559 lines
66 KiB
XML
559 lines
66 KiB
XML
<document id="CE340276416CCACC51FCB07AC254BD43" ID-CLB-Dataset="38930" ID-DOI="10.11646/zootaxa.4118.1.1" ID-GBIF-Dataset="a4e408c4-2575-4b5c-878b-e1b9010ec619" ID-ISSN="1175-5326" ID-Zenodo-Dep="264597" ID-ZooBank="CFA586DA-10EE-468B-80EE-35351E3845FD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="guilherme" checkinTime="1465242077321" checkinUser="plazi" docAuthor="C, Lees David" docDate="2016" docId="038747324C54C6711EB72966FF2221A3" docLanguage="en" docName="zootaxa.4118.pdf.imf" docOrigin="Zootaxa 4118 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Heteropsis westwoodi Lees, sp. nov." docType="treatment" docVersion="12" lastPageNumber="38" masterDocId="FFBE3F4A4C76C6571E202D2DFFD32174" masterDocTitle="Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision" masterLastPageNumber="97" masterPageNumber="1" pageNumber="35" updateTime="1735917000942" updateUser="ExternalLinkService">
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<mods:mods id="62A7E1B685A6D93F8766EBFE1925759F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="EAEBF6D266007FAE17440A05DFD49802">
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<mods:title id="DA6D0D4ACE85DB0D22D6FF68BEEB2334">Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision</mods:title>
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</mods:titleInfo>
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<mods:name id="EAE823E7831F32302E606C1DB479F5A5" type="personal">
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<mods:roleTerm id="D275C6A0C658900886347B787719A0FF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="694ECA3867510EA734800A1D383F5BCB">C, Lees David</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="8D6749D642176AE3A5612C6A9D4AA965">text</mods:typeOfResource>
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<mods:titleInfo id="B084D4B16F848E34C0D02CB9F5C1F50E">
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<mods:title id="AF3ADB188B310398AECFE48EA656D9D6">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="EA5FC1CDE28531E1F87282FE18F5437C">
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<mods:date id="91DA4BE5285521F0B9A804BEB627EE72">2016</mods:date>
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<mods:detail id="993071195475A8BF8544FFF67694D4F4" type="volume">
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<mods:number id="60062E5466A421F0C4050A69057F9440">4118</mods:number>
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</mods:detail>
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<mods:detail id="CEB78964F66EE7DCB197E6C600ED240E" type="issue">
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<mods:number id="66CB12A5E40219F78048668EA99E64CB">1</mods:number>
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<mods:extent id="4B1DFEA83C45C3AFB49D7136B7355E53" unit="page">
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<mods:start id="10046931A897DF191A034C746F35C618">1</mods:start>
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<mods:end id="37DB71FB91200007CA3B14DA8FB188F9">97</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="9CA2AB236E95AD62A14E513318DDA246">journal article</mods:classification>
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<mods:identifier id="ABA8C03EA6B98FDE201CAE3A82C77EA5" type="CLB-Dataset">38930</mods:identifier>
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<mods:identifier id="44795459B52DA66B2FD93BB176FF86C8" type="DOI">10.11646/zootaxa.4118.1.1</mods:identifier>
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<mods:identifier id="4069D47F5039A1F47109F6E3B7EF334F" type="GBIF-Dataset">a4e408c4-2575-4b5c-878b-e1b9010ec619</mods:identifier>
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<mods:identifier id="24946380AFA26349C0CB0A117A163824" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="441F130B19C5574D6788EB6E158BEB32" type="Zenodo-Dep">264597</mods:identifier>
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<mods:identifier id="A5085564617F16C8F45B7191A0570755" type="ZooBank">CFA586DA-10EE-468B-80EE-35351E3845FD</mods:identifier>
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</mods:mods>
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<treatment id="038747324C54C6711EB72966FF2221A3" ID-DOI="http://doi.org/10.5281/zenodo.6086422" ID-GBIF-Taxon="120817122" ID-Zenodo-Dep="6086422" LSID="urn:lsid:plazi:treatment:038747324C54C6711EB72966FF2221A3" httpUri="http://treatment.plazi.org/id/038747324C54C6711EB72966FF2221A3" lastPageId="38" lastPageNumber="38" pageId="34" pageNumber="35" scope_family="Nymphalidae" scope_subFamily="Satyrinae" scope_tribe="Satyrini">
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<subSubSection id="C334A5AF4C54C6751EB72966FBC425F2" pageId="34" pageNumber="35" type="nomenclature">
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<paragraph id="8B91F6244C54C6751EB72966FD942511" blockId="34.[151,1047,1099,1158]" box="[151,583,1099,1125]" pageId="34" pageNumber="35">
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<heading id="D0D941484C54C6751EB72966FD942511" bold="true" box="[151,583,1099,1125]" fontSize="11" level="1" pageId="34" pageNumber="35" reason="1">
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<emphasis id="B95A2A364C54C6751EB72966FD942511" bold="true" box="[151,583,1099,1125]" pageId="34" pageNumber="35">
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<taxonomicName id="4C2E8DA74C54C6751EB72966FE0E2511" authority="Lees" authorityName="Lees" box="[151,477,1099,1125]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="westwoodi" status="sp. nov.">
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<emphasis id="B95A2A364C54C6751EB72966FE482511" bold="true" box="[151,411,1099,1125]" italics="true" pageId="34" pageNumber="35">Heteropsis westwoodi</emphasis>
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Lees
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</taxonomicName>
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,
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<taxonomicNameLabel id="A269974D4C54C6751FC92966FD942511" box="[489,583,1099,1125]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8B91F6244C54C6751EB72943FBC425F2" blockId="34.[151,1047,1099,1158]" box="[151,1047,1133,1158]" pageId="34" pageNumber="35">LSID: urn:lsid:zoobank.org:act:422E5526-8FBB-4AD6-A47D-EC95FF54E16F</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C334A5AF4C54C6751EB72998FF662586" pageId="34" pageNumber="35" type="description">
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||
<paragraph id="8B91F6244C54C6751EB72998FF662586" blockId="34.[151,1437,1205,2022]" pageId="34" pageNumber="35">
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<emphasis id="B95A2A364C54C6751EB72998FE8425BA" bold="true" box="[151,343,1205,1230]" pageId="34" pageNumber="35">Prior references</emphasis>
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: sp. 13B (
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<bibRefCitation id="EFBF8BD54C54C6751FEA299BFDA725BA" author="Lees" box="[458,628,1205,1230]" pageId="34" pageNumber="35" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997: 64</bibRefCitation>
|
||
). “KA522_Heteopsis_
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<taxonomicName id="4C2E8DA74C54C6751D502998FBD325BA" box="[880,1024,1205,1230]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="andravahana">andravahana</taxonomicName>
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” [
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<emphasis id="B95A2A364C54C6751A3A299AFBEA25BA" box="[1050,1081,1207,1230]" italics="true" pageId="34" pageNumber="35">sic</emphasis>
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] (Aduse-Poku
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<emphasis id="B95A2A364C54C6751AC4299AFAF125BA" box="[1252,1314,1205,1230]" italics="true" pageId="34" pageNumber="35">et al.,</emphasis>
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2015:
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<figureCitation id="1315EAA14C54C6751B502998FF772586" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." figureDoi="http://doi.org/10.5281/zenodo.264598" httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="34" pageNumber="35">Fig. 1</figureCitation>
|
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).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C334A5AF4C54C6751EE729D0FC8A249B" pageId="34" pageNumber="35" type="materials_examined">
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||
<paragraph id="8B91F6244C54C6751EE729D0FE9A24F0" blockId="34.[151,1437,1205,2022]" pageId="34" pageNumber="35">
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||
<emphasis id="B95A2A364C54C6751EE729D0FD0C2462" bold="true" box="[199,735,1277,1302]" pageId="34" pageNumber="35">
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<typeStatus id="549548864C54C6751EE729D0FED12462" box="[199,258,1277,1302]" pageId="34" pageNumber="35">Type</typeStatus>
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||
material., Deposition
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<collectionCode id="ED3F6EE14C54C6751C2A29D0FDB62462" box="[522,613,1277,1302]" pageId="34" pageNumber="35">BMNH</collectionCode>
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||
:
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<typeStatus id="549548864C54C6751C5229D0FD0C2462" box="[626,735,1277,1302]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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: ♂ (
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<figureCitation id="1315EAA14C54C6751D3229D0FCB02462" box="[786,867,1277,1302]" captionStart="FIGURE 10" captionStartId="36.[151,250,1737,1759]" captionTargetBox="[272,1316,212,1704]" captionTargetId="figure@36.[258,1329,193,1715]" captionTargetPageId="36" captionText="FIGURE 10. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis westwoodi Lees, sp. nov. (Vohitrasiva, Andringitra; NHMUK 010289130). B: referred ♀ of Ht. westwoodi (Andohahela, Antananandava, Andohahela; BMNH (E) # 674842). C: HT ♂ of Ht. pauliani Lees, sp. nov. (Tsaratanana; DCL-DB- 2966; MNHN). D: PT ♀ of Ht. pauliani (Tsaratanana; DCL-DB- 2975; MNHN). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264607" httpUri="https://zenodo.org/record/264607/files/figure.png" pageId="34" pageNumber="35">Fig. 10</figureCitation>
|
||
A),
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||
<collectingCountry id="F339B6B44C54C6751DAE29D0FBC62462" box="[910,1045,1277,1302]" name="Madagascar" pageId="34" pageNumber="35">Madagascar</collectingCountry>
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||
SE, Vohitrasiva, near ridge, "
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||
<quantity id="4CD65BC14C54C6751B4129D3FF622448" metricMagnitude="3" metricUnit="m" metricValue="1.39" pageId="34" pageNumber="35" unit="m" value="1390.0">1390 m</quantity>
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||
", Andringitra, [mountaintop on way to Vohipia (
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||
<quantity id="4CD65BC14C54C6751CED2809FCF02448" box="[717,803,1316,1341]" metricMagnitude="3" metricUnit="m" metricValue="1.76" pageId="34" pageNumber="35" unit="m" value="1760.0">1760 m</quantity>
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||
pass)],
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||
<geoCoordinate id="EE1A90E34C54C6751D5A2809FC3E2448" box="[890,1005,1312,1341]" direction="south" orientation="latitude" pageId="34" pageNumber="35" precision="5" value="-22.1684">22.1684o S</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE1A90E34C54C6751DDA2809FBBE2448" box="[1018,1133,1312,1340]" direction="east" orientation="longitude" pageId="34" pageNumber="35" precision="5" value="47.0289">47.0289o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C54C6751ABB2809FB382449" box="[1179,1259,1316,1341]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="34" pageNumber="35" unit="km" value="0.1">0.1 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C54C6751AD62809FA442448" box="[1270,1431,1316,1341]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="34" pageNumber="35" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">1450 +/- 50 m</quantity>
|
||
,
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||
<date id="FF90D0E44C54C6751EB7286AFED82414" box="[151,267,1351,1376]" pageId="34" pageNumber="35" value="1995-02-17">17/2/1995</date>
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||
, 11:40–13:15, D.C. Lees:
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||
<collectionCode id="ED3F6EE14C54C6751C16286AFD8D2414" box="[566,606,1351,1376]" pageId="34" pageNumber="35">DL</collectionCode>
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||
95-0002; KAP20 [=KA-P20;
|
||
<collectionCode id="ED3F6EE14C54C6751D8C2865FC3A2414" box="[940,1001,1352,1376]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="34" pageNumber="35">DNA</collectionCode>
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||
extract number],
|
||
<collectionCode id="ED3F6EE14C54C6751A942865FACD2414" box="[1204,1310,1352,1376]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="34" pageNumber="35">NHMUK</collectionCode>
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010289130 [
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||
<collectingCountry id="F339B6B44C54C6751EBD2841FF0B24F0" box="[157,216,1388,1412]" name="Qatar" pageId="34" pageNumber="35">QTR</collectingCountry>
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||
barcode].
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||
</paragraph>
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||
<paragraph id="8B91F6244C54C6751EE728A2FC8A249B" blockId="34.[151,1437,1205,2022]" pageId="34" pageNumber="35">
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||
<emphasis id="B95A2A364C54C6751EE728A2FDFF24DC" bold="true" box="[199,556,1423,1448]" pageId="34" pageNumber="35">
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||
<typeStatus id="549548864C54C6751EE728A2FE9024DC" box="[199,323,1423,1448]" pageId="34" pageNumber="35" type="paratype">Paratypes</typeStatus>
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: Deposition
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<collectionCode id="ED3F6EE14C54C6751FF528A2FDFF24DC" box="[469,556,1423,1448]" pageId="34" pageNumber="35">BMNH</collectionCode>
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||
</emphasis>
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: ♂, data as HT but
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<collectionCode id="ED3F6EE14C54C6751D2628A2FCFD24DC" box="[774,814,1423,1448]" pageId="34" pageNumber="35">DL</collectionCode>
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||
95-0001, IA288 [isotope voucher], KA751 [=KA-P751;
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<collectionCode id="ED3F6EE14C54C6751EB7289EFF0724BF" box="[151,212,1459,1483]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="34" pageNumber="35">DNA</collectionCode>
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||
extract number],
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||
<collectionCode id="ED3F6EE14C54C6751F8C289EFDC424BF" box="[428,535,1459,1483]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="34" pageNumber="35">NHMUK</collectionCode>
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||
010289131 [
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||
<collectingCountry id="F339B6B44C54C6751C83289EFD0D24BF" box="[675,734,1459,1483]" name="Qatar" pageId="34" pageNumber="35">QTR</collectingCountry>
|
||
barcode]; ♂, data as HT but
|
||
<collectionCode id="ED3F6EE14C54C6751A16289FFB8D24BF" box="[1078,1118,1458,1483]" pageId="34" pageNumber="35">DL</collectionCode>
|
||
95-0003, 236
|
||
<collectionCode id="ED3F6EE14C54C6751AD3289FFACA24BF" box="[1267,1305,1458,1483]" pageId="34" pageNumber="35">DL</collectionCode>
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||
[genitalia], IA290 [isotope voucher],
|
||
<collectionCode id="ED3F6EE14C54C6751F9828FAFDF1249B" box="[440,546,1495,1519]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="34" pageNumber="35">NHMUK</collectionCode>
|
||
010289182 [
|
||
<collectingCountry id="F339B6B44C54C6751C8D28FAFD3B249B" box="[685,744,1495,1519]" name="Qatar" pageId="34" pageNumber="35">QTR</collectingCountry>
|
||
barcode].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C54C6751EE728D7FC0F270A" pageId="34" pageNumber="35" type="description">
|
||
<paragraph id="8B91F6244C54C6751EE728D7FEB6272E" blockId="34.[151,1437,1205,2022]" pageId="34" pageNumber="35">
|
||
<emphasis id="B95A2A364C54C6751EE728D7FE662767" bold="true" box="[199,437,1530,1555]" pageId="34" pageNumber="35">Deposition MNHN:</emphasis>
|
||
♂,
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||
<collectingCountry id="F339B6B44C54C6751FCA28D7FDA02767" box="[490,627,1530,1555]" name="Madagascar" pageId="34" pageNumber="35">Madagascar</collectingCountry>
|
||
, forêt d’Anjavidilava, Andringitra Oriental,
|
||
<quantity id="4CD65BC14C54C6751AA028D7FB0E2766" box="[1152,1245,1530,1555]" metricMagnitude="3" metricUnit="m" metricValue="2.005" pageId="34" pageNumber="35" unit="m" value="2005.0">2005 m</quantity>
|
||
, 1/
|
||
<date id="FF90D0E44C54C6751B2228D7FAA42767" box="[1282,1399,1530,1555]" pageId="34" pageNumber="35" value="1971-01-15">15-I-1971</date>
|
||
, P. Griveaud et P. Soga|DCL-DB-2923; six ♂♂, data as above but: representing the rest of series from DCL-DB-2919 to DCL-DB-2925.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C54C6751EE72B48FC0F270A" blockId="34.[151,1437,1205,2022]" box="[199,988,1637,1662]" pageId="34" pageNumber="35">
|
||
<emphasis id="B95A2A364C54C6751EE72B48FE6D270A" bold="true" box="[199,446,1637,1662]" pageId="34" pageNumber="35">Deposition summary</emphasis>
|
||
: BMNH (HT ♂, 2 PT ♂♂), MNHN (7 PT ♂♂).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C54C6751EE72BA1FA8F27D1" box="[199,1372,1672,1701]" pageId="34" pageNumber="35" type="materials_examined">
|
||
<paragraph id="8B91F6244C54C6751EE72BA1FA8F27D1" blockId="34.[151,1437,1205,2022]" box="[199,1372,1672,1701]" pageId="34" pageNumber="35">
|
||
<emphasis id="B95A2A364C54C6751EE72BA1FEB627D1" bold="true" box="[199,357,1676,1701]" pageId="34" pageNumber="35">
|
||
<typeStatus id="549548864C54C6751EE72BA1FED127D1" box="[199,258,1676,1701]" pageId="34" pageNumber="35">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="F339B6B44C54C6751F4C2BA1FE2027D1" box="[364,499,1676,1701]" name="Madagascar" pageId="34" pageNumber="35">Madagascar</collectingCountry>
|
||
E, Vohitrasiva, Andringitra,
|
||
<geoCoordinate id="EE1A90E34C54C6751D172BA1FC7827D0" box="[823,939,1672,1701]" direction="south" orientation="latitude" pageId="34" pageNumber="35" precision="5" value="-22.1684">22.1684o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C54C6751D972BA1FBF827D0" box="[951,1067,1672,1700]" direction="east" orientation="longitude" pageId="34" pageNumber="35" precision="5" value="47.0289">47.0289o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C54C6751A7A2BA1FB7827D1" box="[1114,1195,1676,1701]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="34" pageNumber="35" unit="km" value="0.1">0.1 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C54C6751A962BA1FA8727D0" box="[1206,1364,1676,1701]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="34" pageNumber="35" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">
|
||
1450 +/-
|
||
<specimenCount id="9D283DAD4C54C6751B3D2BA1FA8727D0" box="[1309,1364,1676,1701]" pageId="34" pageNumber="35" type="generic">50 m</specimenCount>
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C54C6751EE72B82FE2D2640" pageId="34" pageNumber="35" type="diagnosis">
|
||
<paragraph id="8B91F6244C54C6751EE72B82FE2D2640" blockId="34.[151,1437,1205,2022]" pageId="34" pageNumber="35">
|
||
<emphasis id="B95A2A364C54C6751EE72B82FE9227BC" bold="true" box="[199,321,1711,1736]" pageId="34" pageNumber="35">Diagnosis.</emphasis>
|
||
There is only one particularly similar species to
|
||
<taxonomicName id="4C2E8DA74C54C6751D452B9CFBD527BC" box="[869,1030,1711,1736]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="westwoodi">
|
||
<emphasis id="B95A2A364C54C6751D452B9CFBD527BC" box="[869,1030,1711,1736]" italics="true" pageId="34" pageNumber="35">Ht. westwoodi</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<taxonomicName id="4C2E8DA74C54C6751A352B9CFB5727BC" box="[1045,1156,1713,1736]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="viettei">
|
||
<emphasis id="B95A2A364C54C6751A352B9CFB5727BC" box="[1045,1156,1713,1736]" italics="true" pageId="34" pageNumber="35">Ht. viettei</emphasis>
|
||
</taxonomicName>
|
||
has a similar white band distad of the HWV Mb, but is generally smaller and much more widely distributed (sympatric in Fianarantsoa province) in
|
||
<collectingCountry id="F339B6B44C54C6751F0C2BDAFE652664" box="[300,438,1783,1808]" name="Madagascar" pageId="34" pageNumber="35">Madagascar</collectingCountry>
|
||
. Among related species,
|
||
<taxonomicName id="4C2E8DA74C54C6751CFA2BD5FC882664" box="[730,859,1783,1808]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B95A2A364C54C6751CFA2BD5FC882664" box="[730,859,1783,1808]" italics="true" pageId="34" pageNumber="35">Ht. harveyi</emphasis>
|
||
</taxonomicName>
|
||
♂♂ have yellow rather than whitish highlighting distad of the Mb in some forms.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C54C6741EE72A13FC98248A" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="description">
|
||
<paragraph id="8B91F6244C54C6741EE72A13FCCA2264" blockId="34.[151,1437,1205,2022]" lastBlockId="35.[151,1437,151,1999]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
|
||
<emphasis id="B95A2A364C54C6751EE72A13FE782623" bold="true" box="[199,427,1854,1879]" pageId="34" pageNumber="35">Description. Wings</emphasis>
|
||
: dorsal surface fairly uniform dark brown, darker in area of cell and above towards costa. FWD space-CuA1 ocellus expressed, roughly circular, its black iris spanning inter-vein distance CuA1-CuA2, with a narrow concentric dark orange ring. HWD space-cuA1 ocellus elliptic, with a black iris spanning more than half intervein distance CuA1-CuA2, and a dark orange ring relatively slightly broader than in the FWD and nearly spanning the intervein distance. No other ocelli expressed on FWD. A dark Sml quite closely hugs the margin in both wings and both surfaces. HW margin moderately and evenly crenulate, with rather blunt dark blackish brown tails on the otherwise violet-grey-brown fringe. Ventral surface with a light grey-violet cast, especially on the HW. FWV ocellus space-CuA1 is circular and same size as on the FWD with a concentric quite narrow orange ring. M1 ocellus FWV fairly small and circular with black iris and faint narrow orange ring. A small ocellus with very small black iris is expressed just below the M1 ocellus FWV. In the HWV the space-CuA1 ocellus is as on the HWD and space-R5-M3 ocelli are expressed as very small white points. Mb irregular, strongly concave from the margin to M2 where it forms a cusp, then strongly and irregularly indented and maximally concave in space-CuA1, convex again in space-CuA2 and concave again to vein 1A+2A where it terminates. The Mb HWV delineates a relatively darker brown irrorated area proximad to the PMb that only slightly repeats the shape of the Mb. Distad of the Mb from veins M2-1A is a clear white band, also irregular, with prominence in space M2 and maximum width proximad of space-CuA1 ocellus, tapering to 1A+2A with a slight expansion in Space-CuA2. Distad of this white band is a slightly paler brown area irrorated less densely with darker brown than the band in the more basal area. Patterning on the FWV is fairly uniform, darker proximad of the Mb that is concave around the space-CuA1 ocellus and recurves back towards mid-costa, with indications of three or four paler stripes above the cell. There is slightly paler brown highlighting in premarginal area just proximad of the dark Sml.
|
||
<emphasis id="B95A2A364C55C6741A442FA1FB0823D1" bold="true" box="[1124,1243,652,677]" pageId="35" pageNumber="36">Variation.</emphasis>
|
||
A PT ♂ (DL95- 0001) has a much stronger violet-grey cast, especially to the HWV. Sexually dimorphic (
|
||
<figureCitation id="1315EAA14C55C6741A542F82FB1623BC" box="[1140,1221,687,712]" captionStart="FIGURE 10" captionStartId="36.[151,250,1737,1759]" captionTargetBox="[272,1316,212,1704]" captionTargetId="figure@36.[258,1329,193,1715]" captionTargetPageId="36" captionText="FIGURE 10. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis westwoodi Lees, sp. nov. (Vohitrasiva, Andringitra; NHMUK 010289130). B: referred ♀ of Ht. westwoodi (Andohahela, Antananandava, Andohahela; BMNH (E) # 674842). C: HT ♂ of Ht. pauliani Lees, sp. nov. (Tsaratanana; DCL-DB- 2966; MNHN). D: PT ♀ of Ht. pauliani (Tsaratanana; DCL-DB- 2975; MNHN). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264607" httpUri="https://zenodo.org/record/264607/files/figure.png" pageId="35" pageNumber="36">Fig. 10</figureCitation>
|
||
A–B), ♀ larger and lighter in referred specimen BMNH(E) #674842 (
|
||
<figureCitation id="1315EAA14C55C6741CE32FFEFCC52398" box="[707,790,723,748]" captionStart="FIGURE 10" captionStartId="36.[151,250,1737,1759]" captionTargetBox="[272,1316,212,1704]" captionTargetId="figure@36.[258,1329,193,1715]" captionTargetPageId="36" captionText="FIGURE 10. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis westwoodi Lees, sp. nov. (Vohitrasiva, Andringitra; NHMUK 010289130). B: referred ♀ of Ht. westwoodi (Andohahela, Antananandava, Andohahela; BMNH (E) # 674842). C: HT ♂ of Ht. pauliani Lees, sp. nov. (Tsaratanana; DCL-DB- 2966; MNHN). D: PT ♀ of Ht. pauliani (Tsaratanana; DCL-DB- 2975; MNHN). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264607" httpUri="https://zenodo.org/record/264607/files/figure.png" pageId="35" pageNumber="36">Fig. 10</figureCitation>
|
||
B), but similar in ventral patterning, although with a more contrasting darker band between the HWV PMb and Mb.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C55C6741EE72E36FC0F2223" blockId="35.[151,1437,151,1999]" pageId="35" pageNumber="36">
|
||
<emphasis id="B95A2A364C55C6741EE72E36FEBF2240" bold="true" box="[199,364,795,820]" pageId="35" pageNumber="36">Wingspan/fwl</emphasis>
|
||
: range 37.5–40.3/
|
||
<quantity id="4CD65BC14C55C6741C1C2E36FD0C2247" box="[572,735,795,820]" metricMagnitude="-2" metricUnit="m" metricValue="2.02" metricValueMax="2.07" metricValueMin="1.97" pageId="35" pageNumber="36" unit="mm" value="20.2" valueMax="20.7" valueMin="19.7">19.7–20.7 mm</quantity>
|
||
(n=3
|
||
<typeStatus id="549548864C55C6741D042E31FC872240" box="[804,852,796,820]" pageId="35" pageNumber="36">type</typeStatus>
|
||
♂♂); mean = 37.4 +/- 1.1 SD/20.7 +/- 0.3 SD mm (n=3 ♂♂), including HT ♂ 38.6/
|
||
<quantity id="4CD65BC14C55C6741C272E13FDBE2223" box="[519,621,830,855]" metricMagnitude="-2" metricUnit="m" metricValue="2.07" pageId="35" pageNumber="36" unit="mm" value="20.7">20.7 mm</quantity>
|
||
. 44.1/
|
||
<quantity id="4CD65BC14C55C6741C902E12FCC72223" box="[688,788,831,855]" metricMagnitude="-2" metricUnit="m" metricValue="2.28" pageId="35" pageNumber="36" unit="mm" value="22.8">22.8 mm</quantity>
|
||
(n=1 referred ♀).
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C55C6741EE72E4FFED322EB" blockId="35.[151,1437,151,1999]" pageId="35" pageNumber="36">
|
||
<emphasis id="B95A2A364C55C6741EE72E4FFE87220F" bold="true" box="[199,340,866,891]" pageId="35" pageNumber="36">Androconia</emphasis>
|
||
: HWD discocellular brush light brown. HWD discocellular patch ‘bullet’-shaped, peach pink coloured.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C55C6741EE72E87FC2A2293" blockId="35.[151,1437,151,1999]" pageId="35" pageNumber="36">
|
||
<emphasis id="B95A2A364C55C6741EE72E87FEDB22B7" bold="true" box="[199,264,938,963]" pageId="35" pageNumber="36">Palps</emphasis>
|
||
: outside face ochreous with distinct dark brown medial strip, fringed with thin blackish scales except where palps brush eye, and sandwiching ochreous hair scales away from eye.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C55C6741EE72EDCFC98248A" blockId="35.[151,1437,151,1999]" pageId="35" pageNumber="36">
|
||
<emphasis id="B95A2A364C55C6741EE72EDCFE99257E" bold="true" box="[199,330,1009,1034]" pageId="35" pageNumber="36">♂ genitalia</emphasis>
|
||
: 236DL, PT ♂ from Vohitraseva (
|
||
<bibRefCitation id="EFBF8BD54C55C6741CEC2EDFFB02257E" author="Lees" box="[716,1233,1009,1034]" pageId="35" pageNumber="36" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997" yearSuffix="i">
|
||
Lees, 1997: 109,
|
||
<figureCitation id="1315EAA14C55C6741DB02EDCFC00257E" box="[912,979,1009,1034]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." figureDoi="http://doi.org/10.5281/zenodo.264604" httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="35" pageNumber="36">
|
||
Fig.
|
||
<date id="FF90D0E44C55C6741DE42EDCFC00257E" box="[964,979,1009,1034]" pageId="35" pageNumber="36">7</date>
|
||
</figureCitation>
|
||
i, “13B”; see
|
||
<figureCitation id="1315EAA14C55C6741A4A2EDCFB6A257E" box="[1130,1209,1009,1034]" captionStart="FIGURE 11" captionStartId="37.[151,250,1177,1199]" captionTargetBox="[264,1363,233,1142]" captionTargetId="figure@37.[212,1379,190,1160]" captionTargetPageId="37" captionText="FIGURE 11. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis westwoodi Lees, sp. nov. (Antanandava, Andohahela; 236 DL). B: Ht. pauliani, ♂ genitalia, PT ♂ (Tsaratanana; 97 DL)." figureDoi="http://doi.org/10.5281/zenodo.264608" httpUri="https://zenodo.org/record/264608/files/figure.png" pageId="35" pageNumber="36">Fig. 11</figureCitation>
|
||
A
|
||
</bibRefCitation>
|
||
). From LV, a very shallowly domed tegumen with a rather straight dorsal edge profile and slight brow leads to a ‘hand-scythe’-shaped uncus, which is fairly evenly deep except towards the tip of the hook, where it features more of a ‘head’ dorsad; very thin from DV. Tegumen with wide ‘U’-shaped proximad notch, from LV proximad profile bending round sharply to a constriction at junction with vinculum. This ‘waist’ is over half the maximum dimension of the tegumen (and thus the tegumen does not have a quadrate shape from LV). Valve with a very prominent convex dorsal ‘shoulder’ making up part of an oblong shape which leads to the narrow quite tapering and fairly straight valve arms, which are ventrally slightly elbowed and with a very slightly expanded club extending beyond the uncus tip, with a dorsal tooth and with a slightly incurved, rather blunt and serrate distal region (pointed and somewhat incurved from DV); this club covered in an array of spinoid setae on its mesad face. Gnathos from rightangled base pointing in line with the uncus tip in its porrect position, relatively straight and tapered to pointed tip and with negligible serration; from DV slightly sinuate and inrecurved at tip. Saccus moderately extended and markedly inflated proximad, less than a third of total valve length. Aedeagus very thin, dorsoventrally flattened and uprecurved in distad half, with a long ostium proximad, proximad tip rather semicircular from DV. Juxta quite prominent proximad and keel-like from LV, bilobed from DV.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C55C6741EE72B24FD45271E" pageId="35" pageNumber="36" type="etymology">
|
||
<paragraph id="8B91F6244C55C6741EE72B24FD45271E" blockId="35.[151,1437,151,1999]" pageId="35" pageNumber="36">
|
||
<emphasis id="B95A2A364C55C6741EE72B24FE9D2756" bold="true" box="[199,334,1545,1570]" pageId="35" pageNumber="36">Etymology.</emphasis>
|
||
After John Obadiah Westwood, founder of the genus
|
||
<taxonomicName id="4C2E8DA74C55C6741D942B27FBFF2755" box="[948,1068,1546,1569]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A2A364C55C6741D942B27FBFF2755" box="[948,1068,1546,1569]" italics="true" pageId="35" pageNumber="36">Heteropsis</emphasis>
|
||
</taxonomicName>
|
||
and describer of the remarkable
|
||
<taxonomicName id="4C2E8DA74C55C6741EB72B03FEA02732" box="[151,371,1581,1606]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="drepana">
|
||
<emphasis id="B95A2A364C55C6741EB72B03FEA02732" box="[151,371,1581,1606]" italics="true" pageId="35" pageNumber="36">Heteropsis drepana</emphasis>
|
||
</taxonomicName>
|
||
. The Hope Department of Entomology at Oxford, that Westwood founded, contains a number of other interesting species from
|
||
<collectingCountry id="F339B6B44C55C6741C282B7CFD41271E" box="[520,658,1617,1642]" name="Madagascar" pageId="35" pageNumber="36">Madagascar</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C55C6741EE72B59FCF026BB" pageId="35" pageNumber="36" type="discussion">
|
||
<paragraph id="8B91F6244C55C6741EE72B59FAE42610" blockId="35.[151,1437,151,1999]" pageId="35" pageNumber="36">
|
||
<emphasis id="B95A2A364C55C6741EE72B59FE9F27F9" bold="true" box="[199,332,1652,1677]" pageId="35" pageNumber="36">Discussion.</emphasis>
|
||
No historical museum material is known and the first field specimens of this species were recognized by C. Kremen in
|
||
<quantity id="4CD65BC14C55C6741FCA2BB4FD5B27C5" box="[490,648,1688,1713]" metricMagnitude="1" metricUnit="m" metricValue="5.05333" metricValueMax="5.05714" metricValueMin="5.04952" pageId="35" pageNumber="36" unit="in" value="1989.5" valueMax="1991.0" valueMin="1988.0">
|
||
<date id="FF90D0E44C55C6741FCA2BB4FDBB27C5" box="[490,616,1689,1713]" pageId="35" pageNumber="36" value="1988" valueMax="1991">1988–1991</date>
|
||
in
|
||
</quantity>
|
||
what is now Ranomafana National Park (
|
||
<bibRefCitation id="EFBF8BD54C55C6741A522BB4FAF027C5" author="Lees" box="[1138,1315,1688,1713]" pageId="35" pageNumber="36" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997: 64</bibRefCitation>
|
||
). The sole surviving ST ♀ (in BMNH; now a LT) of
|
||
<taxonomicName id="4C2E8DA74C55C6741CAC2B91FC4827A1" box="[652,923,1724,1749]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="andravahana">
|
||
<emphasis id="B95A2A364C55C6741CAC2B91FC4827A1" box="[652,923,1724,1749]" italics="true" pageId="35" pageNumber="36">Mycalesis andravahana</emphasis>
|
||
</taxonomicName>
|
||
was examined and belongs to a not closely related species, as discussed above. The HT of
|
||
<taxonomicName id="4C2E8DA74C55C6741CE92BCCFBFB278C" authority="Butler, 1879" authorityName="Butler" authorityYear="1879" box="[713,1064,1759,1784]" class="Insecta" family="Nymphalidae" genus="Henotesia" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="wardii">
|
||
<emphasis id="B95A2A364C55C6741CE92BCCFC5D278C" box="[713,910,1759,1784]" italics="true" pageId="35" pageNumber="36">Henotesia wardii</emphasis>
|
||
<bibRefCitation id="EFBF8BD54C55C6741DB92BF2FBFB278C" author="Butler" box="[921,1064,1759,1784]" pageId="35" pageNumber="36" refString="Butler, A. G. (1879) Descriptions of new species of Lepidoptera from Madagascar, with notes on some of the forms already described. Annals and Magazine of Natural History, Series 5, 4, 227 - 246. http: // dx. doi. org / 10.1080 / 00222937908679820" type="journal article" year="1879">Butler, 1879</bibRefCitation>
|
||
</taxonomicName>
|
||
was also examined (BMNH(E) #674838) together with its genitalic slide dissection (P. Viette no. 4847; see also above) and this species (now known as
|
||
<taxonomicName id="4C2E8DA74C55C6741F362A05FDA02634" authority="Lees, 2003" authorityName="Lees" authorityYear="2003" box="[278,627,1832,1856]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="viettei">
|
||
<emphasis id="B95A2A364C55C6741F362A05FE32264B" box="[278,481,1832,1855]" italics="true" pageId="35" pageNumber="36">Heteropsis viettei</emphasis>
|
||
Lees, 2003
|
||
</taxonomicName>
|
||
) is clearly not conspecific, despite the superficial similarity to
|
||
<taxonomicName id="4C2E8DA74C55C6741B592A05FEDE2610" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="westwoodi">
|
||
<emphasis id="B95A2A364C55C6741B592A05FEDE2610" italics="true" pageId="35" pageNumber="36">Ht. westwoodi</emphasis>
|
||
</taxonomicName>
|
||
of Pl. 5,
|
||
<figureCitation id="1315EAA14C55C6741F542A66FE122610" box="[372,449,1867,1892]" captionStart="FIGURE 6. A" captionStartId="24.[151,250,1718,1740]" captionTargetBox="[210,1375,193,1675]" captionTargetId="figure@24.[210,1377,193,1694]" captionTargetPageId="24" captionText="FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9." figureDoi="http://doi.org/10.5281/zenodo.264603" httpUri="https://zenodo.org/record/264603/files/figure.png" pageId="35" pageNumber="36">Figs. 6</figureCitation>
|
||
,
|
||
<figureCitation id="1315EAA14C55C6741FE62A66FE062610" box="[454,469,1867,1892]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." figureDoi="http://doi.org/10.5281/zenodo.264604" httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="35" pageNumber="36">7</figureCitation>
|
||
of
|
||
<bibRefCitation id="EFBF8BD54C55C6741FDA2A66FD7C2610" box="[506,687,1867,1892]" pageId="35" pageNumber="36" refString="Mabille, P. (1885 - 1887) Histoire, Physique, Naturelle et Politique de Madagascar. Vol. 18 - 19. Imprimerie Nationale, Paris, 364 pp., 55 pls. [Grandidier, A. [1885 - 1887] (Ed.)] http: // dx. doi. org / 10.5962 / bhl. title. 1599" type="book chapter">Mabille ([1885]</bibRefCitation>
|
||
). The specimen represents
|
||
<taxonomicName id="4C2E8DA74C55C6741DFF2A61FB9E2617" box="[991,1101,1868,1891]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="viettei">
|
||
<emphasis id="B95A2A364C55C6741DFF2A61FB9E2617" box="[991,1101,1868,1891]" italics="true" pageId="35" pageNumber="36">Ht. viettei</emphasis>
|
||
</taxonomicName>
|
||
, as discussed above.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C55C6741EE72A43FCF026BB" blockId="35.[151,1437,151,1999]" pageId="35" pageNumber="36">
|
||
<emphasis id="B95A2A364C55C6741EE72A43FD3526F3" bold="true" box="[199,742,1902,1927]" pageId="35" pageNumber="36">Additional information. DNA divergences</emphasis>
|
||
: COI-5P cluster number BOLD:ACW4998 (exemplars BMAD134-15 DL06-980 from RS Kalambatritra and BMAD243-15 CK92-0005 from PN Andohahela; also KA- P510 (see Aduse-Poku
|
||
<emphasis id="B95A2A364C55C6741FBD2A9AFE0F26BB" box="[413,476,1974,1999]" italics="true" pageId="35" pageNumber="36">et al.,</emphasis>
|
||
2015) from Ambondrombe).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF51A6AC4C52C6731EB72BE4FE61264E" ID-DOI="http://doi.org/10.5281/zenodo.264607" httpUri="https://zenodo.org/record/264607/files/figure.png" pageId="36" pageNumber="37" targetBox="[272,1316,212,1704]" targetPageId="36">
|
||
<paragraph id="8B91F6244C52C6731EB72BE4FE61264E" blockId="36.[151,1436,1737,1850]" pageId="36" pageNumber="37">
|
||
<emphasis id="B95A2A364C52C6731EB72BE4FEF127AA" bold="true" box="[151,290,1737,1759]" pageId="36" pageNumber="37">FIGURE 10.</emphasis>
|
||
Adults in DV (left), VV (right).
|
||
<emphasis id="B95A2A364C52C6731CA42BE4FD4527AA" bold="true" box="[644,662,1737,1758]" pageId="36" pageNumber="37">A</emphasis>
|
||
: HT ♂ of
|
||
<taxonomicName id="4C2E8DA74C52C6731D2C2BE7FBFA27AB" authority="Lees" authorityName="Lees" box="[780,1065,1737,1759]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="westwoodi" status="sp. nov.">
|
||
<emphasis id="B95A2A364C52C6731D2C2BE7FC3D27AB" box="[780,1006,1737,1759]" italics="true" pageId="36" pageNumber="37">Heteropsis westwoodi</emphasis>
|
||
Lees
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A269974D4C52C6731A162BE4FB5927AA" box="[1078,1162,1737,1758]" pageId="36" pageNumber="37" rank="species">
|
||
<emphasis id="B95A2A364C52C6731A162BE4FB5927AA" bold="true" box="[1078,1162,1737,1758]" pageId="36" pageNumber="37">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(Vohitrasiva, Andringitra; NHMUK010289130).
|
||
<emphasis id="B95A2A364C52C6731F5D2BC5FE5D2789" bold="true" box="[381,398,1768,1789]" pageId="36" pageNumber="37">B</emphasis>
|
||
: referred ♀ of
|
||
<taxonomicName id="4C2E8DA74C52C6731C052BC4FD652789" box="[549,694,1767,1790]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="westwoodi">
|
||
<emphasis id="B95A2A364C52C6731C052BC4FD652789" box="[549,694,1767,1790]" italics="true" pageId="36" pageNumber="37">Ht. westwoodi</emphasis>
|
||
</taxonomicName>
|
||
(Andohahela, Antananandava, Andohahela; BMNH(E) #674842).
|
||
<emphasis id="B95A2A364C52C6731B7B2BCAFABE2789" bold="true" box="[1371,1389,1767,1789]" pageId="36" pageNumber="37">C</emphasis>
|
||
: HT ♂ of
|
||
<taxonomicName id="4C2E8DA74C52C6731EEB2A2AFEAD2668" authority="Lees" authorityName="Lees" box="[203,382,1798,1820]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="pauliani" status="sp. nov.">
|
||
<emphasis id="B95A2A364C52C6731EEB2A2AFE952668" box="[203,326,1798,1820]" italics="true" pageId="36" pageNumber="37">Ht. pauliani</emphasis>
|
||
Lees
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A269974D4C52C6731FA82A2AFE0B2668" box="[392,472,1799,1820]" pageId="36" pageNumber="37" rank="species">
|
||
<emphasis id="B95A2A364C52C6731FA82A2AFE0B2668" bold="true" box="[392,472,1799,1820]" pageId="36" pageNumber="37">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(Tsaratanana; DCL-DB-2966; MNHN).
|
||
<emphasis id="B95A2A364C52C6731D552A2AFC542668" bold="true" box="[885,903,1799,1820]" pageId="36" pageNumber="37">D</emphasis>
|
||
: PT ♀ of
|
||
<taxonomicName id="4C2E8DA74C52C6731DCD2A2AFBBB2668" box="[1005,1128,1798,1820]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="pauliani">
|
||
<emphasis id="B95A2A364C52C6731DCD2A2AFBBB2668" box="[1005,1128,1798,1820]" italics="true" pageId="36" pageNumber="37">Ht. pauliani</emphasis>
|
||
</taxonomicName>
|
||
(Tsaratanana; DCL-DB-2975; MNHN). Scale bar: 30 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C334A5AF4C52C6721EE72A4AFE0424EA" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="description">
|
||
<paragraph id="8B91F6244C52C6731EE72A4AFB5E269F" blockId="36.[151,1436,1895,2028]" pageId="36" pageNumber="37">
|
||
<emphasis id="B95A2A364C52C6731EE72A4AFE3426F4" bold="true" box="[199,487,1895,1920]" pageId="36" pageNumber="37">Phylogeny/sister species</emphasis>
|
||
: apparently most closely related to
|
||
<taxonomicName id="4C2E8DA74C52C6731DA72A45FBDB26F4" box="[903,1032,1895,1920]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B95A2A364C52C6731DA72A45FBDB26F4" box="[903,1032,1895,1920]" italics="true" pageId="36" pageNumber="37">Ht. harveyi</emphasis>
|
||
</taxonomicName>
|
||
(cluster number BOLD:ACW5694 and to
|
||
<taxonomicName id="4C2E8DA74C52C6731EC82AA0FE8A26D0" box="[232,345,1933,1956]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="viettei">
|
||
<emphasis id="B95A2A364C52C6731EC82AA0FE8A26D0" box="[232,345,1933,1956]" italics="true" pageId="36" pageNumber="37">Ht. viettei</emphasis>
|
||
</taxonomicName>
|
||
(BOLDAAK5839).
|
||
<bibRefCitation id="EFBF8BD54C52C6731C662AA1FD1C26D0" author="Lees" box="[582,719,1932,1956]" pageId="36" pageNumber="37" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees (1997)</bibRefCitation>
|
||
recovered
|
||
<taxonomicName id="4C2E8DA74C52C6731D712AA0FC2026D0" box="[849,1011,1931,1956]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="westwoodi">
|
||
<emphasis id="B95A2A364C52C6731D712AA0FC2026D0" box="[849,1011,1931,1956]" italics="true" pageId="36" pageNumber="37">Ht. westwoodi</emphasis>
|
||
</taxonomicName>
|
||
(as ‘THRTB’) as sister to
|
||
<taxonomicName id="4C2E8DA74C52C6731B0B2AA0FA4F26D0" box="[1323,1436,1933,1956]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="viettei">
|
||
<emphasis id="B95A2A364C52C6731B0B2AA0FA4F26D0" box="[1323,1436,1933,1956]" italics="true" pageId="36" pageNumber="37">Ht. viettei</emphasis>
|
||
</taxonomicName>
|
||
(‘THRTN’), based on all morphological evidence, genitalic characters, and non-genitalic characters, but without any jackknife support.
|
||
<bibRefCitation id="EFBF8BD54C52C6731FB72AFEFD602698" author="Aduse-Poku" box="[407,691,2003,2028]" pageId="36" pageNumber="37" refString="Aduse-Poku, K., Lees, D. C., Brattstrom, O., Kodandaramaiah, U., Collins, S. & Brakefield, P. M., (2016) Molecular phylogeny and generic-level taxonomy of the widespread palaeotropical ' Heteropsis clade' (Nymphalidae: Satyrinae: Mycalesina). Systematic Entomology, in press. http: // dx. doi. org / syen. 12183" type="book" year="2016">
|
||
Aduse-Poku
|
||
<emphasis id="B95A2A364C52C6731C092AF8FDBB2698" box="[553,616,2003,2028]" italics="true" pageId="36" pageNumber="37">et al.,</emphasis>
|
||
(2016
|
||
</bibRefCitation>
|
||
, in press) confirm it as sister to
|
||
<taxonomicName id="4C2E8DA74C52C6731A382AF8FB552698" box="[1048,1158,2005,2028]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="viettei">
|
||
<emphasis id="B95A2A364C52C6731A382AF8FB552698" box="[1048,1158,2005,2028]" italics="true" pageId="36" pageNumber="37">Ht. viettei</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF51A6AC4C53C6721EB729B4FC6025BA" ID-DOI="http://doi.org/10.5281/zenodo.264608" httpUri="https://zenodo.org/record/264608/files/figure.png" pageId="37" pageNumber="38" targetBox="[264,1363,233,1142]" targetPageId="37">
|
||
<paragraph id="8B91F6244C53C6721EB729B4FC6025BA" blockId="37.[151,1436,1177,1230]" pageId="37" pageNumber="38">
|
||
<emphasis id="B95A2A364C53C6721EB729B4FEF225DA" bold="true" box="[151,289,1177,1199]" pageId="37" pageNumber="38">FIGURE 11.</emphasis>
|
||
♂ genitalia in LV (left), DV (right). A: ♂ genitalia of
|
||
<taxonomicName id="4C2E8DA74C53C6721D5B29B7FB4A25DB" authority="Lees" authorityName="Lees" box="[891,1177,1177,1199]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="westwoodi" status="sp. nov.">
|
||
<emphasis id="B95A2A364C53C6721D5B29B7FB8E25DB" box="[891,1117,1177,1199]" italics="true" pageId="37" pageNumber="38">Heteropsis westwoodi</emphasis>
|
||
Lees
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A269974D4C53C6721A8829B7FB2E25DB" box="[1192,1277,1178,1199]" pageId="37" pageNumber="38" rank="species">
|
||
<emphasis id="B95A2A364C53C6721A8829B7FB2E25DB" bold="true" box="[1192,1277,1178,1199]" pageId="37" pageNumber="38">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(Antanandava, Andohahela; 236DL). B:
|
||
<taxonomicName id="4C2E8DA74C53C6721FB92994FDC725BA" box="[409,532,1208,1230]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pauliani">
|
||
<emphasis id="B95A2A364C53C6721FB92994FDC725BA" box="[409,532,1208,1230]" italics="true" pageId="37" pageNumber="38">Ht. pauliani</emphasis>
|
||
</taxonomicName>
|
||
, ♂ genitalia, PT ♂ (Tsaratanana; 97DL).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B91F6244C53C6721EE729D8FE25247A" blockId="37.[151,1437,1269,2035]" box="[199,502,1269,1294]" pageId="37" pageNumber="38">
|
||
<emphasis id="B95A2A364C53C6721EE729D8FE25247A" bold="true" box="[199,502,1269,1294]" pageId="37" pageNumber="38">Ecology and distribution.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C53C6721EE72834FC722446" blockId="37.[151,1437,1269,2035]" box="[199,929,1305,1330]" pageId="37" pageNumber="38">
|
||
<emphasis id="B95A2A364C53C6721EE72834FEF02446" bold="true" box="[199,291,1305,1330]" pageId="37" pageNumber="38">Habitat</emphasis>
|
||
: montane rainforest with fine-leaved climbing bamboos.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C53C6721EE72810FD3A2422" blockId="37.[151,1437,1269,2035]" box="[199,745,1341,1366]" pageId="37" pageNumber="38">
|
||
<emphasis id="B95A2A364C53C6721EE72810FE902422" bold="true" box="[199,323,1341,1366]" pageId="37" pageNumber="38">Behaviour</emphasis>
|
||
: ♂♂ fly quite rapidly in the substory.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C53C6721EE7284CFC89240E" blockId="37.[151,1437,1269,2035]" box="[199,858,1377,1402]" pageId="37" pageNumber="38">
|
||
<emphasis id="B95A2A364C53C6721EE7284CFEEE240E" bold="true" box="[199,317,1377,1402]" pageId="37" pageNumber="38">Hostplant</emphasis>
|
||
: suspected to be fine-leaved climbing bamboos.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C53C6721EE728A8FE0424EA" blockId="37.[151,1437,1269,2035]" box="[199,471,1413,1438]" pageId="37" pageNumber="38">
|
||
<emphasis id="B95A2A364C53C6721EE728A8FE8B24EA" bold="true" box="[199,344,1413,1438]" pageId="37" pageNumber="38">Early stages</emphasis>
|
||
: unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C53C6721EE72884FB35277E" pageId="37" pageNumber="38" type="distribution">
|
||
<paragraph id="8B91F6244C53C6721EE72884FBB82492" blockId="37.[151,1437,1269,2035]" pageId="37" pageNumber="38">
|
||
<emphasis id="B95A2A364C53C6721EE72884FE8A24B6" bold="true" box="[199,345,1449,1474]" pageId="37" pageNumber="38">Distribution</emphasis>
|
||
: endemic to the southeastern mountains of
|
||
<collectingCountry id="F339B6B44C53C6721D6F2884FC0524B6" box="[847,982,1449,1474]" name="Madagascar" pageId="37" pageNumber="38">Madagascar</collectingCountry>
|
||
from Andohahela north to Andringitra, Mt. Ambondrombe and Mt Maharira, Ranomafana National Park (
|
||
<figureCitation id="1315EAA14C53C6721DA128E0FC012492" box="[897,978,1485,1510]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="37" pageNumber="38">Fig. 30</figureCitation>
|
||
A, pink dots).
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C53C6721EE728DCFB35277E" blockId="37.[151,1437,1269,2035]" box="[199,1254,1521,1546]" pageId="37" pageNumber="38">
|
||
<emphasis id="B95A2A364C53C6721EE728DCFE4A277E" bold="true" box="[199,409,1521,1546]" pageId="37" pageNumber="38">Elevational range</emphasis>
|
||
:
|
||
<quantity id="4CD65BC14C53C6721F8828DCFDEB277D" box="[424,568,1521,1546]" metricMagnitude="3" metricUnit="m" metricValue="1.301" metricValueMax="1.8169999999999997" metricValueMin="0.7849999999999999" pageId="37" pageNumber="38" unit="m" value="1301.0" valueMax="1817.0" valueMin="785.0">785–1817 m</quantity>
|
||
. (n=27 including referred specimens and observational data).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C53C6711EE72B35FF2221A3" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" type="materials_examined">
|
||
<paragraph id="8B91F6244C53C6711EE72B35FF2221A3" blockId="37.[151,1437,1269,2035]" lastBlockId="38.[151,1436,151,215]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
|
||
<emphasis id="B95A2A364C53C6721EE72B35FE692745" bold="true" box="[199,442,1560,1585]" pageId="37" pageNumber="38">Referred specimens.</emphasis>
|
||
♂,
|
||
<collectingCountry id="F339B6B44C53C6721FC72B35FDBD2745" box="[487,622,1560,1585]" name="Madagascar" pageId="37" pageNumber="38">Madagascar</collectingCountry>
|
||
E, Ambondrombe,
|
||
<geoCoordinate id="EE1A90E34C53C6721D6F2B34FC102745" box="[847,963,1557,1585]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="5" value="-21.8782">21.8782o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C53C6721DF12B34FB962744" box="[977,1093,1557,1585]" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="5" value="47.2474">47.2474o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C53C6721A562B35FB1B2745" box="[1142,1224,1560,1585]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="38" unit="km" value="0.5">0.5 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C53C6721AF52B35FAF62745" box="[1237,1317,1560,1585]" metricMagnitude="3" metricUnit="m" metricValue="1.6039999999999999" pageId="37" pageNumber="38" unit="m" value="1604.0">1604m</quantity>
|
||
, +/-
|
||
<quantity id="4CD65BC14C53C6721B7B2B35FA442744" box="[1371,1431,1560,1585]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="38" unit="m" value="50.0">50 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C53C6721EB72B10FEDF2721" box="[151,268,1597,1621]" pageId="37" pageNumber="38" value="2004-03-22">22/3/2004</date>
|
||
, D.C. Lees et al.: DL-4-882, 638 [=
|
||
<collectionCode id="ED3F6EE14C53C6721CEC2B11FD272721" box="[716,756,1596,1621]" pageId="37" pageNumber="38">DL</collectionCode>
|
||
0638;
|
||
<collectionCode id="ED3F6EE14C53C6721D622B10FCAC2721" box="[834,895,1597,1621]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="37" pageNumber="38">DNA</collectionCode>
|
||
extract number],
|
||
<collectionCode id="ED3F6EE14C53C6721A7C2B10FB602721" box="[1116,1203,1597,1621]" pageId="37" pageNumber="38">BMNH</collectionCode>
|
||
(E) #672003; ♂, E, Ambondrombe,
|
||
<geoCoordinate id="EE1A90E34C53C6721F6F2B49FE022708" box="[335,465,1632,1660]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="1" value="-21.87874">21.87874o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C53C6721FFF2B4EFDB3270F" box="[479,608,1632,1660]" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="1" value="47.24969">47.24969o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C53C6721CAD2B49FD152708" box="[653,710,1635,1660]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="38" unit="km" value="1.0">1 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C53C6721CF22B49FCBA270F" box="[722,873,1635,1660]" metricMagnitude="2" metricUnit="m" metricValue="1.7" metricValueMax="2.2" metricValueMin="1.2" pageId="37" pageNumber="38" unit="m" value="170.0" valueMax="220.0" valueMin="120.0">170 +/- 50 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C53C6721D562B4EFC392708" box="[886,1002,1635,1660]" pageId="37" pageNumber="38" value="1995-02-17">17/2/1995</date>
|
||
, D.C. Lees, DL-4-863, KA522 [=KA- P522; extract number; sequenced specimen in Aduse-Poku
|
||
<emphasis id="B95A2A364C53C6721D192BA5FCAA27D4" box="[825,889,1671,1696]" italics="true" pageId="37" pageNumber="38">et al.,</emphasis>
|
||
2015]; ♀, E, Andringitra, ridge,
|
||
<quantity id="4CD65BC14C53C6721AD02BA5FA9A27EB" box="[1264,1353,1672,1696]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="37" pageNumber="38" unit="m" value="1800.0">1800 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C53C6721B762BA5FF0227B0" pageId="37" pageNumber="38" value="2011-11-20">20/11/ 2011</date>
|
||
, D.C.Lees: DL-11-0001, IA645 [isotope voucher]; ♂, SE, Andohahela, Antananandava,
|
||
<quantity id="4CD65BC14C53C6721A9D2B81FAC627B7" box="[1213,1301,1708,1732]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="37" pageNumber="38" unit="m" value="1200.0">1200 m</quantity>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C53C6721B022B81FAAB27B0" box="[1314,1400,1707,1732]" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="48.9">48 54'E</geoCoordinate>
|
||
<geoCoordinate id="EE1A90E34C53C6721BA02B81FF19279C" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="-24.566668">24 34'S</geoCoordinate>
|
||
, [
|
||
<geoCoordinate id="EE1A90E34C53C6721EC32BFDFE8D279C" box="[227,350,1743,1769]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="5" value="-24.5735">24.5735°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C53C6721F522BFDFE13279C" box="[370,448,1743,1769]" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="5555" value="46.2">46.2°E</geoCoordinate>
|
||
+/-km,
|
||
<quantity id="4CD65BC14C53C6721C092BFDFD5A279C" box="[553,649,1744,1768]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="37" pageNumber="38" unit="m" value="1200.0">1200 m</quantity>
|
||
],
|
||
<date id="FF90D0E44C53C6721C842BFDFCF5279C" box="[676,806,1744,1768]" pageId="37" pageNumber="38" value="1992-11-29">29/11/1992</date>
|
||
; ♂, data as above but
|
||
<collectionCode id="ED3F6EE14C53C6721A6B2BFDFB71279C" box="[1099,1186,1744,1768]" pageId="37" pageNumber="38">BMNH</collectionCode>
|
||
(E) #674844; ♂, SE, Andohahela, Antananandava, site S10,
|
||
<quantity id="4CD65BC14C53C6721C732BD5FD78267B" box="[595,683,1784,1808]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="37" pageNumber="38" unit="m" value="1200.0">1200 m</quantity>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C53C6721C982BD5FCDE2664" box="[696,781,1783,1808]" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="48.9">48 54'E</geoCoordinate>
|
||
<geoCoordinate id="EE1A90E34C53C6721D352BD5FCBF2664" box="[789,876,1784,1808]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="-24.566668">24 34'S</geoCoordinate>
|
||
, [
|
||
<geoCoordinate id="EE1A90E34C53C6721D5F2BDAFC262664" box="[895,1013,1780,1808]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="5" value="-24.5735">24.5735o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C53C6721A232BDAFB9E267B" box="[1027,1101,1780,1808]" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="5555" value="46.2">46.2o E</geoCoordinate>
|
||
<quantity id="4CD65BC14C53C6721A742BD5FB7D267B" box="[1108,1198,1784,1808]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="37" pageNumber="38" unit="m" value="1200.0">1200 m</quantity>
|
||
],
|
||
<date id="FF90D0E44C53C6721AE22BD5FA972664" box="[1218,1348,1784,1808]" pageId="37" pageNumber="38" value="1992-11-29">29/11/1992</date>
|
||
, CK92- 0 0 0 5,
|
||
<collectionCode id="ED3F6EE14C53C6721EFC2A36FEE72640" box="[220,308,1819,1844]" pageId="37" pageNumber="38">BMAD</collectionCode>
|
||
243-15 [
|
||
<collectionCode id="ED3F6EE14C53C6721FAC2A31FE1F2640" box="[396,460,1820,1844]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="37" pageNumber="38">DNA</collectionCode>
|
||
barcode number]; ♀ (
|
||
<figureCitation id="1315EAA14C53C6721CE32A36FCC62640" box="[707,789,1819,1844]" captionStart="FIGURE 10" captionStartId="36.[151,250,1737,1759]" captionTargetBox="[272,1316,212,1704]" captionTargetId="figure@36.[258,1329,193,1715]" captionTargetPageId="36" captionText="FIGURE 10. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis westwoodi Lees, sp. nov. (Vohitrasiva, Andringitra; NHMUK 010289130). B: referred ♀ of Ht. westwoodi (Andohahela, Antananandava, Andohahela; BMNH (E) # 674842). C: HT ♂ of Ht. pauliani Lees, sp. nov. (Tsaratanana; DCL-DB- 2966; MNHN). D: PT ♀ of Ht. pauliani (Tsaratanana; DCL-DB- 2975; MNHN). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264607" httpUri="https://zenodo.org/record/264607/files/figure.png" pageId="37" pageNumber="38">Fig. 10</figureCitation>
|
||
B), SE, PN Andohahela, Antananandava,
|
||
<quantity id="4CD65BC14C53C6721AC62A31FAEF2647" box="[1254,1340,1820,1844]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="37" pageNumber="38" unit="m" value="1200.0">1200 m</quantity>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C53C6721B672A31FA4F2640" box="[1351,1436,1819,1844]" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="48.9">48 54'E</geoCoordinate>
|
||
<geoCoordinate id="EE1A90E34C53C6721EB72A6EFF25262F" box="[151,246,1859,1883]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="-24.566668">24 34'S</geoCoordinate>
|
||
, [
|
||
<geoCoordinate id="EE1A90E34C53C6721F302A6FFE54262F" box="[272,391,1855,1883]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="5" value="-24.5735">24.5735o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C53C6721FBC2A6FFE3B262E" box="[412,488,1855,1883]" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="5555" value="46.2">46.2o E</geoCoordinate>
|
||
,
|
||
<quantity id="4CD65BC14C53C6721FDC2A6EFD8D262E" box="[508,606,1859,1883]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="37" pageNumber="38" unit="m" value="1200.0">1200 m</quantity>
|
||
],
|
||
<date id="FF90D0E44C53C6721C5A2A6EFD2F262F" box="[634,764,1859,1883]" pageId="37" pageNumber="38" value="1992-11-29">29/11/1992</date>
|
||
,
|
||
<collectionCode id="ED3F6EE14C53C6721D2F2A6EFCB5262F" box="[783,870,1859,1883]" pageId="37" pageNumber="38">BMNH</collectionCode>
|
||
(E) #674842; ♂, SE, Andohahela, C. Kremen,
|
||
<collectionCode id="ED3F6EE14C53C6721EB72A4AFF3D260B" box="[151,238,1895,1919]" pageId="37" pageNumber="38">BMNH</collectionCode>
|
||
(E) #697865, 238BDL [genitalia], IA636 [isotope voucher]; specimen, SE, Befarara, RS Kalambatritra (western),
|
||
<geoCoordinate id="EE1A90E34C53C6721F332AA3FEAC26D2" box="[275,383,1930,1958]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="-23.417">23.417 o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C53C6721FB02AA0FE2326D1" box="[400,496,1930,1958]" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="555" value="46.43">46.43 o E</geoCoordinate>
|
||
,
|
||
<quantity id="4CD65BC14C53C6721C212AA0FD3326D1" box="[513,736,1933,1958]" metricMagnitude="3" metricUnit="m" metricValue="1.507" metricValueMax="1.6469999999999998" metricValueMin="1.367" pageId="37" pageNumber="38" unit="m" value="1507.0">1507 m +/- 140 m</quantity>
|
||
, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C53C6721D592AA0FC7326D2" box="[889,928,1933,1958]" pageId="37" pageNumber="38">DL</collectionCode>
|
||
06-980,
|
||
<date id="FF90D0E44C53C6721A202AA0FB4B26D2" box="[1024,1176,1933,1958]" pageId="37" pageNumber="38" value="2006-12-15">15-Dec-2006</date>
|
||
: 11:59,
|
||
<collectionCode id="ED3F6EE14C53C6721AD82AA0FA8226D2" box="[1272,1361,1933,1958]" pageId="37" pageNumber="38">BMAD</collectionCode>
|
||
134-15 [
|
||
<collectionCode id="ED3F6EE14C53C6721EBD2A99FF0E26B8" box="[157,221,1972,1996]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="37" pageNumber="38">DNA</collectionCode>
|
||
barcode voucher]; ♂, SE, Befarara, RS Kalambatritra (western),
|
||
<geoCoordinate id="EE1A90E34C53C6721D9B2A99FBFD26B8" box="[955,1070,1968,1997]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="5" value="-23.4186">23.4186o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C53C6721A1B2A99FB6E26B8" box="[1083,1213,1968,1997]" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="1" value="46.44025">46.44025o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C53C6721ACC2A99FA8A26B9" box="[1260,1369,1972,1997]" metricMagnitude="1" metricUnit="m" metricValue="4.2" pageId="37" pageNumber="38" unit="km" value="0.042">0.042 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C53C6721B442A99FF2F2686" metricMagnitude="3" metricUnit="m" metricValue="1.569" metricValueMax="1.594" metricValueMin="1.544" pageId="37" pageNumber="38" unit="m" value="1569.0" valueMax="1594.0" valueMin="1544.0">1569 +/- 25 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C53C6721F282AF7FE5F2687" box="[264,396,2010,2035]" pageId="37" pageNumber="38" value="2006-12-12">12/12/2006</date>
|
||
: 11:47, R. Ranaivosolo:
|
||
<collectionCode id="ED3F6EE14C53C6721C832AF7FD182687" box="[675,715,2010,2035]" pageId="37" pageNumber="38">DL</collectionCode>
|
||
06-767; ♂, SE, Befarara, RS Kalambatritra (western),
|
||
<geoCoordinate id="EE1A90E34C53C6721B112AF6FA452687" box="[1329,1430,2007,2035]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="-23.418">23.418o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C50C6711EB72DB7FF3D21C6" box="[151,238,151,179]" direction="east" orientation="longitude" pageId="38" pageNumber="39" precision="555" value="46.44">46.44o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C50C6711F022DB7FE5021C7" box="[290,387,154,179]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="38" pageNumber="39" unit="km" value="0.15">0.15 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C50C6711FB02DB7FD9121C6" box="[400,578,154,179]" metricMagnitude="3" metricUnit="m" metricValue="1.557" metricValueMax="1.5819999999999999" metricValueMin="1.5319999999999998" pageId="38" pageNumber="39" unit="m" value="1557.0" valueMax="1582.0" valueMin="1532.0">1557 +/- 25 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C50C6711C702DB7FD0121C7" box="[592,722,154,179]" pageId="38" pageNumber="39" value="2006-12-12">12/12/2006</date>
|
||
, M. Linares:
|
||
<collectionCode id="ED3F6EE14C50C6711D4E2DB7FC4521C7" box="[878,918,154,179]" pageId="38" pageNumber="39">DL</collectionCode>
|
||
06-811, IA161 [isotope voucher]; right wings imaged.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |