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<document id="4468AD885170DB362DD62D2EF85BD2EA" ID-CLB-Dataset="38930" ID-DOI="10.11646/zootaxa.4118.1.1" ID-GBIF-Dataset="a4e408c4-2575-4b5c-878b-e1b9010ec619" ID-ISSN="1175-5326" ID-Zenodo-Dep="264597" ID-ZooBank="CFA586DA-10EE-468B-80EE-35351E3845FD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="guilherme" checkinTime="1465242077321" checkinUser="plazi" docAuthor="C, Lees David" docDate="2016" docId="038747324C5EC67C1EB7292BFD3E27E9" docLanguage="en" docName="zootaxa.4118.pdf.imf" docOrigin="Zootaxa 4118 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Heteropsis imerina Lees, sp. nov." docType="treatment" docVersion="12" lastPageNumber="43" masterDocId="FFBE3F4A4C76C6571E202D2DFFD32174" masterDocTitle="Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision" masterLastPageNumber="97" masterPageNumber="1" pageNumber="41" updateTime="1735917000942" updateUser="ExternalLinkService">
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<mods:title id="828D5633C66D766B6F15BE4E8B4C8CA4">Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision</mods:title>
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<mods:name id="24EEF955E66173C65F654025C20E8D8E" type="personal">
<mods:role id="AB2EF08C64ABC2E0AF8C3BF0CA91F5DF">
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<mods:namePart id="9B47101F227068B078C15FB4E4D1BD42">C, Lees David</mods:namePart>
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<mods:typeOfResource id="2AB11AB45153BB2A124A518AFD0EAB61">text</mods:typeOfResource>
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<mods:title id="5AFF961F4D841D71A0FB05445EC659DE">Zootaxa</mods:title>
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<mods:part id="037BE1F18AA15FEB6AD727A84E4E9E27">
<mods:date id="BEBAAC4B16E62831FA449A8FBE8D4A81">2016</mods:date>
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<mods:number id="D3F2296A23854D0F9032A25D19E6A976">4118</mods:number>
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<mods:classification id="19D69E10FF3FF17C09D7F72A051C7447">journal article</mods:classification>
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<mods:identifier id="35B9D8CC0CB52CA3F9DC00645F16A18F" type="DOI">10.11646/zootaxa.4118.1.1</mods:identifier>
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<mods:identifier id="CE3F965948DF7A015DE25555BC764B4A" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
<treatment id="038747324C5EC67C1EB7292BFD3E27E9" ID-DOI="http://doi.org/10.5281/zenodo.6086428" ID-GBIF-Taxon="120817089" ID-Zenodo-Dep="6086428" LSID="urn:lsid:plazi:treatment:038747324C5EC67C1EB7292BFD3E27E9" httpUri="http://treatment.plazi.org/id/038747324C5EC67C1EB7292BFD3E27E9" lastPageId="43" lastPageNumber="43" pageId="40" pageNumber="41" scope_family="Nymphalidae" scope_subFamily="Satyrinae" scope_tribe="Satyrini">
<subSubSection id="C334A5AF4C5EC67F1EB7292BFBCA2535" pageId="40" pageNumber="41" type="nomenclature">
<paragraph id="8B91F6244C5EC67F1EB7292BFDF92554" blockId="40.[151,1049,1030,1089]" box="[151,554,1030,1056]" pageId="40" pageNumber="41">
<heading id="D0D941484C5EC67F1EB7292BFDF92554" bold="true" box="[151,554,1030,1056]" fontSize="11" level="1" pageId="40" pageNumber="41" reason="1">
<emphasis id="B95A2A364C5EC67F1EB7292BFDF92554" bold="true" box="[151,554,1030,1056]" pageId="40" pageNumber="41">
<taxonomicName id="4C2E8DA74C5EC67F1EB7292BFE6C2554" authority="Lees" authorityName="Lees" box="[151,447,1030,1056]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="imerina" status="sp. nov.">
<emphasis id="B95A2A364C5EC67F1EB7292BFEAE2554" bold="true" box="[151,381,1030,1056]" italics="true" pageId="40" pageNumber="41">Heteropsis imerina</emphasis>
Lees
</taxonomicName>
,
<taxonomicNameLabel id="A269974D4C5EC67F1FEC292BFDF92554" box="[460,554,1030,1056]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B91F6244C5EC67F1EB72904FBCA2535" blockId="40.[151,1049,1030,1089]" box="[151,1049,1064,1089]" pageId="40" pageNumber="41">LSID: urn:lsid:zoobank.org:act:988AC583-72F8-4CA4-824A-157D9F7C5AEC</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C5EC67F1EB7295DFDE725D9" pageId="40" pageNumber="41" type="description">
<paragraph id="8B91F6244C5EC67F1EB7295DFC8A25FD" blockId="40.[151,1436,1136,2019]" box="[151,857,1136,1161]" pageId="40" pageNumber="41">
<emphasis id="B95A2A364C5EC67F1EB7295DFE8425FD" bold="true" box="[151,343,1136,1161]" pageId="40" pageNumber="41">Prior references</emphasis>
: sp. 22 (
<bibRefCitation id="EFBF8BD54C5EC67F1F98295CFDE425FD" author="Lees" box="[440,567,1137,1161]" pageId="40" pageNumber="41" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997</bibRefCitation>
;
<bibRefCitation id="EFBF8BD54C5EC67F1C63295CFCC025FD" author="Torres" box="[579,787,1136,1161]" pageId="40" pageNumber="41" refString="Torres, E., Lees, D. C., Vane-Wright, R. I., Kremen, C., Leonard, J. A. &amp; Wayne, R. K. (2001) Examining monophyly in a large radiation of Madagascan butterflies (Lepidoptera: Satyrinae: Mycalesina) based on mitochondrial DNA data. Molecular Phylogenetics and Evolution, 20, 460 - 473. http: // dx. doi. org / 10.1006 / mpev. 2001.0984" type="journal article" year="2001">
Torres
<emphasis id="B95A2A364C5EC67F1CB1295FFD0325FD" box="[657,720,1136,1161]" italics="true" pageId="40" pageNumber="41">et al.,</emphasis>
2001
</bibRefCitation>
: 462);
</paragraph>
<paragraph id="8B91F6244C5EC67F1EE729B8FDE725D9" blockId="40.[151,1436,1136,2019]" box="[199,564,1172,1197]" pageId="40" pageNumber="41">
Sp. 76 (
<bibRefCitation id="EFBF8BD54C5EC67F1F3D29B8FE3C25D9" author="Torres" box="[285,495,1172,1197]" pageId="40" pageNumber="41" refString="Torres, E., Lees, D. C., Vane-Wright, R. I., Kremen, C., Leonard, J. A. &amp; Wayne, R. K. (2001) Examining monophyly in a large radiation of Madagascan butterflies (Lepidoptera: Satyrinae: Mycalesina) based on mitochondrial DNA data. Molecular Phylogenetics and Evolution, 20, 460 - 473. http: // dx. doi. org / 10.1006 / mpev. 2001.0984" type="journal article" year="2001">
Torres
<emphasis id="B95A2A364C5EC67F1F4D29BBFE7F25D9" box="[365,428,1172,1197]" italics="true" pageId="40" pageNumber="41">et al.,</emphasis>
2001
</bibRefCitation>
: 462).
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C5EC67F1EE72995FC8C24F0" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="8B91F6244C5EC67F1EE72995FDE4246C" blockId="40.[151,1436,1136,2019]" pageId="40" pageNumber="41">
<emphasis id="B95A2A364C5EC67F1EE72995FD3B25A5" bold="true" box="[199,744,1208,1233]" pageId="40" pageNumber="41">
<typeStatus id="549548864C5EC67F1EE72995FED125A5" box="[199,258,1208,1233]" pageId="40" pageNumber="41">Type</typeStatus>
material., Deposition
<collectionCode id="ED3F6EE14C5EC67F1C312995FDBF25A5" box="[529,620,1208,1233]" pageId="40" pageNumber="41">BMNH</collectionCode>
:
<typeStatus id="549548864C5EC67F1C5B2995FD3B25A5" box="[635,744,1208,1233]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
</emphasis>
: ♂ (
<figureCitation id="1315EAA14C5EC67F1D002995FCB525A5" box="[800,870,1208,1233]" captionStart="FIGURE 9" captionStartId="32.[151,250,1768,1790]" captionTargetBox="[262,1325,218,1725]" captionTargetId="figure@32.[252,1334,193,1747]" captionTargetPageId="32" captionText="FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264606" httpUri="https://zenodo.org/record/264606/files/figure.png" pageId="40" pageNumber="41">Fig. 9</figureCitation>
C),
<collectingCountry id="F339B6B44C5EC67F1DB22995FBCA25A5" box="[914,1049,1208,1233]" name="Madagascar" pageId="40" pageNumber="41">Madagascar</collectingCountry>
E, Mantadia National Park, Jofa, near bridge,
<geoCoordinate id="EE1A90E34C5EC67F1F1929F1FE612581" box="[313,434,1244,1269]" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="5" value="-18.7674">18.7674°S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5EC67F1FE829F1FDEC2580" box="[456,575,1244,1268]" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="5" value="48.4324">48.4324°E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5EC67F1CA429F1FD0C2581" box="[644,735,1244,1269]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="40" pageNumber="41" unit="km" value="0.4">0.4 km</quantity>
,
<quantity id="4CD65BC14C5EC67F1CD529F1FC782580" box="[757,939,1244,1269]" metricMagnitude="2" metricUnit="m" metricValue="9.45" metricValueMax="9.6" metricValueMin="9.3" pageId="40" pageNumber="41" unit="m" value="945.0" valueMax="960.0" valueMin="930.0">
945 +/-
<specimenCount id="9D283DAD4C5EC67F1D4429F1FC782580" box="[868,939,1244,1269]" pageId="40" pageNumber="41" type="generic">15 m</specimenCount>
</quantity>
.
<date id="FF90D0E44C5EC67F1DE029F1FB972581" box="[960,1092,1244,1269]" pageId="40" pageNumber="41" value="2014-01-16">16/01/2014</date>
. D.C. Lees:
<collectionCode id="ED3F6EE14C5EC67F1ACE29F1FAC42581" box="[1262,1303,1244,1269]" pageId="40" pageNumber="41">DL</collectionCode>
16114-22-1,
<collectionCode id="ED3F6EE14C5EC67F1EB7282DFED2246C" box="[151,257,1280,1304]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="40" pageNumber="41">NHMUK</collectionCode>
010289132 [
<collectingCountry id="F339B6B44C5EC67F1FAC282DFE14246C" box="[396,455,1280,1304]" name="Qatar" pageId="40" pageNumber="41">QTR</collectingCountry>
barcode].
</paragraph>
<paragraph id="8B91F6244C5EC67F1EE7280EFC8C24F0" blockId="40.[151,1436,1136,2019]" pageId="40" pageNumber="41">
<typeStatus id="549548864C5EC67F1EE7280EFEEC2448" box="[199,319,1315,1340]" pageId="40" pageNumber="41" type="paratype">
<emphasis id="B95A2A364C5EC67F1EE7280EFEEC2448" bold="true" box="[199,319,1315,1340]" pageId="40" pageNumber="41">Paratypes</emphasis>
</typeStatus>
:
<emphasis id="B95A2A364C5EC67F1F70280EFDFC2448" bold="true" box="[336,559,1315,1340]" pageId="40" pageNumber="41">
Deposition
<collectionCode id="ED3F6EE14C5EC67F1FF8280EFDFC2448" box="[472,559,1315,1340]" pageId="40" pageNumber="41">BMNH</collectionCode>
</emphasis>
: ♂, data as HT but
<collectionCode id="ED3F6EE14C5EC67F1D3A280EFC912448" box="[794,834,1315,1340]" pageId="40" pageNumber="41">DL</collectionCode>
16114-22-2,
<collectionCode id="ED3F6EE14C5EC67F1DF12809FBE82448" box="[977,1083,1316,1340]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="40" pageNumber="41">NHMUK</collectionCode>
010289133 [
<collectingCountry id="F339B6B44C5EC67F1AE92809FAD72448" box="[1225,1284,1316,1340]" name="Qatar" pageId="40" pageNumber="41">QTR</collectingCountry>
barcode]; ♂,
<collectingCountry id="F339B6B44C5EC67F1EB7286AFECD2414" box="[151,286,1351,1376]" name="Madagascar" pageId="40" pageNumber="41">Madagascar</collectingCountry>
E., Mantadia National Park, Mitsinjo site PK 17.5 from Vakona Forest Lodge,
<quantity id="4CD65BC14C5EC67F1A92286AFAD9242B" box="[1202,1290,1351,1375]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="40" pageNumber="41" unit="m" value="1100.0">1100 m</quantity>
,
<date id="FF90D0E44C5EC67F1B38286AFA4B242B" box="[1304,1432,1351,1375]" pageId="40" pageNumber="41" value="2011-12-11">11/12/2011</date>
, D.C. Lees:
<collectionCode id="ED3F6EE14C5EC67F1F362846FEED24F7" box="[278,318,1387,1411]" pageId="40" pageNumber="41">DL</collectionCode>
11M-0001,
<collectionCode id="ED3F6EE14C5EC67F1F9F2846FDFA24F7" box="[447,553,1387,1411]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="40" pageNumber="41">NHMUK</collectionCode>
010289134 [
<collectingCountry id="F339B6B44C5EC67F1C942846FD3C24F7" box="[692,751,1387,1411]" name="Qatar" pageId="40" pageNumber="41">QTR</collectingCountry>
barcode];
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C5EC67F1EE728A3FC2E2766" pageId="40" pageNumber="41" type="description">
<paragraph id="8B91F6244C5EC67F1EE728A3FD83249B" blockId="40.[151,1436,1136,2019]" pageId="40" pageNumber="41">
<emphasis id="B95A2A364C5EC67F1EE728A3FE6124D3" bold="true" box="[199,434,1422,1447]" pageId="40" pageNumber="41">Deposition MNHN:</emphasis>
♂,
<collectingCountry id="F339B6B44C5EC67F1FC128A3FDBB24D3" box="[481,616,1422,1447]" name="Madagascar" pageId="40" pageNumber="41">Madagascar</collectingCountry>
E. route dAnosibe
<date id="FF90D0E44C5EC67F1D7428A3FC6324D3" box="[852,944,1422,1447]" pageId="40" pageNumber="41" value="1966-06">VI-1966</date>
/MUSEUM PARIS COLL. G. DUJARDIN DEL. ET R. VIOSSAT|DCL-DB-2908; ♀ (
<figureCitation id="1315EAA14C5EC67F1CA3289FFD1B24BF" box="[643,712,1458,1483]" captionStart="FIGURE 9" captionStartId="32.[151,250,1768,1790]" captionTargetBox="[262,1325,218,1725]" captionTargetId="figure@32.[252,1334,193,1747]" captionTargetPageId="32" captionText="FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264606" httpUri="https://zenodo.org/record/264606/files/figure.png" pageId="40" pageNumber="41">Fig. 9</figureCitation>
D),
<collectingCountry id="F339B6B44C5EC67F1CD5289FFCAF24BF" box="[757,892,1458,1483]" name="Madagascar" pageId="40" pageNumber="41">Madagascar</collectingCountry>
Est, route de Lakato km 15, Ankasoka
<quantity id="4CD65BC14C5EC67F1B67289EFA4F24BE" box="[1351,1436,1459,1483]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="40" pageNumber="41" unit="m" value="1100.0">1100 m</quantity>
17/
<date id="FF90D0E44C5EC67F1E9A28FBFEE6249B" box="[186,309,1494,1519]" pageId="40" pageNumber="41" value="1963-10-21">21-X-1963</date>
P. Viette|DCL-DB-2906.
</paragraph>
<paragraph id="8B91F6244C5EC67F1EE728D7FC2E2766" blockId="40.[151,1436,1136,2019]" box="[199,1021,1530,1555]" pageId="40" pageNumber="41">
<emphasis id="B95A2A364C5EC67F1EE728D7FE6D2767" bold="true" box="[199,446,1530,1555]" pageId="40" pageNumber="41">Deposition summary</emphasis>
: BMNH (HT ♂, 2 PT ♂♂), MNHN (PT ♂, PT ♀).
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C5EC67F1EE72B30FEE3272D" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="8B91F6244C5EC67F1EE72B30FEE3272D" blockId="40.[151,1436,1136,2019]" pageId="40" pageNumber="41">
<emphasis id="B95A2A364C5EC67F1EE72B30FEB42742" bold="true" box="[199,359,1565,1590]" pageId="40" pageNumber="41">
<typeStatus id="549548864C5EC67F1EE72B30FED12742" box="[199,258,1565,1590]" pageId="40" pageNumber="41">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F339B6B44C5EC67F1F502B30FE242742" box="[368,503,1565,1590]" name="Madagascar" pageId="40" pageNumber="41">Madagascar</collectingCountry>
E., Mantadia National Park, Jofa, near bridge,
<geoCoordinate id="EE1A90E34C5EC67F1A362B30FB5D2742" box="[1046,1166,1565,1590]" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="5" value="-18.7674">18.7674°S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5EC67F1ABB2B33FAC02742" box="[1179,1299,1566,1590]" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="5" value="48.4324">48.4324°E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5EC67F1B652B33FA4B2742" box="[1349,1432,1565,1590]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="40" pageNumber="41" unit="km" value="0.4">0.4 km</quantity>
,
<quantity id="4CD65BC14C5EC67F1EB72B6CFEFF272D" box="[151,300,1601,1626]" metricMagnitude="2" metricUnit="m" metricValue="9.45" metricValueMax="9.6" metricValueMin="9.3" pageId="40" pageNumber="41" unit="m" value="945.0" valueMax="960.0" valueMin="930.0">
945 +/-
<specimenCount id="9D283DAD4C5EC67F1ED02B6CFEFF272D" box="[240,300,1601,1626]" pageId="40" pageNumber="41" type="generic">15 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C5EC67F1EE72B48FB94260C" pageId="40" pageNumber="41" type="diagnosis">
<paragraph id="8B91F6244C5EC67F1EE72B48FB94260C" blockId="40.[151,1436,1136,2019]" pageId="40" pageNumber="41">
<emphasis id="B95A2A364C5EC67F1EE72B48FE92270A" bold="true" box="[199,321,1637,1662]" pageId="40" pageNumber="41">Diagnosis.</emphasis>
Very distinct by its shorter HW tails from the somewhat similar
<taxonomicName id="4C2E8DA74C5EC67F1A6F2B4BFB09270A" box="[1103,1242,1637,1662]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="pauliani" status="sp. nov.">
<emphasis id="B95A2A364C5EC67F1A6F2B4BFB09270A" box="[1103,1242,1637,1662]" italics="true" pageId="40" pageNumber="41">Ht. pauliani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269974D4C5EC67F1AC62B48FA96270A" box="[1254,1349,1637,1662]" pageId="40" pageNumber="41" rank="species">
<emphasis id="B95A2A364C5EC67F1AC62B48FA96270A" bold="true" box="[1254,1349,1637,1662]" pageId="40" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
, where potentially (see discussion) sympatric (from RS Anjanaharibe Sud to RNI Tsaratanana).
<taxonomicName id="4C2E8DA74C5EC67F1AA22BA7FA8427D5" box="[1154,1367,1672,1697]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="harveyi">
<emphasis id="B95A2A364C5EC67F1AA22BA7FA8427D5" box="[1154,1367,1672,1697]" italics="true" pageId="40" pageNumber="41">Heteropsis harveyi</emphasis>
</taxonomicName>
is the species most likely to be confused in the main central-eastern rainforest block, from which
<taxonomicName id="4C2E8DA74C5EC67F1A852B83FAF427B0" box="[1189,1319,1709,1733]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="imerina">
<emphasis id="B95A2A364C5EC67F1A852B83FAF427B0" box="[1189,1319,1709,1733]" italics="true" pageId="40" pageNumber="41">Ht. imerina</emphasis>
</taxonomicName>
differs by lack of an androconial patch on the FWV, by the proximad inflexion of the HWV medial band around ocellus space-CuA1 being rather sharply angled, and by the more evenly pronounced HW tails of the former, as for the other small dark species (in the case of ♂♂).
<taxonomicName id="4C2E8DA74C5EC67F1C8B2A34FC8D2644" box="[683,862,1815,1840]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="vanewrighti" status="sp. nov.">
<emphasis id="B95A2A364C5EC67F1C8B2A34FC8D2644" box="[683,862,1815,1840]" italics="true" pageId="40" pageNumber="41">Ht. vanewrighti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269974D4C5EC67F1D472A3AFC172644" box="[871,964,1815,1840]" pageId="40" pageNumber="41" rank="species">
<emphasis id="B95A2A364C5EC67F1D472A3AFC172644" bold="true" box="[871,964,1815,1840]" pageId="40" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="4C2E8DA74C5EC67F1DF42A34FB962644" box="[980,1093,1817,1840]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="viettei">
<emphasis id="B95A2A364C5EC67F1DF42A34FB962644" box="[980,1093,1817,1840]" italics="true" pageId="40" pageNumber="41">Ht. viettei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2E8DA74C5EC67F1AA22A34FAF72644" box="[1154,1316,1815,1840]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="westwoodi">
<emphasis id="B95A2A364C5EC67F1AA22A34FAF72644" box="[1154,1316,1815,1840]" italics="true" pageId="40" pageNumber="41">Ht. westwoodi</emphasis>
</taxonomicName>
are easily distinguished from
<taxonomicName id="4C2E8DA74C5EC67F1F532A10FE252620" box="[371,502,1853,1876]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="imerina">
<emphasis id="B95A2A364C5EC67F1F532A10FE252620" box="[371,502,1853,1876]" italics="true" pageId="40" pageNumber="41">Ht. imerina</emphasis>
</taxonomicName>
by their whitish highlighting distad of the HWV Mb, which in
<taxonomicName id="4C2E8DA74C5EC67F1AEC2A10FA9D2620" box="[1228,1358,1853,1876]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="imerina">
<emphasis id="B95A2A364C5EC67F1AEC2A10FA9D2620" box="[1228,1358,1853,1876]" italics="true" pageId="40" pageNumber="41">Ht. imerina</emphasis>
</taxonomicName>
♂♂ is yellowish, sometimes with a reddish suffusion proximad of the space-CuA1 ocellus.
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C5EC67E1EE72AAEFD14255A" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="description">
<paragraph id="8B91F6244C5EC67E1EE72AAEFD3E2223" blockId="40.[151,1436,1136,2019]" lastBlockId="41.[151,1437,151,2037]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
<emphasis id="B95A2A364C5EC67F1EE72AAEFE7E26E8" bold="true" box="[199,429,1923,1948]" pageId="40" pageNumber="41">Description. Wings</emphasis>
: upperside uniform mid to dark brown (costad area not much darker) with space-CuA1 ocellus well expressed on FWD, with a round concentric orange-reddish ring, its black iris almost spanning or spanning CuA1-CuA2, and the Orng compromising these veins and moderately narrow and eccentric. FWD space- M1 ocellus not evident. HWD space-CuA1 ocellus also hardly evident. FWV space-CuA1 ocellus similar size to FWD and marginally more eccentric with mid orange ring slightly skewed to tergum. Space-M1 ocellus much smaller and with a small black iris and a faint mid orange ring; that in space-M2 almost reduced to a small white spot. HWV space-CuA1 ocellus relatively small compared with others of
<taxonomicName id="4C2E8DA74C5FC67E1DF42C29FB882068" box="[980,1115,259,284]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="drepana">
<emphasis id="B95A2A364C5FC67E1DF42C29FB882068" box="[980,1115,259,284]" italics="true" pageId="41" pageNumber="42">Ht. drepana</emphasis>
</taxonomicName>
group and subelliptic with a narrow mid orange ring. Other HWV ocelli, particularly those of spaces Rs-M3, also of space-CuA2-1A, expressed as white spots. Ventral wing background colour with a distinct dark reddish brown cast. FWV more uniform than HWV but paler towards tergal angle with a pale ochreous mark near costa towards apex, HWV lighter brown distad of Mb. This dark russet brown Mb is irregularly snaking, concave around its intersection with M1, convex around that with M2, curving back to the deepest concavity in space-CuA1 and convex again near vein CuA2 before terminating at 1A. This line is highlighted with ochreous admixture with reddish brown scales distad of it and sombrely shaded proximad of it, with a more consistent irroration with russet brown contrasting with more ochreous brown scales in the basal areas; the basal HWV areas are thus relatively dark in the ♂, while the PMb is indistinct and basad of it is also ochreous highlighted. Before the margin in both wings is a slightly more ochreous highlighting and a double thin russet-brown Pml (pre-marginal line), the outer of which follows the margin, which is lightly crenate. The slightly more prominent but short dark brown tail is at the end of M3, which can be a field character distinguishing from similar members of the
<taxonomicName id="4C2E8DA74C5FC67E1CCD2F9CFCA023BC" box="[749,883,687,712]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="drepana">
<emphasis id="B95A2A364C5FC67E1CCD2F9CFCA023BC" box="[749,883,687,712]" italics="true" pageId="41" pageNumber="42">Ht. drepana</emphasis>
</taxonomicName>
group, particularly in the ♀ (
<figureCitation id="1315EAA14C5FC67E1A972F82FB2923BC" box="[1207,1274,687,712]" captionStart="FIGURE 9" captionStartId="32.[151,250,1768,1790]" captionTargetBox="[262,1325,218,1725]" captionTargetId="figure@32.[252,1334,193,1747]" captionTargetPageId="32" captionText="FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264606" httpUri="https://zenodo.org/record/264606/files/figure.png" pageId="41" pageNumber="42">Fig. 9</figureCitation>
D).
<emphasis id="B95A2A364C5FC67E1B042F82FA4F23BC" bold="true" box="[1316,1436,687,712]" pageId="41" pageNumber="42">Variation.</emphasis>
♂♂ vary quite a lot in their underside colouration, and the HWD space-CuA1 ocellus is sometimes expressed. White spots not always evident on HWV. ♀ is similar but larger, lighter brown on upperside and with more ochreous cast on upperside with PMb and Mb russet brown on HWV, thus more pronounced. In apparent dry seasonal forms, the HWV ocelli can be very reduced.
</paragraph>
<paragraph id="8B91F6244C5FC67E1EE72E4FFD7722EB" blockId="41.[151,1437,151,2037]" pageId="41" pageNumber="42">
<emphasis id="B95A2A364C5FC67E1EE72E4FFE8E220F" bold="true" box="[199,349,866,891]" pageId="41" pageNumber="42">Androconia:</emphasis>
discocellular brush HWD mid brown, to ochreous brown towards tip. Underlying patch lenticular, pale ochreous. No other androconia.
</paragraph>
<paragraph id="8B91F6244C5FC67E1EE72E87FED8257E" blockId="41.[151,1437,151,2037]" pageId="41" pageNumber="42">
<emphasis id="B95A2A364C5FC67E1EE72E87FEA522B7" bold="true" box="[199,374,938,963]" pageId="41" pageNumber="42">Wingspan/fwl:</emphasis>
35.2 (n=1)/
<quantity id="4CD65BC14C5FC67E1FD92E87FD7222B6" box="[505,673,938,962]" metricMagnitude="-2" metricUnit="m" metricValue="1.8850000000000002" metricValueMax="1.9300000000000002" metricValueMin="1.8399999999999999" pageId="41" pageNumber="42" unit="mm" value="18.85" valueMax="19.3" valueMin="18.4">18.419.3 mm</quantity>
(n=20); 35.438.1/19.722.0 mm (n=4); mean = 33.8 +/- 1.44 SD/ 17.87+/- 0.54 SD (n=6 ♂♂, including HT, 35.4/
<quantity id="4CD65BC14C5FC67E1CE12EE0FCF92292" box="[705,810,973,998]" metricMagnitude="-2" metricUnit="m" metricValue="1.8399999999999999" pageId="41" pageNumber="42" unit="mm" value="18.4">18.4 mm</quantity>
); mean = 36.2 +/- 3.03 SD (n=4) /18.75 +/- 1 SD mm (n=4) (♀).
</paragraph>
<paragraph id="8B91F6244C5FC67E1EE72938FD14255A" blockId="41.[151,1437,151,2037]" box="[199,711,1045,1070]" pageId="41" pageNumber="42">
<emphasis id="B95A2A364C5FC67E1EE72938FEC2255A" bold="true" box="[199,273,1045,1070]" pageId="41" pageNumber="42">Palps:</emphasis>
mid brown without clear pale striping.
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C5FC67E1EE72914FF0825EC" pageId="41" pageNumber="42" type="etymology">
<paragraph id="8B91F6244C5FC67E1EE72914FF0825EC" blockId="41.[151,1437,151,2037]" pageId="41" pageNumber="42">
<emphasis id="B95A2A364C5FC67E1EE72914FE9D2526" bold="true" box="[199,334,1081,1106]" pageId="41" pageNumber="42">Etymology.</emphasis>
Refers to its potentially wide distribution across the Madagascan
<emphasis id="B95A2A364C5FC67E1A742914FB232526" box="[1108,1264,1081,1106]" italics="true" pageId="41" pageNumber="42">haut-plateaux</emphasis>
also including surrounding mountain ranges, as for the principal Merian tribe of the
<collectingCountry id="F339B6B44C5FC67E1D802971FBDE2501" box="[928,1037,1116,1141]" name="Madagascar" pageId="41" pageNumber="42">Malagasy</collectingCountry>
people, which gives this species its name.
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C5FC67E1EE72989FBC226F3" pageId="41" pageNumber="42" type="discussion">
<paragraph id="8B91F6244C5FC67E1EE72989FA8E248A" blockId="41.[151,1437,151,2037]" pageId="41" pageNumber="42">
<emphasis id="B95A2A364C5FC67E1EE72989FE9F25C9" bold="true" box="[199,332,1188,1213]" pageId="41" pageNumber="42">Discussion.</emphasis>
There is a certain amount of historical material, but perhaps surprisingly the species is not implicated in any pre-existing descriptions. This species was first recognized in the field as new as morphospecies 22 at Ankazomivady forest on
<date id="FF90D0E44C5FC67E1FD229C1FDA12470" box="[498,626,1260,1284]" pageId="41" pageNumber="42" value="1992-01-12">12/01/1992</date>
by Claire Kremen (
<bibRefCitation id="EFBF8BD54C5FC67E1D7229C1FC292470" author="Lees" box="[850,1018,1259,1284]" pageId="41" pageNumber="42" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997: 64</bibRefCitation>
) and as sp. 76 by the same worker in
<date id="FF90D0E44C5FC67E1EB72822FEC6245C" box="[151,277,1295,1320]" pageId="41" pageNumber="42" value="1995" valueMax="1996">19951996</date>
at Anjanaharibe Sud, but there were earlier collections, mostly represented in MNHN. The morphospecies sp. 76 (Anjanaharibe Sud) is definitely referable to
<taxonomicName id="4C2E8DA74C5FC67E1DBA2819FBC8243F" box="[922,1051,1332,1355]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="imerina">
<emphasis id="B95A2A364C5FC67E1DBA2819FBC8243F" box="[922,1051,1332,1355]" italics="true" pageId="41" pageNumber="42">Ht. imerina</emphasis>
</taxonomicName>
, although its ♂ genitalia had been considered distinct in
<bibRefCitation id="EFBF8BD54C5FC67E1FAC2875FDEC241B" author="Lees" box="[396,575,1366,1391]" pageId="41" pageNumber="42" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees (1997: 65)</bibRefCitation>
; spp. 22 and 76 were recovered anyway as sister taxa in
<bibRefCitation id="EFBF8BD54C5FC67E1AF72875FA4B241B" author="Lees" box="[1239,1432,1366,1391]" pageId="41" pageNumber="42" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees (1997: 156)</bibRefCitation>
, but the recent discovery by myself of additional populations with adult phenotype similar to
<taxonomicName id="4C2E8DA74C5FC67E1AF62851FA8824E7" box="[1238,1371,1404,1427]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="imerina">
<emphasis id="B95A2A364C5FC67E1AF62851FA8824E7" box="[1238,1371,1404,1427]" italics="true" pageId="41" pageNumber="42">Ht. imerina</emphasis>
</taxonomicName>
from Manongarivo through Tsaratanana to Marojejy and Anjanaharibe Sud suggest the species is indeed quite widespread. A deeper study of variation in the genitalic variation would be interesting, especially if backed up by molecular sequences; the study of
<bibRefCitation id="EFBF8BD54C5FC67E1C3928CAFD2B248A" box="[537,760,1509,1534]" pageId="41" pageNumber="42" refString="Torres, E., Lees, D. C., Vane-Wright, R. I., Kremen, C., Leonard, J. A. &amp; Wayne, R. K. (2001) Examining monophyly in a large radiation of Madagascan butterflies (Lepidoptera: Satyrinae: Mycalesina) based on mitochondrial DNA data. Molecular Phylogenetics and Evolution, 20, 460 - 473. http: // dx. doi. org / 10.1006 / mpev. 2001.0984" type="journal article">
Torres
<emphasis id="B95A2A364C5FC67E1C4728CAFD75248A" box="[615,678,1509,1534]" italics="true" pageId="41" pageNumber="42">et al.,</emphasis>
(2001)
</bibRefCitation>
also showed minimal differences in COII (see below).
</paragraph>
<paragraph id="8B91F6244C5FC67E1EE72B24FBC226F3" blockId="41.[151,1437,151,2037]" pageId="41" pageNumber="42">
<emphasis id="B95A2A364C5FC67E1EE72B24FDA92756" bold="true" box="[199,634,1545,1570]" pageId="41" pageNumber="42">Additional information. ♂ genitalia</emphasis>
: 154DL (not in
<typeStatus id="549548864C5FC67E1D612B27FCA22756" box="[833,881,1546,1570]" pageId="41" pageNumber="42">type</typeStatus>
series,
<figureCitation id="1315EAA14C5FC67E1DEF2B24FBC72756" box="[975,1044,1545,1570]" captionStart="FIGURE 8" captionStartId="26.[151,250,1929,1951]" captionTargetBox="[192,1379,232,1885]" captionTargetId="figure@26.[161,1426,193,1908]" captionTargetPageId="26" captionText="FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264605" httpUri="https://zenodo.org/record/264605/files/figure.png" pageId="41" pageNumber="42">Fig. 8</figureCitation>
C): from LV, tegumen with fairly straight dorsal profile (strong v-shaped notch proximad from DV), leading to narrowly scythe-hooked uncus without a strong ventral dewlap. Gnathos relatively thin and sinuate, tapering, first downcurved and then upcurved (also sinuate from DV). Constriction in division with vinculum about half maximum dimension of tegumen. Vinculum not strongly bowed and broadening towards base of saccus. Valves relatively long, with distinctly angled shoulder, and stretched out valve base narrowing gradually to valve arm which is slightly longer than valve base with distinct ventral elbow, leading to spatulate tips with a limited number of spinoid setae on inner face/at tip (valve arms gently incurved from DV without a strong terminal spine but with a small number of teeth on mesad edge towards tip). Valve tips protrude considerably proud of maximum extension of uncus tip (
<bibRefCitation id="EFBF8BD54C5FC67E1B462A05FEA12610" author="Lees" pageId="41" pageNumber="42" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997" yearSuffix="i">
Lees 1997: 109,
<figureCitation id="1315EAA14C5FC67E1F012A66FEB92610" box="[289,362,1867,1892]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees &amp; Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced tail on HW vein M 3." figureDoi="http://doi.org/10.5281/zenodo.264604" httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="41" pageNumber="42">
Fig.
<date id="FF90D0E44C5FC67E1F7A2A66FEB92610" box="[346,362,1867,1892]" pageId="41" pageNumber="42">7</date>
</figureCitation>
i
</bibRefCitation>
). Aedeagus very thin and gently recurved distad of ostium, as long as valve. Saccus not particularly long and bulbous. Juxta prominent and narrowly cupped proximad.
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C5FC67D1EE72ABFFD6022B7" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="description">
<paragraph id="8B91F6244C5FC67D1EE72ABFFC5420DE" blockId="41.[151,1437,151,2037]" lastBlockId="42.[151,1437,151,2030]" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
<emphasis id="B95A2A364C5FC67E1EE72ABFFE4B26DF" bold="true" box="[199,408,1938,1963]" pageId="41" pageNumber="42">DNA divergences</emphasis>
: the COI-5P cluster number (BOLD:AAE4110) includes the same specimen as sequenced for cytochrome b as “sp. 22” in the Monaghan
<emphasis id="B95A2A364C5FC67E1C822A9AFD3226BB" box="[674,737,1974,1999]" italics="true" pageId="41" pageNumber="42">et al.,</emphasis>
(2001) study (DL-4-515, BMNH(E) #676683, BMAD046-09, Analamay [
<geoCoordinate id="EE1A90E34C5FC67E1F002AF0FE5A2681" box="[288,393,2009,2037]" direction="south" orientation="latitude" pageId="41" pageNumber="42" precision="55" value="-18.828">18.828o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5FC67E1FBE2AF0FDD52681" box="[414,518,2009,2037]" direction="east" orientation="longitude" pageId="41" pageNumber="42" precision="55" value="48.339">48.339o E</geoCoordinate>
,
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]), as well as exemplars BMAD231-15 from RNI Tsaratanana and BMAD246-15 from RS Anjanaharibe Sud. This last exemplar sp. 76 has an identical sequence to that from Analamay. The 357 bp cytochrome b sequence of the last specimen (BMNH(E) #676683
<accessionNumber id="947D6BC74C5CC67D1A852D96FAC421A0" box="[1189,1303,187,212]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819266" pageId="42" pageNumber="43">FJ819266</accessionNumber>
, BMNH(E) #676685
<accessionNumber id="947D6BC74C5CC67D1F212DF2FEA2218C" box="[257,369,223,248]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819267" pageId="42" pageNumber="43">FJ819267</accessionNumber>
) is about 4.2% pairwise divergent to those of
<taxonomicName id="4C2E8DA74C5CC67D1D5D2DCDFC2F218C" box="[893,1020,223,248]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="harveyi">
<emphasis id="B95A2A364C5CC67D1D5D2DCDFC2F218C" box="[893,1020,223,248]" italics="true" pageId="42" pageNumber="43">Ht. harveyi</emphasis>
</taxonomicName>
described here (BMNH(E) #671648
<accessionNumber id="947D6BC74C5CC67D1EB72C2EFEDB2068" box="[151,264,259,284]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819453" pageId="42" pageNumber="43">FJ819453</accessionNumber>
, BMNH(E) #676467
<accessionNumber id="947D6BC74C5CC67D1FDB2C2EFDBD2068" box="[507,622,259,284]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819454" pageId="42" pageNumber="43">FJ819454</accessionNumber>
;
<bibRefCitation id="EFBF8BD54C5CC67D1C582C2EFCAA206F" author="Monaghan" box="[632,889,259,284]" pageId="42" pageNumber="43" refString="Monaghan, M. T., Wild, R., Elliot, E., Fujisawa, T., Balke, M, Inward, D. J. G., Lees, D. C., Ranaivosolo, R., Eggleton, P., Barraclough, T. G. &amp; Vogler, A. P. (2009) Accelerated species inventory on Madagascar using coalescent-based models of species delineation. Systematic Biology, 58, 298 - 311. http: // dx. doi. org / 10.1093 / sysbio / syp 027" type="journal article" year="2009">
Monaghan
<emphasis id="B95A2A364C5CC67D1CD72C29FCFC2068" box="[759,815,259,284]" italics="true" pageId="42" pageNumber="43">et al.</emphasis>
, 2009
</bibRefCitation>
; additional members of the
<taxonomicName id="4C2E8DA74C5CC67D1A902C29FAE42068" box="[1200,1335,259,284]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="drepana">
<emphasis id="B95A2A364C5CC67D1A902C29FAE42068" box="[1200,1335,259,284]" italics="true" pageId="42" pageNumber="43">Ht. drepana</emphasis>
</taxonomicName>
group in that study include
<emphasis id="B95A2A364C5CC67D1F582C05FCB1204B" box="[376,866,294,319]" italics="true" pageId="42" pageNumber="43">
<taxonomicName id="4C2E8DA74C5CC67D1F582C05FDE3204B" box="[376,560,294,319]" class="Insecta" family="Nymphalidae" genus="Erebia" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="passandava">Ht. passandava</taxonomicName>
,
<taxonomicName id="4C2E8DA74C5CC67D1C622C05FD10204B" box="[578,707,296,319]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="narova">Ht. narova</taxonomicName>
,
<taxonomicName id="4C2E8DA74C5CC67D1CF52C05FCB1204B" box="[725,866,294,319]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="difficilis">Ht. difficilis</taxonomicName>
</emphasis>
and
<taxonomicName id="4C2E8DA74C5CC67D1D852C05FBC5204B" box="[933,1046,296,319]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="strato">
<emphasis id="B95A2A364C5CC67D1D852C05FBC5204B" box="[933,1046,296,319]" italics="true" pageId="42" pageNumber="43">Ht. strato</emphasis>
</taxonomicName>
). A 706-bp relatively conserved 28SDDFF sequence (BMNH(E) #676640,
<accessionNumber id="947D6BC74C5CC67D1CA82C67FD2B2017" box="[648,760,330,355]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ817836" pageId="42" pageNumber="43">FJ817836</accessionNumber>
) as aligned is identical to those of
<taxonomicName id="4C2E8DA74C5CC67D1A802C61FADC2017" box="[1184,1295,332,355]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="strato">
<emphasis id="B95A2A364C5CC67D1A802C61FADC2017" box="[1184,1295,332,355]" italics="true" pageId="42" pageNumber="43">Ht. strato</emphasis>
</taxonomicName>
(BMNH(E) #671544
<accessionNumber id="947D6BC74C5CC67D1F292C42FEA920F3" box="[265,378,367,391]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ817810" pageId="42" pageNumber="43">FJ817810</accessionNumber>
) and
<taxonomicName id="4C2E8DA74C5CC67D1FEA2C42FD4120F3" box="[458,658,366,391]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="andravahana">
<emphasis id="B95A2A364C5CC67D1FEA2C42FD4120F3" box="[458,658,366,391]" italics="true" pageId="42" pageNumber="43">Ht. andravahana</emphasis>
</taxonomicName>
(BMNH(E) #671565
<accessionNumber id="947D6BC74C5CC67D1D862C43FBC420F3" box="[934,1047,366,391]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ817806" pageId="42" pageNumber="43">FJ817806</accessionNumber>
, BMNH(E) #676629
<accessionNumber id="947D6BC74C5CC67D1B062C42FA4B20F3" box="[1318,1432,367,391]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ817807" pageId="42" pageNumber="43">FJ817807</accessionNumber>
, BMNH(E) #676641
<accessionNumber id="947D6BC74C5CC67D1F5E2CBFFE2220DE" box="[382,497,402,426]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ817808" pageId="42" pageNumber="43">FJ817808</accessionNumber>
;
<taxonomicName id="4C2E8DA74C5CC67D1C252CBFFDB220DF" authority="Monaghan" authorityName="Monaghan" authorityYear="2009" box="[517,609,402,427]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="difficilis">
<emphasis id="B95A2A364C5CC67D1C252CBFFDB220DF" box="[517,609,402,427]" italics="true" pageId="42" pageNumber="43">difficilis</emphasis>
</taxonomicName>
:
<bibRefCitation id="EFBF8BD54C5CC67D1C592CBFFCA820DE" author="Monaghan" box="[633,891,402,427]" pageId="42" pageNumber="43" refString="Monaghan, M. T., Wild, R., Elliot, E., Fujisawa, T., Balke, M, Inward, D. J. G., Lees, D. C., Ranaivosolo, R., Eggleton, P., Barraclough, T. G. &amp; Vogler, A. P. (2009) Accelerated species inventory on Madagascar using coalescent-based models of species delineation. Systematic Biology, 58, 298 - 311. http: // dx. doi. org / 10.1093 / sysbio / syp 027" type="journal article" year="2009">
Monaghan
<emphasis id="B95A2A364C5CC67D1CD82CBEFCE320DF" box="[760,816,402,427]" italics="true" pageId="42" pageNumber="43">et al.</emphasis>
, 2009
</bibRefCitation>
).
</paragraph>
<paragraph id="8B91F6244C5CC67D1EE72C98FEC023B3" blockId="42.[151,1437,151,2030]" pageId="42" pageNumber="43">
<emphasis id="B95A2A364C5CC67D1EE72C98FE3720BA" bold="true" box="[199,484,437,462]" pageId="42" pageNumber="43">Phylogeny/sister species</emphasis>
: Based on parsimony analysis of COII sequences,
<bibRefCitation id="EFBF8BD54C5CC67D1A022C9AFB2320BA" author="Torres" box="[1058,1264,437,462]" pageId="42" pageNumber="43" refString="Torres, E., Lees, D. C., Vane-Wright, R. I., Kremen, C., Leonard, J. A. &amp; Wayne, R. K. (2001) Examining monophyly in a large radiation of Madagascan butterflies (Lepidoptera: Satyrinae: Mycalesina) based on mitochondrial DNA data. Molecular Phylogenetics and Evolution, 20, 460 - 473. http: // dx. doi. org / 10.1006 / mpev. 2001.0984" type="journal article" year="2001">
Torres
<emphasis id="B95A2A364C5CC67D1A512C9AFB6320BA" box="[1137,1200,437,462]" italics="true" pageId="42" pageNumber="43">et al.,</emphasis>
2001
</bibRefCitation>
had sp. 76 as sister to
<taxonomicName id="4C2E8DA74C5CC67D1EDB2CF7FEAC2085" box="[251,383,474,497]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="imerina">
<emphasis id="B95A2A364C5CC67D1EDB2CF7FEAC2085" box="[251,383,474,497]" italics="true" pageId="42" pageNumber="43">Ht. imerina</emphasis>
</taxonomicName>
(their
<emphasis id="B95A2A364C5CC67D1FF82CF7FDD42085" box="[472,519,474,497]" italics="true" pageId="42" pageNumber="43">Hen</emphasis>
. sp. 22) with 100% bootstrap support. In their dataset, with 417 bp compared, ignoring two differences at the 5 end,
<taxonomicName id="4C2E8DA74C5CC67D1C6F2CD3FD032361" box="[591,720,510,533]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="imerina">
<emphasis id="B95A2A364C5CC67D1C6F2CD3FD032361" box="[591,720,510,533]" italics="true" pageId="42" pageNumber="43">Ht. imerina</emphasis>
</taxonomicName>
(
<accessionNumber id="947D6BC74C5CC67D1CC02CD0FC882362" box="[736,859,509,534]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AY040150" pageId="42" pageNumber="43">
AY
<date id="FF90D0E44C5CC67D1D272CD0FC882362" box="[775,859,509,534]" pageId="42" pageNumber="43" value="50-04-01">040150</date>
</accessionNumber>
based on 2 ♂♂ from Ankazomivady Forest
<quantity id="4CD65BC14C5CC67D1B752CD3FA4F2362" box="[1365,1436,509,534]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="42" pageNumber="43" unit="km" value="32.0">32 km</quantity>
S. Ambositra, see correction below) is only 2 bp (0.48% pairwise divergent) from their
<emphasis id="B95A2A364C5CC67D1A5F2F0FFB7D234D" box="[1151,1198,546,569]" italics="true" pageId="42" pageNumber="43">Hen</emphasis>
. sp. 76 (
<accessionNumber id="947D6BC74C5CC67D1B392F0CFA4B234D" box="[1305,1432,545,569]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AY040174" pageId="42" pageNumber="43">
AY
<date id="FF90D0E44C5CC67D1B622F0CFA4B234D" box="[1346,1432,545,569]" pageId="42" pageNumber="43" value="74-04-01">040174</date>
</accessionNumber>
, based on 2 ♂♂ from RS Anjanaharibe Sud). According to the total evidence morphological analysis of
<bibRefCitation id="EFBF8BD54C5CC67D1B472F68FEB923F5" author="Lees" pageId="42" pageNumber="43" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">
Lees (1997: 156,
<figureCitation id="1315EAA14C5CC67D1F3F2F45FEB323F5" box="[287,352,616,641]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." figureDoi="http://doi.org/10.5281/zenodo.264598" httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="42" pageNumber="43">Fig. 1</figureCitation>
)
</bibRefCitation>
, there was 0.963 jackknife support for a sister relationship of these exemplars with
<taxonomicName id="4C2E8DA74C5CC67D1B342F44FA4823F5" box="[1300,1435,616,641]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="B95A2A364C5CC67D1B342F44FA4823F5" box="[1300,1435,616,641]" italics="true" pageId="42" pageNumber="43">Ht. pauliani</emphasis>
</taxonomicName>
(“sp. 37”). There is some support for
<taxonomicName id="4C2E8DA74C5CC67D1C152FA0FD6523D0" box="[565,694,653,676]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="imerina">
<emphasis id="B95A2A364C5CC67D1C152FA0FD6523D0" box="[565,694,653,676]" italics="true" pageId="42" pageNumber="43">Ht. imerina</emphasis>
</taxonomicName>
as the sister of
<taxonomicName id="4C2E8DA74C5CC67D1D422FA0FC3A23D1" box="[866,1001,652,677]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="B95A2A364C5CC67D1D422FA0FC3A23D1" box="[866,1001,652,677]" italics="true" pageId="42" pageNumber="43">Ht. pauliani</emphasis>
</taxonomicName>
in the combined tree of
<bibRefCitation id="EFBF8BD54C5CC67D1AD72FA1FEDF23BC" pageId="42" pageNumber="43" refString="Aduse-Poku, K., Lees, D. C., Brattstrom, O., Kodandaramaiah, U., Collins, S. &amp; Brakefield, P. M., (2016) Molecular phylogeny and generic-level taxonomy of the widespread palaeotropical ' Heteropsis clade' (Nymphalidae: Satyrinae: Mycalesina). Systematic Entomology, in press. http: // dx. doi. org / syen. 12183" type="book">
Aduse-Poku
<emphasis id="B95A2A364C5CC67D1BA82FA0FF6823BC" italics="true" pageId="42" pageNumber="43">et al.,</emphasis>
(2016)
</bibRefCitation>
.
</paragraph>
<paragraph id="8B91F6244C5CC67D1EE72FFEFE252398" blockId="42.[151,1437,151,2030]" box="[199,502,723,748]" pageId="42" pageNumber="43">
<emphasis id="B95A2A364C5CC67D1EE72FFEFE252398" bold="true" box="[199,502,723,748]" pageId="42" pageNumber="43">Ecology and distribution.</emphasis>
</paragraph>
<paragraph id="8B91F6244C5CC67D1EE72FDAFE2F2223" blockId="42.[151,1437,151,2030]" pageId="42" pageNumber="43">
<emphasis id="B95A2A364C5CC67D1EE72FDAFEF02264" bold="true" box="[199,291,759,784]" pageId="42" pageNumber="43">Habitat</emphasis>
: primary rainforest. This species was (remarkably) once found in Parc de Tsimbazaza in 1951 (MNHN), so it may persist or may have persisted until recently in other central habitats well separated from the current eastern rainforest block.
</paragraph>
<paragraph id="8B91F6244C5CC67D1EE72E4FFABF220F" blockId="42.[151,1437,151,2030]" box="[199,1388,866,891]" pageId="42" pageNumber="43">
<emphasis id="B95A2A364C5CC67D1EE72E4FFE90220F" bold="true" box="[199,323,866,891]" pageId="42" pageNumber="43">Behaviour</emphasis>
: Low flying, found mostly along ridges but also in riparian areas, perching low off the ground.
</paragraph>
<paragraph id="8B91F6244C5CC67D1EE72EABFBCC22EA" blockId="42.[151,1437,151,2030]" box="[199,1055,902,927]" pageId="42" pageNumber="43">
<emphasis id="B95A2A364C5CC67D1EE72EABFE9522EB" bold="true" box="[199,326,902,927]" pageId="42" pageNumber="43">Hostplant:</emphasis>
occurs among native climbing bamboos, which it might feed on.
</paragraph>
<paragraph id="8B91F6244C5CC67D1EE72E87FD6022B7" blockId="42.[151,1437,151,2030]" box="[199,691,938,963]" pageId="42" pageNumber="43">
<emphasis id="B95A2A364C5CC67D1EE72E87FE8B22B7" bold="true" box="[199,344,938,963]" pageId="42" pageNumber="43">Early stages</emphasis>
: expressed egg pearly whitish.
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C5CC67D1EE72EE0FDC72594" pageId="42" pageNumber="43" type="distribution">
<paragraph id="8B91F6244C5CC67D1EE72EE0FAE125ED" blockId="42.[151,1437,151,2030]" pageId="42" pageNumber="43">
<emphasis id="B95A2A364C5CC67D1EE72EE0FE8A2292" bold="true" box="[199,345,973,998]" pageId="42" pageNumber="43">Distribution</emphasis>
: widespread in the principal eastern rainforest belt and also in high plateau relics notably Ankazomivady south of Ambositra (
<figureCitation id="1315EAA14C5CC67D1C1D2EDCFD42257E" box="[573,657,1009,1034]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees &amp; Kremen, sp. nov. Light blue: Ht. hazovola Lees &amp; Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees &amp; Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees &amp; Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees &amp; Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees &amp; Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees &amp; Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui &amp; Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees &amp; Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="42" pageNumber="43">Fig. 30</figureCitation>
A, turquoise dots). The locality was incorrectly given in Table 1 of
<bibRefCitation id="EFBF8BD54C5CC67D1EB7293BFEB4255A" author="Torres" box="[151,359,1045,1070]" pageId="42" pageNumber="43" refString="Torres, E., Lees, D. C., Vane-Wright, R. I., Kremen, C., Leonard, J. A. &amp; Wayne, R. K. (2001) Examining monophyly in a large radiation of Madagascan butterflies (Lepidoptera: Satyrinae: Mycalesina) based on mitochondrial DNA data. Molecular Phylogenetics and Evolution, 20, 460 - 473. http: // dx. doi. org / 10.1006 / mpev. 2001.0984" type="journal article" year="2001">
Torres
<emphasis id="B95A2A364C5CC67D1EC6293BFEF3255A" box="[230,288,1045,1070]" italics="true" pageId="42" pageNumber="43">et al.</emphasis>
, 2001
</bibRefCitation>
(which contains a formatting glitch) as Anjanaharibe Sud, 1240 m instead of the line below,
<quantity id="4CD65BC14C5CC67D1EBC2914FF342526" box="[156,231,1081,1106]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="42" pageNumber="43" unit="km" value="32.0">32 km</quantity>
S. of Ambositra. The species extends to the northern mountain range of the rainforest biome. Sp. 76 from RS Anjanaharibe Sud is doubtless referable to
<taxonomicName id="4C2E8DA74C5CC67D1CF72973FC8C2500" box="[727,863,1117,1141]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="imerina">
<emphasis id="B95A2A364C5CC67D1CF72973FC8C2500" box="[727,863,1117,1141]" italics="true" pageId="42" pageNumber="43">Ht. imerina</emphasis>
</taxonomicName>
, as for material (sp. 65) from Bekolosy (RS Manongarivo) and RNI Tsaratanana, while this is a species for which phylogeographic study is desirable.
</paragraph>
<paragraph id="8B91F6244C5CC67D1EE72989FDC72594" blockId="42.[151,1437,151,2030]" pageId="42" pageNumber="43">
<emphasis id="B95A2A364C5CC67D1EE72989FE4F25C9" bold="true" box="[199,412,1188,1213]" pageId="42" pageNumber="43">Elevational range</emphasis>
:
<quantity id="4CD65BC14C5CC67D1F8E2989FD9125C8" box="[430,578,1188,1213]" metricMagnitude="3" metricUnit="m" metricValue="1.359" metricValueMax="1.932" metricValueMin="0.7859999999999999" pageId="42" pageNumber="43" unit="m" value="1359.0" valueMax="1932.0" valueMin="786.0">7861932 m</quantity>
. (n=118; DCL-DB-2901 from Tamatave-Anivorano [estimated at
<quantity id="4CD65BC14C5CC67D1B1A2988FAAB25C8" box="[1338,1400,1189,1213]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="42" pageNumber="43" unit="m" value="80.0">80 m</quantity>
] is discounted as an unlikely outlier).
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C5CC67C1EE729C6FD3E27E9" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="8B91F6244C5CC67C1EE729C6FD3E27E9" blockId="42.[151,1437,151,2030]" lastBlockId="43.[151,1437,151,1693]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
<emphasis id="B95A2A364C5CC67D1EE729C6FE6A2470" bold="true" box="[199,441,1259,1284]" pageId="42" pageNumber="43">Referred specimens</emphasis>
: ♂,
<collectingCountry id="F339B6B44C5CC67D1FD929C6FD532470" box="[505,640,1259,1284]" name="Madagascar" pageId="42" pageNumber="43">Madagascar</collectingCountry>
E, Bemoara camp (going out from to Ambavala), Zahamena PN [northwestern sector],
<geoCoordinate id="EE1A90E34C5CC67D1FB6283FFDCB245F" box="[406,536,1295,1323]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="-17.51875">17.51875o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5CC67D1C06283FFD7B245E" box="[550,680,1295,1323]" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="48.72455">48.72455o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5CC67D1CFA283EFCE9245F" box="[730,826,1298,1323]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="42" pageNumber="43" unit="km" value="0.27">0.27 km</quantity>
,
<quantity id="4CD65BC14C5CC67D1D66283EFC3F245E" box="[838,1004,1298,1323]" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.125" metricValueMin="1.075" pageId="42" pageNumber="43" unit="m" value="1100.0" valueMax="1125.0" valueMin="1075.0">1100 +/- 25 m</quantity>
,
<date id="FF90D0E44C5CC67D1DD9283FFBBF245F" box="[1017,1132,1298,1323]" pageId="42" pageNumber="43" value="2006-11-04">4/11/2006</date>
, D.C. Lees:
<collectionCode id="ED3F6EE14C5CC67D1ADB283FFAF0245F" box="[1275,1315,1298,1323]" pageId="42" pageNumber="43">DL</collectionCode>
06-023; ♂,
<taxonomicName id="4C2E8DA74C5CC67D1EB72818FF3D243A" box="[151,238,1333,1358]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="93" phylum="Arthropoda" rank="species" species="imerina">Imerina</taxonomicName>
<collectingCountry id="F339B6B44C5CC67D1EDA2818FE55243A" box="[250,390,1333,1358]" name="Madagascar" pageId="42" pageNumber="43">Madagascar</collectingCountry>
|1910 E. La Moult |
<collectionCode id="ED3F6EE14C5CC67D1C4A281BFD69243A" box="[618,698,1334,1358]" pageId="42" pageNumber="43">COLL</collectionCode>
. BOULLET MUSEUM
<collectionCode id="ED3F6EE14C5CC67D1DFC281BFBE3243A" box="[988,1072,1334,1358]" pageId="42" pageNumber="43">PARIS</collectionCode>
|
<taxonomicName id="4C2E8DA74C5CC67D1A11281BFB30243A" box="[1073,1251,1333,1358]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="andravahana">H. andravahana</taxonomicName>
ab. marmorata, Auriv. [sic]|DCL-DB-2897; ♀,
<taxonomicName id="4C2E8DA74C5CC67D1FDB2874FD812406" box="[507,594,1369,1394]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="93" phylum="Arthropoda" rank="species" species="imerina">Imerina</taxonomicName>
<collectingCountry id="F339B6B44C5CC67D1C7A2874FD352406" box="[602,742,1369,1394]" name="Madagascar" pageId="42" pageNumber="43">Madagascar</collectingCountry>
|1910 E. La Moult |DCL-DB-2911; ♂ [
<collectionCode id="ED3F6EE14C5CC67D1ABC2877FB292405" box="[1180,1274,1370,1393]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="42" pageNumber="43">MNHN</collectionCode>
], Centre, forêt d'Andranobe, route d'Andriamena, O. of Lac Alaotra [
<geoCoordinate id="EE1A90E34C5CC67D1CD228ADFC8924EC" box="[754,858,1405,1433]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="55" value="-17.755">17.755o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5CC67D1D4728ADFC6D24EC" box="[871,958,1405,1432]" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="555" value="47.98">47.98o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5CC67D1DCD28ADFBEE24ED" box="[1005,1085,1408,1433]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="42" pageNumber="43" unit="km" value="1.5">1.5 km</quantity>
,
<quantity id="4CD65BC14C5CC67D1A6928ADFB7224EC" box="[1097,1185,1408,1433]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="42" pageNumber="43" unit="m" value="1250.0">1250 m</quantity>
],
<date id="FF90D0E44C5CC67D1A9428ADFAF324EC" box="[1204,1312,1408,1432]" pageId="42" pageNumber="43" value="1972-11-01">1/11/1972</date>
|DCL-DB- 2914; ♂ [
<collectionCode id="ED3F6EE14C5CC67D1EDF2888FE8E24C8" box="[255,349,1445,1468]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="42" pageNumber="43">MNHN</collectionCode>
], data as above but:
<date id="FF90D0E44C5CC67D1C1C2889FD6D24C9" box="[572,702,1444,1469]" pageId="42" pageNumber="43" value="1972-09-01" valueMax="1972-09-06" valueMin="1972-09-01">16/9/1972</date>
, A. Peyrieras|DCL-DB-2915; ♀, Centre, forêt d'Andranobe, route d'Andriamena O. du lac Alaotra [ca.
<geoCoordinate id="EE1A90E34C5CC67D1C1128E7FD452497" box="[561,662,1479,1507]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="55" value="-17.755">17.755o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5CC67D1C8328E6FD292496" box="[675,762,1479,1507]" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="555" value="47.98">47.98o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5CC67D1D0828E7FCAA2497" box="[808,889,1482,1507]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="42" pageNumber="43" unit="km" value="1.5">1.5 km</quantity>
,
<quantity id="4CD65BC14C5CC67D1DA328E7FC0F2496" box="[899,988,1482,1507]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="42" pageNumber="43" unit="m" value="1250.0">1250 m</quantity>
], 29-1/
<date id="FF90D0E44C5CC67D1A0928E7FB4B2497" box="[1065,1176,1482,1507]" pageId="42" pageNumber="43" value="1970-02-06">6-II-1970</date>
, P. Griveaud|DCL-DB- 2913; ♀, data as above but DCL-DB-2913; ♀, [
<collectionCode id="ED3F6EE14C5CC67D1CEC28C2FCF92772" box="[716,810,1519,1542]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="42" pageNumber="43">MNHN</collectionCode>
], data as above but|DCL-DB-2916; ♂, E, Tamatave et Anivorano, [ca.
<geoCoordinate id="EE1A90E34C5CC67D1F762B39FE1A2759" box="[342,457,1553,1581]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="5" value="-18.4585">18.4585o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5CC67D1FFA2B38FD9D2758" box="[474,590,1553,1581]" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="5" value="49.1832">49.1832o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5CC67D1CA52B38FD002759" box="[645,723,1556,1581]" metricMagnitude="4" metricUnit="m" metricValue="3.9" pageId="42" pageNumber="43" unit="km" value="39.0">39 km</quantity>
,
<quantity id="4CD65BC14C5CC67D1CC32B38FCF12759" box="[739,802,1556,1581]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="42" pageNumber="43" unit="m" value="80.0">80 m</quantity>
]|1915|MUSEUM
<collectionCode id="ED3F6EE14C5CC67D1DD32B38FB932759" box="[1011,1088,1557,1581]" pageId="42" pageNumber="43">PARIS</collectionCode>
DON.
<collectingCountry id="F339B6B44C5CC67D1ABB2B38FB132758" box="[1179,1216,1557,1580]" name="South Georgia and the South Sandwich Islands" pageId="42" pageNumber="43">DE</collectingCountry>
H. UNGEMACH/ GENITALIA m. P. VIETTE
<collectionCode id="ED3F6EE14C5CC67D1FC52B14FDFA2724" box="[485,553,1593,1616]" pageId="42" pageNumber="43">PREP</collectionCode>
. NO. 4864|DCL-DB-2901; ♂, E, Analamay, 18.82795oS, 48.3388oE +/-
<quantity id="4CD65BC14C5CC67D1B592B14FF6D2701" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="42" pageNumber="43" unit="km" value="2.1">2.1 km</quantity>
,
<quantity id="4CD65BC14C5CC67D1EEE2B70FEA02700" box="[206,371,1628,1653]" metricMagnitude="2" metricUnit="m" metricValue="9.78" metricValueMax="10.28" metricValueMin="9.28" pageId="42" pageNumber="43" unit="m" value="978.0" valueMax="1028.0" valueMin="928.0">978 +/- 50 m</quantity>
,
<date id="FF90D0E44C5CC67D1FA42B70FE2B2701" box="[388,504,1629,1653]" pageId="42" pageNumber="43" value="2004-03-11">11/3/2004</date>
, D.C. Lees: DL-4-515,
<collectionCode id="ED3F6EE14C5CC67D1D362B70FCBE2701" box="[790,877,1629,1653]" pageId="42" pageNumber="43">BMNH</collectionCode>
(E) #676683,
<collectionCode id="ED3F6EE14C5CC67D1A302B71FBB42701" box="[1040,1127,1628,1653]" pageId="42" pageNumber="43">BMAD</collectionCode>
046-09, CCDB-02225-D10 [
<collectionCode id="ED3F6EE14C5CC67D1EBD2BAEFF0E27EF" box="[157,221,1667,1691]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="42" pageNumber="43">DNA</collectionCode>
barcode; cluster number
<collectionCode id="ED3F6EE14C5CC67D1C262BAEFD8727EF" box="[518,596,1667,1691]" pageId="42" pageNumber="43">BOLD</collectionCode>
:AAE4110; ♂, E, Ambatovy-Berano,
<geoCoordinate id="EE1A90E34C5CC67D1A272BAFFB4527EF" box="[1031,1174,1663,1691]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="-18.848845">18.848845o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5CC67D1A862BAFFAE627EE" box="[1190,1333,1663,1691]" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="48.337055">48.337055o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5CC67D1B4B2BAFFF6D27B6" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="42" pageNumber="43" unit="km" value="0.65">0.65 km</quantity>
,
<quantity id="4CD65BC14C5CC67D1EEA2B84FE8027B5" box="[202,339,1705,1730]" metricMagnitude="2" metricUnit="m" metricValue="9.65" metricValueMax="9.72" metricValueMin="9.58" pageId="42" pageNumber="43" unit="m" value="965.0" valueMax="972.0" valueMin="958.0">965 +/- 7 m</quantity>
,
<date id="FF90D0E44C5CC67D1F402B87FE0527B6" box="[352,470,1706,1730]" pageId="42" pageNumber="43" value="2004-03-12">12/3/2004</date>
: 10:43, R. Ranaivosolo: DL-4-643; ♀, E, Ambatovy,
<geoCoordinate id="EE1A90E34C5CC67D1A172B84FB6B27B6" box="[1079,1208,1702,1730]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="-18.84935">18.84935o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5CC67D1AE62B87FA9427B5" box="[1222,1351,1702,1730]" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="48.32399">48.32399o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5CC67D1B592B87FF6D279C" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="42" pageNumber="43" unit="km" value="0.3">0.3 km</quantity>
,
<quantity id="4CD65BC14C5CC67D1EEA2BFDFEA22793" box="[202,369,1743,1768]" metricMagnitude="3" metricUnit="m" metricValue="1.024" metricValueMax="1.049" metricValueMin="0.999" pageId="42" pageNumber="43" unit="m" value="1024.0" valueMax="1049.0" valueMin="999.0">1024 +/- 25 m</quantity>
,
<date id="FF90D0E44C5CC67D1F5E2BE2FE26279C" box="[382,501,1743,1768]" pageId="42" pageNumber="43" value="2004-02-25">25/2/2004</date>
: 15:33, R. Ranaivosolo: DL-4-99; ♂, E, Ambatovy-Berano,
<geoCoordinate id="EE1A90E34C5CC67D1A872BE2FB2D279C" box="[1191,1278,1740,1768]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="555" value="-18.85">18.85o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5CC67D1B2C2BE2FAA12793" box="[1292,1394,1740,1768]" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="55" value="48.335">48.335o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5CC67D1EB72BD9FF3F2678" box="[151,236,1779,1804]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="42" pageNumber="43" unit="km" value="1.7">1.7 km</quantity>
,
<quantity id="4CD65BC14C5CC67D1EDD2BD9FE7D267F" box="[253,430,1780,1804]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.09" metricValueMin="0.9099999999999999" pageId="42" pageNumber="43" unit="m" value="1000.0" valueMax="1090.0" valueMin="910.0">1000 +/- 90 m</quantity>
,
<date id="FF90D0E44C5CC67D1F9F2BD9FDE72678" box="[447,564,1780,1804]" pageId="42" pageNumber="43" value="2004-03-12">12/3/2004</date>
, D.C. Lees: DL-4-590, IA27 [isotope voucher]; ♂, data as above but: R. Ranaivosolo: DL-4-666; ♂, E, Ambatovy second site,
<geoCoordinate id="EE1A90E34C5CC67D1D392A37FC582646" box="[793,907,1814,1843]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="5" value="-18.8511">18.8511o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5CC67D1DBA2A37FBDE2646" box="[922,1037,1814,1842]" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="5" value="48.3221">48.3221o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5CC67D1A602A37FB732647" box="[1088,1184,1818,1843]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="42" pageNumber="43" unit="km" value="0.15">0.15 km</quantity>
,
<quantity id="4CD65BC14C5CC67D1A8D2A37FA862646" box="[1197,1365,1818,1843]" metricMagnitude="3" metricUnit="m" metricValue="1.055" metricValueMax="1.08" metricValueMin="1.03" pageId="42" pageNumber="43" unit="m" value="1055.0" valueMax="1080.0" valueMin="1030.0">1055 +/- 25 m</quantity>
,
<date id="FF90D0E44C5CC67D1B422A37FF072622" pageId="42" pageNumber="43" value="2004-02-25">25/2/ 2004</date>
: after 12:00, D.C. Lees: DL-4-139, 547 [=
<collectionCode id="ED3F6EE14C5CC67D1C992A10FD322622" box="[697,737,1853,1878]" pageId="42" pageNumber="43">DL</collectionCode>
0547;
<collectionCode id="ED3F6EE14C5CC67D1D0A2A13FCB42622" box="[810,871,1854,1878]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="42" pageNumber="43">DNA</collectionCode>
extract number],
<collectionCode id="ED3F6EE14C5CC67D1A142A13FB582622" box="[1076,1163,1854,1878]" pageId="42" pageNumber="43">BMNH</collectionCode>
(E) #671912; ♀, data as above but:
<date id="FF90D0E44C5CC67D1F332A4FFE5A260E" box="[275,393,1890,1914]" pageId="42" pageNumber="43" value="2004-03-21">21/3/2004</date>
: 09:29, V. Rakotomalala: VFR_1329 [photo, verified
<collectionCode id="ED3F6EE14C5CC67D1DC12A4FFBCC260E" box="[993,1055,1890,1914]" pageId="42" pageNumber="43">DCL</collectionCode>
]; ♀, data as above but:
<date id="FF90D0E44C5CC67D1B032A4FFA4A260E" box="[1315,1433,1890,1914]" pageId="42" pageNumber="43" value="2004-03-19">19/3/2004</date>
: 09:54, V. Rakotomalala: VFR_1254 [photo, verified
<collectionCode id="ED3F6EE14C5CC67D1CD82AA5FCE526D4" box="[760,822,1928,1952]" pageId="42" pageNumber="43">DCL</collectionCode>
]; ♂, E, Ambatovy,
<geoCoordinate id="EE1A90E34C5CC67D1A392AA5FB5F26D4" box="[1049,1164,1925,1953]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="5" value="-18.8522">18.8522o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5CC67D1ABC2AA5FADC26D4" box="[1180,1295,1925,1952]" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="5" value="48.3177">48.3177o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5CC67D1B642AA5FA4B26D5" box="[1348,1432,1928,1953]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="42" pageNumber="43" unit="km" value="0.5">0.5 km</quantity>
,
<quantity id="4CD65BC14C5CC67D1EB72A83FEEF26B2" box="[151,316,1966,1991]" metricMagnitude="3" metricUnit="m" metricValue="1.085" metricValueMax="1.115" metricValueMin="1.055" pageId="42" pageNumber="43" unit="m" value="1085.0" valueMax="1115.0" valueMin="1055.0">1085 +/- 30 m</quantity>
,
<date id="FF90D0E44C5CC67D1F692A83FE6C26B3" box="[329,447,1966,1991]" pageId="42" pageNumber="43" value="2004-02-25">25/2/2004</date>
: 12:59, R. Ranaivosolo: DL-4-84; ♂, E, Ambatovy-Sahavarina,
<geoCoordinate id="EE1A90E34C5CC67D1AB92A83FADF26B3" box="[1177,1292,1963,1991]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="5" value="-18.8577">18.8577o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5CC67D1B3A2A83FA4F26B2" box="[1306,1436,1963,1991]" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="48.32579">48.32579o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5CC67D1E9F2AFBFEC3269A" box="[191,272,2005,2030]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="42" pageNumber="43" unit="km" value="0.3">0.3 km</quantity>
,
<quantity id="4CD65BC14C5CC67D1F3C2AFBFE6C2699" box="[284,447,2006,2030]" metricMagnitude="3" metricUnit="m" metricValue="1.09" metricValueMax="1.13" metricValueMin="1.05" pageId="42" pageNumber="43" unit="m" value="1090.0" valueMax="1130.0" valueMin="1050.0">1090 +/- 40 m</quantity>
,
<date id="FF90D0E44C5CC67D1FEB2AFBFD92269A" box="[459,577,2006,2030]" pageId="42" pageNumber="43" value="2004-03-13">13/3/2004</date>
: 09:23, R. Ranaivosolo: DL-4-707; ♀, E, Ambatovy-Sahavarina,
<geoCoordinate id="EE1A90E34C5CC67D1B032AF8FA45269A" box="[1315,1430,2002,2030]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="5" value="-18.859">18.8590o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1EB72DB7FED921C6" box="[151,266,151,179]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5" value="48.3252">48.3252o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1F1C2DB6FE5D21C7" box="[316,398,154,179]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="43" pageNumber="44" unit="km" value="0.4">0.4 km</quantity>
,
<quantity id="4CD65BC14C5DC67C1FBA2DB6FD9321C6" box="[410,576,154,179]" metricMagnitude="3" metricUnit="m" metricValue="1.09" metricValueMax="1.135" metricValueMin="1.045" pageId="43" pageNumber="44" unit="m" value="1090.0" valueMax="1135.0" valueMin="1045.0">1090 +/- 45 m</quantity>
,
<date id="FF90D0E44C5DC67C1C6D2DB6FD1021C7" box="[589,707,155,179]" pageId="43" pageNumber="44" value="2004-03-13">13/3/2004</date>
: 10:40, R. Ranaivosolo: DL-4-681; ♂, data as above but: DL-4- 682; ♀, E, Ambatovy- Sahamaloto-Torotorofotsy path, S. to Sahavarina,
<geoCoordinate id="EE1A90E34C5DC67C1DF22DECFB8721AD" box="[978,1108,190,218]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="1" value="-18.86313">18.86313o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1A412DECFB0621AD" box="[1121,1237,190,218]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5" value="48.3425">48.3425o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1B262DECFAB521AE" box="[1286,1382,193,218]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="43" pageNumber="44" unit="km" value="0.15">0.15 km</quantity>
,
<quantity id="4CD65BC14C5DC67C1B522DECFED9218B" metricMagnitude="2" metricUnit="m" metricValue="9.76" metricValueMax="10.01" metricValueMin="9.51" pageId="43" pageNumber="44" unit="m" value="976.0" valueMax="1001.0" valueMin="951.0">976 +/- 25 m</quantity>
,
<date id="FF90D0E44C5DC67C1F3E2DC5FE462074" box="[286,405,232,256]" pageId="43" pageNumber="44" value="2004-03-14">14/3/2004</date>
: 10:39, D.C. Lees:
<collectionCode id="ED3F6EE14C5DC67C1CAE2DCAFD162074" box="[654,709,231,256]" country="USA" httpUri="http://grbio.org/cool/s9x3-awht" name="Dicty Stock Center" pageId="43" pageNumber="44">DSC</collectionCode>
_0496-7 [photo]; ♂, E, Torotorofotsy-Sahavarina,
<geoCoordinate id="EE1A90E34C5DC67C1B032DCAFA452074" box="[1315,1430,228,256]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="5" value="-18.8657">18.8657o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1EB72C23FECB2052" box="[151,280,267,295]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="1" value="48.34547">48.34547o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1F732C23FE642053" box="[339,439,270,295]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="43" pageNumber="44" unit="km" value="0.25">0.25 km</quantity>
,
<quantity id="4CD65BC14C5DC67C1FE82C23FDBE2052" box="[456,621,270,295]" metricMagnitude="2" metricUnit="m" metricValue="9.690000000000001" metricValueMax="9.940000000000001" metricValueMin="9.440000000000001" pageId="43" pageNumber="44" unit="m" value="969.0" valueMax="994.0" valueMin="944.0">969 +/- 25 m</quantity>
,
<date id="FF90D0E44C5DC67C1C5E2C22FD262053" box="[638,757,271,295]" pageId="43" pageNumber="44" value="2004-03-14">14/3/2004</date>
: 10:5213:54, D.C. Lees: DL-4-730; ♀, E, Torotorofotsy (Sahavarina), ilot,
<geoCoordinate id="EE1A90E34C5DC67C1F502C19FE372039" box="[368,484,305,333]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="5" value="-18.8665">18.8665o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1FD52C18FDBA2039" box="[501,617,305,333]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5" value="48.3492">48.3492o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1C812C19FCC02039" box="[673,787,308,333]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="43" pageNumber="44" unit="km" value="0.015">0.015 km</quantity>
,
<quantity id="4CD65BC14C5DC67C1D022C19FC172039" box="[802,964,308,333]" metricMagnitude="2" metricUnit="m" metricValue="9.59" metricValueMax="9.84" metricValueMin="9.34" pageId="43" pageNumber="44" unit="m" value="959.0" valueMax="984.0" valueMin="934.0">959 +/- 25 m</quantity>
,
<date id="FF90D0E44C5DC67C1DF52C18FB9A2039" box="[981,1097,309,333]" pageId="43" pageNumber="44" value="2004-03-14">14/3/2004</date>
, D.C. Lees: DL-4-769, 2485 [=
<collectionCode id="ED3F6EE14C5DC67C1E8D2C75FF052005" box="[173,214,344,369]" pageId="43" pageNumber="44">DL</collectionCode>
2485;
<collectionCode id="ED3F6EE14C5DC67C1F3D2C74FE892005" box="[285,346,345,369]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="43" pageNumber="44">DNA</collectionCode>
extract number],
<collectionCode id="ED3F6EE14C5DC67C1C042C74FDA82005" box="[548,635,345,369]" pageId="43" pageNumber="44">BMNH</collectionCode>
(E) #676685; ♀, data as above but: 14:30, D.C. Lees: DL-4-774; ♂, E, Ambatovy-Sahavarina,
<geoCoordinate id="EE1A90E34C5DC67C1F812C52FDC720E3" box="[417,532,380,408]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="5" value="-18.862">18.8620o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1C012C52FD4620E3" box="[545,661,380,407]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5" value="48.3408">48.3408o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1CE32C52FCC020EC" box="[707,787,383,408]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="43" pageNumber="44" unit="km" value="0.2">0.2 km</quantity>
,
<quantity id="4CD65BC14C5DC67C1D3F2C52FC1220E3" box="[799,961,383,407]" metricMagnitude="3" metricUnit="m" metricValue="1.004" metricValueMax="1.032" metricValueMin="0.976" pageId="43" pageNumber="44" unit="m" value="1004.0" valueMax="1032.0" valueMin="976.0">1004 +/- 28 m</quantity>
,
<date id="FF90D0E44C5DC67C1DED2C52FB9020E3" box="[973,1091,383,407]" pageId="43" pageNumber="44" value="2004-03-14">14/3/2004</date>
: 10:30, R. Ranaivosolo: DL-4- 737; ♂ [
<collectionCode id="ED3F6EE14C5DC67C1EDA2C8BFE8B20C9" box="[250,344,422,445]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="43" pageNumber="44">MNHN</collectionCode>
], C, Tsimbazaza, [ca.
<geoCoordinate id="EE1A90E34C5DC67C1C7D2C8BFD1120CA" box="[605,706,418,446]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="55" value="-18.933">18.933o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1CF32C88FCEB20C9" box="[723,824,418,446]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="55" value="47.533">47.533o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1D4F2C8BFC1020CA" box="[879,963,421,446]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="43" pageNumber="44" unit="km" value="0.9">0.9 km</quantity>
,
<quantity id="4CD65BC14C5DC67C1DF32C8BFBE320C9" box="[979,1072,422,446]" metricMagnitude="3" metricUnit="m" metricValue="1.3169999999999997" pageId="43" pageNumber="44" unit="m" value="1317.0">1317 m</quantity>
],
<date id="FF90D0E44C5DC67C1A672C88FB7E20CA" box="[1095,1197,421,446]" pageId="43" pageNumber="44" value="1951-01-07">7/1/1951</date>
, Villette; ♂ [
<collectionCode id="ED3F6EE14C5DC67C1B692C8BFA4320CA" box="[1353,1424,422,446]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="43" pageNumber="44">CMN</collectionCode>
], Ambatolaona [ca.
<geoCoordinate id="EE1A90E34C5DC67C1F7D2CE0FE062091" box="[349,469,457,485]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="5" value="-18.9333">18.9333o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1FC52CE0FDFD2090" box="[485,558,457,485]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5555" value="47.9">47.9o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1C442CE1FD622091" box="[612,689,460,485]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="43" pageNumber="44" unit="km" value="15.0">15 km</quantity>
,
<quantity id="4CD65BC14C5DC67C1C9F2CE0FCCF2090" box="[703,796,461,485]" metricMagnitude="3" metricUnit="m" metricValue="1.3719999999999999" pageId="43" pageNumber="44" unit="m" value="1372.0">1372 m</quantity>
], 12/1932, M.E.H. Fountaine; ♂ [
<collectionCode id="ED3F6EE14C5DC67C1A852CE0FAD02090" box="[1189,1283,461,484]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="43" pageNumber="44">MNHN</collectionCode>
], CE, Perinet ANALAMAZAOTRA, [ca.
<geoCoordinate id="EE1A90E34C5DC67C1FC32CDEFD85237F" box="[483,598,495,523]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="5" value="-18.9347">18.9347o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1C4A2CDFFD0E237E" box="[618,733,495,523]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5" value="48.4305">48.4305o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1D3A2CDFFC53237F" box="[794,896,498,523]" metricMagnitude="3" metricUnit="m" metricValue="1.68" pageId="43" pageNumber="44" unit="km" value="1.68">1.68 km</quantity>
,
<quantity id="4CD65BC14C5DC67C1DB12CDFFBE8237E" box="[913,1083,498,523]" metricMagnitude="2" metricUnit="m" metricValue="9.25" metricValueMax="9.45" metricValueMin="9.05" pageId="43" pageNumber="44" unit="m" value="925.0" valueMax="945.0" valueMin="905.0">925 +/- 20 m</quantity>
], 7/1967, P. Griveaud; 4 ♂♂ [
<collectionCode id="ED3F6EE14C5DC67C1EBD2F37FF2A2345" box="[157,249,538,561]" pageId="43" pageNumber="44">BMNH</collectionCode>
], CE, Perinet [ca.
<geoCoordinate id="EE1A90E34C5DC67C1FF22F34FD962345" box="[466,581,534,561]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="5" value="-18.9347">18.9347o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1C762F34FD1A2345" box="[598,713,534,562]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5" value="48.4305">48.4305o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1CDF2F34FCB12346" box="[767,866,537,562]" metricMagnitude="3" metricUnit="m" metricValue="1.68" pageId="43" pageNumber="44" unit="km" value="1.68">1.68 km</quantity>
,
<quantity id="4CD65BC14C5DC67C1D502F34FBC22345" box="[880,1041,537,562]" metricMagnitude="2" metricUnit="m" metricValue="9.25" metricValueMax="9.75" metricValueMin="8.75" pageId="43" pageNumber="44" unit="m" value="925.0" valueMax="975.0" valueMin="875.0">925 +/- 50 m</quantity>
], 4/1973, A. E. Wright; 3 ♂♂, E, Perinet, forêt de l'Est,
<collectingCountry id="F339B6B44C5DC67C1FBD2F12FDF7232C" box="[413,548,575,600]" name="Madagascar" pageId="43" pageNumber="44">Madagascar</collectingCountry>
1910. E. Le Moult [ca.
<geoCoordinate id="EE1A90E34C5DC67C1D632F6DFC64232C" box="[835,951,572,600]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="5" value="-18.9347">18.9347o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1DE72F12FBE92323" box="[967,1082,572,600]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5" value="48.4305">48.4305o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1A502F12FB07232C" box="[1136,1236,575,600]" metricMagnitude="3" metricUnit="m" metricValue="1.68" pageId="43" pageNumber="44" unit="km" value="1.68">1.68 km</quantity>
], DCL-DB-
<date id="FF90D0E44C5DC67C1B7C2F6DFF00230B" pageId="43" pageNumber="44" value="2899" valueMax="2901">2899- 2901</date>
; 1 ♀, E, Perinet, forêt de l'Est,
<collectingCountry id="F339B6B44C5DC67C1C0F2F4BFD65230B" box="[559,694,614,639]" name="Madagascar" pageId="43" pageNumber="44">Madagascar</collectingCountry>
1910. E. Le Moult [ca.
<geoCoordinate id="EE1A90E34C5DC67C1DE42F4AFBE4230B" box="[964,1079,611,639]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="5" value="-18.9347">18.9347o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1A642F4BFB6B230A" box="[1092,1208,611,639]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5" value="48.4305">48.4305o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1AC72F4BFA9B230B" box="[1255,1352,614,639]" metricMagnitude="3" metricUnit="m" metricValue="1.68" pageId="43" pageNumber="44" unit="km" value="1.68">1.68 km</quantity>
], DCL- DB-2912; ♂, E, Mantadia, Jofa,
<geoCoordinate id="EE1A90E34C5DC67C1C282FA0FDB323D1" box="[520,608,649,677]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="555" value="-18.78">18.78o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1C4D2FA0FD1623D1" box="[621,709,649,677]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="555" value="48.43">48.43o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1CD52FA0FCE123D1" box="[757,818,652,677]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="43" pageNumber="44" unit="km" value="1.0">1 km</quantity>
,
<quantity id="4CD65BC14C5DC67C1D1D2FA1FC0723D1" box="[829,980,652,677]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="43" pageNumber="44" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">950 +/- 50 m</quantity>
,
<date id="FF90D0E44C5DC67C1DC02FA1FB8523D1" box="[992,1110,652,677]" pageId="43" pageNumber="44" value="2014-01-16">16/1/2014</date>
: 12:29, D.C. Lees:
<collectionCode id="ED3F6EE14C5DC67C1B112FA0FAA523D1" box="[1329,1398,653,677]" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="43" pageNumber="44">MAD</collectionCode>
121 [pheromone voucher], IA374 [isotope voucher]; ♂, E, Maromizaha,
<geoCoordinate id="EE1A90E34C5DC67C1DB22F9EFBD623B8" box="[914,1029,688,716]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="5" value="-18.9668">18.9668o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1A312F9EFB5623BF" box="[1041,1157,688,716]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5" value="48.4547">48.4547o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1A932F99FAC223B8" box="[1203,1297,691,716]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="43" pageNumber="44" unit="km" value="1.49">1.49 km</quantity>
,
<quantity id="4CD65BC14C5DC67C1B3B2F99FF7C2386" metricMagnitude="3" metricUnit="m" metricValue="1.109" metricValueMax="1.182" metricValueMin="1.036" pageId="43" pageNumber="44" unit="m" value="1109.0" valueMax="1182.0" valueMin="1036.0">1109 +/- 73 m</quantity>
,
<date id="FF90D0E44C5DC67C1E9B2FF7FEE32386" box="[187,304,730,754]" pageId="43" pageNumber="44" value="2002-02-24">24/2/2002</date>
, D.C. Lees; ♂, CE, Antanandava: Sandrangato Forest,
<geoCoordinate id="EE1A90E34C5DC67C1DBA2FF7FBDE2386" box="[922,1037,726,754]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="5" value="-19.1083">19.1083o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1A3A2FF7FB5E2386" box="[1050,1165,726,754]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5" value="48.2417">48.2417o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1A9C2FF7FACF2387" box="[1212,1308,730,755]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="43" pageNumber="44" unit="km" value="0.44">0.44 km</quantity>
,
<quantity id="4CD65BC14C5DC67C1B072FF7FF7C226C" metricMagnitude="2" metricUnit="m" metricValue="8.8" metricValueMax="9.3" metricValueMin="8.3" pageId="43" pageNumber="44" unit="m" value="880.0" valueMax="930.0" valueMin="830.0">880 +/- 50 m</quantity>
,
<date id="FF90D0E44C5DC67C1E9A2E2DFEFE226D" box="[186,301,768,793]" pageId="43" pageNumber="44" value="1994-11-06">6/11/1994</date>
, D.C. Lees; ♂ [
<collectionCode id="ED3F6EE14C5DC67C1FF92E2CFDE4226C" box="[473,567,769,792]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="43" pageNumber="44">MNHN</collectionCode>
], CE, Anosibe, route d' km 45 [ca.
<geoCoordinate id="EE1A90E34C5DC67C1D9B2E2CFBC1226D" box="[955,1042,765,793]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="555" value="-19.13">19.13o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1A3E2E2CFBA5226C" box="[1054,1142,765,793]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="555" value="48.34">48.34o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1A842E2DFB0C226D" box="[1188,1247,768,793]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="43" pageNumber="44" unit="km" value="5.0">5 km</quantity>
,
<quantity id="4CD65BC14C5DC67C1ACA2E2CFA90226C" box="[1258,1347,769,793]" metricMagnitude="3" metricUnit="m" metricValue="1.041" pageId="43" pageNumber="44" unit="m" value="1041.0">1041 m</quantity>
],
<date id="FF90D0E44C5DC67C1B752E2DFF012249" pageId="43" pageNumber="44" value="1966-11-25">25/11/ 1966</date>
, P. Viette|DCL-DB-2907; 2 ♂♂ [
<collectionCode id="ED3F6EE14C5DC67C1C592E08FD042248" box="[633,727,805,828]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="43" pageNumber="44">MNHN</collectionCode>
], C, Pays Betsileo route du Sud, km 292; ♂ [
<collectionCode id="ED3F6EE14C5DC67C1B142E08FA412248" box="[1332,1426,805,828]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="43" pageNumber="44">MNHN</collectionCode>
],
<collectingCountry id="F339B6B44C5DC67C1EB72E66FE8D2217" box="[151,350,843,867]" name="Madagascar" pageId="43" pageNumber="44">MADAGASCAR</collectingCountry>
CENTRE,
<quantity id="4CD65BC14C5DC67C1FC12E66FDEA2216" box="[481,569,843,867]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="43" pageNumber="44" unit="m" value="1700.0">1700 m</quantity>
. [ca.
<geoCoordinate id="EE1A90E34C5DC67C1C562E67FD3A2217" box="[630,745,839,867]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="5" value="-20.7667">20.7667o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1CD62E66FCB92216" box="[758,874,839,867]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5" value="47.1833">47.1833o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1DBA2E67FC042217" box="[922,983,842,867]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="43" pageNumber="44" unit="km" value="5.0">5 km</quantity>
]
<date id="FF90D0E44C5DC67C1DC42E66FB8B2217" box="[996,1112,843,867]" pageId="43" pageNumber="44" value="1974-02-14">14/2/1974</date>
, P. Viette et A. Peyrieras; ♂, E, Amboditavy, Ankazomivady,
<geoCoordinate id="EE1A90E34C5DC67C1C3C2E5FFD4D22FE" box="[540,670,878,906]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="1" value="-20.77188">20.77188o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1C922E5CFCE722FD" box="[690,820,878,906]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="1" value="47.16522">47.16522o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1D522E5FFC6522FE" box="[882,950,881,906]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="43" pageNumber="44" unit="km" value="1.0">1 km</quantity>
,
<quantity id="4CD65BC14C5DC67C1DE82E5CFB5222FD" box="[968,1153,881,906]" metricMagnitude="3" metricUnit="m" metricValue="1.76" metricValueMax="1.81" metricValueMin="1.71" pageId="43" pageNumber="44" unit="m" value="1760.0" valueMax="1810.0" valueMin="1710.0">1760 +/- 50 m</quantity>
,
<date id="FF90D0E44C5DC67C1AB52E5FFADA22FE" box="[1173,1289,882,906]" pageId="43" pageNumber="44" value="2003-03-21">21/3/2003</date>
, D.C. Lees:
<collectionCode id="ED3F6EE14C5DC67C1EB72EB8FF3D22D9" box="[151,238,917,941]" pageId="43" pageNumber="44">BMNH</collectionCode>
(E) #697284 [
<collectionCode id="ED3F6EE14C5DC67C1FA82EB8FE1B22D9" box="[392,456,917,941]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="43" pageNumber="44">DNA</collectionCode>
voucher; cytochrome b], KA535 [=KA-P535;
<collectionCode id="ED3F6EE14C5DC67C1DFF2EB8FBCF22D9" box="[991,1052,917,941]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="43" pageNumber="44">DNA</collectionCode>
extract voucher]; ♂, C,
<quantity id="4CD65BC14C5DC67C1B142EB8FAAE22D9" box="[1332,1405,916,941]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="43" pageNumber="44" unit="km" value="32.0">32 km</quantity>
S. Ambositra,
<quantity id="4CD65BC14C5DC67C1F3C2E91FEA622A7" box="[284,373,956,980]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="43" pageNumber="44" unit="m" value="1400.0">1400 m</quantity>
. [should be 1600+],
<geoCoordinate id="EE1A90E34C5DC67C1C402E96FD0722A0" box="[608,724,952,980]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="5" value="-20.8167">20.8167o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1CC22E91FC8622A7" box="[738,853,952,980]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5" value="47.1833">47.1833o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1DA62E91FC3422A0" box="[902,999,955,980]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="43" pageNumber="44" unit="km" value="0.87">0.87 km</quantity>
,
<quantity id="4CD65BC14C5DC67C1DD32E96FB9822A7" box="[1011,1099,955,980]" metricMagnitude="3" metricUnit="m" metricValue="1.67" pageId="43" pageNumber="44" unit="m" value="1670.0">1670 m</quantity>
,
<date id="FF90D0E44C5DC67C1A782E91FB1F22A0" box="[1112,1228,956,980]" pageId="43" pageNumber="44" value="1992-01-12">12/1/1992</date>
, C. Kremen et al.: Genitalia 162; ♂, C, Ankazomivady,
<geoCoordinate id="EE1A90E34C5DC67C1C1A2ECFFD7E228E" box="[570,685,990,1019]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="5" value="-20.8167">20.8167o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1C9A2ECFFCFD228E" box="[698,814,990,1018]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5" value="47.1833">47.1833o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1D7D2ECFFC6F228F" box="[861,956,994,1019]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="43" pageNumber="44" unit="km" value="0.87">0.87 km</quantity>
,
<quantity id="4CD65BC14C5DC67C1DE82ECFFBCC228E" box="[968,1055,994,1019]" metricMagnitude="3" metricUnit="m" metricValue="1.67" pageId="43" pageNumber="44" unit="m" value="1670.0">1670 m</quantity>
,
<date id="FF90D0E44C5DC67C1A0B2ECFFB7E228E" box="[1067,1197,994,1018]" pageId="43" pageNumber="44" value="1992-10-29">29/10/1992</date>
, C. Kremen: CK816; ♂, C, Ankazomivady,
<geoCoordinate id="EE1A90E34C5DC67C1FBA2925FDD32555" box="[410,512,1029,1057]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="55" value="-20.769">20.769o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1C2F2924FDA62554" box="[527,629,1029,1057]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="55" value="47.182">47.182o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1C8A2925FCD82555" box="[682,779,1032,1057]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="43" pageNumber="44" unit="km" value="0.25">0.25 km</quantity>
,
<quantity id="4CD65BC14C5DC67C1D392925FC172554" box="[793,964,1032,1057]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="43" pageNumber="44" unit="m" value="1750.0" valueMax="1800.0" valueMin="1700.0">1750 +/- 50 m</quantity>
,
<date id="FF90D0E44C5DC67C1DF22924FB9A2555" box="[978,1097,1033,1057]" pageId="43" pageNumber="44" value="2003-03-22">22/3/2003</date>
: 10:55; WG25 [wing image]; IA117 [isotope voucher]; ♂, C, Ankazomivady,
<quantity id="4CD65BC14C5DC67C1C982902FD2C253C" box="[696,767,1071,1096]" metricMagnitude="4" metricUnit="m" metricValue="3.3" pageId="43" pageNumber="44" unit="km" value="33.0">33 km</quantity>
S Ambositra,
<geoCoordinate id="EE1A90E34C5DC67C1D832902FBDA2533" box="[931,1033,1068,1096]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="55" value="-20.769">20.769o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1A362902FBAF2533" box="[1046,1148,1068,1095]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="55" value="47.182">47.182o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1A8C2902FADF253C" box="[1196,1292,1071,1096]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="43" pageNumber="44" unit="km" value="0.25">0.25 km</quantity>
,
<quantity id="4CD65BC14C5DC67C1B372902FF7C251F" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="43" pageNumber="44" unit="m" value="1750.0" valueMax="1800.0" valueMin="1700.0">1750 +/- 50 m</quantity>
,
<date id="FF90D0E44C5DC67C1E9F297EFEE02518" box="[191,307,1107,1132]" pageId="43" pageNumber="44" value="1995-02-21">21/2/1995</date>
; D.C. Lees:
<collectionCode id="ED3F6EE14C5DC67C1FEB297EFDE32518" box="[459,560,1107,1132]" pageId="43" pageNumber="44">DLKAZ</collectionCode>
95-5A, IA185 [isotope voucher]; ♀, C, Ankazomivady,
<quantity id="4CD65BC14C5DC67C1A99297EFB292518" box="[1209,1274,1107,1132]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="43" pageNumber="44" unit="km" value="35.0">35km</quantity>
S Ambositra,
<geoCoordinate id="EE1A90E34C5DC67C1EB72954FF2E25E6" box="[151,253,1142,1170]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="55" value="-20.769">20.769o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1F292957FEBD25E5" box="[265,366,1142,1170]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="55" value="47.182">47.182o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1FBD2954FE2825E6" box="[413,507,1145,1170]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="43" pageNumber="44" unit="km" value="0.25">0.25 km</quantity>
,
<quantity id="4CD65BC14C5DC67C1C272957FD8E25E5" box="[519,605,1146,1170]" metricMagnitude="3" metricUnit="m" metricValue="1.73" pageId="43" pageNumber="44" unit="m" value="1730.0">1730 m</quantity>
,
<date id="FF90D0E44C5DC67C1C492954FD1C25E6" box="[617,719,1145,1170]" pageId="43" pageNumber="44" value="1995-03-05">5/3/1995</date>
, D.C. Lees:
<collectionCode id="ED3F6EE14C5DC67C1D7A2954FC6C25E6" box="[858,959,1145,1170]" pageId="43" pageNumber="44">DLKAZ</collectionCode>
95-26;
<specimenCount id="9D283DAD4C5DC67C1A282957FB9525E6" box="[1032,1094,1146,1170]" pageId="43" pageNumber="44" type="egg">1 egg</specimenCount>
expressed 5/3, IA186 [isotope voucher]; ♀, C, Ankazomivady,
<geoCoordinate id="EE1A90E34C5DC67C1C2F29B2FDA725CC" box="[527,628,1180,1208]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="55" value="-20.769">20.769o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1CA3298DFD3A25C3" box="[643,745,1180,1208]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="55" value="47.182">47.182o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1D3D29B2FCAD25CC" box="[797,894,1183,1208]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="43" pageNumber="44" unit="km" value="0.25">0.25 km</quantity>
,
<quantity id="4CD65BC14C5DC67C1DAC29B2FBE625C3" box="[908,1077,1183,1208]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="43" pageNumber="44" unit="m" value="1750.0" valueMax="1800.0" valueMin="1700.0">1750 +/- 50 m</quantity>
,
<date id="FF90D0E44C5DC67C1A64298DFB6B25CC" box="[1092,1208,1184,1208]" pageId="43" pageNumber="44" value="2004-12-08">8/12/2004</date>
, DL-05-497, IA202 [isotope voucher]; ♀, data as above but DL-05-505, IA203 [isotope voucher]; ♀, E, Ranomafana National Park: Sahamalaotra,
<geoCoordinate id="EE1A90E34C5DC67C1F6229C7FE662476" box="[322,437,1254,1283]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="5" value="-21.2357">21.2357o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1FE629C7FDEF2476" box="[454,572,1254,1282]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5" value="47.3977">47.3977o E</geoCoordinate>
,
<quantity id="4CD65BC14C5DC67C1C6C29C7FD282476" box="[588,763,1258,1283]" metricMagnitude="3" metricUnit="m" metricValue="1.154" metricValueMax="1.204" metricValueMin="1.104" pageId="43" pageNumber="44" unit="m" value="1154.0" valueMax="1204.0" valueMin="1104.0">1154 +/- 50 m</quantity>
,
<date id="FF90D0E44C5DC67C1D2B29C7FC5E2476" box="[779,909,1258,1282]" pageId="43" pageNumber="44" value="2004-12-20">20/12/2004</date>
, D.C. Lees: DL-05-922,
<collectionCode id="ED3F6EE14C5DC67C1A9729C7FADD2476" box="[1207,1294,1258,1282]" pageId="43" pageNumber="44">BMNH</collectionCode>
(E) #676640 [
<collectionCode id="ED3F6EE14C5DC67C1EBD283CFF0E245D" box="[157,221,1297,1321]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="43" pageNumber="44">DNA</collectionCode>
voucher]; ♀, NW, Bekolosy,
<geoCoordinate id="EE1A90E34C5DC67C1C07283DFD48245D" box="[551,667,1293,1321]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="5" value="-14.0458">14.0458o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1C87283DFCCD245C" box="[679,798,1293,1321]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5" value="48.2962">48.2962o E</geoCoordinate>
,
<quantity id="4CD65BC14C5DC67C1D08283DFCAD245C" box="[808,894,1296,1321]" metricMagnitude="3" metricUnit="m" metricValue="1.1629999999999998" pageId="43" pageNumber="44" unit="m" value="1163.0">1163 m</quantity>
,
<date id="FF90D0E44C5DC67C1DAA283CFBDF245D" box="[906,1036,1297,1321]" pageId="43" pageNumber="44" value="1994-12-22">22/12/1994</date>
, D.C. Lees:
<collectionCode id="ED3F6EE14C5DC67C1AB6283CFB2B245D" box="[1174,1272,1297,1321]" pageId="43" pageNumber="44">DLBEK</collectionCode>
94_183, IA334 [isotope voucher]; ♀, data as above but:
<quantity id="4CD65BC14C5DC67C1CB12818FD3C2438" box="[657,751,1333,1357]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="43" pageNumber="44" unit="m" value="1130.0">1130 m</quantity>
,
<date id="FF90D0E44C5DC67C1D222819FC572439" box="[770,900,1332,1357]" pageId="43" pageNumber="44" value="1994-12-15">15/12/1994</date>
, D.C. Lees:
<collectionCode id="ED3F6EE14C5DC67C1A052818FB552439" box="[1061,1158,1333,1357]" pageId="43" pageNumber="44">DLBEK</collectionCode>
94_170, IA335 [isotope voucher]; ♂, N, Tsaratanana,
<date id="FF90D0E44C5DC67C1C212875FDA62404" box="[513,629,1368,1392]" pageId="43" pageNumber="44" value="2013-10-07">7/10/2013</date>
, Ravo: DCL-14-028, IA330 [isotope voucher]; ♂, N, Tsaratanana,
<geoCoordinate id="EE1A90E34C5DC67C1EB72853FF2E24E3" box="[151,253,1403,1431]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="55" value="-14.158">14.158o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1F2E2853FE5724E3" box="[270,388,1403,1431]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5" value="48.9525">48.9525o E</geoCoordinate>
,
<quantity id="4CD65BC14C5DC67C1FB52852FE2224E3" box="[405,497,1407,1431]" metricMagnitude="3" metricUnit="m" metricValue="1.932" pageId="43" pageNumber="44" unit="m" value="1932.0">1932 m</quantity>
,
<date id="FF90D0E44C5DC67C1C212852FD5024E3" box="[513,643,1407,1431]" pageId="43" pageNumber="44" value="2004-12-22">22/12/2004</date>
, D.C. Lees:
<collectionCode id="ED3F6EE14C5DC67C1D3E2852FCA624E3" box="[798,885,1407,1431]" pageId="43" pageNumber="44">BMNH</collectionCode>
(E) #671601; IA331 [isotope voucher]; ♀, NE, Anjanaharibe Sud,
<quantity id="4CD65BC14C5DC67C1F4D288FFE1024CE" box="[365,451,1442,1467]" metricMagnitude="3" metricUnit="m" metricValue="1.26" pageId="43" pageNumber="44" unit="m" value="1260.0">1260 m</quantity>
,
<date id="FF90D0E44C5DC67C1FEE288FFDE624CF" box="[462,565,1442,1467]" pageId="43" pageNumber="44" value="1996-02-04">4/2/1996</date>
, C. Kremen: CK703, IA135 [isotope voucher]; ♂, NE, Anjanaharibe Sud,
<date id="FF90D0E44C5DC67C1B51288EFF0124AB" pageId="43" pageNumber="44" value="1996-02-03">3/2/ 1996</date>
. H. Raharitsimba:
<collectingCountry id="F339B6B44C5DC67C1F8628EBFE1F24AB" box="[422,460,1478,1503]" name="Chile" pageId="43" pageNumber="44">TR</collectingCountry>
511, IA171 [isotope voucher]; ♀, NE, Anjanaharibe Sud,
<date id="FF90D0E44C5DC67C1A7728EBFB6D24AB" box="[1111,1214,1478,1503]" pageId="43" pageNumber="44" value="1996-02-04">4/2/1996</date>
: 09:15, C. Kremen: CK666,
<specimenCount id="9D283DAD4C5DC67C1EDA28C0FEED2771" box="[250,318,1517,1541]" pageId="43" pageNumber="44" type="egg">1 egg</specimenCount>
, IA172 [isotope voucher]; ♂, NE, Anjanaharibe Sud,
<geoCoordinate id="EE1A90E34C5DC67C1D9928C0FBFF2771" box="[953,1068,1513,1541]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="5" value="-14.7482">14.7482o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1A1D28C1FB602770" box="[1085,1203,1513,1541]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5" value="49.4659">49.4659o E</geoCoordinate>
, 1320,
<date id="FF90D0E44C5DC67C1B2D28C1FAA02771" box="[1293,1395,1516,1541]" pageId="43" pageNumber="44" value="1996-02-03">3/2/1996</date>
, C. Kremen: CK662;
<collectionCode id="ED3F6EE14C5DC67C1F4D2B3EFE402758" box="[365,403,1555,1580]" pageId="43" pageNumber="44">DL</collectionCode>
9667 [
<collectionCode id="ED3F6EE14C5DC67C1FFE2B39FDCD2758" box="[478,542,1556,1580]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="43" pageNumber="44">DNA</collectionCode>
voucher], IA332 [isotope voucher]; ♂, NE, Anjanaharibe Sud,
<geoCoordinate id="EE1A90E34C5DC67C1B032B3EFA452758" box="[1315,1430,1552,1580]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="5" value="-14.7524">14.7524o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1EB72B17FEDF2725" box="[151,268,1590,1618]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5" value="49.4777">49.4777o E</geoCoordinate>
,
<quantity id="4CD65BC14C5DC67C1F382B17FEBD2725" box="[280,366,1594,1618]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="43" pageNumber="44" unit="m" value="1130.0">1130 m</quantity>
,
<date id="FF90D0E44C5DC67C1F5A2B14FE332726" box="[378,480,1593,1618]" pageId="43" pageNumber="44" value="1996-02-04">4/2/1996</date>
, C. Kremen: CK689, IA333 [isotope voucher]; ♂, NE, Anjanaharibe Sud,
<geoCoordinate id="EE1A90E34C5DC67C1B032B14FA452726" box="[1315,1430,1590,1618]" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="5" value="-14.7454">14.7454o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C5DC67C1EB72B4DFED9270C" box="[151,266,1629,1657]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5" value="49.4632">49.4632o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C5DC67C1F632B4CFE4B270D" box="[323,408,1632,1657]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="43" pageNumber="44" unit="km" value="0.4">0.4 km</quantity>
,
<quantity id="4CD65BC14C5DC67C1F882B4CFD84270C" box="[424,599,1632,1657]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.45" metricValueMin="1.35" pageId="43" pageNumber="44" unit="m" value="1400.0" valueMax="1450.0" valueMin="1350.0">1400 +/- 50 m</quantity>
, C. Kremen: CK732,
<collectionCode id="ED3F6EE14C5DC67C1D7E2B4CFC66270D" box="[862,949,1633,1657]" pageId="43" pageNumber="44">BMNH</collectionCode>
(E) 697885, 471 [=
<collectionCode id="ED3F6EE14C5DC67C1AB42B4CFB6E270D" box="[1172,1213,1633,1657]" pageId="43" pageNumber="44">DL</collectionCode>
0471,
<collectionCode id="ED3F6EE14C5DC67C1B272B4CFA97270D" box="[1287,1348,1633,1657]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="43" pageNumber="44">DNA</collectionCode>
extract number], KA581 [=KA-P581,
<collectionCode id="ED3F6EE14C5DC67C1FCF2BA9FDFF27E8" box="[495,556,1668,1692]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="43" pageNumber="44">DNA</collectionCode>
extract number].
</paragraph>
</subSubSection>
</treatment>
</document>