697 lines
82 KiB
XML
697 lines
82 KiB
XML
<document id="B8307C3D8467AB414A449BFB39ECE57B" ID-CLB-Dataset="38930" ID-DOI="10.11646/zootaxa.4118.1.1" ID-GBIF-Dataset="a4e408c4-2575-4b5c-878b-e1b9010ec619" ID-ISSN="1175-5326" ID-Zenodo-Dep="264597" ID-ZooBank="CFA586DA-10EE-468B-80EE-35351E3845FD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="guilherme" checkinTime="1465242077321" checkinUser="plazi" docAuthor="C, Lees David" docDate="2016" docId="038747324C2EC60C1EB72DBAFC8B2602" docLanguage="en" docName="zootaxa.4118.pdf.imf" docOrigin="Zootaxa 4118 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Heteropsis vertigo Lees & Raharitsimba, sp. nov." docType="treatment" docVersion="12" lastPageNumber="92" masterDocId="FFBE3F4A4C76C6571E202D2DFFD32174" masterDocTitle="Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision" masterLastPageNumber="97" masterPageNumber="1" pageNumber="89" updateTime="1735917000942" updateUser="ExternalLinkService">
|
||
<mods:mods id="FFE7C7BA9C6613F6FDFC3A04CD00698F" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="4B3E3C089676B4F940D8F5A623CB09A3">
|
||
<mods:title id="D0A3421C9400D9D7FFFA6D154D59BB7E">Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="5309871E20F296326FE3359E57F04B18" type="personal">
|
||
<mods:role id="9FBEBF9B934D5A6D32D0D6107879EC0B">
|
||
<mods:roleTerm id="9491F27EE354DD932E1BD020779C36E5">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="25E69DD312658384C501D131B05E36CB">C, Lees David</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="684F6AD02EAA671ABE0ABAFA90D1565C">text</mods:typeOfResource>
|
||
<mods:relatedItem id="F2AE61E9D0372E8D5A98BD69D02EA930" type="host">
|
||
<mods:titleInfo id="8B9D3703142D3ABF3413A4F4155BD3D6">
|
||
<mods:title id="ACA0DA11A10F8CA56AD27C3F463D9924">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="113207ED52FED9A9A6E47F82648338F0">
|
||
<mods:date id="16E4556805D4B5F76A3623CC4CFDB46B">2016</mods:date>
|
||
<mods:detail id="F185184A9A97AA2D9FC20475511D7AAA" type="volume">
|
||
<mods:number id="65F4B888EBC83724BE43BE5E7FDECF56">4118</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="2D244FDF4D4F093F442348079E8BB67E" type="issue">
|
||
<mods:number id="56001052420A02F2481F20907406286E">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="7B0B908E0A0E0B0DEF5EA25C0ECE1E83" unit="page">
|
||
<mods:start id="E49128C79E78FB956915E0ADB2B7CBA3">1</mods:start>
|
||
<mods:end id="EAC0E61E3508695B71872C42853785D4">97</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="E8249BAA66C7472AC0D48FB8F5A99985">journal article</mods:classification>
|
||
<mods:identifier id="18CB7A7261CBC85A3A01BFB5C0A7143F" type="CLB-Dataset">38930</mods:identifier>
|
||
<mods:identifier id="7DD97CA58E87DEAD9412B3654FAEC636" type="DOI">10.11646/zootaxa.4118.1.1</mods:identifier>
|
||
<mods:identifier id="39CB45E6B3497F9507F8B737ABDF89B1" type="GBIF-Dataset">a4e408c4-2575-4b5c-878b-e1b9010ec619</mods:identifier>
|
||
<mods:identifier id="432245CECD02A3BCD74B74CFFE1474C5" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="CE9BC5BEF0418E077B026F3F8A503311" type="Zenodo-Dep">264597</mods:identifier>
|
||
<mods:identifier id="D376D76407CAB06B80E9C2934515AFB8" type="ZooBank">CFA586DA-10EE-468B-80EE-35351E3845FD</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="038747324C2EC60C1EB72DBAFC8B2602" ID-DOI="http://doi.org/10.5281/zenodo.6086464" ID-GBIF-Taxon="120817149" ID-Zenodo-Dep="6086464" LSID="urn:lsid:plazi:treatment:038747324C2EC60C1EB72DBAFC8B2602" httpUri="http://treatment.plazi.org/id/038747324C2EC60C1EB72DBAFC8B2602" lastPageId="91" lastPageNumber="92" pageId="88" pageNumber="89" scope_family="Nymphalidae" scope_subFamily="Satyrinae" scope_tribe="Satyrini">
|
||
<subSubSection id="C334A5AF4C2EC60F1EB72DBAFBCE21A7" pageId="88" pageNumber="89" type="nomenclature">
|
||
<paragraph id="8B91F6244C2EC60F1EB72DBAFD2121C6" blockId="88.[151,1053,151,211]" box="[151,754,151,178]" pageId="88" pageNumber="89">
|
||
<heading id="D0D941484C2EC60F1EB72DBAFD2121C6" bold="true" box="[151,754,151,178]" fontSize="11" level="1" pageId="88" pageNumber="89" reason="1">
|
||
<emphasis id="B95A2A364C2EC60F1EB72DBAFD2121C6" bold="true" box="[151,754,151,178]" pageId="88" pageNumber="89">
|
||
<taxonomicName id="4C2E8DA74C2EC60F1EB72DBAFD5421C6" authority="Lees & Raharitsimba" authorityName="Lees & Raharitsimba" box="[151,647,151,178]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="vertigo" status="sp. nov.">
|
||
<emphasis id="B95A2A364C2EC60F1EB72DBAFEA121C5" bold="true" box="[151,370,151,177]" italics="true" pageId="88" pageNumber="89">Heteropsis vertigo</emphasis>
|
||
Lees & Raharitsimba
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A269974D4C2EC60F1CB42DB5FD2121C6" box="[660,754,152,178]" pageId="88" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C2EC60F1EB72D96FBCE21A7" blockId="88.[151,1053,151,211]" box="[151,1053,186,211]" pageId="88" pageNumber="89">LSID: urn:lsid:zoobank.org:act:38AC77EE-A10A-4DCA-9444-F55C845FFA23</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C2EC60F1EB72C2FFA89206F" box="[151,1370,258,283]" pageId="88" pageNumber="89" type="description">
|
||
<paragraph id="8B91F6244C2EC60F1EB72C2FFA89206F" blockId="88.[151,1437,258,2028]" box="[151,1370,258,283]" pageId="88" pageNumber="89">
|
||
<emphasis id="B95A2A364C2EC60F1EB72C2FFE84206F" bold="true" box="[151,343,258,283]" pageId="88" pageNumber="89">Prior references</emphasis>
|
||
: sp. 61 (on envelopes and/or datalabels but not mentioned in the list of
|
||
<bibRefCitation id="EFBF8BD54C2EC60F1A592C2EFB2B206F" author="Lees" box="[1145,1272,259,283]" pageId="88" pageNumber="89" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997</bibRefCitation>
|
||
: 64–65).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C2EC60F1EE72C08FDE422BD" pageId="88" pageNumber="89" type="materials_examined">
|
||
<paragraph id="8B91F6244C2EC60F1EE72C08FA8520F3" blockId="88.[151,1437,258,2028]" pageId="88" pageNumber="89">
|
||
<emphasis id="B95A2A364C2EC60F1EE72C08FCC3204A" bold="true" box="[199,784,293,318]" pageId="88" pageNumber="89">
|
||
<typeStatus id="549548864C2EC60F1EE72C08FED1204A" box="[199,258,293,318]" pageId="88" pageNumber="89">Type</typeStatus>
|
||
material., Deposition
|
||
<collectionCode id="ED3F6EE14C2EC60F1C082C08FD50204A" box="[552,643,293,318]" pageId="88" pageNumber="89">BMNH</collectionCode>
|
||
:
|
||
<typeStatus id="549548864C2EC60F1CBA2C08FCD8204A" box="[666,779,293,318]" pageId="88" pageNumber="89" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
♂ (
|
||
<figureCitation id="1315EAA14C2EC60F1D702C08FC7F204A" box="[848,940,293,318]" captionStart="FIGURE 28" captionStartId="89.[151,250,1865,1887]" captionTargetBox="[278,1298,207,1823]" captionTargetId="figure@89.[278,1309,193,1844]" captionTargetPageId="89" captionText="FIGURE 28. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; NHMUK 010289173). B: PT ♀ of Ht. vertigo (Mt. Bekolosy; NHMUK 010289175). C: LT ♂ of Culapa turbans Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674780). D: LT ♂ of Culapa curvatula Oberthür, 1916 (‘ Antsianaka et Lac Alaotra’; BMNH (E) # 674784). E: HT ♂ of Culapa sabas Oberthür, 1923 (‘ Tamatave’; BMNH (E) # 674789). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264625" httpUri="https://zenodo.org/record/264625/files/figure.png" pageId="88" pageNumber="89">Fig. 28</figureCitation>
|
||
A),
|
||
<collectingCountry id="F339B6B44C2EC60F1DC12C08FBBB204A" box="[993,1128,293,318]" name="Madagascar" pageId="88" pageNumber="89">Madagascar</collectingCountry>
|
||
NW, Bekolosy Mt., RS Manongarivo, c.
|
||
<quantity id="4CD65BC14C2EC60F1F782C66FE7C2017" box="[344,431,331,356]" metricMagnitude="3" metricUnit="m" metricValue="1.1629999999999998" pageId="88" pageNumber="89" unit="m" value="1163.0">1163 m</quantity>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C2EC60F1F9C2C66FDEE2010" box="[444,573,328,356]" direction="south" orientation="latitude" pageId="88" pageNumber="89" precision="1" value="-14.04575">14.04575o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C2EC60F1C6B2C66FD1F2017" box="[587,716,328,356]" direction="east" orientation="longitude" pageId="88" pageNumber="89" precision="1" value="48.29623">48.29623o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C2EC60F1CDD2C66FC9C2010" box="[765,847,331,356]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="88" pageNumber="89" unit="km" value="0.5">0.5 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C2EC60F1D7B2C61FC2D2017" box="[859,1022,331,356]" metricMagnitude="3" metricUnit="m" metricValue="1.13" metricValueMax="1.165" metricValueMin="1.095" pageId="88" pageNumber="89" unit="m" value="1130.0" valueMax="1165.0" valueMin="1095.0">1130 +/- 35 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C2EC60F1A2B2C66FB5E2010" box="[1035,1165,331,356]" pageId="88" pageNumber="89" value="1994-12-16">16/12/1994</date>
|
||
, soleil/lo [fruit trap], H. Raharitsimba:
|
||
<collectionCode id="ED3F6EE14C2EC60F1F1B2C42FE4820F3" box="[315,411,367,391]" pageId="88" pageNumber="89">DLBEK</collectionCode>
|
||
94_77, KA2049 [=KA-P2049; extract number],
|
||
<collectionCode id="ED3F6EE14C2EC60F1D952C42FBCC20F3" box="[949,1055,367,391]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="88" pageNumber="89">NHMUK</collectionCode>
|
||
010289173 [
|
||
<collectingCountry id="F339B6B44C2EC60F1A8A2C42FB3620F3" box="[1194,1253,367,391]" name="Qatar" pageId="88" pageNumber="89">QTR</collectingCountry>
|
||
barcode].
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C2EC60F1EE72CBFFDE422BD" blockId="88.[151,1437,258,2028]" pageId="88" pageNumber="89">
|
||
<typeStatus id="549548864C2EC60F1EE72CBFFEEC20DF" box="[199,319,402,427]" pageId="88" pageNumber="89" type="paratype">
|
||
<emphasis id="B95A2A364C2EC60F1EE72CBFFEEC20DF" bold="true" box="[199,319,402,427]" pageId="88" pageNumber="89">Paratypes</emphasis>
|
||
</typeStatus>
|
||
:
|
||
<emphasis id="B95A2A364C2EC60F1F6D2CBFFDF820DF" bold="true" box="[333,555,402,427]" pageId="88" pageNumber="89">
|
||
Deposition
|
||
<collectionCode id="ED3F6EE14C2EC60F1FF42CBFFDF820DF" box="[468,555,402,427]" pageId="88" pageNumber="89">BMNH</collectionCode>
|
||
</emphasis>
|
||
: ♂, data as HT but
|
||
<date id="FF90D0E44C2EC60F1D292CBFFC6520DE" box="[777,950,402,426]" pageId="88" pageNumber="89" value="1994-12-12" valueMax="1994-12-22" valueMin="1994-12-12">12–22/12/1994</date>
|
||
, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C2EC60F1A612CBFFB7120DE" box="[1089,1186,402,426]" pageId="88" pageNumber="89">DLBEK</collectionCode>
|
||
94_83, IA354 [isotope voucher];
|
||
<collectionCode id="ED3F6EE14C2EC60F1F342C98FEBE20BA" box="[276,365,437,462]" pageId="88" pageNumber="89">BMAD</collectionCode>
|
||
241-15 [
|
||
<collectionCode id="ED3F6EE14C2EC60F1FF02C9BFDC320BA" box="[464,528,438,462]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="88" pageNumber="89">DNA</collectionCode>
|
||
barcoded specimen],
|
||
<collectionCode id="ED3F6EE14C2EC60F1D062C9BFC4320BA" box="[806,912,438,462]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="88" pageNumber="89">NHMUK</collectionCode>
|
||
010289174 [
|
||
<collectingCountry id="F339B6B44C2EC60F1A062C9BFBB220BA" box="[1062,1121,438,462]" name="Qatar" pageId="88" pageNumber="89">QTR</collectingCountry>
|
||
barcode]; ♀ (
|
||
<figureCitation id="1315EAA14C2EC60F1B3E2C98FAAF20BA" box="[1310,1404,437,462]" captionStart="FIGURE 28" captionStartId="89.[151,250,1865,1887]" captionTargetBox="[278,1298,207,1823]" captionTargetId="figure@89.[278,1309,193,1844]" captionTargetPageId="89" captionText="FIGURE 28. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; NHMUK 010289173). B: PT ♀ of Ht. vertigo (Mt. Bekolosy; NHMUK 010289175). C: LT ♂ of Culapa turbans Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674780). D: LT ♂ of Culapa curvatula Oberthür, 1916 (‘ Antsianaka et Lac Alaotra’; BMNH (E) # 674784). E: HT ♂ of Culapa sabas Oberthür, 1923 (‘ Tamatave’; BMNH (E) # 674789). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264625" httpUri="https://zenodo.org/record/264625/files/figure.png" pageId="88" pageNumber="89">Fig. 28</figureCitation>
|
||
B),
|
||
<collectingCountry id="F339B6B44C2EC60F1EB72CF6FECD2080" box="[151,286,475,500]" name="Madagascar" pageId="88" pageNumber="89">Madagascar</collectingCountry>
|
||
E, Bekolosy Mt., RS Manongarivo,
|
||
<collectingCountry id="F339B6B44C2EC60F1CEF2CF6FC852080" box="[719,854,475,500]" name="Madagascar" pageId="88" pageNumber="89">Madagascar</collectingCountry>
|
||
NW,
|
||
<quantity id="4CD65BC14C2EC60F1DBE2CF1FC2A2087" box="[926,1017,476,500]" metricMagnitude="3" metricUnit="m" metricValue="1.31" pageId="88" pageNumber="89" unit="m" value="1310.0">1310 m</quantity>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C2EC60F1A282CF6FBA82080" box="[1032,1147,472,500]" direction="south" orientation="latitude" pageId="88" pageNumber="89" precision="5" value="-14.0385">14.0385o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C2EC60F1AAB2CF1FB2C2087" box="[1163,1279,472,500]" direction="east" orientation="longitude" pageId="88" pageNumber="89" precision="5" value="48.3073">48.3073o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C2EC60F1B152CF6FA4B2080" box="[1333,1432,475,500]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="88" pageNumber="89" unit="km" value="0.55">0.55 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C2EC60F1EB72CD2FE9D2362" box="[151,334,510,535]" metricMagnitude="3" metricUnit="m" metricValue="1.31" metricValueMax="1.3599999999999999" metricValueMin="1.26" pageId="88" pageNumber="89" unit="m" value="1310.0" valueMax="1360.0" valueMin="1260.0">1310 +/- 50 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C2EC60F1F422CD2FE372363" box="[354,484,511,535]" pageId="88" pageNumber="89" value="1994-12-21">21/12/1994</date>
|
||
, lo lt 1310 [fruit trap], D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C2EC60F1DBE2CD2FC2C2363" box="[926,1023,511,535]" pageId="88" pageNumber="89">DLBEK</collectionCode>
|
||
94_129, IA 356 [isotope voucher],
|
||
<collectionCode id="ED3F6EE14C2EC60F1EB72F0CFED2234E" box="[151,257,545,570]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="88" pageNumber="89">NHMUK</collectionCode>
|
||
010289175 [
|
||
<collectingCountry id="F339B6B44C2EC60F1FAC2F0FFE14234E" box="[396,455,546,570]" name="Qatar" pageId="88" pageNumber="89">QTR</collectingCountry>
|
||
barcode]; ♂ (
|
||
<figureCitation id="1315EAA14C2EC60F1C412F0CFD61234E" box="[609,690,545,570]" captionStart="FIGURE 29" captionStartId="90.[151,250,1420,1442]" captionTargetBox="[179,1434,202,1356]" captionTargetId="figure@90.[161,1455,193,1385]" captionTargetPageId="90" captionText="FIGURE 29. ♂ genitalia in LV (left), DV (right). A: Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 159, 366 DL). B: Ht. turbans (Oberthür, 1916) (Zahamena; 370 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264626" httpUri="https://zenodo.org/record/264626/files/figure.png" pageId="88" pageNumber="89">Fig. 29</figureCitation>
|
||
A, genitalia),
|
||
<collectingCountry id="F339B6B44C2EC60F1D6A2F0CFC02234E" box="[842,977,545,570]" name="Madagascar" pageId="88" pageNumber="89">Madagascar</collectingCountry>
|
||
NW, Bekolosy, RS Manongarivo, ~
|
||
<quantity id="4CD65BC14C2EC60F1B432F0FFF7C232B" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="88" pageNumber="89" unit="m" value="1200.0">1200 m</quantity>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C2EC60F1E9B2F6AFEFD2314" box="[187,302,580,608]" direction="south" orientation="latitude" pageId="88" pageNumber="89" precision="5" value="-14.0436">14.0436o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C2EC60F1F1B2F65FE72232B" box="[315,417,580,608]" direction="east" orientation="longitude" pageId="88" pageNumber="89" precision="55" value="48.299">48.299o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C2EC60F1FF02F65FDFD2314" box="[464,558,583,608]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="88" pageNumber="89" unit="km" value="0.43">0.43 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C2EC60F1C1A2F65FD38232B" box="[570,747,583,608]" metricMagnitude="2" metricUnit="m" metricValue="1.2" metricValueMax="1.7" metricValueMin="0.7" pageId="88" pageNumber="89" unit="m" value="120.0">120 m +/- 50 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C2EC60F1CD72F65FCAA2314" box="[759,889,584,608]" pageId="88" pageNumber="89" value="1994-12-13">13/12/1994</date>
|
||
, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C2EC60F1A242F6AFBB62314" box="[1028,1125,583,608]" pageId="88" pageNumber="89">DLBEK</collectionCode>
|
||
94_159, DCLW-0104 [wing prep.], KA3035 [=KA-P3035,
|
||
<collectionCode id="ED3F6EE14C2EC60F1FDA2F46FDE423F7" box="[506,567,619,643]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="88" pageNumber="89">DNA</collectionCode>
|
||
extract number]; IA355 [isotope voucher];
|
||
<collectionCode id="ED3F6EE14C2EC60F1A172F46FB5D23F0" box="[1079,1166,619,644]" pageId="88" pageNumber="89">BMAD</collectionCode>
|
||
240-15 [
|
||
<collectionCode id="ED3F6EE14C2EC60F1ACB2F46FAF823F7" box="[1259,1323,619,643]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="88" pageNumber="89">DNA</collectionCode>
|
||
barcoded specimen] 366
|
||
<collectionCode id="ED3F6EE14C2EC60F1F602FA3FEB523D3" box="[320,358,654,679]" pageId="88" pageNumber="89">DL</collectionCode>
|
||
[genitalia],
|
||
<collectionCode id="ED3F6EE14C2EC60F1FDA2FA3FDB723D3" box="[506,612,654,679]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="88" pageNumber="89">NHMUK</collectionCode>
|
||
010289176 [
|
||
<collectingCountry id="F339B6B44C2EC60F1CD52FA2FCE323D3" box="[757,816,655,679]" name="Qatar" pageId="88" pageNumber="89">QTR</collectingCountry>
|
||
barcode]; ♀, NW, Bekolosy Mt., RS Manongarivo,
|
||
<collectingCountry id="F339B6B44C2EC60F1EB72F99FECD23B9" box="[151,286,692,717]" name="Madagascar" pageId="88" pageNumber="89">Madagascar</collectingCountry>
|
||
NW,>
|
||
<quantity id="4CD65BC14C2EC60F1FA02F99FE3323B8" box="[384,480,692,717]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="88" pageNumber="89" unit="m" value="1250.0">1250 m</quantity>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C2EC60F1FD42F98FDB423B9" box="[500,615,689,717]" direction="south" orientation="latitude" pageId="88" pageNumber="89" precision="5" value="-14.0427">14.0427o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C2EC60F1C5C2F98FD2323B8" box="[636,752,689,717]" direction="east" orientation="longitude" pageId="88" pageNumber="89" precision="5" value="48.3028">48.3028o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C2EC60F1D112F99FC5A23B9" box="[817,905,692,717]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="88" pageNumber="89" unit="km" value="0.5">0.5 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C2EC60F1DBD2F99FBA323B8" box="[925,1136,692,717]" metricMagnitude="2" metricUnit="m" metricValue="1.25" metricValueMax="1.75" metricValueMin="0.75" pageId="88" pageNumber="89" unit="m" value="125.0">125 m +/- 50 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C2EC60F1AA52F98FAD423B9" box="[1157,1287,693,717]" pageId="88" pageNumber="89" value="1994-12-19">19/12/1994</date>
|
||
, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C2EC60F1EB72FF5FF242384" box="[151,247,728,752]" pageId="88" pageNumber="89">DLBEK</collectionCode>
|
||
94_110, 1035 [=
|
||
<collectionCode id="ED3F6EE14C2EC60F1F952FFAFE0D2384" box="[437,478,727,752]" pageId="88" pageNumber="89">DL</collectionCode>
|
||
1035;
|
||
<collectionCode id="ED3F6EE14C2EC60F1C082FF5FDB62384" box="[552,613,728,752]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="88" pageNumber="89">DNA</collectionCode>
|
||
extract number],
|
||
<collectionCode id="ED3F6EE14C2EC60F1D1C2FF5FC402384" box="[828,915,728,752]" pageId="88" pageNumber="89">BMNH</collectionCode>
|
||
(E) #672400 [
|
||
<collectionCode id="ED3F6EE14C2EC60F1A142FF5FBA72384" box="[1076,1140,728,752]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="88" pageNumber="89">DNA</collectionCode>
|
||
voucher; cytochrome b], 372
|
||
<collectionCode id="ED3F6EE14C2EC60F1EE02FD6FF352267" box="[192,230,763,787]" pageId="88" pageNumber="89">DL</collectionCode>
|
||
[♀ genitalia]; ‘leg for KAP’; IA357 [isotope voucher]; ♀, NW, Bekolosy Mt., RS Manongarivo,
|
||
<quantity id="4CD65BC14C2EC60F1B762FD6FF23224C" metricMagnitude="3" metricUnit="m" metricValue="1.225" metricValueMax="1.25" metricValueMin="1.2" pageId="88" pageNumber="89" unit="m" value="1225.0" valueMax="1250.0" valueMin="1200.0">1200– 1250 m</quantity>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C2EC60F1EDD2E0CFEAC224D" box="[253,383,797,825]" direction="south" orientation="latitude" pageId="88" pageNumber="89" precision="1" value="-14.04334">14.04334o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C2EC60F1FAD2E0CFDD2224C" box="[397,513,797,825]" direction="east" orientation="longitude" pageId="88" pageNumber="89" precision="5" value="48.3013">48.3013o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C2EC60F1C142E0CFDA1224D" box="[564,626,800,825]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="88" pageNumber="89" unit="km" value="1.0">1 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C2EC60F1C5F2E0DFCF4224C" box="[639,807,800,825]" metricMagnitude="3" metricUnit="m" metricValue="1.225" metricValueMax="1.275" metricValueMin="1.175" pageId="88" pageNumber="89" unit="m" value="1225.0" valueMax="1275.0" valueMin="1175.0">1225 +/- 50 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C2EC60F1D152E0CFC64224D" box="[821,951,801,825]" pageId="88" pageNumber="89" value="1994-12-12">12/12/1994</date>
|
||
, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C2EC60F1A682E0CFB7A224D" box="[1096,1193,801,825]" pageId="88" pageNumber="89">DLBEK</collectionCode>
|
||
94_23, Egg exp. [egg expression voucher], IA353 [isotope voucher],
|
||
<collectionCode id="ED3F6EE14C2EC60F1C982E69FCF12228" box="[696,802,836,860]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="88" pageNumber="89">NHMUK</collectionCode>
|
||
010289177 [
|
||
<collectingCountry id="F339B6B44C2EC60F1D902E69FC382228" box="[944,1003,836,860]" name="Qatar" pageId="88" pageNumber="89">QTR</collectingCountry>
|
||
barcode]; ♀, NW, Bekolosy Mt., RS Manongarivo, c.
|
||
<quantity id="4CD65BC14C2EC60F1F7E2E44FE6B22F5" box="[350,440,873,898]" metricMagnitude="3" metricUnit="m" metricValue="1.2449999999999999" pageId="88" pageNumber="89" unit="m" value="1245.0">1245 m</quantity>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C2EC60F1FE82E47FDE822F6" box="[456,571,870,898]" direction="south" orientation="latitude" pageId="88" pageNumber="89" precision="5" value="-14.0427">14.0427o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C2EC60F1C6C2E47FD6C22F5" box="[588,703,870,898]" direction="east" orientation="longitude" pageId="88" pageNumber="89" precision="5" value="48.3028">48.3028o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C2EC60F1CD62E44FC9922F6" box="[758,842,873,898]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="88" pageNumber="89" unit="km" value="0.5">0.5 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C2EC60F1D792E44FC6722F5" box="[857,948,873,898]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="88" pageNumber="89" unit="m" value="1250.0">1250 m</quantity>
|
||
, +/-
|
||
<quantity id="4CD65BC14C2EC60F1DCF2E44FBFE22F5" box="[1007,1069,873,898]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="88" pageNumber="89" unit="m" value="50.0">50 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C2EC60F1A1D2E47FB6C22F6" box="[1085,1215,874,898]" pageId="88" pageNumber="89" value="1994-12-12">12/12/1994</date>
|
||
, 10:40, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C2EC60F1EB72EA0FF2622D1" box="[151,245,909,933]" pageId="88" pageNumber="89">DLBEK</collectionCode>
|
||
94_22 [Egg voucher]; IA352 [isotope voucher], KA2050 [=KA-P2050; extract number],
|
||
<collectionCode id="ED3F6EE14C2EC60F1EB72E9DFED222BC" box="[151,257,944,968]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="88" pageNumber="89">NHMUK</collectionCode>
|
||
010289178 [
|
||
<collectingCountry id="F339B6B44C2EC60F1FAC2E9DFE1422BC" box="[396,455,944,968]" name="Qatar" pageId="88" pageNumber="89">QTR</collectingCountry>
|
||
barcode].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C2EC60F1EE72EFEFCBE2298" box="[199,877,979,1004]" pageId="88" pageNumber="89" type="description">
|
||
<paragraph id="8B91F6244C2EC60F1EE72EFEFCBE2298" blockId="88.[151,1437,258,2028]" box="[199,877,979,1004]" pageId="88" pageNumber="89">
|
||
<emphasis id="B95A2A364C2EC60F1EE72EFEFE6D2298" bold="true" box="[199,446,979,1004]" pageId="88" pageNumber="89">Deposition summary</emphasis>
|
||
: BMNH (HT ♂, 2 PT ♂♂, 4 PT ♀♀).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C2EC60F1EE72ED4FA522565" box="[199,1409,1014,1042]" pageId="88" pageNumber="89" type="materials_examined">
|
||
<paragraph id="8B91F6244C2EC60F1EE72ED4FA522565" blockId="88.[151,1437,258,2028]" box="[199,1409,1014,1042]" pageId="88" pageNumber="89">
|
||
<emphasis id="B95A2A364C2EC60F1EE72ED4FEB62566" bold="true" box="[199,357,1017,1042]" pageId="88" pageNumber="89">
|
||
<typeStatus id="549548864C2EC60F1EE72ED4FED12566" box="[199,258,1017,1042]" pageId="88" pageNumber="89">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
Bekolosy Mt., RS Manongarivo, c.
|
||
<quantity id="4CD65BC14C2EC60F1CDB2ED4FC822565" box="[763,849,1017,1042]" metricMagnitude="3" metricUnit="m" metricValue="1.1629999999999998" pageId="88" pageNumber="89" unit="m" value="1163.0">1163 m</quantity>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C2EC60F1D7D2ED4FC032566" box="[861,976,1014,1042]" direction="south" orientation="latitude" pageId="88" pageNumber="89" precision="5" value="-14.0458">14.0458o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C2EC60F1DFD2ED4FB832565" box="[989,1104,1014,1042]" direction="east" orientation="longitude" pageId="88" pageNumber="89" precision="5" value="48.2962">48.2962o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C2EC60F1A5F2ED4FB1C2566" box="[1151,1231,1017,1042]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="88" pageNumber="89" unit="km" value="0.5">0.5 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C2EC60F1AFB2ED7FAAE2565" box="[1243,1405,1017,1042]" metricMagnitude="3" metricUnit="m" metricValue="1.13" metricValueMax="1.165" metricValueMin="1.095" pageId="88" pageNumber="89" unit="m" value="1130.0" valueMax="1165.0" valueMin="1095.0">
|
||
1130 +/-
|
||
<specimenCount id="9D283DAD4C2EC60F1B612ED4FAAE2565" box="[1345,1405,1017,1042]" pageId="88" pageNumber="89" type="generic">35 m</specimenCount>
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C2EC60F1EE72931FE01247C" pageId="88" pageNumber="89" type="diagnosis">
|
||
<paragraph id="8B91F6244C2EC60F1EE72931FE01247C" blockId="88.[151,1437,258,2028]" pageId="88" pageNumber="89">
|
||
<emphasis id="B95A2A364C2EC60F1EE72931FE922541" bold="true" box="[199,321,1052,1077]" pageId="88" pageNumber="89">Diagnosis.</emphasis>
|
||
In its translucence, the extended FWD space-CuA1 Orng of
|
||
<taxonomicName id="4C2E8DA74C2EC60F1DC92933FBB12540" box="[1001,1122,1053,1077]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="vertigo">
|
||
<emphasis id="B95A2A364C2EC60F1DC92933FBB12540" box="[1001,1122,1053,1077]" italics="true" pageId="88" pageNumber="89">Ht. vertigo</emphasis>
|
||
</taxonomicName>
|
||
is intermediate between
|
||
<taxonomicName id="4C2E8DA74C2EC60F1B592933FF3E252C" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="turbans">
|
||
<emphasis id="B95A2A364C2EC60F1B592933FF3E252C" italics="true" pageId="88" pageNumber="89">Ht. turbans</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2E8DA74C2EC60F1F03296CFE5E252C" box="[291,397,1087,1112]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sabas">
|
||
<emphasis id="B95A2A364C2EC60F1F03296CFE5E252C" box="[291,397,1087,1112]" italics="true" pageId="88" pageNumber="89">Ht. sabas</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C2E8DA74C2EC60F1FBB296CFDC7252C" box="[411,532,1089,1112]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="vertigo">
|
||
<emphasis id="B95A2A364C2EC60F1FBB296CFDC7252C" box="[411,532,1089,1112]" italics="true" pageId="88" pageNumber="89">Ht. vertigo</emphasis>
|
||
</taxonomicName>
|
||
in wing pattern resembles
|
||
<taxonomicName id="4C2E8DA74C2EC60F1D63296CFC11252C" box="[835,962,1087,1112]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="turbans">
|
||
<emphasis id="B95A2A364C2EC60F1D63296CFC11252C" box="[835,962,1087,1112]" italics="true" pageId="88" pageNumber="89">Ht. turbans</emphasis>
|
||
</taxonomicName>
|
||
more than it does
|
||
<taxonomicName id="4C2E8DA74C2EC60F1AB2296CFB2F252C" box="[1170,1276,1087,1112]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sabas">
|
||
<emphasis id="B95A2A364C2EC60F1AB2296CFB2F252C" box="[1170,1276,1087,1112]" italics="true" pageId="88" pageNumber="89">Ht. sabas</emphasis>
|
||
</taxonomicName>
|
||
, but the FWD space-CuA1 Orng is largely suffused with orange and only pale whitish-translucent in the anteriad half, especially proximad, rather than throughout it, as in
|
||
<taxonomicName id="4C2E8DA74C2EC60F1CA529AAFD2025EB" box="[645,755,1158,1183]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sabas">
|
||
<emphasis id="B95A2A364C2EC60F1CA529AAFD2025EB" box="[645,755,1158,1183]" italics="true" pageId="88" pageNumber="89">Ht. sabas</emphasis>
|
||
</taxonomicName>
|
||
, or largely orange as in
|
||
<emphasis id="B95A2A364C2EC60F1A3829AAFAF925EA" box="[1048,1322,1158,1183]" italics="true" pageId="88" pageNumber="89">
|
||
<taxonomicName id="4C2E8DA74C2EC60F1A3829AAFB4C25EB" box="[1048,1183,1158,1183]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="turbans">Ht. turbans</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C2E8DA74C2EC60F1A8D29AAFAF925EA" box="[1197,1322,1159,1182]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="vertigo">Ht. vertigo</taxonomicName>
|
||
</emphasis>
|
||
lacks the extreme ear-like cupping of the gnathos seen in
|
||
<taxonomicName id="4C2E8DA74C2EC60F1CF32987FBC825B6" authority="Lees, 1997: 107" authorityName="Lees" authorityPageNumber="107" authorityYear="1997" box="[723,1051,1193,1218]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sabas">
|
||
<emphasis id="B95A2A364C2EC60F1CF32987FC9125B6" box="[723,834,1193,1218]" italics="true" pageId="88" pageNumber="89">Ht. sabas</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBF8BD54C2EC60F1D752987FBC025B6" author="Lees" box="[853,1043,1194,1218]" pageId="88" pageNumber="89" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997: 107</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2E8DA74C2EC60F1A7B2987FB0C25B6" box="[1115,1247,1193,1218]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="turbans">
|
||
<emphasis id="B95A2A364C2EC60F1A7B2987FB0C25B6" box="[1115,1247,1193,1218]" italics="true" pageId="88" pageNumber="89">Ht. turbans</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1315EAA14C2EC60F1AD32984FA9825B6" box="[1267,1355,1193,1218]" captionStart="FIGURE 29" captionStartId="90.[151,250,1420,1442]" captionTargetBox="[179,1434,202,1356]" captionTargetId="figure@90.[161,1455,193,1385]" captionTargetPageId="90" captionText="FIGURE 29. ♂ genitalia in LV (left), DV (right). A: Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 159, 366 DL). B: Ht. turbans (Oberthür, 1916) (Zahamena; 370 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264626" httpUri="https://zenodo.org/record/264626/files/figure.png" pageId="88" pageNumber="89">Fig. 29</figureCitation>
|
||
B; NW Zahamena) as well as some related species (e.g.
|
||
<emphasis id="B95A2A364C2EC60F1C9129E3FC0E2591" box="[689,989,1228,1253]" italics="true" pageId="88" pageNumber="89">
|
||
<taxonomicName id="4C2E8DA74C2EC60F1C9129E3FCBB2591" box="[689,872,1228,1253]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="angulifascia">Ht. angulifascia</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2E8DA74C2EC60F1D5229E3FC0E2591" box="[882,989,1228,1253]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="borgo">Ht. borgo</taxonomicName>
|
||
</emphasis>
|
||
), and the HWV Mb is much less kinked than in any of these species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C2EC60C1EE7283EFC88218C" lastPageId="91" lastPageNumber="92" pageId="88" pageNumber="89" type="description">
|
||
<paragraph id="8B91F6244C2EC60F1EE7283EFB6E26D1" blockId="88.[151,1437,258,2028]" pageId="88" pageNumber="89">
|
||
<emphasis id="B95A2A364C2EC60F1EE7283EFE7E2458" bold="true" box="[199,429,1299,1324]" pageId="88" pageNumber="89">Description. Wings</emphasis>
|
||
: Similar to
|
||
<taxonomicName id="4C2E8DA74C2EC60F1C162839FCDF2458" box="[566,780,1299,1324]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="turbans">
|
||
<emphasis id="B95A2A364C2EC60F1C162839FCDF2458" box="[566,780,1299,1324]" italics="true" pageId="88" pageNumber="89">Heteropsis turbans</emphasis>
|
||
</taxonomicName>
|
||
. Upperside uniform mid brown, except with conspicuous orange Mf/crescent (as in a few other of the
|
||
<taxonomicName id="4C2E8DA74C2EC60F1CB9281AFCCF243B" box="[665,796,1334,1359]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="strigula">
|
||
<emphasis id="B95A2A364C2EC60F1CB9281AFCCF243B" box="[665,796,1334,1359]" italics="true" pageId="88" pageNumber="89">Ht. strigula</emphasis>
|
||
</taxonomicName>
|
||
-group, notably
|
||
<taxonomicName id="4C2E8DA74C2EC60F1DF4281AFB84243B" box="[980,1111,1334,1359]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="strigula">
|
||
<emphasis id="B95A2A364C2EC60F1DF4281AFB84243B" box="[980,1111,1334,1359]" italics="true" pageId="88" pageNumber="89">Ht. strigula</emphasis>
|
||
</taxonomicName>
|
||
) at base of HWD space-M2 following curve at base of vein M3. FW space-CuA1 ocellus with orange to pale yellow/translucent ‘ring,’ more like a hexagon that has been compressed along a diagonal running from mid-costa to tergal angle. FWD space-M1 ocellus quite small and elliptic with narrow orange ring. HWD space-CuA1 ocellus the only one expressed there and slightly elliptic, with narrow orange ring. Darkish brown, diffuse, slightly wavy, uprecurved hair brush emanates mainly from space-1A as far as mid-vein. Underside greyish brown, similar to
|
||
<taxonomicName id="4C2E8DA74C2EC60F1A5628CAFB26248B" box="[1142,1269,1510,1535]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="turbans">
|
||
<emphasis id="B95A2A364C2EC60F1A5628CAFB26248B" box="[1142,1269,1510,1535]" italics="true" pageId="88" pageNumber="89">Ht. turbans</emphasis>
|
||
</taxonomicName>
|
||
but with a less kinked HWV Mb, and limited irroration, especially in basal area. Space-CuA1 ocellus with orange ring yellowing proximad following reddish brown concave curve of Mb before it bends back to mid-costa. Space-M1 ocellus fairly elliptic, with white pupil and wide narrow black iris, but narrow orange ring. HWV space-CuA1 ocellus large and round to elliptic with narrow pale orange ring. HWV space-Rs-M2 ocelli generally not expressed and space- CuA2 ocellus small and elliptic with faint orange ring. HWV Mb reddish brown and fairly straight but outflexed around M2, generally with a yellow patch (Mf) distad of it in space-M2, and curved back towards the anal angle in space-CuA2. Slight, wavy brown Pml tracks the wing margin and Sml more closely follows the fairly crenate (notably much less so than in the related
|
||
<taxonomicName id="4C2E8DA74C2EC60F1C592A2CFD36266C" box="[633,741,1791,1816]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sabas">
|
||
<emphasis id="B95A2A364C2EC60F1C592A2CFD36266C" box="[633,741,1791,1816]" italics="true" pageId="88" pageNumber="89">Ht. sabas</emphasis>
|
||
</taxonomicName>
|
||
) HW margin. FWV cell area with four darker brown rather equally spaced transverse Cbs, the outermost of which flank area that is paler than for the inner pair. Basal areas not much darker than areas distad of the Mb, and darker brown PMb weakly represented.
|
||
<emphasis id="B95A2A364C2EC60F1A5B2A6BFB3C262B" bold="true" box="[1147,1263,1862,1887]" pageId="88" pageNumber="89">Variation.</emphasis>
|
||
♂♂ similar to each other, but seasonal variation is not known, as the entire
|
||
<typeStatus id="549548864C2EC60F1D442A47FC4726F6" box="[868,916,1898,1922]" pageId="88" pageNumber="89">type</typeStatus>
|
||
series was caught near the start of the rainy season in mid December. ♀♀ similar, but dorsally lighter and larger with more rounded wings.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C2EC60F1EE72A82FEAB269F" blockId="88.[151,1437,258,2028]" pageId="88" pageNumber="89">
|
||
<emphasis id="B95A2A364C2EC60F1EE72A82FEBF26BC" bold="true" box="[199,364,1967,1992]" pageId="88" pageNumber="89">Wingspan/fwl</emphasis>
|
||
: range 32.2–35.6/17.3–19.6 (n=2 ♂♂), including HT ♂ 35.2/
|
||
<quantity id="4CD65BC14C2EC60F1A052A82FB5826B3" box="[1061,1163,1967,1992]" metricMagnitude="-2" metricUnit="m" metricValue="1.96" pageId="88" pageNumber="89" unit="mm" value="19.6">19.6 mm</quantity>
|
||
. Range 35.3–36.3/
|
||
<quantity id="4CD65BC14C2EC60F1B7D2A82FF28269F" metricMagnitude="-2" metricUnit="m" metricValue="1.9449999999999998" metricValueMax="1.95" metricValueMin="1.94" pageId="88" pageNumber="89" unit="mm" value="19.45" valueMax="19.5" valueMin="19.4">19.4– 19.5 mm</quantity>
|
||
(n=2 ♀♀).
|
||
</paragraph>
|
||
<caption id="DF51A6AC4C2FC60E1EB72A64FB0126CF" ID-DOI="http://doi.org/10.5281/zenodo.264625" httpUri="https://zenodo.org/record/264625/files/figure.png" pageId="89" pageNumber="90" targetBox="[278,1298,207,1823]" targetPageId="89">
|
||
<paragraph id="8B91F6244C2FC60E1EB72A64FB0126CF" blockId="89.[151,1436,1865,1979]" pageId="89" pageNumber="90">
|
||
<emphasis id="B95A2A364C2FC60E1EB72A64FEF2262A" bold="true" box="[151,289,1865,1887]" pageId="89" pageNumber="90">FIGURE 28.</emphasis>
|
||
Adults in DV (left), VV (right).
|
||
<emphasis id="B95A2A364C2FC60E1C582A64FD59262A" bold="true" box="[632,650,1865,1886]" pageId="89" pageNumber="90">A</emphasis>
|
||
: HT ♂ of
|
||
<taxonomicName id="4C2E8DA74C2FC60E1CDA2A67FB4F262B" authority="Lees & Raharitsimba" authorityName="Lees & Raharitsimba" box="[762,1180,1865,1887]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="vertigo" status="sp. nov.">
|
||
<emphasis id="B95A2A364C2FC60E1CDA2A67FC64262B" box="[762,951,1866,1887]" italics="true" pageId="89" pageNumber="90">Heteropsis vertigo</emphasis>
|
||
Lees & Raharitsimba
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A269974D4C2FC60E1A872A64FB2A262A" box="[1191,1273,1865,1886]" pageId="89" pageNumber="90" rank="species">
|
||
<emphasis id="B95A2A364C2FC60E1A872A64FB2A262A" bold="true" box="[1191,1273,1865,1886]" pageId="89" pageNumber="90">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(Mt. Bekolosy; NHMUK010289173).
|
||
<emphasis id="B95A2A364C2FC60E1F5D2A45FE5D2609" bold="true" box="[381,398,1896,1917]" pageId="89" pageNumber="90">B</emphasis>
|
||
: PT ♀ of
|
||
<taxonomicName id="4C2E8DA74C2FC60E1FD22A44FD8C260A" box="[498,607,1897,1918]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="vertigo">
|
||
<emphasis id="B95A2A364C2FC60E1FD22A44FD8C260A" box="[498,607,1897,1918]" italics="true" pageId="89" pageNumber="90">Ht. vertigo</emphasis>
|
||
</taxonomicName>
|
||
(Mt. Bekolosy; NHMUK010289175).
|
||
<emphasis id="B95A2A364C2FC60E1DCA2A45FC2F260A" bold="true" box="[1002,1020,1896,1918]" pageId="89" pageNumber="90">C</emphasis>
|
||
: LT ♂ of
|
||
<taxonomicName id="4C2E8DA74C2FC60E1A7F2A45FA4F260A" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" box="[1119,1436,1896,1918]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="turbans">
|
||
<emphasis id="B95A2A364C2FC60E1A7F2A45FB79260A" box="[1119,1194,1896,1918]" italics="true" pageId="89" pageNumber="90">Culapa</emphasis>
|
||
<emphasis id="B95A2A364C2FC60E1A8F2A45FB2F260A" box="[1199,1276,1896,1918]" italics="true" pageId="89" pageNumber="90">turbans</emphasis>
|
||
Oberthür, 1916
|
||
</taxonomicName>
|
||
(‘Antsianaka’; BMNH(E) #674780).
|
||
<emphasis id="B95A2A364C2FC60E1C362AAAFDFB26E8" bold="true" box="[534,552,1927,1948]" pageId="89" pageNumber="90">D</emphasis>
|
||
: LT ♂ of
|
||
<taxonomicName id="4C2E8DA74C2FC60E1CB62AABFC3C26E8" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" box="[662,1007,1926,1948]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="curvatula">
|
||
<emphasis id="B95A2A364C2FC60E1CB62AABFD3226E8" box="[662,737,1926,1948]" italics="true" pageId="89" pageNumber="90">Culapa</emphasis>
|
||
<emphasis id="B95A2A364C2FC60E1CC92AABFC9926E8" box="[745,842,1926,1948]" italics="true" pageId="89" pageNumber="90">curvatula</emphasis>
|
||
Oberthür, 1916
|
||
</taxonomicName>
|
||
(‘Antsianaka et Lac Alaotra’; BMNH(E) #674784).
|
||
<emphasis id="B95A2A364C2FC60E1F242A8BFEC626CF" bold="true" box="[260,277,1958,1979]" pageId="89" pageNumber="90">E</emphasis>
|
||
: HT ♂ of
|
||
<taxonomicName id="4C2E8DA74C2FC60E1F5F2A88FD7926CF" authority="Oberthur, 1923" authorityName="Oberthur" authorityYear="1923" box="[383,682,1957,1979]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="sabas">
|
||
<emphasis id="B95A2A364C2FC60E1F5F2A88FE1926CF" box="[383,458,1957,1979]" italics="true" pageId="89" pageNumber="90">Culapa</emphasis>
|
||
<emphasis id="B95A2A364C2FC60E1FF02A88FDD926CF" box="[464,522,1957,1979]" italics="true" pageId="89" pageNumber="90">sabas</emphasis>
|
||
Oberthür, 1923
|
||
</taxonomicName>
|
||
(‘Tamatave’; BMNH(E) #674789). Scale bar: 30 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF51A6AC4C2CC60D1EB728A1FBA124B4" ID-DOI="http://doi.org/10.5281/zenodo.264626" httpUri="https://zenodo.org/record/264626/files/figure.png" pageId="90" pageNumber="91" targetBox="[179,1434,202,1356]" targetPageId="90">
|
||
<paragraph id="8B91F6244C2CC60D1EB728A1FBA124B4" blockId="90.[151,1436,1420,1472]" pageId="90" pageNumber="91">
|
||
<emphasis id="B95A2A364C2CC60D1EB728A1FEF724D5" bold="true" box="[151,292,1420,1442]" pageId="90" pageNumber="91">FIGURE 29.</emphasis>
|
||
♂ genitalia in LV (left), DV (right).
|
||
<emphasis id="B95A2A364C2CC60D1C9D28A1FD1C24D5" bold="true" box="[701,719,1420,1441]" pageId="90" pageNumber="91">A</emphasis>
|
||
:
|
||
<taxonomicName id="4C2E8DA74C2CC60D1CC128A0FB5C24D6" authority="Lees & Raharitsimba" authorityName="Lees & Raharitsimba" box="[737,1167,1420,1442]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="vertigo" status="sp. nov.">
|
||
<emphasis id="B95A2A364C2CC60D1CC128A0FC9D24D6" box="[737,846,1421,1442]" italics="true" pageId="90" pageNumber="91">Heteropsis</emphasis>
|
||
<emphasis id="B95A2A364C2CC60D1D7928A0FC7224D6" box="[857,929,1421,1442]" italics="true" pageId="90" pageNumber="91">vertigo</emphasis>
|
||
Lees & Raharitsimba
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A269974D4C2CC60D1ABD28A1FB2024D5" box="[1181,1267,1420,1441]" pageId="90" pageNumber="91" rank="species">
|
||
<emphasis id="B95A2A364C2CC60D1ABD28A1FB2024D5" bold="true" box="[1181,1267,1420,1441]" pageId="90" pageNumber="91">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(Mt. Bekolosy; DLBEK94_159, 366DL).
|
||
<emphasis id="B95A2A364C2CC60D1F802886FE6224B4" bold="true" box="[416,433,1451,1472]" pageId="90" pageNumber="91">B</emphasis>
|
||
:
|
||
<taxonomicName id="4C2E8DA74C2CC60D1F9F2886FD3024B4" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" box="[447,739,1450,1472]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="turbans">
|
||
<emphasis id="B95A2A364C2CC60D1F9F2886FE0C24B4" box="[447,479,1451,1472]" italics="true" pageId="90" pageNumber="91">Ht.</emphasis>
|
||
<emphasis id="B95A2A364C2CC60D1FC52887FDE124B4" box="[485,562,1450,1472]" italics="true" pageId="90" pageNumber="91">turbans</emphasis>
|
||
(Oberthür, 1916)
|
||
</taxonomicName>
|
||
(Zahamena; 370DL). Scale bar: 3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B91F6244C2CC60D1EE728CAFDD427E4" blockId="90.[151,1437,1511,2005]" pageId="90" pageNumber="91">
|
||
<emphasis id="B95A2A364C2CC60D1EE728CAFE872774" bold="true" box="[199,340,1511,1536]" pageId="90" pageNumber="91">Androconia</emphasis>
|
||
: 1A+2A has a narrowly tapered inflation and 3A is much more swollen distad, from base to midvein, covered in thin grey-brown scales. As for
|
||
<taxonomicName id="4C2E8DA74C2CC60D1C972B20FCEB2751" box="[695,824,1548,1573]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="turbans">
|
||
<emphasis id="B95A2A364C2CC60D1C972B20FCEB2751" box="[695,824,1548,1573]" italics="true" pageId="90" pageNumber="91">Ht. turbans</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2E8DA74C2CC60D1D522B20FC0E2751" box="[882,989,1548,1573]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="sabas">
|
||
<emphasis id="B95A2A364C2CC60D1D522B20FC0E2751" box="[882,989,1548,1573]" italics="true" pageId="90" pageNumber="91">Ht. sabas</emphasis>
|
||
</taxonomicName>
|
||
, along their length HW veins M2, M3, CuA1 and CuA2 are also narrowly inflated and have specialized scales on the dorsal surface. Abdominal black androconia just visible ventro-laterally around A4–A5. HWD Sdb dark brown, Sdp grey, small, lenticular, composed of narrow grey scales.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C2CC60D1EE72BB1FE962789" blockId="90.[151,1437,1511,2005]" pageId="90" pageNumber="91">
|
||
<emphasis id="B95A2A364C2CC60D1EE72BB1FEDB27C1" bold="true" box="[199,264,1692,1717]" pageId="90" pageNumber="91">Palps</emphasis>
|
||
: ochreous brown with darker brown thin medial stripe and with dark brown border away from compound eye and with dark brown scales above where potentially brushing eye, sandwiching ochreous scales away from eye.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C2CC60C1EE72A2AFC88218C" blockId="90.[151,1437,1511,2005]" lastBlockId="91.[151,1437,151,1910]" lastPageId="91" lastPageNumber="92" pageId="90" pageNumber="91">
|
||
<emphasis id="B95A2A364C2CC60D1EE72A2AFE9D2654" bold="true" box="[199,334,1799,1824]" pageId="90" pageNumber="91">♂ genitalia</emphasis>
|
||
: 366DL, PT (
|
||
<figureCitation id="1315EAA14C2CC60D1FD82A2AFD832654" box="[504,592,1799,1824]" captionStart="FIGURE 29" captionStartId="90.[151,250,1420,1442]" captionTargetBox="[179,1434,202,1356]" captionTargetId="figure@90.[161,1455,193,1385]" captionTargetPageId="90" captionText="FIGURE 29. ♂ genitalia in LV (left), DV (right). A: Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 159, 366 DL). B: Ht. turbans (Oberthür, 1916) (Zahamena; 370 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264626" httpUri="https://zenodo.org/record/264626/files/figure.png" pageId="90" pageNumber="91">Fig. 29</figureCitation>
|
||
A): from LV, uncus fairly straight in line with tegumen in non-deflexed position and recurved down to a sharply hooked point, with a distinct ‘head’ before tip; tegumen about same length as uncus and constriction at fusion line with vinculum only moderate, such that tegumen from LV tapers only slightly from a rectangular frame; tegumen features wide and shallow proximad notch from DV. Gnathos on angled base, flattened distinctly before tip into broadened structure (not nearly as earlike as for example in
|
||
<taxonomicName id="4C2E8DA74C2CC60D1ACE2AB4FABF26C4" box="[1262,1388,1943,1968]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="turbans">
|
||
<emphasis id="B95A2A364C2CC60D1ACE2AB4FABF26C4" box="[1262,1388,1943,1968]" italics="true" pageId="90" pageNumber="91">Ht. turbans</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2E8DA74C2CC60D1EB72A90FE8026A1" box="[151,339,1980,2005]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="angulifascia">
|
||
<emphasis id="B95A2A364C2CC60D1EB72A90FE8026A1" box="[151,339,1980,2005]" italics="true" pageId="90" pageNumber="91">Ht. angulifascia</emphasis>
|
||
</taxonomicName>
|
||
), with drawn out, tapered and pointed tip, most noticeable in DV. Valves fairly stout and uprecurved arms fairly short leading to a ‘bird’s head’ (rather ‘cassowary head’-like, from DV) tip with spinoid setae and with inwardly projecting ‘beak’. Saccus quite short and inflated towards proximad tip and aedeagus about 1/5 longer than valve. Juxta protruding considerably proximad.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C2DC60C1EE72C29FD4D2081" pageId="91" pageNumber="92" type="etymology">
|
||
<paragraph id="8B91F6244C2DC60C1EE72C29FD4D2081" blockId="91.[151,1437,151,1910]" pageId="91" pageNumber="92">
|
||
<emphasis id="B95A2A364C2DC60C1EE72C29FE9D2069" bold="true" box="[199,334,260,285]" pageId="91" pageNumber="92">Etymology.</emphasis>
|
||
A direct reference to the vertiginous waterfall directly below the camp at Bekolosy Mt. (
|
||
<geoCoordinate id="EE1A90E34C2DC60C1EBF2C0AFEF52034" box="[159,294,295,320]" direction="south" orientation="latitude" pageId="91" pageNumber="92" precision="1" value="-14.04514">14.04514°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C2DC60C1F162C0AFE6D2034" box="[310,446,295,320]" direction="east" orientation="longitude" pageId="91" pageNumber="92" precision="1" value="48.29616">48.29616°E</geoCoordinate>
|
||
,
|
||
<quantity id="4CD65BC14C2DC60C1FEE2C05FDFB204B" box="[462,552,296,320]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="91" pageNumber="92" unit="m" value="1130.0">1130 m</quantity>
|
||
, from Google Earth) that marks the lower elevational limit of this species. Returning after a storm at the summit, I narrowly avoided falling into this cascade, when jumping between two rocks, my net bag caught the strength of the current, and it was thanks to the alertness of the camp assistant, who rushed over, and holding on to my leg, ensured I reached the rock on other side (netless, however). The net was even found in the morning among the rocks at the top of the fall. Alludes also to the marvellous Hitchcock film of this name, with its subliminal spiralling motif.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C2DC60C1EE72CD2FF3B2593" pageId="91" pageNumber="92" type="discussion">
|
||
<paragraph id="8B91F6244C2DC60C1EE72CD2FBB52541" blockId="91.[151,1437,151,1910]" pageId="91" pageNumber="92">
|
||
<emphasis id="B95A2A364C2DC60C1EE72CD2FE9F236C" bold="true" box="[199,332,511,536]" pageId="91" pageNumber="92">Discussion.</emphasis>
|
||
No historical museum collections of
|
||
<taxonomicName id="4C2E8DA74C2DC60C1D2C2F2CFC58236C" box="[780,907,513,536]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="vertigo">
|
||
<emphasis id="B95A2A364C2DC60C1D2C2F2CFC58236C" box="[780,907,513,536]" italics="true" pageId="91" pageNumber="92">Ht. vertigo</emphasis>
|
||
</taxonomicName>
|
||
are known. When first found in the field in
|
||
<date id="FF90D0E44C2DC60C1EB72F09FE9F2348" box="[151,332,548,573]" pageId="91" pageNumber="92" value="1994-12">December 1994</date>
|
||
, the species was given a morphospecies number (sp. 61) but was nevertheless thought to represent
|
||
<taxonomicName id="4C2E8DA74C2DC60C1EB72F64FECA2314" box="[151,281,583,608]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="turbans">
|
||
<emphasis id="B95A2A364C2DC60C1EB72F64FECA2314" box="[151,281,583,608]" italics="true" pageId="91" pageNumber="92">Ht. turbans</emphasis>
|
||
</taxonomicName>
|
||
(indeed
|
||
<bibRefCitation id="EFBF8BD54C2DC60C1FA02F65FE282314" author="Lees" box="[384,507,584,608]" pageId="91" pageNumber="92" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees 1997</bibRefCitation>
|
||
: 225
|
||
<figureCitation id="1315EAA14C2DC60C1C1C2F6AFDAD2314" box="[572,638,583,608]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="91" pageNumber="92">Fig. 3</figureCitation>
|
||
K erroneously shows the distribution of that species extending to the northwest of
|
||
<collectingCountry id="F339B6B44C2DC60C1F0C2F41FE6A23F1" box="[300,441,620,645]" name="Madagascar" pageId="91" pageNumber="92">Madagascar</collectingCountry>
|
||
).
|
||
<taxonomicName id="4C2E8DA74C2DC60C1FEB2F40FD9723F0" box="[459,580,621,644]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="vertigo">
|
||
<emphasis id="B95A2A364C2DC60C1FEB2F40FD9723F0" box="[459,580,621,644]" italics="true" pageId="91" pageNumber="92">Ht. vertigo</emphasis>
|
||
</taxonomicName>
|
||
has since been found on the adjacent mountain of Antsatrotro in
|
||
<date id="FF90D0E44C2DC60C1B062F41FF0123DC" pageId="91" pageNumber="92" value="2011-11">November 2011</date>
|
||
. The species is confusable with a number of described taxa. Examined were the STs of
|
||
<taxonomicName id="4C2E8DA74C2DC60C1AC92FA2FEEF23B9" authority="Oberthur 1916" authorityName="Oberthur" authorityYear="1916" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="turbans">
|
||
<emphasis id="B95A2A364C2DC60C1AC92FA2FA4823DC" box="[1257,1435,655,680]" italics="true" pageId="91" pageNumber="92">Culapa turbans</emphasis>
|
||
<bibRefCitation id="EFBF8BD54C2DC60C1EB72F99FEEF23B9" author="Oberthur" box="[151,316,692,717]" pageId="91" pageNumber="92" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür 1916</bibRefCitation>
|
||
</taxonomicName>
|
||
(LT ♂, here designated,
|
||
<figureCitation id="1315EAA14C2DC60C1C7B2F99FD7D23B8" box="[603,686,692,717]" captionStart="FIGURE 28" captionStartId="89.[151,250,1865,1887]" captionTargetBox="[278,1298,207,1823]" captionTargetId="figure@89.[278,1309,193,1844]" captionTargetPageId="89" captionText="FIGURE 28. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; NHMUK 010289173). B: PT ♀ of Ht. vertigo (Mt. Bekolosy; NHMUK 010289175). C: LT ♂ of Culapa turbans Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674780). D: LT ♂ of Culapa curvatula Oberthür, 1916 (‘ Antsianaka et Lac Alaotra’; BMNH (E) # 674784). E: HT ♂ of Culapa sabas Oberthür, 1923 (‘ Tamatave’; BMNH (E) # 674789). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264625" httpUri="https://zenodo.org/record/264625/files/figure.png" pageId="91" pageNumber="92">Fig. 28</figureCitation>
|
||
C, specimen bearing labels “
|
||
<collectingCountry id="F339B6B44C2DC60C1DD02F99FBA923B9" box="[1008,1146,692,717]" name="Madagascar" pageId="91" pageNumber="92">Madagascar</collectingCountry>
|
||
Antsianaka Perrot Freres 2e Semestre 1893|
|
||
<taxonomicName id="4C2E8DA74C2DC60C1F5E2FFAFD422384" authority="Obthr" authorityName="Obthr" box="[382,657,727,752]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="turbans">Culapa turbans Obthr</taxonomicName>
|
||
♂
|
||
<typeStatus id="549548864C2DC60C1CF32FF5FCDA2384" box="[723,777,728,752]" pageId="91" pageNumber="92">type</typeStatus>
|
||
|P.E.L. Viette det. 1968
|
||
<taxonomicName id="4C2E8DA74C2DC60C1A692FFAFA4E2384" authority="Ch. Obthr." authorityName="Ch. Obthr." box="[1097,1437,727,752]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="turbans">
|
||
Culapa turbans
|
||
<collectingCountry id="F339B6B44C2DC60C1B372FFAFAE82384" box="[1303,1339,727,752]" name="Switzerland" pageId="91" pageNumber="92">Ch</collectingCountry>
|
||
. Obthr.
|
||
</taxonomicName>
|
||
<typeStatus id="549548864C2DC60C1EB72FD1FEE82261" box="[151,315,764,789]" pageId="91" pageNumber="92" type="lectotype">LECTOTYPE</typeStatus>
|
||
|supposed “
|
||
<typeStatus id="549548864C2DC60C1F982FD1FE3F2260" box="[440,492,764,788]" pageId="91" pageNumber="92">type</typeStatus>
|
||
” of
|
||
<taxonomicName id="4C2E8DA74C2DC60C1C012FD1FC932261" authority="Oberthur" authorityName="Oberthur" box="[545,832,764,789]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="turbans">Culapa turbans Oberthür</taxonomicName>
|
||
|[copy of
|
||
<bibRefCitation id="EFBF8BD54C2DC60C1D8A2FD1FB872261" author="Oberthur" box="[938,1108,764,789]" pageId="91" pageNumber="92" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür 1916</bibRefCitation>
|
||
, Pl. 356, f. 3050]|BMNH(E) #674780”) and of
|
||
<taxonomicName id="4C2E8DA74C2DC60C1F522E32FD25224C" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" box="[370,758,799,824]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="curvatula">
|
||
<emphasis id="B95A2A364C2DC60C1F522E32FDEF224C" box="[370,572,799,824]" italics="true" pageId="91" pageNumber="92">Culapa curvatula</emphasis>
|
||
<bibRefCitation id="EFBF8BD54C2DC60C1C672E32FD25224C" author="Oberthur" box="[583,758,799,824]" pageId="91" pageNumber="92" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür, 1916</bibRefCitation>
|
||
</taxonomicName>
|
||
(LT, here designated,
|
||
<figureCitation id="1315EAA14C2DC60C1A222E32FB84224C" box="[1026,1111,799,824]" captionStart="FIGURE 28" captionStartId="89.[151,250,1865,1887]" captionTargetBox="[278,1298,207,1823]" captionTargetId="figure@89.[278,1309,193,1844]" captionTargetPageId="89" captionText="FIGURE 28. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; NHMUK 010289173). B: PT ♀ of Ht. vertigo (Mt. Bekolosy; NHMUK 010289175). C: LT ♂ of Culapa turbans Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674780). D: LT ♂ of Culapa curvatula Oberthür, 1916 (‘ Antsianaka et Lac Alaotra’; BMNH (E) # 674784). E: HT ♂ of Culapa sabas Oberthür, 1923 (‘ Tamatave’; BMNH (E) # 674789). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264625" httpUri="https://zenodo.org/record/264625/files/figure.png" pageId="91" pageNumber="92">Fig. 28</figureCitation>
|
||
D, specimen bearing labels: “Antsianaka et lac Alaotra 2e Trimestre 1889 Perrot Freres|[copy of
|
||
<bibRefCitation id="EFBF8BD54C2DC60C1D8C2E69FB862229" author="Oberthur" box="[940,1109,836,861]" pageId="91" pageNumber="92" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür 1916</bibRefCitation>
|
||
, Pl. 356, f. 3057]|BMNH(E) #674784; automatically then, PLT ♂: BMNH(E) #674786; PLT ♀♀: BMNH(E) #674785) and BMNH(E) #674787, with same locality data).
|
||
<taxonomicName id="4C2E8DA74C2DC60C1C3E2EA0FD6022D1" box="[542,691,908,933]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="curvatula">
|
||
<emphasis id="B95A2A364C2DC60C1C3E2EA0FD6022D1" box="[542,691,908,933]" italics="true" pageId="91" pageNumber="92">Ht. curvatula</emphasis>
|
||
</taxonomicName>
|
||
was considered a synonym of
|
||
<taxonomicName id="4C2E8DA74C2DC60C1A282EA0FB5422D1" box="[1032,1159,908,933]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="turbans">
|
||
<emphasis id="B95A2A364C2DC60C1A282EA0FB5422D1" box="[1032,1159,908,933]" italics="true" pageId="91" pageNumber="92">Ht. turbans</emphasis>
|
||
</taxonomicName>
|
||
by Lees
|
||
<emphasis id="B95A2A364C2DC60C1ACC2EA0FAF922D1" box="[1260,1322,908,933]" italics="true" pageId="91" pageNumber="92">et al.,</emphasis>
|
||
2003 (see also below). Also examined (
|
||
<figureCitation id="1315EAA14C2DC60C1FCC2E82FD9322BC" box="[492,576,943,968]" captionStart="FIGURE 28" captionStartId="89.[151,250,1865,1887]" captionTargetBox="[278,1298,207,1823]" captionTargetId="figure@89.[278,1309,193,1844]" captionTargetPageId="89" captionText="FIGURE 28. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; NHMUK 010289173). B: PT ♀ of Ht. vertigo (Mt. Bekolosy; NHMUK 010289175). C: LT ♂ of Culapa turbans Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674780). D: LT ♂ of Culapa curvatula Oberthür, 1916 (‘ Antsianaka et Lac Alaotra’; BMNH (E) # 674784). E: HT ♂ of Culapa sabas Oberthür, 1923 (‘ Tamatave’; BMNH (E) # 674789). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264625" httpUri="https://zenodo.org/record/264625/files/figure.png" pageId="91" pageNumber="92">Fig. 28</figureCitation>
|
||
E) was the HT ♂ of
|
||
<taxonomicName id="4C2E8DA74C2DC60C1D182E82FB5F22BC" authority="Oberthur, 1923" authorityName="Oberthur" authorityYear="1923" box="[824,1164,943,968]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="sabas">
|
||
<emphasis id="B95A2A364C2DC60C1D182E82FC0722BC" box="[824,980,943,968]" italics="true" pageId="91" pageNumber="92">Culapa sabas</emphasis>
|
||
<bibRefCitation id="EFBF8BD54C2DC60C1DFE2E82FB5F22BC" author="Oberthur" box="[990,1164,943,968]" pageId="91" pageNumber="92" refString="Oberthur, C. (1923) Lepidopteres de Madagascar et d'Afrique tropicale. Etudes de Lepidopterologie Comparee, 21, 119 - 155." type="journal article" year="1923">Oberthür, 1923</bibRefCitation>
|
||
</taxonomicName>
|
||
(‘Tamatave’|BMNH(E) #674789), in addition to extensive field material of
|
||
<taxonomicName id="4C2E8DA74C2DC60C1CFA2EF8FC892299" box="[730,858,980,1005]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="turbans">
|
||
<emphasis id="B95A2A364C2DC60C1CFA2EF8FC892299" box="[730,858,980,1005]" italics="true" pageId="91" pageNumber="92">Ht. turbans</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2E8DA74C2DC60C1DB12EF8FC282299" box="[913,1019,980,1005]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="sabas">
|
||
<emphasis id="B95A2A364C2DC60C1DB12EF8FC282299" box="[913,1019,980,1005]" italics="true" pageId="91" pageNumber="92">Ht. sabas</emphasis>
|
||
</taxonomicName>
|
||
across their ranges. As well as being allopatric, the species is clearly separated from
|
||
<typeStatus id="549548864C2DC60C1CE72ED5FCD12564" box="[711,770,1016,1040]" pageId="91" pageNumber="92">types</typeStatus>
|
||
of both species by a consistently differing FWD ocellus colour pattern and in field collected material of those species, by divergent ♂ genitalia.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C2DC60C1EE72912FF3B2593" blockId="91.[151,1437,151,1910]" pageId="91" pageNumber="92">
|
||
<emphasis id="B95A2A364C2DC60C1EE72912FD03252C" bold="true" box="[199,720,1087,1112]" pageId="91" pageNumber="92">Additional information. DNA divergences:</emphasis>
|
||
COI-5P barcode cluster number BOLD:ACD8579. Bekolosy and Antsatrotro specimens have an identical haplotype; the mountains are not far apart. The COI-5P barcode is about 1.06% divergent to
|
||
<taxonomicName id="4C2E8DA74C2DC60C1F9D29A4FDED25D4" box="[445,574,1159,1184]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="turbans">
|
||
<emphasis id="B95A2A364C2DC60C1F9D29A4FDED25D4" box="[445,574,1159,1184]" italics="true" pageId="91" pageNumber="92">Ht. turbans</emphasis>
|
||
</taxonomicName>
|
||
and about 1.7% divergent to
|
||
<taxonomicName id="4C2E8DA74C2DC60C1DB029A4FC2F25D4" box="[912,1020,1159,1184]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="sabas">
|
||
<emphasis id="B95A2A364C2DC60C1DB029A4FC2F25D4" box="[912,1020,1159,1184]" italics="true" pageId="91" pageNumber="92">Ht. sabas</emphasis>
|
||
</taxonomicName>
|
||
with six apparently fixed nucleotide differences to
|
||
<taxonomicName id="4C2E8DA74C2DC60C1F172980FE6425B1" box="[311,439,1196,1221]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="turbans">
|
||
<emphasis id="B95A2A364C2DC60C1F172980FE6425B1" box="[311,439,1196,1221]" italics="true" pageId="91" pageNumber="92">Ht. turbans</emphasis>
|
||
</taxonomicName>
|
||
; all three species share the same cluster number. Sister to
|
||
<taxonomicName id="4C2E8DA74C2DC60C1A642980FF32259C" authority="in Aduse-Poku et al., (2016)" authorityName="in Aduse-Poku et al." authorityYear="2016" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="sabas">
|
||
<emphasis id="B95A2A364C2DC60C1A642980FB7425B1" box="[1092,1191,1196,1221]" italics="true" pageId="91" pageNumber="92">H. sabas</emphasis>
|
||
in
|
||
<bibRefCitation id="EFBF8BD54C2DC60C1AEB2981FF32259C" pageId="91" pageNumber="92" refString="Aduse-Poku, K., Lees, D. C., Brattstrom, O., Kodandaramaiah, U., Collins, S. & Brakefield, P. M., (2016) Molecular phylogeny and generic-level taxonomy of the widespread palaeotropical ' Heteropsis clade' (Nymphalidae: Satyrinae: Mycalesina). Systematic Entomology, in press. http: // dx. doi. org / syen. 12183" type="book">
|
||
Aduse-Poku
|
||
<emphasis id="B95A2A364C2DC60C1B7C2980FA4F25B1" box="[1372,1436,1196,1221]" italics="true" pageId="91" pageNumber="92">et al.,</emphasis>
|
||
(2016)
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C2DC60C1EE729D9FE042759" pageId="91" pageNumber="92" type="description">
|
||
<paragraph id="8B91F6244C2DC60C1EE729D9FCB62403" blockId="91.[151,1437,151,1910]" pageId="91" pageNumber="92">
|
||
<emphasis id="B95A2A364C2DC60C1EE729D9FE382479" bold="true" box="[199,491,1268,1293]" pageId="91" pageNumber="92">Phylogeny/sister species</emphasis>
|
||
: sister species unclear, but closely related both to
|
||
<taxonomicName id="4C2E8DA74C2DC60C1A7A29D8FB182479" box="[1114,1227,1268,1293]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="sabas">
|
||
<emphasis id="B95A2A364C2DC60C1A7A29D8FB182479" box="[1114,1227,1268,1293]" italics="true" pageId="91" pageNumber="92">Ht. sabas</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2E8DA74C2DC60C1B2E29D8FA472479" box="[1294,1428,1268,1293]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="turbans">
|
||
<emphasis id="B95A2A364C2DC60C1B2E29D8FA472479" box="[1294,1428,1268,1293]" italics="true" pageId="91" pageNumber="92">Ht. turbans</emphasis>
|
||
</taxonomicName>
|
||
, according to the COI-5P barcode. A ♀ of the ‘
|
||
<taxonomicName id="4C2E8DA74C2DC60C1CF0283AFCEF2444" box="[720,828,1303,1328]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="curvatula">
|
||
<emphasis id="B95A2A364C2DC60C1CF0283AFCEF2444" box="[720,828,1303,1328]" italics="true" pageId="91" pageNumber="92">curvatula</emphasis>
|
||
</taxonomicName>
|
||
’ phenotype (DL06-1099, BMAD259-15) from NW Zahamena was one nucleotide different (COI-5P) to other barcoded members of
|
||
<taxonomicName id="4C2E8DA74C2DC60C1A7C2810FB092421" box="[1116,1242,1340,1365]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="turbans">
|
||
<emphasis id="B95A2A364C2DC60C1A7C2810FB092421" box="[1116,1242,1340,1365]" italics="true" pageId="91" pageNumber="92">H. turbans</emphasis>
|
||
</taxonomicName>
|
||
(cluster number BOLD:ACD8579), and so the above synonymy appears correct.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C2DC60C1EE728A9FE2524E9" blockId="91.[151,1437,151,1910]" box="[199,502,1412,1437]" pageId="91" pageNumber="92">
|
||
<emphasis id="B95A2A364C2DC60C1EE728A9FE2524E9" bold="true" box="[199,502,1412,1437]" pageId="91" pageNumber="92">Ecology and distribution.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C2DC60C1EE7288AFDB824B4" blockId="91.[151,1437,151,1910]" box="[199,619,1447,1472]" pageId="91" pageNumber="92">
|
||
<emphasis id="B95A2A364C2DC60C1EE7288AFEF024B4" bold="true" box="[199,291,1447,1472]" pageId="91" pageNumber="92">Habitat</emphasis>
|
||
: primary montane rainforest.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C2DC60C1EE728E1FBD02490" blockId="91.[151,1437,151,1910]" box="[199,1027,1484,1509]" pageId="91" pageNumber="92">
|
||
<emphasis id="B95A2A364C2DC60C1EE728E1FE902491" bold="true" box="[199,323,1484,1509]" pageId="91" pageNumber="92">Behaviour</emphasis>
|
||
: one was caught in a low hung fruit trap at the waterfall camp.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C2DC60C1EE728C2FE6F277C" blockId="91.[151,1437,151,1910]" box="[199,444,1519,1544]" pageId="91" pageNumber="92">
|
||
<emphasis id="B95A2A364C2DC60C1EE728C2FEEE277C" bold="true" box="[199,317,1519,1544]" pageId="91" pageNumber="92">Hostplant</emphasis>
|
||
: unknown.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C2DC60C1EE72B39FE042759" blockId="91.[151,1437,151,1910]" box="[199,471,1556,1581]" pageId="91" pageNumber="92">
|
||
<emphasis id="B95A2A364C2DC60C1EE72B39FE8B2759" bold="true" box="[199,344,1556,1581]" pageId="91" pageNumber="92">Early stages</emphasis>
|
||
: unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C2DC60C1EE72B1AFEDA27C9" pageId="91" pageNumber="92" type="distribution">
|
||
<paragraph id="8B91F6244C2DC60C1EE72B1AFE5C2701" blockId="91.[151,1437,151,1910]" pageId="91" pageNumber="92">
|
||
<emphasis id="B95A2A364C2DC60C1EE72B1AFE8A2724" bold="true" box="[199,345,1591,1616]" pageId="91" pageNumber="92">Distribution</emphasis>
|
||
: as far as is known endemic to Réserve Speciale de Manongarivo, Mts. Bekolosy and Antsatrotro (
|
||
<figureCitation id="1315EAA14C2DC60C1EBF2B71FF212700" box="[159,242,1628,1653]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="91" pageNumber="92">Fig. 30</figureCitation>
|
||
B, fawn dots).
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C2DC60C1EE72B52FEDA27C9" blockId="91.[151,1437,151,1910]" pageId="91" pageNumber="92">
|
||
<emphasis id="B95A2A364C2DC60C1EE72B52FE4827EC" bold="true" box="[199,411,1663,1688]" pageId="91" pageNumber="92">Elevational range</emphasis>
|
||
:
|
||
<quantity id="4CD65BC14C2DC60C1F8C2BADFD9F27E3" box="[428,588,1664,1688]" metricMagnitude="3" metricUnit="m" metricValue="1.27" metricValueMax="1.41" metricValueMin="1.13" pageId="91" pageNumber="92" unit="m" value="1270.0" valueMax="1410.0" valueMin="1130.0">1130–1410 m</quantity>
|
||
. (n=45; including referred specimens and observational data, such as fruit trap data).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C2DC60C1EE72BE7FC8B2602" pageId="91" pageNumber="92" type="materials_examined">
|
||
<paragraph id="8B91F6244C2DC60C1EE72BE7FC8B2602" blockId="91.[151,1437,151,1910]" pageId="91" pageNumber="92">
|
||
<emphasis id="B95A2A364C2DC60C1EE72BE7FE692797" bold="true" box="[199,442,1738,1763]" pageId="91" pageNumber="92">Referred specimens.</emphasis>
|
||
♂,
|
||
<collectingCountry id="F339B6B44C2DC60C1FC62BE7FDBE2797" box="[486,621,1738,1763]" name="Madagascar" pageId="91" pageNumber="92">Madagascar</collectingCountry>
|
||
NW, RS Manongarivo, Bekolosy,
|
||
<quantity id="4CD65BC14C2DC60C1DD92BE7FB822796" box="[1017,1105,1738,1763]" metricMagnitude="3" metricUnit="m" metricValue="1.215" pageId="91" pageNumber="92" unit="m" value="1215.0">1215 m</quantity>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C2DC60C1A7F2BE6FB172797" box="[1119,1220,1735,1763]" direction="south" orientation="latitude" pageId="91" pageNumber="92" precision="55" value="-14.034">14.034o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C2DC60C1AF22BE6FAFA2796" box="[1234,1321,1735,1763]" direction="east" orientation="longitude" pageId="91" pageNumber="92" precision="555" value="48.31">48.31o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C2DC60C1B7B2BE6FA4B2797" box="[1371,1432,1738,1763]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="91" pageNumber="92" unit="km" value="1.0">1 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C2DC60C1EB72BC3FE902673" box="[151,323,1774,1799]" metricMagnitude="3" metricUnit="m" metricValue="1.215" metricValueMax="1.265" metricValueMin="1.165" pageId="91" pageNumber="92" unit="m" value="1215.0" valueMax="1265.0" valueMin="1165.0">1215 +/- 50 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C2DC60C1F722BC2FE072673" box="[338,468,1775,1799]" pageId="91" pageNumber="92" value="1994-12-18">18/12/1994</date>
|
||
, 09:30, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C2DC60C1C9A2BC3FCC82673" box="[698,795,1774,1799]" pageId="91" pageNumber="92">DLBEK</collectionCode>
|
||
94_165, 46; ♂, NW, Antsatrotro Mt., RS Manongarivo, [1250] m, near camp,
|
||
<geoCoordinate id="EE1A90E34C2DC60C1FB82A3BFDDF265A" box="[408,524,1810,1838]" direction="south" orientation="latitude" pageId="91" pageNumber="92" precision="5" value="-14.0824">14.0824o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C2DC60C1C3B2A38FD5D2659" box="[539,654,1810,1838]" direction="east" orientation="longitude" pageId="91" pageNumber="92" precision="5" value="48.3663">48.3663o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C2DC60C1CE22A38FCF0265A" box="[706,803,1813,1838]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="91" pageNumber="92" unit="km" value="0.35">0.35 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C2DC60C1D112A3BFC092659" box="[817,986,1813,1838]" metricMagnitude="3" metricUnit="m" metricValue="1.24" metricValueMax="1.265" metricValueMin="1.215" pageId="91" pageNumber="92" unit="m" value="1240.0" valueMax="1265.0" valueMin="1215.0">1240 +/- 25 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C2DC60C1DC82A3BFB8F265A" box="[1000,1116,1814,1838]" pageId="91" pageNumber="92" value="2011-12-02">2/12/2011</date>
|
||
, 14:51, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C2DC60C1B1E2A3BFAB5265A" box="[1342,1382,1814,1838]" pageId="91" pageNumber="92">DL</collectionCode>
|
||
11A- 0001/
|
||
<collectionCode id="ED3F6EE14C2DC60C1EF62A14FEFD2626" box="[214,302,1849,1874]" pageId="91" pageNumber="92">BMAD</collectionCode>
|
||
245-15 [
|
||
<collectionCode id="ED3F6EE14C2DC60C1FA92A17FE1A2626" box="[393,457,1850,1874]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="91" pageNumber="92">DNA</collectionCode>
|
||
barcoded specimen], IA609 [isotope voucher]; ♀, data as above but:
|
||
<date id="FF90D0E44C2DC60C1AC72A17FA892626" box="[1255,1370,1850,1874]" pageId="91" pageNumber="92" value="2011-12-03">3/12/2011</date>
|
||
, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C2DC60C1EFB2A73FED12602" box="[219,258,1886,1910]" pageId="91" pageNumber="92">DL</collectionCode>
|
||
11A-0002/
|
||
<collectionCode id="ED3F6EE14C2DC60C1F582A70FE1C2602" box="[376,463,1885,1910]" pageId="91" pageNumber="92">BMAD</collectionCode>
|
||
244-15 [
|
||
<collectionCode id="ED3F6EE14C2DC60C1C092A73FDBA2602" box="[553,617,1886,1910]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="91" pageNumber="92">DNA</collectionCode>
|
||
barcoded specimen].
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |