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<document ID-DOI="10.25221/fee.448.1" ID-ISSN="2713-2196" ID-Zenodo-Dep="7166846" approvalRequired="57" approvalRequired_for_textStreams="57" checkinTime="1665256722461" checkinUser="felipe" docAuthor="Khachikov, E. A., Háva, J. &amp; Poushkova, S. V." docDate="2022" docId="03F88337FFF6FFBCFF2F7C484E41F9FE" docLanguage="en" docName="FarEastEntomol.448.1-10.pdf" docOrigin="Far Eastern Entomologist 448" docSource="http://dx.doi.org/10.25221/fee.448.1" docStyle="DocumentStyle:68641B8F4A21450A5F9A74164B730F50.1:FarEastEntomol.2017-.journal_article" docStyleId="68641B8F4A21450A5F9A74164B730F50" docStyleName="FarEastEntomol.2017-.journal_article" docStyleVersion="1" docTitle="Phradonoma villosulum" docType="treatment" docVersion="2" lastPageNumber="4" masterDocId="FFC1FB4FFFF7FFBFFFCC792B4A1CFFD2" masterDocTitle="New records of the skin beetles (Coleoptera: Dermestidae) from Russia, Central Asia and Caucasus, with description of a new species from Kazakhstan" masterLastPageNumber="10" masterPageNumber="1" pageNumber="2" updateTime="1665271042267" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New records of the skin beetles (Coleoptera: Dermestidae) from Russia, Central Asia and Caucasus, with description of a new species from Kazakhstan</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Khachikov, E. A.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Háva, J.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Poushkova, S. V.</mods:namePart>
</mods:name>
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<mods:titleInfo>
<mods:title>Far Eastern Entomologist</mods:title>
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<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-02-01</mods:number>
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<mods:detail type="volume">
<mods:number>448</mods:number>
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<mods:start>1</mods:start>
<mods:end>10</mods:end>
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<mods:url>http://dx.doi.org/10.25221/fee.448.1</mods:url>
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<mods:identifier type="DOI">10.25221/fee.448.1</mods:identifier>
<mods:identifier type="ISSN">2713-2196</mods:identifier>
<mods:identifier type="Zenodo-Dep">7166846</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:03F88337FFF6FFBCFF2F7C484E41F9FE" httpUri="http://treatment.plazi.org/id/03F88337FFF6FFBCFF2F7C484E41F9FE" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
<subSubSection box="[227,718,1379,1403]" pageId="1" pageNumber="2" type="nomenclature">
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<heading box="[227,718,1379,1403]" centered="true" fontSize="9" level="2" pageId="1" pageNumber="2" reason="2">
<taxonomicName authority="(Duftschmid, 1825)" baseAuthorityName="Duftschmid" baseAuthorityYear="1825" box="[227,718,1379,1403]" class="Insecta" family="Dermestidae" genus="Phradonoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="villosulum">
<emphasis bold="true" box="[227,718,1379,1403]" pageId="1" pageNumber="2">
<emphasis bold="true" box="[227,492,1379,1403]" italics="true" pageId="1" pageNumber="2">Phradonoma villosulum</emphasis>
(Duftschmid, 1825)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[227,506,1409,1434]" pageId="1" pageNumber="2" type="description">
<paragraph blockId="1.[227,718,1379,1434]" box="[227,506,1409,1434]" pageId="1" pageNumber="2">
<figureCitation box="[227,388,1409,1434]" captionStart="Figs 112" captionStartId="2.[265,306,1472,1494]" captionTargetBox="[274,1079,415,1454]" captionTargetId="figure-46@2.[257,1096,407,1457]" captionTargetPageId="2" captionText="Figs 112. Antennae and male genitalia of Phradonoma spp. 13, 7, 9, 10 Ph. villosulum:" figureDoi="http://doi.org/10.5281/zenodo.7166850" httpUri="https://zenodo.org/record/7166850/files/figure.png" pageId="1" pageNumber="2">Figs 13, 9, 10</figureCitation>
,
<figureCitation box="[399,469,1410,1434]" captionStart="Figs 1318" captionStartId="3.[265,306,1531,1553]" captionTargetBox="[243,1098,636,1483]" captionTargetId="figure-100@3.[227,1114,625,1498]" captionTargetPageId="3" captionText="Figs 1318. Phradonoma villosulum, ♀. 13 IX genital segment; 14 VIIVIII ster-" figureDoi="http://doi.org/10.5281/zenodo.7166852" httpUri="https://zenodo.org/record/7166852/files/figure.png" pageId="1" pageNumber="2">1318</figureCitation>
,
<figureCitation box="[479,506,1410,1434]" captionStart="Figs 1921" captionStartId="5.[265,306,1552,1574]" captionTargetBox="[260,1084,647,1500]" captionTargetId="figure-122@5.[227,1111,625,1519]" captionTargetPageId="5" captionText="Figs 1921. Phradonoma spp. 19, 20 Ph. beskalensis sp. n.: 19 dorsal aspect of body;" figureDoi="http://doi.org/10.5281/zenodo.7166854" httpUri="https://zenodo.org/record/7166854/files/figure.png" pageId="1" pageNumber="2">21</figureCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="materials_examined">
<paragraph blockId="1.[227,1127,1471,1587]" box="[265,1126,1471,1495]" pageId="1" pageNumber="2">
<materialsCitation box="[265,1126,1471,1495]" collectionCode="MATERIAL" country="Russia" location="Krasnokutskii distr." pageId="1" pageNumber="2" specimenCount="1" stateProvince="Saratov">
<collectionCode box="[265,403,1471,1495]" pageId="1" pageNumber="2">MATERIAL</collectionCode>
EXAMINED.
<emphasis bold="true" box="[565,650,1471,1495]" pageId="1" pageNumber="2">
<collectingCountry box="[565,645,1471,1495]" name="Russia" pageId="1" pageNumber="2">Russia</collectingCountry>
:
</emphasis>
<collectingRegion box="[657,824,1471,1495]" country="Russia" name="Saratov" pageId="1" pageNumber="2">Saratov region</collectingRegion>
:
<location LSID="urn:lsid:plazi:treatment:03F88337FFF6FFBCFF2F7C484E41F9FE:8E8E64FAFFF6FFBEFC8F7C944E3FFA05" box="[835,1059,1471,1495]" country="Russia" name="Krasnokutskii distr." pageId="1" pageNumber="2" stateProvince="Saratov">Krasnokutskii distr.</location>
, Dya-
</materialsCitation>
</paragraph>
<paragraph blockId="1.[227,1127,1471,1587]" box="[227,1127,1501,1526]" pageId="1" pageNumber="2">
kovka vill. env.,
<date box="[416,587,1501,1526]" pageId="1" pageNumber="2" value="2012-06-25" valueMax="2012-06-28" valueMin="2012-06-25">
2528.
<collectionCode box="[491,522,1501,1525]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="1" pageNumber="2">VI</collectionCode>
2012
</date>
,
<specimenCount box="[599,635,1502,1526]" pageId="1" pageNumber="2" type="male">2♂</specimenCount>
, leg. D. Astakhov (
<collectionCode box="[853,905,1502,1526]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="2" type="Museum">ZIN</collectionCode>
);
<collectingRegion box="[925,1123,1501,1525]" country="Russia" name="Volgograd" pageId="1" pageNumber="2">Volgograd region</collectingRegion>
:
</paragraph>
<paragraph blockId="1.[227,1127,1471,1587]" box="[227,1127,1532,1556]" pageId="1" pageNumber="2">
Sarepta [=now part of
<collectingRegion box="[512,632,1532,1556]" country="Russia" name="Volgograd" pageId="1" pageNumber="2">Volgograd</collectingRegion>
],
<specimenCount box="[660,693,1532,1556]" pageId="1" pageNumber="2" type="female">1♀</specimenCount>
, leg. Bekker (
<collectionCode box="[878,928,1532,1556]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="2" type="Museum">ZIN</collectionCode>
);
<collectingRegion box="[957,1123,1532,1556]" country="Russia" name="Rostov" pageId="1" pageNumber="2">Rostov region</collectingRegion>
:
</paragraph>
<paragraph blockId="1.[227,1127,1471,1587]" box="[227,1118,1563,1587]" pageId="1" pageNumber="2">
<materialsCitation box="[227,1114,1563,1587]" collectingDate="2001-07-10" collectionCode="CZ" collectorName="E. Khachikov" location="Sholokhovskii distr." municipality="Veshenskaya" pageId="1" pageNumber="2" specimenCount="1" specimenCount-female="1">
<location LSID="urn:lsid:plazi:treatment:03F88337FFF6FFBCFF2F7C484E41F9FE:8E8E64FAFFF6FFBEFF2F7F304BDCF9E1" box="[227,448,1563,1587]" municipality="Veshenskaya" name="Sholokhovskii distr." pageId="1" pageNumber="2">Sholokhovskii distr.</location>
,
<collectingMunicipality box="[459,602,1563,1587]" pageId="1" pageNumber="2">Veshenskaya</collectingMunicipality>
vill.,
<date box="[665,801,1563,1587]" pageId="1" pageNumber="2" value="2001-07-10">
<collectingDate box="[665,801,1563,1587]" pageId="1" pageNumber="2" value="2001-07-10">10.VII 2001</collectingDate>
</date>
,
<specimenCount box="[811,844,1563,1587]" pageId="1" pageNumber="2" type="female">1♀</specimenCount>
, leg.
<collectorName box="[904,1052,1563,1587]" pageId="1" pageNumber="2">E. Khachikov</collectorName>
(
<collectionCode box="[1068,1107,1563,1587]" pageId="1" pageNumber="2">CZ</collectionCode>
)
</materialsCitation>
;
</paragraph>
<paragraph blockId="2.[227,1127,262,378]" box="[227,1127,262,287]" pageId="2" pageNumber="3">
Tsimlyanskii distr., reserve &quot;Tsimlyansky”, sector 7,
<geoCoordinate box="[797,932,262,287]" degrees="47" direction="north" minutes="55" orientation="latitude" pageId="2" pageNumber="3" precision="15" seconds="59" value="47.933056">47°55ʹ59ʺ N</geoCoordinate>
,
<geoCoordinate box="[943,1075,262,287]" degrees="42" direction="east" minutes="13" orientation="longitude" pageId="2" pageNumber="3" precision="15" seconds="11" value="42.219723">42°13ʹ11ʺ E</geoCoordinate>
, 21
</paragraph>
<paragraph blockId="2.[227,1127,262,378]" pageId="2" pageNumber="3">
<materialsCitation box="[227,961,293,317]" collectionCode="CZ" collectorName="E. Khachikov &amp; S. Poushkova" pageId="2" pageNumber="3" specimenCode="VI 2019" specimenCount="29" specimenCount-female="15" specimenCount-male="14">
25.
<specimenCode box="[258,356,293,317]" collectionCode="VI" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="2" pageNumber="3">VI 2019</specimenCode>
,
<specimenCount box="[371,419,293,317]" pageId="2" pageNumber="3" type="male">14♂</specimenCount>
,
<specimenCount box="[436,483,293,317]" pageId="2" pageNumber="3" type="female">15♀</specimenCount>
, leg.
<collectorName box="[551,702,293,317]" pageId="2" pageNumber="3">E. Khachikov</collectorName>
&amp;
<collectorName box="[745,895,293,317]" pageId="2" pageNumber="3">S. Poushkova</collectorName>
(
<collectionCode box="[914,953,293,317]" pageId="2" pageNumber="3">CZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectionCode="CZ" location="Ust-Donetskii distr." pageId="2" pageNumber="3" specimenCount="1">
<location LSID="urn:lsid:plazi:treatment:03F88337FFF6FFBCFF2F7C484E41F9FE:8E8E64FAFFF5FFBDFC1D780E4B00FE89" name="Ust-Donetskii distr." pageId="2" pageNumber="3">Ust-Donetskii distr.</location>
, Nizhne-Kundruchenskaya vill.,
<date box="[653,711,323,347]" pageId="2" pageNumber="3" value="2019-05-09" valueMax="2019-05-11" valueMin="2019-05-09">911.</date>
</materialsCitation>
<materialsCitation box="[710,1127,323,347]" collectionCode="V" collectorName="E. Khachikov" pageId="2" pageNumber="3" specimenCount="5" specimenCount-female="1" specimenCount-male="4">
<collectionCode box="[710,732,323,347]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="3" type="Museum">V</collectionCode>
2019,
<specimenCount box="[817,852,323,347]" pageId="2" pageNumber="3" type="male">4♂</specimenCount>
,
<specimenCount box="[870,903,323,347]" pageId="2" pageNumber="3" type="female">1♀</specimenCount>
, leg.
<collectorName box="[974,1127,323,347]" pageId="2" pageNumber="3">E. Khachikov</collectorName>
</materialsCitation>
</paragraph>
<paragraph blockId="2.[227,1127,262,378]" box="[227,1123,353,378]" pageId="2" pageNumber="3">
(
<collectionCode box="[235,274,353,377]" pageId="2" pageNumber="3">CZ</collectionCode>
);
<materialsCitation box="[293,775,353,377]" collectingDate="1905-07-18" collectionCode="V" location="Yevpatoria" pageId="2" pageNumber="3" specimenCount="1" specimenCount-female="1" stateProvince="Crimea">
<collectingRegion box="[293,376,353,377]" country="Ukraine" name="Crimea" pageId="2" pageNumber="3">Crimea</collectingRegion>
:
<location LSID="urn:lsid:plazi:treatment:03F88337FFF6FFBCFF2F7C484E41F9FE:8E8E64FAFFF5FFBDFE4D784A4BE6FEAB" box="[385,506,353,377]" name="Yevpatoria" pageId="2" pageNumber="3" stateProvince="Crimea">Yevpatoria</location>
,
<date box="[517,650,353,377]" pageId="2" pageNumber="3" value="1905-07-18">
<collectingDate box="[517,650,353,377]" pageId="2" pageNumber="3" value="1905-07-18">18.VII 1905</collectingDate>
</date>
,
<specimenCount box="[661,694,353,377]" pageId="2" pageNumber="3" type="female">1♀</specimenCount>
, leg.
<collectionCode box="[753,775,354,377]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="3" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation box="[779,801,354,377]" collectingDate="1905-07-18" collectionCode="V" isTailEnumeration="true" location="Yevpatoria" pageId="2" pageNumber="3" specimenCount="1" stateProvince="Crimea">
<collectionCode box="[779,801,354,377]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="3" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation box="[812,1119,353,378]" collectionCode="ZIN" location="Yakovlev" pageId="2" pageNumber="3" specimenCount="1" stateProvince="Crimea">
<location LSID="urn:lsid:plazi:treatment:03F88337FFF6FFBCFF2F7C484E41F9FE:8E8E64FAFFF5FFBDFCE0784A4988FEAB" box="[812,916,353,377]" name="Yakovlev" pageId="2" pageNumber="3" stateProvince="Crimea">Yakovlev</location>
(
<collectionCode box="[931,980,354,378]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="2" pageNumber="3" type="Museum">ZIN</collectionCode>
); Theodosia
</materialsCitation>
,
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7166850" ID-Zenodo-Dep="7166850" box="[265,1127,1472,1494]" httpUri="https://zenodo.org/record/7166850/files/figure.png" pageId="2" pageNumber="3" startId="2.[265,306,1472,1494]" targetBox="[274,1079,415,1454]" targetPageId="2">
<paragraph blockId="2.[227,1127,1472,1629]" box="[265,1127,1472,1494]" pageId="2" pageNumber="3">
Figs 112. Antennae and male genitalia of
<taxonomicName box="[670,852,1472,1494]" class="Insecta" family="Dermestidae" genus="Phradonoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis box="[670,793,1472,1493]" italics="true" pageId="2" pageNumber="3">Phradonoma</emphasis>
spp. 1
</taxonomicName>
3, 7, 9, 10
<taxonomicName baseAuthorityName="Duftschmid" baseAuthorityYear="1825" box="[985,1120,1472,1493]" class="Insecta" family="Dermestidae" genus="Phradonoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="villosulum">
<emphasis box="[985,1120,1472,1493]" italics="true" pageId="2" pageNumber="3">Ph. villosulum</emphasis>
</taxonomicName>
:
</paragraph>
</caption>
<paragraph blockId="2.[227,1127,1472,1629]" box="[227,1127,1499,1521]" pageId="2" pageNumber="3">
1 antennae of male from
<collectingCountry box="[479,551,1499,1521]" name="Austria" pageId="2" pageNumber="3">Austria</collectingCountry>
; 2 the same, Rostov region, 3 the same,
<collectingCountry box="[977,1086,1499,1521]" name="Uzbekistan" pageId="2" pageNumber="3">Uzbekistan</collectingCountry>
; 7
</paragraph>
<paragraph blockId="2.[227,1127,1472,1629]" box="[227,1127,1526,1548]" pageId="2" pageNumber="3">
antennae of female; 9 male genitalia, ventrally; 10 the same, dorsally; 10 4, 12
<emphasis box="[1094,1127,1526,1547]" italics="true" pageId="2" pageNumber="3">Ph.</emphasis>
</paragraph>
<paragraph blockId="2.[227,1127,1472,1629]" box="[227,1126,1553,1575]" pageId="2" pageNumber="3">
<taxonomicName authorityName="Zhantiev" authorityYear="1976" box="[227,326,1553,1574]" class="Insecta" family="Dermestidae" genus="Phradonoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="parthicum">
<emphasis box="[227,326,1553,1574]" italics="true" pageId="2" pageNumber="3">parthicum</emphasis>
</taxonomicName>
: 4 antennae of male; 12 male genitalia; 5, 6, 11
<taxonomicName authority="Khachikov &amp; Háva &amp; Poushkova, 2022" authorityName="Khachikov &amp; Háva &amp; Poushkova" authorityYear="2022" box="[868,1018,1553,1574]" class="Insecta" family="Dermestidae" genus="Phradonoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="beskalensis" status="sp. n.">
<emphasis box="[868,1018,1553,1574]" italics="true" pageId="2" pageNumber="3">Ph. beskalensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1024,1080,1553,1574]" pageId="2" pageNumber="3">
<taxonomicNameLabel box="[1024,1080,1553,1574]" pageId="2" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
: 5
</paragraph>
<paragraph blockId="2.[227,1127,1472,1629]" pageId="2" pageNumber="3">
antennae of male; 6 the same, phase-contrast; 11 male genitalia; 8
<taxonomicName authorityName="Hava" authorityYear="2006" box="[900,1006,1580,1602]" class="Insecta" family="Dermestidae" genus="Phradonoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="nigrum">
<emphasis box="[900,1006,1580,1602]" italics="true" pageId="2" pageNumber="3">Ph. nigrum</emphasis>
</taxonomicName>
, antennae of female.
</paragraph>
<paragraph blockId="3.[227,1127,263,563]" pageId="3" pageNumber="4">
Dvuyakornaya Bay,
<date box="[450,571,263,287]" pageId="3" pageNumber="4" value="1902-07-01">1.VII 1902</date>
,
<specimenCount box="[583,616,263,287]" pageId="3" pageNumber="4" type="male">1♂</specimenCount>
(
<collectionCode box="[632,721,263,287]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="3" pageNumber="4" type="Museum">ZMMU</collectionCode>
);
<materialsCitation collectingDate="1932-07-04" collectorName="Gussakovskii" country="Uzbekistan" location="Aman-Kutan" pageId="3" pageNumber="4" specimenCount="1" specimenCount-male="1">
<emphasis bold="true" box="[741,868,263,287]" pageId="3" pageNumber="4">
<collectingCountry box="[741,868,263,287]" name="Uzbekistan" pageId="3" pageNumber="4">Uzbekistan</collectingCountry>
</emphasis>
:
<location LSID="urn:lsid:plazi:treatment:03F88337FFF6FFBCFF2F7C484E41F9FE:8E8E64FAFFF4FFBCFCB8782C4E1EFECD" box="[884,1026,263,287]" country="Uzbekistan" name="Aman-Kutan" pageId="3" pageNumber="4">Aman-Kutan</location>
,
<locationDeviation location="Samarkand" pageId="3" pageNumber="4">S of Samarkand</locationDeviation>
,
<date box="[337,461,294,318]" pageId="3" pageNumber="4" value="1932-07-04">
<collectingDate box="[337,461,294,318]" pageId="3" pageNumber="4" value="1932-07-04">4.VII 1932</collectingDate>
</date>
,
<specimenCount box="[476,510,294,318]" pageId="3" pageNumber="4" type="male">1♂</specimenCount>
, leg.
<collectorName box="[576,720,294,318]" pageId="3" pageNumber="4">Gussakovskii</collectorName>
</materialsCitation>
(labelled as
<taxonomicName authorityName="Zhantiev" authorityYear="1976" box="[867,1127,294,318]" class="Insecta" family="Dermestidae" genus="Phradonoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="parthicum">
<emphasis box="[867,1127,294,318]" italics="true" pageId="3" pageNumber="4">Phradonoma parthicum</emphasis>
</taxonomicName>
</paragraph>
<paragraph blockId="3.[227,1127,263,563]" box="[227,1127,324,349]" pageId="3" pageNumber="4">
<materialsCitation box="[227,1123,324,349]" collectionCode="ZIN" country="Germany" determinerName="Zhantiev &amp; M. Mroczkowski" location="Germany" pageId="3" pageNumber="4" specimenCount="3" specimenCount-female="3">
Zhant.,
<determinerName box="[312,407,324,348]" pageId="3" pageNumber="4">Zhantiev</determinerName>
det.) (
<collectionCode box="[481,531,325,349]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="3" pageNumber="4" type="Museum">ZIN</collectionCode>
);
<emphasis bold="true" box="[552,660,324,348]" pageId="3" pageNumber="4">
<collectingCountry box="[552,660,324,348]" name="Germany" pageId="3" pageNumber="4">Germany</collectingCountry>
</emphasis>
:
<specimenCount box="[677,711,325,349]" pageId="3" pageNumber="4" type="female">3♀</specimenCount>
, (
<determinerName box="[734,921,324,349]" pageId="3" pageNumber="4">M. Mroczkowski</determinerName>
det., 1958) (
<collectionCode box="[1065,1115,325,349]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="3" pageNumber="4" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
;
</paragraph>
<paragraph blockId="3.[227,1127,263,563]" box="[227,895,355,379]" pageId="3" pageNumber="4">
<materialsCitation box="[227,891,355,379]" collectionCode="ZIN" country="Austria" determinerName="M. Mroczkowski" location="Kraatz" pageId="3" pageNumber="4" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<emphasis bold="true" box="[227,313,355,379]" pageId="3" pageNumber="4">
<collectingCountry box="[227,313,355,379]" name="Austria" pageId="3" pageNumber="4">Austria</collectingCountry>
</emphasis>
:
<location LSID="urn:lsid:plazi:treatment:03F88337FFF6FFBCFF2F7C484E41F9FE:8E8E64FAFFF4FFBCFE8B784F4B8DFEA9" box="[327,401,356,379]" country="Austria" name="Kraatz" pageId="3" pageNumber="4">Kraatz</location>
,
<specimenCount box="[411,446,355,379]" pageId="3" pageNumber="4" type="male">1♂</specimenCount>
,
<specimenCount box="[458,491,355,379]" pageId="3" pageNumber="4" type="female">1♀</specimenCount>
, (
<determinerName box="[513,698,355,379]" pageId="3" pageNumber="4">M. Mroczkowski</determinerName>
det., 1958) (
<collectionCode box="[833,883,355,379]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="3" pageNumber="4" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="3.[227,1127,263,563]" box="[265,1120,386,410]" pageId="3" pageNumber="4">
DISTRIBUTION. Europe,
<collectingCountry box="[568,639,386,410]" name="Russia" pageId="3" pageNumber="4">Russia</collectingCountry>
(European part, Caucasus),
<collectingCountry box="[967,1047,386,410]" name="Turkey" pageId="3" pageNumber="4">Turkey</collectingCountry>
,
<collectingCountry box="[1064,1117,386,410]" name="Syria" pageId="3" pageNumber="4">Syria</collectingCountry>
,
</paragraph>
<paragraph blockId="3.[227,1127,263,563]" pageId="3" pageNumber="4">
<collectingCountry box="[227,273,417,440]" name="Iran" pageId="3" pageNumber="4">Iran</collectingCountry>
,
<collectingCountry box="[284,407,417,441]" name="Kyrgyzstan" pageId="3" pageNumber="4">Kyrgyzstan</collectingCountry>
(Pushkin, 2002, 2019; Háva, 2006, 2015; Khachikov, 2008). New record for
<collectingCountry box="[340,451,449,471]" name="Uzbekistan" pageId="3" pageNumber="4">Uzbekistan</collectingCountry>
.
</paragraph>
<paragraph blockId="3.[227,1127,263,563]" box="[265,1127,478,502]" pageId="3" pageNumber="4">
REMARKS. Specimens recorded from
<collectingRegion box="[687,821,480,502]" country="Russia" name="Rostov" pageId="3" pageNumber="4">Rostov region</collectingRegion>
(Rostov-on-Don and Nedvi-
</paragraph>
<paragraph blockId="3.[227,1127,263,563]" pageId="3" pageNumber="4">
govka) by Pushkin (2019) are absent in the collection of the Department of Zoology of the Academy of Biology and Biotechnology (
<collectionCode box="[742,781,539,563]" pageId="3" pageNumber="4">CZ</collectionCode>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7166852" ID-Zenodo-Dep="7166852" box="[265,1127,1531,1553]" httpUri="https://zenodo.org/record/7166852/files/figure.png" pageId="3" pageNumber="4" startId="3.[265,306,1531,1553]" targetBox="[243,1098,636,1483]" targetPageId="3">
<paragraph blockId="3.[227,1127,1531,1580]" box="[265,1127,1531,1553]" pageId="3" pageNumber="4">
Figs 1318.
<taxonomicName baseAuthorityName="Duftschmid" baseAuthorityYear="1825" box="[391,625,1531,1552]" class="Insecta" family="Dermestidae" genus="Phradonoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="villosulum">
<emphasis box="[391,625,1531,1552]" italics="true" pageId="3" pageNumber="4">Phradonoma villosulum</emphasis>
</taxonomicName>
, ♀. 13 IX genital segment; 14 VIIVIII ster-
</paragraph>
</caption>
<paragraph blockId="3.[227,1127,1531,1580]" box="[227,1117,1558,1580]" pageId="3" pageNumber="4">nites; 15 IX sternite; 16 VIII tergite; 17 sclerites in bursa copulatrix; 18 spermatheca.</paragraph>
</subSubSection>
</treatment>
</document>