206 lines
33 KiB
XML
206 lines
33 KiB
XML
<document id="33CB8B678AB47A69A85B7D535094583A" ID-CLB-Dataset="38930" ID-DOI="10.11646/zootaxa.4118.1.1" ID-GBIF-Dataset="a4e408c4-2575-4b5c-878b-e1b9010ec619" ID-ISSN="1175-5326" ID-Zenodo-Dep="264597" ID-ZooBank="CFA586DA-10EE-468B-80EE-35351E3845FD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="guilherme" checkinTime="1465242077321" checkinUser="plazi" docAuthor="C, Lees David" docDate="2016" docId="038747324C78C6471EB729E2FC722384" docLanguage="en" docName="zootaxa.4118.pdf.imf" docOrigin="Zootaxa 4118 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Gallienia alaokola" docType="treatment" docVersion="12" lastPageNumber="15" masterDocId="FFBE3F4A4C76C6571E202D2DFFD32174" masterDocTitle="Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision" masterLastPageNumber="97" masterPageNumber="1" pageNumber="15" updateTime="1735917000942" updateUser="ExternalLinkService">
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<mods:title id="A289CB6842787D8F32B42C0588805FCC">Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision</mods:title>
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<mods:namePart id="2A460F6A48776CC8F17F6916017EC55F">C, Lees David</mods:namePart>
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<mods:title id="3D07A7F091DE3DFB22F5957050D1FB54">Zootaxa</mods:title>
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<mods:date id="C2C8B7A1BF2662B14C1296B991F496D5">2016</mods:date>
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<mods:number id="D0DEB92108D0FBE77E06EF6FE917E7A0">4118</mods:number>
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<mods:classification id="0326F5097D95C77FD8C21F2EBA793EFE">journal article</mods:classification>
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<mods:identifier id="52DE3DDBF42C08FB81104EB5DCF15863" type="CLB-Dataset">38930</mods:identifier>
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<mods:identifier id="9B9E6A01A558CA464C7B2638751F0C48" type="DOI">10.11646/zootaxa.4118.1.1</mods:identifier>
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<treatment id="038747324C78C6471EB729E2FC722384" ID-DOI="http://doi.org/10.5281/zenodo.6086412" ID-GBIF-Taxon="120817077" ID-Zenodo-Dep="6086412" LSID="urn:lsid:plazi:treatment:038747324C78C6471EB729E2FC722384" httpUri="http://treatment.plazi.org/id/038747324C78C6471EB729E2FC722384" lastPageId="16" lastPageNumber="15" pageId="14" pageNumber="15" scope_family="Nymphalidae" scope_subFamily="Satyrinae" scope_tribe="Satyrini">
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<subSubSection id="C334A5AF4C78C6471EB729E2FC722384" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15" type="nomenclature">
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<paragraph id="8B91F6244C78C6591EB729E2FE34259E" blockId="14.[151,487,1231,1258]" box="[151,487,1231,1258]" pageId="14" pageNumber="15">
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<heading id="D0D941484C78C6591EB729E2FE34259E" bold="true" box="[151,487,1231,1258]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
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<emphasis id="B95A2A364C78C6591EB729E2FE34259E" bold="true" box="[151,487,1231,1258]" pageId="14" pageNumber="15">
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<taxonomicName id="4C2E8DA74C78C6591EB729E2FEA5259D" ID-CoL="953DN" box="[151,374,1231,1257]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="alaokola">
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<emphasis id="B95A2A364C78C6591EB729E2FEA5259D" bold="true" box="[151,374,1231,1257]" italics="true" pageId="14" pageNumber="15">Gallienia alaokola</emphasis>
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</taxonomicName>
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problem
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B91F6244C78C6471EB7283BFC722384" blockId="14.[151,1436,1302,2014]" lastBlockId="16.[151,1437,151,752]" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15">
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The left image (#3043) in
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<figureCitation id="1315EAA14C78C6591F9E283BFE2D245B" box="[446,510,1302,1327]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="14" pageNumber="15">Fig. 3</figureCitation>
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A represents the specimen (BMNH(E) 672621) that P. Viette intended to represent the LT of
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<taxonomicName id="4C2E8DA74C78C6591F282817FD402427" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" box="[264,659,1338,1363]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="alaokola">
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<emphasis id="B95A2A364C78C6591F282817FE0E2427" box="[264,477,1338,1363]" italics="true" pageId="14" pageNumber="15">Gallienia alaokola</emphasis>
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<bibRefCitation id="EFBF8BD54C78C6591FC42817FD402427" author="Oberthur" box="[484,659,1338,1363]" pageId="14" pageNumber="15" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür, 1916</bibRefCitation>
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</taxonomicName>
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, according to a label under the specimen (but it is certain that Viette never published this). That on the right (#3044) represents the specimen that
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<bibRefCitation id="EFBF8BD54C78C6591DD02873FB732403" author="d'Abrera" box="[1008,1184,1374,1399]" pageId="14" pageNumber="15" refString="d'Abrera, B. L. (1980) Butterflies of the Afrotropical Region. Lansdowne Editions, Melbourne, 593 pp." type="book" year="1980">d’Abrera (1980</bibRefCitation>
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,
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<bibRefCitation id="EFBF8BD54C78C6591A8A2872FB302403" author="d'Abrera" box="[1194,1251,1375,1399]" pageId="14" pageNumber="15" refString="d'Abrera, B. L. (1997) Butterflies of the Afrotropical Region. Part 1. Papilionidae, Pieridae, Acraeidae, Danaidae, Satyridae. Hill House Publishers, Melbourne & London, 263 pp." type="book" year="1997">1997</bibRefCitation>
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) illustrated as [a ♀ of] ‘
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<emphasis id="B95A2A364C78C6591ECA28A9FEDA24EF" box="[234,265,1412,1435]" italics="true" pageId="14" pageNumber="15">M.</emphasis>
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?
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<taxonomicName id="4C2E8DA74C78C6591F0828AFFE5924EF" box="[296,394,1410,1435]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="alaokola">
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<emphasis id="B95A2A364C78C6591F0828AFFE5924EF" box="[296,394,1410,1435]" italics="true" pageId="14" pageNumber="15">alaokola</emphasis>
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</taxonomicName>
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’. I contend, considering dorsal colouration, ocellus expression in space-M3 and relatively more pointed HW tails, that both specimens in fact represent
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<taxonomicName id="4C2E8DA74C78C6591D6F288AFC0524CB" box="[847,982,1446,1471]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="mabillei">
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<emphasis id="B95A2A364C78C6591D6F288AFC0524CB" box="[847,982,1446,1471]" italics="true" pageId="14" pageNumber="15">Ht. mabillei</emphasis>
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</taxonomicName>
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(both surfaces of the HT ♂ of
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<taxonomicName id="4C2E8DA74C78C6591B18288BFE582497" authority="Butler, 1879" authorityName="Butler" authorityYear="1879" class="Insecta" family="Nymphalidae" genus="Strabena" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="mabillei">
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<emphasis id="B95A2A364C78C6591B18288BFF202497" italics="true" pageId="14" pageNumber="15">Strabena mabillei</emphasis>
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<bibRefCitation id="EFBF8BD54C78C6591EDC28E7FE582497" author="Butler" box="[252,395,1482,1507]" pageId="14" pageNumber="15" refString="Butler, A. G. (1879) Descriptions of new species of Lepidoptera from Madagascar, with notes on some of the forms already described. Annals and Magazine of Natural History, Series 5, 4, 227 - 246. http: // dx. doi. org / 10.1080 / 00222937908679820" type="journal article" year="1879">Butler, 1879</bibRefCitation>
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</taxonomicName>
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, BMNH(E) 672624, are figured in
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<figureCitation id="1315EAA14C78C6591D0028E7FCB72497" box="[800,868,1482,1507]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="14" pageNumber="15">Fig. 3</figureCitation>
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C). Moreover, although
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<bibRefCitation id="EFBF8BD54C78C6591A5828E7FAF82497" author="d'Abrera" box="[1144,1323,1482,1507]" pageId="14" pageNumber="15" refString="d'Abrera, B. L. (1980) Butterflies of the Afrotropical Region. Lansdowne Editions, Melbourne, 593 pp." type="book" year="1980">d’Abrera (1980</bibRefCitation>
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: 190) had placed a red label next to photograph of “
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<taxonomicName id="4C2E8DA74C78C6591C4C28C2FD2F2773" box="[620,764,1518,1543]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="alaokola">
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<emphasis id="B95A2A364C78C6591C4C28C2FD2F2773" box="[620,764,1518,1543]" italics="true" pageId="14" pageNumber="15">M. alaokola</emphasis>
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</taxonomicName>
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”, the accompanying text says “♀ (?
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<typeStatus id="549548864C78C6591ABB28C2FB032773" box="[1179,1232,1519,1543]" pageId="14" pageNumber="15">type</typeStatus>
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)”, and by his next edition,
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<bibRefCitation id="EFBF8BD54C78C6591ED92B3FFE66275F" box="[249,437,1554,1579]" pageId="14" pageNumber="15" refString="d'Abrera, B. L. (1997) Butterflies of the Afrotropical Region. Part 1. Papilionidae, Pieridae, Acraeidae, Danaidae, Satyridae. Hill House Publishers, Melbourne & London, 263 pp." type="book">d’Abrera (1997)</bibRefCitation>
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had removed the red label. Without DNA data, choice of a ♀
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<typeStatus id="549548864C78C6591A9B2B3FFAF1275F" box="[1211,1314,1554,1579]" pageId="14" pageNumber="15" type="lectotype">lectotype</typeStatus>
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would be extremely unsatisfactory in this difficult-to-distinguish species group, and thus unlikely to promote stability. It appears that d’Abrera’s (1980) action does not, however, represent a valid lectotypification, since it is not an unambiguous selection of the name-bearing
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<typeStatus id="549548864C78C6591CBA2B52FD1E27E3" box="[666,717,1663,1687]" pageId="14" pageNumber="15">type</typeStatus>
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, considering in particular his qualification of use of the word 'type' by a question mark (Art. 74.5). A proportion of Oberthür’s
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<typeStatus id="549548864C78C6591DA92B8EFC3327CF" box="[905,992,1699,1723]" pageId="14" pageNumber="15" type="syntype">syntype</typeStatus>
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specimens of
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<taxonomicName id="4C2E8DA74C78C6591AA72B8FFA8F27CF" box="[1159,1372,1698,1723]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="alaokola">
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<emphasis id="B95A2A364C78C6591AA72B8FFA8F27CF" box="[1159,1372,1698,1723]" italics="true" pageId="14" pageNumber="15">Gallienia alaokola</emphasis>
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</taxonomicName>
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from Fianarantsoa and some also from Antsianaka (the current region around RNI Zahamena) have apparently generally shorter HW tails than in the HT of
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<taxonomicName id="4C2E8DA74C78C6591C032BC7FCEB2677" authority="Butler" authorityName="Butler" box="[547,824,1770,1795]" class="Insecta" family="Nymphalidae" genus="Strabena" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="mabillei">
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<emphasis id="B95A2A364C78C6591C032BC7FD382677" box="[547,747,1770,1795]" italics="true" pageId="14" pageNumber="15">Strabena mabillei</emphasis>
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Butler
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</taxonomicName>
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(
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<figureCitation id="1315EAA14C78C6591D682BC7FC592677" box="[840,906,1770,1795]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="14" pageNumber="15">Fig. 3</figureCitation>
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C) and a much darker upperside. They lack the ocellus expression in both space-M3 and CuA1 of the ♂ HW upperside, with sometimes neither ocellus expressed (this variation in his mixed series is alluded to in Oberthür’s 1916: 205–206 description). Among Oberthür’s 75
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<typeStatus id="549548864C78C6591EB72A7AFF2F261B" box="[151,252,1879,1903]" pageId="14" pageNumber="15" type="syntype">syntypes</typeStatus>
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, a LT ♂ specimen of
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<taxonomicName id="4C2E8DA74C78C6591FC92A7BFD68261B" box="[489,699,1878,1903]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="alaokola">
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<emphasis id="B95A2A364C78C6591FC92A7BFD68261B" box="[489,699,1878,1903]" italics="true" pageId="14" pageNumber="15">Gallienia alaokola</emphasis>
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</taxonomicName>
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is here therefore chosen to resolve the problem. This LT ♂ is here designated as the specimen (
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<figureCitation id="1315EAA14C78C6591FC22A50FDF526E2" box="[482,550,1917,1942]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="14" pageNumber="15">Fig. 3</figureCitation>
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B) bearing data labels “
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<collectingCountry id="F339B6B44C78C6591D182A50FC1726E2" box="[824,964,1917,1942]" name="Madagascar" pageId="14" pageNumber="15">Madagascar</collectingCountry>
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, Fianarantsoa, Perrot Frères, 2e Semestre 1892|Ex Oberthür Coll. Brit. Mus. 1927-3|Photographed by B. d'Abrera 77/78|BMNH(E) 1717102 [
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<collectingCountry id="F339B6B44C78C6591B252A8FFA9326CE" box="[1285,1344,1954,1978]" name="Qatar" pageId="14" pageNumber="15">QTR</collectingCountry>
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label]”. This is also the “
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<taxonomicName id="4C2E8DA74C78C6591F482AEBFE1E26A9" box="[360,461,1990,2013]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="18" phylum="Arthropoda" rank="species" species="masoura">
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<emphasis id="B95A2A364C78C6591F482AEBFE1E26A9" box="[360,461,1990,2013]" italics="true" pageId="14" pageNumber="15">Masoura</emphasis>
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</taxonomicName>
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? sp.” illustrated by
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<bibRefCitation id="EFBF8BD54C78C6591CE32AE8FCA726AA" author="d'Abrera" box="[707,884,1989,2014]" pageId="14" pageNumber="15" refString="d'Abrera, B. L. (1997) Butterflies of the Afrotropical Region. Part 1. Papilionidae, Pieridae, Acraeidae, Danaidae, Satyridae. Hill House Publishers, Melbourne & London, 263 pp." type="book" year="1997">d’Abrera (1997</bibRefCitation>
|
||
: 229) in DV/VV, following preference for a previously illustrated specimen (Recommendation 74B of the code), although in this case, not the one that the original author intended. This action is also in the spirit of the Code as it promotes stability and obviates the need to describe a new species within Oberthür’s syntypic series. The majority of the series of
|
||
<bibRefCitation id="EFBF8BD54C79C6581A8A2ABDFAB926DD" author="Oberthur" box="[1194,1386,1936,1961]" pageId="15" pageNumber="16" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür (1916)</bibRefCitation>
|
||
’s
|
||
<taxonomicName id="4C2E8DA74C79C6581BA42ABCFF2A26BA" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="alaokola">
|
||
<emphasis id="B95A2A364C79C6581BA42ABCFF2A26BA" italics="true" pageId="15" pageNumber="16">G. alaokola</emphasis>
|
||
</taxonomicName>
|
||
STs from “Fianarantsoa” belong to
|
||
<taxonomicName id="4C2E8DA74C79C6581CAA2A9BFCC326BA" box="[650,784,1973,1998]" class="Insecta" family="Nymphalidae" genus="Strabena" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="mabillei">
|
||
<emphasis id="B95A2A364C79C6581CAA2A9BFCC326BA" box="[650,784,1973,1998]" italics="true" pageId="15" pageNumber="16">Ht. mabillei</emphasis>
|
||
</taxonomicName>
|
||
, and are then automatically among the PLTs of Oberthür’s nominal species. Although not illustrated by
|
||
<bibRefCitation id="EFBF8BD54C79C6581C872AF5FCB72685" author="Oberthur" box="[679,868,2008,2033]" pageId="15" pageNumber="16" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür (1916)</bibRefCitation>
|
||
, the chosen LT represents the more widespread/ common and generally darker-upperside species (formerly known as sp. 56;
|
||
<bibRefCitation id="EFBF8BD54C66C6471DD82DB5FBA221C4" author="Lees" box="[1016,1137,152,176]" pageId="16" pageNumber="17" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees 1997</bibRefCitation>
|
||
: 65), which tends strongly either to lack space-M3 ocellus expression, or to lack expression in both space-M3 and CuA1 on HWD. This species also lacks a ventral abdominal black androconial patch (usually bilateral of midline, sometimes obscured by overlying ochreous scales) that is characteristic of
|
||
<taxonomicName id="4C2E8DA74C66C6471CDD2C28FC562069" box="[765,901,260,285]" class="Insecta" family="Nymphalidae" genus="Strabena" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mabillei">
|
||
<emphasis id="B95A2A364C66C6471CDD2C28FC562069" box="[765,901,260,285]" italics="true" pageId="16" pageNumber="17">Ht. mabillei</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFBF8BD54C66C6471DE32C28FB902068" author="Lees" box="[963,1091,260,284]" pageId="16" pageNumber="17" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997</bibRefCitation>
|
||
: 99) and also, more markedly,
|
||
<taxonomicName id="4C2E8DA74C66C6471EB72C04FEF42034" box="[151,295,297,320]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="masoura">
|
||
<emphasis id="B95A2A364C66C6471EB72C04FEF42034" box="[151,295,297,320]" italics="true" pageId="16" pageNumber="17">Ht. masoura</emphasis>
|
||
</taxonomicName>
|
||
. It (“sp. 56”;
|
||
<bibRefCitation id="EFBF8BD54C66C6471FF22C05FD862034" author="Lees" box="[466,597,296,320]" pageId="16" pageNumber="17" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997</bibRefCitation>
|
||
: 105) also has multiply toothed long valves, which do not have such a projected sharply inbent tip, as does the genitalia of Oberthür’s ♂ ‘iconotype’ of
|
||
<taxonomicName id="4C2E8DA74C66C6471A0C2C60FB7D2011" box="[1068,1198,332,357]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alaokola">
|
||
<emphasis id="B95A2A364C66C6471A0C2C60FB7D2011" box="[1068,1198,332,357]" italics="true" pageId="16" pageNumber="17">G. alaokola</emphasis>
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName id="4C2E8DA74C66C6471AEE2C60FA862011" box="[1230,1365,332,357]" class="Insecta" family="Nymphalidae" genus="Strabena" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mabillei">
|
||
<emphasis id="B95A2A364C66C6471AEE2C60FA862011" box="[1230,1365,332,357]" italics="true" pageId="16" pageNumber="17">Ht. mabillei</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<figureCitation id="1315EAA14C66C6471B4E2C61FF7620FC" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="16" pageNumber="17">Fig. 3</figureCitation>
|
||
D). This feature is not in fact evident in the HT specimen of
|
||
<taxonomicName id="4C2E8DA74C66C6471D7F2C42FBAB20FC" authority="Butler" authorityName="Butler" box="[863,1144,367,392]" class="Insecta" family="Nymphalidae" genus="Strabena" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mabillei">
|
||
<emphasis id="B95A2A364C66C6471D7F2C42FBFB20FC" box="[863,1064,367,392]" italics="true" pageId="16" pageNumber="17">Strabena mabillei</emphasis>
|
||
Butler
|
||
</taxonomicName>
|
||
, although the abdomen is worn and may even not be the original., The strong expression of the space-M3 ocellus in ♂♂ (in contrast to its strong reduction or absence) confirms the identity of specimens currently referred to
|
||
<taxonomicName id="4C2E8DA74C66C6471ABD2C94FAF920A4" box="[1181,1322,439,464]" class="Insecta" family="Nymphalidae" genus="Strabena" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mabillei">
|
||
<emphasis id="B95A2A364C66C6471ABD2C94FAF920A4" box="[1181,1322,439,464]" italics="true" pageId="16" pageNumber="17">Ht. mabillei</emphasis>
|
||
</taxonomicName>
|
||
(in fresh specimens,
|
||
<taxonomicName id="4C2E8DA74C66C6471F3E2CF0FE7C2081" box="[286,431,476,501]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alaokola">
|
||
<emphasis id="B95A2A364C66C6471F3E2CF0FE7C2081" box="[286,431,476,501]" italics="true" pageId="16" pageNumber="17">Ht. alaokola</emphasis>
|
||
</taxonomicName>
|
||
, as here fixed, also has a darker dorsal colouration). Note the androconial scales on abdominal sternum third segment and the inwards recurved valve tips characteristic of
|
||
<taxonomicName id="4C2E8DA74C66C6471A532F2CFB29236C" box="[1139,1274,511,536]" class="Insecta" family="Nymphalidae" genus="Strabena" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mabillei">
|
||
<emphasis id="B95A2A364C66C6471A532F2CFB29236C" box="[1139,1274,511,536]" italics="true" pageId="16" pageNumber="17">Ht. mabillei</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1315EAA14C66C6471B2A2CD2FA9E236C" box="[1290,1357,511,536]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="16" pageNumber="17">Fig. 3</figureCitation>
|
||
D) and also (
|
||
<figureCitation id="1315EAA14C66C6471EF42F09FEC62348" box="[212,277,548,573]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="16" pageNumber="17">Fig. 3</figureCitation>
|
||
A,C) that
|
||
<taxonomicName id="4C2E8DA74C66C6471FA62F08FDDF2349" box="[390,524,548,573]" class="Insecta" family="Nymphalidae" genus="Strabena" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mabillei">
|
||
<emphasis id="B95A2A364C66C6471FA62F08FDDF2349" box="[390,524,548,573]" italics="true" pageId="16" pageNumber="17">Ht. mabillei</emphasis>
|
||
</taxonomicName>
|
||
has relatively long tails especially at M3, a lighter brown dorsal colouration, and three or four white stripes on the FWV tending to be more strongly represented as marks on the FWD costa, than the species/specimen here selected to represent
|
||
<taxonomicName id="4C2E8DA74C66C6471C952F40FC9023F1" box="[693,835,620,645]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alaokola">
|
||
<emphasis id="B95A2A364C66C6471C952F40FC9023F1" box="[693,835,620,645]" italics="true" pageId="16" pageNumber="17">Ht. alaokola</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1315EAA14C66C6471D742F41FC4423F0" box="[852,919,620,645]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="16" pageNumber="17">Fig. 3</figureCitation>
|
||
B). It may be noted that samples respectively representing
|
||
<taxonomicName id="4C2E8DA74C66C6471F092FBCFE6323DC" box="[297,432,655,680]" class="Insecta" family="Nymphalidae" genus="Strabena" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mabillei">
|
||
<emphasis id="B95A2A364C66C6471F092FBCFE6323DC" box="[297,432,655,680]" italics="true" pageId="16" pageNumber="17">Ht. mabillei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2E8DA74C66C6471FC72FBCFDA023DC" box="[487,627,655,680]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alaokola">
|
||
<emphasis id="B95A2A364C66C6471FC72FBCFDA023DC" box="[487,627,655,680]" italics="true" pageId="16" pageNumber="17">Ht. alaokola</emphasis>
|
||
</taxonomicName>
|
||
(KAP524 [KA-P524] = DL-4-233, Anjozorobe; KAP537 [KA-P537] = DL-02-20, Makira, in the study of Aduse-Poku
|
||
<emphasis id="B95A2A364C66C6471C8F2F98FD3B23B9" box="[687,744,692,717]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
|
||
, 2015) are shallowly diverged considering the morphological (mainly androconial and genitalic) differences between these species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |