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<document id="227388657A1541F26FF18CA1D7F456D3" ID-CLB-Dataset="7899" ID-DOI="10.11646/zootaxa.4938.2.1" ID-GBIF-Dataset="e1daacda-87e9-479f-abeb-2431532b98d2" ID-ISSN="1175-5326" ID-Zenodo-Dep="4563736" ID-ZooBank="DFCB5C2C-1E70-4628-B65A-4E5322C57F3B" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614324477351" checkinUser="plazi" docAuthor="León-Espinosa, Angel De, León-Gonzalez, Jesus Angel De &amp; Gómez-Gutiérrez, Jaime" docDate="2021" docId="038087FFFFDDC640B6A0816B56BDFA2B" docLanguage="en" docName="zootaxa.4938.2.1.pdf" docOrigin="Zootaxa 4938 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pigrogromitus timsanus Calman 1927" docType="treatment" docVersion="8" lastPageNumber="174" masterDocId="FFB9FF87FFC9C657B63784065265FFCE" masterDocTitle="Pycnogonids from marine docks located along the west coast of the Gulf of California, Mexico" masterLastPageNumber="195" masterPageNumber="151" pageNumber="171" updateTime="1698931148356" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CE66B1EA337175AD413429D98C2E43A7">Pycnogonids from marine docks located along the west coast of the Gulf of California, Mexico</mods:title>
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<mods:namePart id="49A19D32B71A8328E85156B23826CF56">León-Espinosa, Angel De</mods:namePart>
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<mods:namePart id="977CFACD366F4105CE11711F1A0CF76B">León-Gonzalez, Jesus Angel De</mods:namePart>
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<mods:namePart id="156AB91B0B73E2A83D88EB66641132B5">Gómez-Gutiérrez, Jaime</mods:namePart>
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<mods:date id="7263ABAB3CDA107CE5B44D1A14492E0A">2021</mods:date>
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<mods:identifier id="6BE92C4AC989D9ADA4524F046571DB10" type="DOI">10.11646/zootaxa.4938.2.1</mods:identifier>
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<treatment id="038087FFFFDDC640B6A0816B56BDFA2B" ID-DOI="http://doi.org/10.5281/zenodo.4579377" ID-GBIF-Taxon="179987445" ID-Zenodo-Dep="4579377" LSID="urn:lsid:plazi:treatment:038087FFFFDDC640B6A0816B56BDFA2B" httpUri="http://treatment.plazi.org/id/038087FFFFDDC640B6A0816B56BDFA2B" lastPageId="23" lastPageNumber="174" pageId="20" pageNumber="171">
<subSubSection id="C3336562FFDDC643B6A0816B500AFA49" box="[151,623,1389,1415]" pageId="20" pageNumber="171" type="nomenclature">
<paragraph id="8B9636E9FFDDC643B6A0816B500AFA49" blockId="20.[151,623,1389,1451]" box="[151,623,1389,1415]" pageId="20" pageNumber="171">
<heading id="D0DE8185FFDDC643B6A0816B500AFA49" bold="true" box="[151,623,1389,1415]" fontSize="11" level="1" pageId="20" pageNumber="171" reason="1">
<taxonomicName id="4C294D6AFFDDC643B6A0816B500AFA49" ID-CoL="4HSBP" authority="Calman, 1927" authorityName="Calman" authorityYear="1927" box="[151,623,1389,1415]" class="Pycnogonida" family="Callipallenidae" genus="Pigrogromitus" kingdom="Animalia" order="Pantopoda" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="timsanus">
<emphasis id="B95DEAFBFFDDC643B6A0816B500AFA49" bold="true" box="[151,623,1389,1415]" pageId="20" pageNumber="171">
<emphasis id="B95DEAFBFFDDC643B6A0816B53DFFA49" bold="true" box="[151,442,1389,1415]" italics="true" pageId="20" pageNumber="171">Pigrogromitus timsanus</emphasis>
<bibRefCitation id="EFB84B18FFDDC643B7F6816B500AFA49" author="Calman, W. T." box="[449,623,1389,1415]" pageId="20" pageNumber="171" pagination="403 - 410" refId="ref20734" refString="Calman, W. T. (1927) Report on the Pycnogonida. Zoological results of the Cambridge Expedition to the Suez Canal, 1924. Transactions of the Zoological Society of London, 22 (3), 403 - 410. https: // doi. org / 10.1111 / j. 1096 - 3642.1927. tb 00389. x" type="journal article" year="1927">Calman, 1927</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3336562FFDDC643B6A0819753F9FA65" box="[151,412,1425,1451]" pageId="20" pageNumber="171" type="description">
<paragraph id="8B9636E9FFDDC643B6A0819753F9FA65" blockId="20.[151,623,1389,1451]" box="[151,412,1425,1451]" pageId="20" pageNumber="171">
<heading id="D0DE8185FFDDC643B6A0819753F9FA65" bold="true" box="[151,412,1425,1451]" fontSize="11" level="1" pageId="20" pageNumber="171" reason="1">
<emphasis id="B95DEAFBFFDDC643B6A0819753F9FA65" bold="true" box="[151,412,1425,1451]" pageId="20" pageNumber="171">
(
<figureCitation id="13122A6CFFDDC643B6A881975299FA65" box="[159,252,1425,1451]" captionStart="FIGURE 10" captionStartId="19.[151,250,1894,1918]" captionTargetBox="[153,1422,194,1860]" captionTargetId="figure-22@19.[151,1435,181,1870]" captionTargetPageId="19" captionText="FIGURE 10. Pigrogromitus timsanus, UANL-FCB-PYCNO-0066, mature male. A, trunk, dorsal view; B, trunk, lateral view; C, third leg; D, tibia 2, tarsus and propodus of third leg, enlarged; E, oviger; F, terminal articles of the oviger, enlarged. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563760" httpUri="https://zenodo.org/record/4563760/files/figure.png" pageId="20" pageNumber="171">Figs. 10</figureCitation>
A–F; 11 A–I)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3336562FFDDC643B6A081D7572CF9C9" pageId="20" pageNumber="171" type="reference_group">
<paragraph id="8B9636E9FFDDC643B6A081D7572CF9C9" blockId="20.[151,1437,1488,1543]" pageId="20" pageNumber="171">
<treatmentCitationGroup id="AB3911C7FFDDC643B6A081D7572CF9C9" pageId="20" pageNumber="171">
<treatmentCitation id="0A8810F8FFDDC643B6A081D7501EFA27" author="Calman, W. T." box="[151,635,1488,1513]" page="410" pageId="20" pageNumber="171" year="1927">
<taxonomicName id="4C294D6AFFDDC643B6A081D7501EFA27" ID-CoL="4HSBP" authority="Calman, 1927: 408 - 410" authorityName="Calman" authorityPageNumber="408 - 410" authorityYear="1927" box="[151,635,1488,1513]" class="Pycnogonida" family="Callipallenidae" genus="Pigrogromitus" kingdom="Animalia" order="Pantopoda" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="timsanus">
<emphasis id="B95DEAFBFFDDC643B6A081D753E1FA26" box="[151,388,1489,1512]" italics="true" pageId="20" pageNumber="171">Pigrogromitus timsanus</emphasis>
<bibRefCitation id="EFB84B18FFDDC643B7BD81D6501EFA27" author="Calman, W. T." box="[394,635,1488,1513]" pageId="20" pageNumber="171" pagination="403 - 410" refId="ref20734" refString="Calman, W. T. (1927) Report on the Pycnogonida. Zoological results of the Cambridge Expedition to the Suez Canal, 1924. Transactions of the Zoological Society of London, 22 (3), 403 - 410. https: // doi. org / 10.1111 / j. 1096 - 3642.1927. tb 00389. x" type="journal article" year="1927">Calman, 1927: 408–410</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 104 a–f;
<treatmentCitation id="0A8810F8FFDDC643B53281D6566FFA26" author="Hedgpeth, J. W." box="[773,1034,1488,1513]" page="214" pageId="20" pageNumber="171" year="1948">
<bibRefCitation id="EFB84B18FFDDC643B53281D6566FFA26" author="Hedgpeth, J. W." box="[773,1034,1488,1513]" pageId="20" pageNumber="171" pagination="157 - 342" refId="ref21840" refString="Hedgpeth, J. W. (1948) The Pycnogonida of the western North Atlantic and the Caribbean. Proceedings of the United States National Museum, 97 (3216), 157 - 342. https: // doi. org / 10.5479 / si. 00963801.97 - 3216.157" type="journal article" year="1948">Hedgpeth, 1948: 214–216</bibRefCitation>
</treatmentCitation>
, fig. 23;
<treatmentCitation id="0A8810F8FFDDC643B25681D65730FA26" author="Stock, J. H." box="[1121,1365,1488,1512]" page="1015" pageId="20" pageNumber="171" year="1975">
<bibRefCitation id="EFB84B18FFDDC643B25681D65730FA26" author="Stock, J. H." box="[1121,1365,1488,1512]" pageId="20" pageNumber="171" pagination="957 - 1092" refId="ref23938" refString="Stock, J. H. (1975) Pycnogonida from the continental shelf, slope, and deep sea of the tropical Atlantic and East Pacific. Biological results of the University of Miami deep-sea expeditions, 108. Bulletin of Marine Science, 24 (4), 957 - 1092." type="journal article" year="1975">Stock, 1975: 1015–1116</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8810F8FFDDC643B36881D65347F9C9" author="Child, C. A." page="46" pageId="20" pageNumber="171" year="1979">
<bibRefCitation id="EFB84B18FFDDC643B36881D65347F9C9" author="Child, C. A." pageId="20" pageNumber="171" pagination="1 - 86" refId="ref20895" refString="Child, C. A. (1979). Shallow-water Pycnogonida of the Isthmus of Panama and the coasts of middle America. Smithsonian Contributions to Zoology, 293, 1 - 86. https: // doi. org / 10.5479 / si. 00810282.293" type="journal article" year="1979">Child, 1979: 46</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8810F8FFDDC643B71C81E95381F9C9" author="Staples, D. A." box="[299,484,1519,1543]" page="457" pageId="20" pageNumber="171" year="1982">
<bibRefCitation id="EFB84B18FFDDC643B71C81E95381F9C9" author="Staples, D. A." box="[299,484,1519,1543]" pageId="20" pageNumber="171" pagination="455 - 471" refId="ref23796" refString="Staples, D. A. (1982) Pycnogonida of the Calliope River and Auckland Creek, Queensland. Memoirs of the Queensland Museum, 20 (3), 455 - 471." type="journal article" year="1982">Staples, 1982: 457</bibRefCitation>
</treatmentCitation>
, fig. 2 G–J;
<treatmentCitation id="0A8810F8FFDDC643B46981E9509EF9C9" author="Child, C. A." box="[606,763,1519,1543]" page="59" pageId="20" pageNumber="171" year="1992">
<bibRefCitation id="EFB84B18FFDDC643B46981E9509EF9C9" author="Child, C. A." box="[606,763,1519,1543]" pageId="20" pageNumber="171" pagination="1 - 86" refId="ref20980" refString="Child, C. A. (1992) Shallow-water Pycnogonida of the Gulf of Mexico. Florida Marine Research Institute, Memories Hourglass Cruises, 9 (1), 1 - 86." type="journal article" year="1992">Child, 1992: 59</bibRefCitation>
</treatmentCitation>
, 64, fig. 29.
<treatmentCitation id="0A8810F8FFDDC643B54181E95760F9C9" author="Hilton, W. A." box="[886,1285,1519,1543]" page="53" pageId="20" pageNumber="171" year="1942">
<taxonomicName id="4C294D6AFFDDC643B54181E95760F9C9" ID-CoL="WBRS" authority="Hilton, 1942 d: 52 - 53" authorityName="Hilton" authorityPageNumber="52 - 53" authorityYear="1942" box="[886,1285,1519,1543]" class="Pycnogonida" family="Callipallenidae" genus="Clotenopsa" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="prima">
<emphasis id="B95DEAFBFFDDC643B54181E9564CF9C9" box="[886,1065,1519,1543]" italics="true" pageId="20" pageNumber="171">Clotenopsa prima</emphasis>
<bibRefCitation id="EFB84B18FFDDC643B21981E95760F9C9" author="Hilton, W. A." box="[1070,1285,1519,1543]" pageId="20" pageNumber="171" pagination="43 - 55" refId="ref22191" refString="Hilton, W. A. (1942 d) Pycnogonids from Hawaii. Occasional Papers of the Bernice P. Bishop Museum, Hawaii, 17 (3), 43 - 55." type="journal article" year="1942">Hilton, 1942d: 52–53</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 8.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3336562FFDDC643B6A0823650A8F9A0" pageId="20" pageNumber="171" type="materials_examined">
<paragraph id="8B9636E9FFDDC643B6A0823650A8F9A0" blockId="20.[151,1437,1584,1971]" pageId="20" pageNumber="171">
<emphasis id="B95DEAFBFFDDC643B6A0823653E3F984" bold="true" box="[151,390,1584,1610]" pageId="20" pageNumber="171">Material examined.</emphasis>
<materialsCitation id="3B413CB4FFDDC643B7A6823750ADF9A0" ID-GBIF-Occurrence="3045071307" collectingDate="2017-06-26" collectorName="La Paz" latitude="24.154722" location="Baja California Sur" longLatPrecision="21" longitude="-117.32528" pageId="20" pageNumber="171" specimenCount="1" stateProvince="Baja California Sur">
Two specimens: 2 ³, (UANL-FCB-PYCNO-0066),
<collectorName id="26DC533FFFDDC643B5C382375620F985" box="[1012,1093,1585,1611]" pageId="20" pageNumber="171">La Paz</collectorName>
marina,
<collectingRegion id="49EDF80BFFDDC643B284823657FDF984" box="[1203,1432,1584,1610]" country="Mexico" name="Baja California Sur" pageId="20" pageNumber="171">Baja California Sur</collectingRegion>
,
<geoCoordinate id="EE1D502EFFDDC643B6A08252534CF9A0" box="[151,297,1620,1646]" degrees="24" direction="north" minutes="09" orientation="latitude" pageId="20" pageNumber="171" precision="15" seconds="17" value="24.154722">24°09´17´´N</geoCoordinate>
, -
<geoCoordinate id="EE1D502EFFDDC643B70A82525384F9A0" box="[317,481,1620,1646]" degrees="117" direction="west" minutes="19" orientation="longitude" pageId="20" pageNumber="171" precision="15" seconds="31" value="-117.32528">117°19´31´´W</geoCoordinate>
,
<date id="FF971029FFDDC643B7DB82525009F9A0" box="[492,620,1620,1646]" pageId="20" pageNumber="171" value="2017-06-26">
<collectingDate id="EFD3E9C1FFDDC643B7DB82525009F9A0" box="[492,620,1620,1646]" pageId="20" pageNumber="171" value="2017-06-26">26/06/2017</collectingDate>
</date>
(
<figureCitation id="13122A6CFFDDC643B44C825250D9F9A0" box="[635,700,1620,1646]" captionStart="FIGURE 1" captionStartId="3.[151,250,1426,1450]" captionTargetBox="[473,1106,721,1395]" captionTargetId="figure-199@3.[473,1113,718,1400]" captionTargetPageId="3" captionText="FIGURE 1. Area of study showing the nine marine docks located along the peninsular coast of the Gulf of California, Mexico. 1 = Cabo San Lucas; 2 = Puerto Los Cabos; 3 = FONATUR-Fidepaz; 4 = La Paz; 5 = Fiscal Pier, 6 = Palmira, 7 = Cantamar; 8 = Puerto Escondido (located about 50 km south of Loreto), and 9 = FONATUR-Santa Rosalía, Baja California Sur, Mexico. A magnification of the map shown marine docks 3–7 located at Bahía de La Paz." figureDoi="http://doi.org/10.5281/zenodo.4563740" httpUri="https://zenodo.org/record/4563740/files/figure.png" pageId="20" pageNumber="171">Fig. 1</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3336562FFDDC640B6F0827E5631FC6B" lastPageId="23" lastPageNumber="174" pageId="20" pageNumber="171" type="description">
<paragraph id="8B9636E9FFDDC643B6F0827E537CF87D" blockId="20.[151,1437,1584,1971]" pageId="20" pageNumber="171">
<emphasis id="B95DEAFBFFDDC643B6F0827E533CF95C" bold="true" box="[199,345,1656,1682]" pageId="20" pageNumber="171">Description.</emphasis>
Body thick, trunk completely segmented, oval; distal end of the cephalon rounded (
<figureCitation id="13122A6CFFDDC643B2CD827E5700F95C" box="[1274,1381,1656,1682]" captionStart="FIGURE 10" captionStartId="19.[151,250,1894,1918]" captionTargetBox="[153,1422,194,1860]" captionTargetId="figure-22@19.[151,1435,181,1870]" captionTargetPageId="19" captionText="FIGURE 10. Pigrogromitus timsanus, UANL-FCB-PYCNO-0066, mature male. A, trunk, dorsal view; B, trunk, lateral view; C, third leg; D, tibia 2, tarsus and propodus of third leg, enlarged; E, oviger; F, terminal articles of the oviger, enlarged. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563760" httpUri="https://zenodo.org/record/4563760/files/figure.png" pageId="20" pageNumber="171">Fig. 10 A</figureCitation>
;
<figureCitation id="13122A6CFFDDC643B347827E5296F979" captionStart="FIGURE 11" captionStartId="21.[151,250,1861,1885]" captionTargetBox="[151,1435,181,1837]" captionTargetId="figure-22@21.[151,1436,181,1837]" captionTargetPageId="21" captionText="FIGURE 11. Pigrogromitus timsanus, UANL-FCB-PYCNO-0066, mature male:A, cephalon, dorsal view; B, proboscis, frontal view; C, ocular tubercle, latero-dorsal view; D, complete specimen, dorsal view; E, third leg; F, tibia 2, tarsus and propodus, enlarged; G, oviger; H, terminal articles of oviger, enlarged; I, abdomen, dorsal view. Scale bars: D = 1 mm; A, G = 0.5 mm; B, E = 0.3 mm; C, F, I = 0.2 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563762" httpUri="https://zenodo.org/record/4563762/files/figure.png" pageId="20" pageNumber="171">Fig. 11 A, D</figureCitation>
). Each segment with a dorsal tubercle of the same height as ocular tubercle (
<figureCitation id="13122A6CFFDDC643B260829A56A3F978" box="[1111,1222,1692,1718]" captionStart="FIGURE 10" captionStartId="19.[151,250,1894,1918]" captionTargetBox="[153,1422,194,1860]" captionTargetId="figure-22@19.[151,1435,181,1870]" captionTargetPageId="19" captionText="FIGURE 10. Pigrogromitus timsanus, UANL-FCB-PYCNO-0066, mature male. A, trunk, dorsal view; B, trunk, lateral view; C, third leg; D, tibia 2, tarsus and propodus of third leg, enlarged; E, oviger; F, terminal articles of the oviger, enlarged. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563760" httpUri="https://zenodo.org/record/4563760/files/figure.png" pageId="20" pageNumber="171">Fig. 10 A</figureCitation>
). Lateral processes with two setae each, separated by less than ¼ of their own diameters (
<figureCitation id="13122A6CFFDDC643B58682C65644F914" box="[945,1057,1728,1754]" captionStart="FIGURE 10" captionStartId="19.[151,250,1894,1918]" captionTargetBox="[153,1422,194,1860]" captionTargetId="figure-22@19.[151,1435,181,1870]" captionTargetPageId="19" captionText="FIGURE 10. Pigrogromitus timsanus, UANL-FCB-PYCNO-0066, mature male. A, trunk, dorsal view; B, trunk, lateral view; C, third leg; D, tibia 2, tarsus and propodus of third leg, enlarged; E, oviger; F, terminal articles of the oviger, enlarged. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563760" httpUri="https://zenodo.org/record/4563760/files/figure.png" pageId="20" pageNumber="171">Fig. 10 A</figureCitation>
). Ocular tubercle conical, located mid-cephalon, with four eyes near the base and a pair of apical elongated structures (
<figureCitation id="13122A6CFFDDC643B26982E256A9F930" box="[1118,1228,1764,1790]" captionStart="FIGURE 11" captionStartId="21.[151,250,1861,1885]" captionTargetBox="[151,1435,181,1837]" captionTargetId="figure-22@21.[151,1436,181,1837]" captionTargetPageId="21" captionText="FIGURE 11. Pigrogromitus timsanus, UANL-FCB-PYCNO-0066, mature male:A, cephalon, dorsal view; B, proboscis, frontal view; C, ocular tubercle, latero-dorsal view; D, complete specimen, dorsal view; E, third leg; F, tibia 2, tarsus and propodus, enlarged; G, oviger; H, terminal articles of oviger, enlarged; I, abdomen, dorsal view. Scale bars: D = 1 mm; A, G = 0.5 mm; B, E = 0.3 mm; C, F, I = 0.2 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563762" httpUri="https://zenodo.org/record/4563762/files/figure.png" pageId="20" pageNumber="171">Fig. 11 C</figureCitation>
), and a similar tubercle on each of the following somites (
<figureCitation id="13122A6CFFDDC643B469830E50AFF8ED" box="[606,714,1800,1827]" captionStart="FIGURE 10" captionStartId="19.[151,250,1894,1918]" captionTargetBox="[153,1422,194,1860]" captionTargetId="figure-22@19.[151,1435,181,1870]" captionTargetPageId="19" captionText="FIGURE 10. Pigrogromitus timsanus, UANL-FCB-PYCNO-0066, mature male. A, trunk, dorsal view; B, trunk, lateral view; C, third leg; D, tibia 2, tarsus and propodus of third leg, enlarged; E, oviger; F, terminal articles of the oviger, enlarged. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563760" httpUri="https://zenodo.org/record/4563760/files/figure.png" pageId="20" pageNumber="171">Fig. 10 B</figureCitation>
,
<figureCitation id="13122A6CFFDDC643B4E2830E5169F8ED" box="[725,780,1800,1827]" captionStart="FIGURE 11" captionStartId="21.[151,250,1861,1885]" captionTargetBox="[151,1435,181,1837]" captionTargetId="figure-22@21.[151,1436,181,1837]" captionTargetPageId="21" captionText="FIGURE 11. Pigrogromitus timsanus, UANL-FCB-PYCNO-0066, mature male:A, cephalon, dorsal view; B, proboscis, frontal view; C, ocular tubercle, latero-dorsal view; D, complete specimen, dorsal view; E, third leg; F, tibia 2, tarsus and propodus, enlarged; G, oviger; H, terminal articles of oviger, enlarged; I, abdomen, dorsal view. Scale bars: D = 1 mm; A, G = 0.5 mm; B, E = 0.3 mm; C, F, I = 0.2 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563762" httpUri="https://zenodo.org/record/4563762/files/figure.png" pageId="20" pageNumber="171">11 D</figureCitation>
) (
<figureCitation id="13122A6CFFDDC643B513830E5111F8EC" box="[804,884,1800,1826]" captionStart="FIGURE 10" captionStartId="19.[151,250,1894,1918]" captionTargetBox="[153,1422,194,1860]" captionTargetId="figure-22@19.[151,1435,181,1870]" captionTargetPageId="19" captionText="FIGURE 10. Pigrogromitus timsanus, UANL-FCB-PYCNO-0066, mature male. A, trunk, dorsal view; B, trunk, lateral view; C, third leg; D, tibia 2, tarsus and propodus of third leg, enlarged; E, oviger; F, terminal articles of the oviger, enlarged. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563760" httpUri="https://zenodo.org/record/4563760/files/figure.png" pageId="20" pageNumber="171">Fig. 10</figureCitation>
A–B; 11 B, C). Abdomen three times longer than wide, directed backwards in a straight line, with four dorso-distal spines, the distal pair smaller (
<figureCitation id="13122A6CFFDDC643B28D832A576CF888" box="[1210,1289,1836,1862]" captionStart="FIGURE 10" captionStartId="19.[151,250,1894,1918]" captionTargetBox="[153,1422,194,1860]" captionTargetId="figure-22@19.[151,1435,181,1870]" captionTargetPageId="19" captionText="FIGURE 10. Pigrogromitus timsanus, UANL-FCB-PYCNO-0066, mature male. A, trunk, dorsal view; B, trunk, lateral view; C, third leg; D, tibia 2, tarsus and propodus of third leg, enlarged; E, oviger; F, terminal articles of the oviger, enlarged. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563760" httpUri="https://zenodo.org/record/4563760/files/figure.png" pageId="20" pageNumber="171">Fig. 10</figureCitation>
A–B;
<figureCitation id="13122A6CFFDDC643B378832A52A0F8A5" captionStart="FIGURE 11" captionStartId="21.[151,250,1861,1885]" captionTargetBox="[151,1435,181,1837]" captionTargetId="figure-22@21.[151,1436,181,1837]" captionTargetPageId="21" captionText="FIGURE 11. Pigrogromitus timsanus, UANL-FCB-PYCNO-0066, mature male:A, cephalon, dorsal view; B, proboscis, frontal view; C, ocular tubercle, latero-dorsal view; D, complete specimen, dorsal view; E, third leg; F, tibia 2, tarsus and propodus, enlarged; G, oviger; H, terminal articles of oviger, enlarged; I, abdomen, dorsal view. Scale bars: D = 1 mm; A, G = 0.5 mm; B, E = 0.3 mm; C, F, I = 0.2 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563762" httpUri="https://zenodo.org/record/4563762/files/figure.png" pageId="20" pageNumber="171">Fig. 11 D, I</figureCitation>
). Proboscis cylindrical, thick, gradually widening towards the distal part, approximately 1.8 times longer than wide, directed forward (
<figureCitation id="13122A6CFFDDC643B79F8372539CF840" box="[424,505,1908,1934]" captionStart="FIGURE 10" captionStartId="19.[151,250,1894,1918]" captionTargetBox="[153,1422,194,1860]" captionTargetId="figure-22@19.[151,1435,181,1870]" captionTargetPageId="19" captionText="FIGURE 10. Pigrogromitus timsanus, UANL-FCB-PYCNO-0066, mature male. A, trunk, dorsal view; B, trunk, lateral view; C, third leg; D, tibia 2, tarsus and propodus of third leg, enlarged; E, oviger; F, terminal articles of the oviger, enlarged. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563760" httpUri="https://zenodo.org/record/4563760/files/figure.png" pageId="20" pageNumber="171">Fig. 10</figureCitation>
A–B;
<figureCitation id="13122A6CFFDDC643B473837250D7F840" box="[580,690,1908,1934]" captionStart="FIGURE 11" captionStartId="21.[151,250,1861,1885]" captionTargetBox="[151,1435,181,1837]" captionTargetId="figure-22@21.[151,1436,181,1837]" captionTargetPageId="21" captionText="FIGURE 11. Pigrogromitus timsanus, UANL-FCB-PYCNO-0066, mature male:A, cephalon, dorsal view; B, proboscis, frontal view; C, ocular tubercle, latero-dorsal view; D, complete specimen, dorsal view; E, third leg; F, tibia 2, tarsus and propodus, enlarged; G, oviger; H, terminal articles of oviger, enlarged; I, abdomen, dorsal view. Scale bars: D = 1 mm; A, G = 0.5 mm; B, E = 0.3 mm; C, F, I = 0.2 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563762" httpUri="https://zenodo.org/record/4563762/files/figure.png" pageId="20" pageNumber="171">Fig. 11 A</figureCitation>
). Triangular mouth, upper lip rounded, lateral lips triangle-shaped (
<figureCitation id="13122A6CFFDDC643B697839E536EF87D" box="[160,267,1944,1971]" captionStart="FIGURE 11" captionStartId="21.[151,250,1861,1885]" captionTargetBox="[151,1435,181,1837]" captionTargetId="figure-22@21.[151,1436,181,1837]" captionTargetPageId="21" captionText="FIGURE 11. Pigrogromitus timsanus, UANL-FCB-PYCNO-0066, mature male:A, cephalon, dorsal view; B, proboscis, frontal view; C, ocular tubercle, latero-dorsal view; D, complete specimen, dorsal view; E, third leg; F, tibia 2, tarsus and propodus, enlarged; G, oviger; H, terminal articles of oviger, enlarged; I, abdomen, dorsal view. Scale bars: D = 1 mm; A, G = 0.5 mm; B, E = 0.3 mm; C, F, I = 0.2 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563762" httpUri="https://zenodo.org/record/4563762/files/figure.png" pageId="20" pageNumber="171">Fig. 11 B</figureCitation>
).
</paragraph>
<caption id="DF566661FFDCC642B6A083435036F807" ID-DOI="http://doi.org/10.5281/zenodo.4563762" ID-Zenodo-Dep="4563762" httpUri="https://zenodo.org/record/4563762/files/figure.png" pageId="21" pageNumber="172" startId="21.[151,250,1861,1885]" targetBox="[151,1435,181,1837]" targetPageId="21">
<paragraph id="8B9636E9FFDCC642B6A083435036F807" blockId="21.[151,1437,1861,1993]" pageId="21" pageNumber="172">
<emphasis id="B95DEAFBFFDCC642B6A083435372F893" bold="true" box="[151,279,1861,1885]" pageId="21" pageNumber="172">FIGURE 11</emphasis>
.
<taxonomicName id="4C294D6AFFDCC642B71583405076F893" authorityName="Calman" authorityYear="1927" box="[290,531,1862,1885]" class="Pycnogonida" family="Callipallenidae" genus="Pigrogromitus" kingdom="Animalia" order="Pantopoda" pageId="21" pageNumber="172" phylum="Arthropoda" rank="species" species="timsanus">
<emphasis id="B95DEAFBFFDCC642B71583405076F893" box="[290,531,1862,1885]" italics="true" pageId="21" pageNumber="172">Pigrogromitus timsanus</emphasis>
</taxonomicName>
, UANL-FCB-PYCNO-0066, mature male: A, cephalon, dorsal view; B, proboscis, frontal view; C, ocular tubercle, latero-dorsal view; D, complete specimen, dorsal view; E, third leg; F, tibia 2, tarsus and propodus, enlarged; G, oviger; H, terminal articles of oviger, enlarged; I, abdomen, dorsal view. Scale bars: D = 1 mm; A, G = 0.5 mm; B, E = 0.3 mm; C, F, I = 0.2 mm; H = 0.1 mm.
</paragraph>
</caption>
<caption id="DF566661FFDFC641B6A0835C5289F877" ID-DOI="http://doi.org/10.5281/zenodo.4563764" ID-Zenodo-Dep="4563764" httpUri="https://zenodo.org/record/4563764/files/figure.png" pageId="22" pageNumber="173" startId="22.[151,250,1882,1906]" targetBox="[151,1429,184,1848]" targetPageId="22">
<paragraph id="8B9636E9FFDFC641B6A0835C5289F877" blockId="22.[151,1436,1882,1978]" pageId="22" pageNumber="173">
<emphasis id="B95DEAFBFFDFC641B6A0835C537EF8BC" bold="true" box="[151,283,1882,1906]" pageId="22" pageNumber="173">FIGURE 12</emphasis>
.
<taxonomicName id="4C294D6AFFDFC641B71C835C505EF8BC" baseAuthorityName="Hall" baseAuthorityYear="1913" box="[299,571,1882,1906]" class="Pycnogonida" family="Callipallenidae" genus="Callipallene" kingdom="Animalia" order="Pantopoda" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="californiensis">
<emphasis id="B95DEAFBFFDFC641B71C835C505EF8BC" box="[299,571,1882,1906]" italics="true" pageId="22" pageNumber="173">Callipallene californiensis</emphasis>
</taxonomicName>
, UANL-FCB-PYCNO-0067, male A, trunk, dorsal view; B, trunk, lateral view; C, third leg; D, tarsus, propodus and claws, enlarged; E, oviger; F, chelae, enlarged. Scale bars: C = 1 mm; A–B = 0.5 mm; D–F = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8B9636E9FFDEC640B6F0849152B8FF1B" blockId="23.[151,1437,151,1509]" pageId="23" pageNumber="174">
Short chelifore, two-articled scape, ½ as long as proboscis. Chela with curved fingers (
<figureCitation id="13122A6CFFDEC640B29B8491569AFF7F" box="[1196,1279,151,177]" captionStart="FIGURE 10" captionStartId="19.[151,250,1894,1918]" captionTargetBox="[153,1422,194,1860]" captionTargetId="figure-22@19.[151,1435,181,1870]" captionTargetPageId="19" captionText="FIGURE 10. Pigrogromitus timsanus, UANL-FCB-PYCNO-0066, mature male. A, trunk, dorsal view; B, trunk, lateral view; C, third leg; D, tibia 2, tarsus and propodus of third leg, enlarged; E, oviger; F, terminal articles of the oviger, enlarged. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563760" httpUri="https://zenodo.org/record/4563760/files/figure.png" pageId="23" pageNumber="174">Fig. 10</figureCitation>
A–B;
<figureCitation id="13122A6CFFDEC640B37B849157F9FF7F" box="[1356,1436,151,177]" captionStart="FIGURE 11" captionStartId="21.[151,250,1861,1885]" captionTargetBox="[151,1435,181,1837]" captionTargetId="figure-22@21.[151,1436,181,1837]" captionTargetPageId="21" captionText="FIGURE 11. Pigrogromitus timsanus, UANL-FCB-PYCNO-0066, mature male:A, cephalon, dorsal view; B, proboscis, frontal view; C, ocular tubercle, latero-dorsal view; D, complete specimen, dorsal view; E, third leg; F, tibia 2, tarsus and propodus, enlarged; G, oviger; H, terminal articles of oviger, enlarged; I, abdomen, dorsal view. Scale bars: D = 1 mm; A, G = 0.5 mm; B, E = 0.3 mm; C, F, I = 0.2 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563762" httpUri="https://zenodo.org/record/4563762/files/figure.png" pageId="23" pageNumber="174">Fig. 11</figureCitation>
A–B).
</paragraph>
<paragraph id="8B9636E9FFDEC640B6F084D9533FFF37" blockId="23.[151,1437,151,1509]" box="[199,346,223,249]" pageId="23" pageNumber="174">Palps absent.</paragraph>
<paragraph id="8B9636E9FFDEC640B6F0850550F3FE3B" blockId="23.[151,1437,151,1509]" pageId="23" pageNumber="174">
Oviger with 10 articles.Articles one to five glabrous. First article short. Second and third articles subequal, each two times longer than first one. Fourth and fifth articles longest, 3.5 and 4 times longer than the first article. Sixth article twice as long as the first, with five ventral setae in midline and two latero-distal setae. Seventh article 1.5 times longer than the first, with four dorsal and four ventral setae, plus two latero-distal spines. Eighth article of the same size as the first, with three ventral setae and two lateral setae. Ninth article ½ the size of the first one, with a ventro-distal seta. Tenth article smaller than ninth, with a thick mid-ventral seta. Oviger claw curved, slightly longer than ninth article (
<figureCitation id="13122A6CFFDEC640B75385DD53D0FE3B" box="[356,437,475,501]" captionStart="FIGURE 10" captionStartId="19.[151,250,1894,1918]" captionTargetBox="[153,1422,194,1860]" captionTargetId="figure-22@19.[151,1435,181,1870]" captionTargetPageId="19" captionText="FIGURE 10. Pigrogromitus timsanus, UANL-FCB-PYCNO-0066, mature male. A, trunk, dorsal view; B, trunk, lateral view; C, third leg; D, tibia 2, tarsus and propodus of third leg, enlarged; E, oviger; F, terminal articles of the oviger, enlarged. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563760" httpUri="https://zenodo.org/record/4563760/files/figure.png" pageId="23" pageNumber="174">Fig. 10</figureCitation>
E–F;
<figureCitation id="13122A6CFFDEC640B7CE85DD5022FE3B" box="[505,583,475,501]" captionStart="FIGURE 11" captionStartId="21.[151,250,1861,1885]" captionTargetBox="[151,1435,181,1837]" captionTargetId="figure-22@21.[151,1436,181,1837]" captionTargetPageId="21" captionText="FIGURE 11. Pigrogromitus timsanus, UANL-FCB-PYCNO-0066, mature male:A, cephalon, dorsal view; B, proboscis, frontal view; C, ocular tubercle, latero-dorsal view; D, complete specimen, dorsal view; E, third leg; F, tibia 2, tarsus and propodus, enlarged; G, oviger; H, terminal articles of oviger, enlarged; I, abdomen, dorsal view. Scale bars: D = 1 mm; A, G = 0.5 mm; B, E = 0.3 mm; C, F, I = 0.2 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563762" httpUri="https://zenodo.org/record/4563762/files/figure.png" pageId="23" pageNumber="174">Fig. 11</figureCitation>
G–H).
</paragraph>
<paragraph id="8B9636E9FFDEC640B6F085F95073FD3F" blockId="23.[151,1437,151,1509]" pageId="23" pageNumber="174">
Thick legs. Coxae 1–3 of similar length, 1 and 2 glabrous; coxa 3 with ventro-distal seta. Femur longest, as long as coxae 1 and 2 together, proximally wider, with two mid-ventral setae and a latero-distal seta. Tibia 1 with two dorsal bumps, single seta proximally and distally and two latero-distal setae. Tibia 2 as long as tibia 1, with two smaller dorsal bumps and a seta on each of them, two latero-distal setae, a middle-ventral seta and three ventro-distal setae. Tarsus armed with four ventral spines. Propodus with five ventral sole spines, a latero-distal seta, a mid-dorsal seta and two dorso-distal setae. Main claw ½ the length of the propodus, curved distally, no auxiliary claws (
<figureCitation id="13122A6CFFDEC640B6D186D15352FD3F" box="[230,311,727,753]" captionStart="FIGURE 10" captionStartId="19.[151,250,1894,1918]" captionTargetBox="[153,1422,194,1860]" captionTargetId="figure-22@19.[151,1435,181,1870]" captionTargetPageId="19" captionText="FIGURE 10. Pigrogromitus timsanus, UANL-FCB-PYCNO-0066, mature male. A, trunk, dorsal view; B, trunk, lateral view; C, third leg; D, tibia 2, tarsus and propodus of third leg, enlarged; E, oviger; F, terminal articles of the oviger, enlarged. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563760" httpUri="https://zenodo.org/record/4563760/files/figure.png" pageId="23" pageNumber="174">Fig. 10</figureCitation>
C–D;
<figureCitation id="13122A6CFFDEC640B7B686D153B5FD3F" box="[385,464,727,753]" captionStart="FIGURE 11" captionStartId="21.[151,250,1861,1885]" captionTargetBox="[151,1435,181,1837]" captionTargetId="figure-22@21.[151,1436,181,1837]" captionTargetPageId="21" captionText="FIGURE 11. Pigrogromitus timsanus, UANL-FCB-PYCNO-0066, mature male:A, cephalon, dorsal view; B, proboscis, frontal view; C, ocular tubercle, latero-dorsal view; D, complete specimen, dorsal view; E, third leg; F, tibia 2, tarsus and propodus, enlarged; G, oviger; H, terminal articles of oviger, enlarged; I, abdomen, dorsal view. Scale bars: D = 1 mm; A, G = 0.5 mm; B, E = 0.3 mm; C, F, I = 0.2 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563762" httpUri="https://zenodo.org/record/4563762/files/figure.png" pageId="23" pageNumber="174">Fig. 11</figureCitation>
E–F).
</paragraph>
<paragraph id="8B9636E9FFDEC640B6F086FD532EFC4F" blockId="23.[151,1437,151,1509]" pageId="23" pageNumber="174">
<emphasis id="B95DEAFBFFDEC640B6F086FD5167FCDA" bold="true" box="[199,770,762,789]" pageId="23" pageNumber="174">Measurements of the illustrated specimen (mm)</emphasis>
. Trunk: 1.25 long from the insertion of the proboscis to the insertion of the abdomen, 0.91 wide between the second pair of lateral processes. Proboscis: 0.56 long, 0.32 wide. Third leg: Coxa 1, 0.17; coxa 2, 0.17; coxa 3, 0.17; femur 0.32; tibia 1, 0.26; tibia 2, 0.26; tarsus 0.06; propodus 0.31; claw 0.14.
</paragraph>
<paragraph id="8B9636E9FFDEC640B6A0878D5631FC6B" blockId="23.[151,1437,151,1509]" box="[151,1108,906,934]" pageId="23" pageNumber="174">Oviger: 1) 0.1; 2) 0.2; 3) 0.3; 4) 0.45; 5) 0.5; 6) 0.3; 7) 0.25; 8) 0.1; 9) 0.05; 10) 0.05.</paragraph>
</subSubSection>
<subSubSection id="C3336562FFDEC640B6F087A85357FB6F" pageId="23" pageNumber="174" type="distribution">
<paragraph id="8B9636E9FFDEC640B6F087A85357FB6F" blockId="23.[151,1437,151,1509]" pageId="23" pageNumber="174">
<emphasis id="B95DEAFBFFDEC640B6F087A8533CFC06" bold="true" box="[199,345,942,968]" pageId="23" pageNumber="174">Distribution</emphasis>
.
<taxonomicName id="4C294D6AFFDEC640B75F87A95011FC06" authorityName="Calman" authorityYear="1927" box="[360,628,943,968]" class="Pycnogonida" family="Callipallenidae" genus="Pigrogromitus" kingdom="Animalia" order="Pantopoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="timsanus">
<emphasis id="B95DEAFBFFDEC640B75F87A95011FC06" box="[360,628,943,968]" italics="true" pageId="23" pageNumber="174">Pigrogromitus timsanus</emphasis>
</taxonomicName>
has a circum(sub)tropical distribution (
<bibRefCitation id="EFB84B18FFDEC640B21987A957F4FC07" author="Lucena, R. A. &amp; Christoffersen, M. L." box="[1070,1425,943,969]" pageId="23" pageNumber="174" pagination="135 - 141" refId="ref22583" refString="Lucena, R. A. &amp; Christoffersen, M. L. (2019) New records of Pigrogromitus timsanus Calman, 1927 (Callipallenidae, Pycnogonida) for the South Atlantic. Check List, 15, 135 - 141. https: // doi. org / 10.15560 / 15.1.135" type="journal article" year="2019">Lucena &amp; Christoffersen, 2019</bibRefCitation>
).
<bibRefCitation id="EFB84B18FFDEC640B6A087D55324FC23" author="Calman, W. T." box="[151,321,979,1005]" pageId="23" pageNumber="174" pagination="403 - 410" refId="ref20734" refString="Calman, W. T. (1927) Report on the Pycnogonida. Zoological results of the Cambridge Expedition to the Suez Canal, 1924. Transactions of the Zoological Society of London, 22 (3), 403 - 410. https: // doi. org / 10.1111 / j. 1096 - 3642.1927. tb 00389. x" type="journal article" year="1927">Calman (1927)</bibRefCitation>
described
<taxonomicName id="4C294D6AFFDEC640B78B87D55059FC22" authorityName="Calman" authorityYear="1927" box="[444,572,979,1004]" class="Pycnogonida" family="Callipallenidae" genus="Pigrogromitus" kingdom="Animalia" order="Pantopoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="timsanus">
<emphasis id="B95DEAFBFFDEC640B78B87D55059FC22" box="[444,572,979,1004]" italics="true" pageId="23" pageNumber="174">P. timsanus</emphasis>
</taxonomicName>
from Lake Timsah, on the Nile Delta,
<collectingCountry id="F33E7679FFDEC640B5C287D5565EFC23" box="[1013,1083,979,1005]" name="Egypt" pageId="23" pageNumber="174">Egypt</collectingCountry>
;
<bibRefCitation id="EFB84B18FFDEC640B27087D55762FC23" author="Hedgpeth, J. W." box="[1095,1287,979,1005]" pageId="23" pageNumber="174" pagination="157 - 342" refId="ref21840" refString="Hedgpeth, J. W. (1948) The Pycnogonida of the western North Atlantic and the Caribbean. Proceedings of the United States National Museum, 97 (3216), 157 - 342. https: // doi. org / 10.5479 / si. 00963801.97 - 3216.157" type="journal article" year="1948">Hedgpeth (1948)</bibRefCitation>
expanded its range of distribution to the Gulf of
<collectingCountry id="F33E7679FFDEC640B41487F1501EFBDF" box="[547,635,1015,1041]" name="Mexico" pageId="23" pageNumber="174">Mexico</collectingCountry>
, finding the species in Lake Worth,
<collectingRegion id="49EDF80BFFDEC640B23887F15606FBDF" box="[1039,1123,1015,1041]" country="United States of America" name="Florida" pageId="23" pageNumber="174">Florida</collectingRegion>
,
<collectingCountry id="F33E7679FFDEC640B25887F156CEFBDF" box="[1135,1195,1015,1041]" name="United States of America" pageId="23" pageNumber="174">USA</collectingCountry>
;
<bibRefCitation id="EFB84B18FFDEC640B28187F15722FBDF" author="Stock, J. H." box="[1206,1351,1014,1041]" pageId="23" pageNumber="174" pagination="957 - 1092" refId="ref23938" refString="Stock, J. H. (1975) Pycnogonida from the continental shelf, slope, and deep sea of the tropical Atlantic and East Pacific. Biological results of the University of Miami deep-sea expeditions, 108. Bulletin of Marine Science, 24 (4), 957 - 1092." type="journal article" year="1975">Stock (1975)</bibRefCitation>
reports this species from
<collectingCountry id="F33E7679FFDEC640B753801D506AFBFB" box="[356,527,1051,1077]" name="Curacao Island" pageId="23" pageNumber="174">Curaçao Island</collectingCountry>
and comments that it has a circum-tropical distribution.
<bibRefCitation id="EFB84B18FFDEC640B294801D5750FBFB" author="Child, C. A." box="[1187,1333,1051,1077]" pageId="23" pageNumber="174" pagination="1 - 86" refId="ref20895" refString="Child, C. A. (1979). Shallow-water Pycnogonida of the Isthmus of Panama and the coasts of middle America. Smithsonian Contributions to Zoology, 293, 1 - 86. https: // doi. org / 10.5479 / si. 00810282.293" type="journal article" year="1979">Child (1979)</bibRefCitation>
reported specimens of
<taxonomicName id="4C294D6AFFDEC640B705803953D4FB96" authorityName="Calman" authorityYear="1927" box="[306,433,1087,1112]" class="Pycnogonida" family="Callipallenidae" genus="Pigrogromitus" kingdom="Animalia" order="Pantopoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="timsanus">
<emphasis id="B95DEAFBFFDEC640B705803953D4FB96" box="[306,433,1087,1112]" italics="true" pageId="23" pageNumber="174">P. timsanus</emphasis>
</taxonomicName>
from
<collectingRegion id="49EDF80BFFDEC640B7C1803950B7FB97" box="[502,722,1087,1113]" country="Mexico" name="Baja California Sur" pageId="23" pageNumber="174">Baja California Sur</collectingRegion>
,
<collectingCountry id="F33E7679FFDEC640B4EA80395156FB97" box="[733,819,1087,1113]" name="Mexico" pageId="23" pageNumber="174">Mexico</collectingCountry>
and both coasts of
<collectingCountry id="F33E7679FFDEC640B23A8039560CFB97" box="[1037,1129,1087,1113]" name="Panama" pageId="23" pageNumber="174">Panama</collectingCountry>
. The most recent reports of this species are at Eastern
<collectingCountry id="F33E7679FFDEC640B78C80655047FBB3" box="[443,546,1123,1149]" name="Australia" pageId="23" pageNumber="174">Australia</collectingCountry>
in the Pacific Ocean (
<bibRefCitation id="EFB84B18FFDEC640B52F806551D0FBB3" author="Arango, C. P." box="[792,949,1123,1149]" pageId="23" pageNumber="174" pagination="2723 - 2772" refId="ref20376" refString="Arango, C. P. (2003) Sea spiders (Pycnogonida, Arthropoda) from the Great Barrier Reef, Australia: new species, new records and ecological annotations. Journal of Natural History, 37 (22), 2723 - 2772. https: // doi. org / 10.1080 / 00222930210158771" type="journal article" year="2003">Arango, 2003</bibRefCitation>
) and southern Atlantic (
<bibRefCitation id="EFB84B18FFDEC640B2F780655342FB6F" author="Lucena, R. A. &amp; Christoffersen, M. L." pageId="23" pageNumber="174" pagination="135 - 141" refId="ref22583" refString="Lucena, R. A. &amp; Christoffersen, M. L. (2019) New records of Pigrogromitus timsanus Calman, 1927 (Callipallenidae, Pycnogonida) for the South Atlantic. Check List, 15, 135 - 141. https: // doi. org / 10.15560 / 15.1.135" type="journal article" year="2019">Lucena &amp; Christoffersen, 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3336562FFDEC640B6F080AD56BDFA2B" pageId="23" pageNumber="174" type="discussion">
<paragraph id="8B9636E9FFDEC640B6F080AD56BDFA2B" blockId="23.[151,1437,151,1509]" pageId="23" pageNumber="174">
<emphasis id="B95DEAFBFFDEC640B6F080AD5351FB0B" bold="true" box="[199,308,1195,1221]" pageId="23" pageNumber="174">Remarks</emphasis>
. The current specimens from the Gulf of
<collectingRegion id="49EDF80BFFDEC640B52280AD51E2FB0B" box="[789,903,1195,1221]" country="United States of America" name="California" pageId="23" pageNumber="174">California</collectingRegion>
differ from the
<typeStatus id="5492884BFFDEC640B27680AD56C6FB0B" box="[1089,1187,1195,1221]" pageId="23" pageNumber="174" type="holotype">holotype</typeStatus>
described by
<bibRefCitation id="EFB84B18FFDEC640B37280AD5287FB27" author="Calman, W. T." pageId="23" pageNumber="174" pagination="403 - 410" refId="ref20734" refString="Calman, W. T. (1927) Report on the Pycnogonida. Zoological results of the Cambridge Expedition to the Suez Canal, 1924. Transactions of the Zoological Society of London, 22 (3), 403 - 410. https: // doi. org / 10.1111 / j. 1096 - 3642.1927. tb 00389. x" type="journal article" year="1927">Calman (1927)</bibRefCitation>
in the absence of setae on the first two coxae. Propodus of the
<typeStatus id="5492884BFFDEC640B5AF80C9519FFB27" box="[920,1018,1231,1257]" pageId="23" pageNumber="174" type="holotype">holotype</typeStatus>
presents three times more sole spines than the specimens of the present study. The oviger of the current specimens do not have setae on the first five articles, while the oviger of the
<typeStatus id="5492884BFFDEC640B43681115006FAFF" box="[513,611,1303,1329]" pageId="23" pageNumber="174" type="holotype">holotype</typeStatus>
has four setae on the third article, one on the fourth and five on the fifth article. The rest of the oviger articles of the
<typeStatus id="5492884BFFDEC640B4B1813D508DFA9B" box="[646,744,1339,1365]" pageId="23" pageNumber="174" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C294D6AFFDEC640B539813D51E8FA9A" authorityName="Calman" authorityYear="1927" box="[782,909,1339,1364]" class="Pycnogonida" family="Callipallenidae" genus="Pigrogromitus" kingdom="Animalia" order="Pantopoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="timsanus">
<emphasis id="B95DEAFBFFDEC640B539813D51E8FA9A" box="[782,909,1339,1364]" italics="true" pageId="23" pageNumber="174">P. timsanus</emphasis>
</taxonomicName>
and the specimens from the Gulf of
<collectingRegion id="49EDF80BFFDEC640B31C813D57F8FA9B" box="[1323,1437,1339,1365]" country="United States of America" name="California" pageId="23" pageNumber="174">California</collectingRegion>
do not differ in the number of setae. Another relevant morphological difference is the presence of four spines on the abdomen of the specimens from the Gulf of
<collectingRegion id="49EDF80BFFDEC640B4BF81855098FA53" box="[648,765,1411,1437]" country="United States of America" name="California" pageId="23" pageNumber="174">California</collectingRegion>
; these are absent in the
<typeStatus id="5492884BFFDEC640B2308185560CFA53" box="[1031,1129,1411,1437]" pageId="23" pageNumber="174" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="EFB84B18FFDEC640B24081855776FA53" author="Calman, W. T." box="[1143,1299,1411,1437]" pageId="23" pageNumber="174" pagination="403 - 410" refId="ref20734" refString="Calman, W. T. (1927) Report on the Pycnogonida. Zoological results of the Cambridge Expedition to the Suez Canal, 1924. Transactions of the Zoological Society of London, 22 (3), 403 - 410. https: // doi. org / 10.1111 / j. 1096 - 3642.1927. tb 00389. x" type="journal article" year="1927">Calman 1927</bibRefCitation>
). Except for the setae of the abdomen, all these differences can be attributed to ontogenetic variability among specimens, because the two specimens observed from
<collectingRegion id="49EDF80BFFDEC640B42081CD5095FA2B" box="[535,752,1483,1509]" country="Mexico" name="Baja California Sur" pageId="23" pageNumber="174">Baja California Sur</collectingRegion>
were about 35% smaller than the
<typeStatus id="5492884BFFDEC640B25981CD56B1FA2B" box="[1134,1236,1483,1509]" pageId="23" pageNumber="174" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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