B954EA8EA811FFE0FCA9FBD3FCFBFBFC" box="[831,848,1034,1057]" italics="true" pageId="9" pageNumber="68">E</emphasis>
.
<emphasis id="B954EA8EA811FFE0FCF7FBD3FC56FBFC" box="[865,1021,1034,1057]" italics="true" pageId="9" pageNumber="68">novaeguineae</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFB14B6DA811FFE0FBBAFBD0FB66FBFF" author="Goode" box="[1068,1229,1033,1058]" pageId="9" pageNumber="68" refString="Goode, J. (1967) Freshwater Tortoises of Australia and New Guinea (in the Family Chelidae). Lansdowne Press, Melbourne, 154 pp." type="book" year="1967">Goode (1967)</bibRefCitation>
. That
<taxonomicName id="4C204D1FA811FFE0FA8EFBD3FA37FBFC" box="[1304,1436,1033,1057]" class="Reptilia" family="Chelidae" genus="Emydura" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="68" phylum="Chordata" rank="species" species="schultzei">
<emphasis id="B954EA8EA811FFE0FA8EFBD3FA37FBFC" box="[1304,1436,1033,1057]" italics="true" pageId="9" pageNumber="68">E. schultzei</emphasis>
</taxonomicName>
represents a distinct taxa has been confirmed by recent genetic studies (Georges
<emphasis id="B954EA8EA811FFE0FB8AFBF4FBFEFB98" box="[1052,1109,1069,1093]" italics="true" pageId="9" pageNumber="68">et al.</emphasis>
2014). Therefore we remove this species from the synonymy of
<taxonomicName id="4C204D1FA811FFE0FD89FB89FCA7FBB5" box="[543,780,1104,1128]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="68" phylum="Chordata" rank="species" species="novaeguineae">
<emphasis id="B954EA8EA811FFE0FD89FB89FCA7FBB5" box="[543,780,1104,1128]" italics="true" pageId="9" pageNumber="68">Elseya novaeguineae</emphasis>
</taxonomicName>
recognizing it as a valid taxon.
</paragraph>
</subSubSection>
</treatment>
</document>