301 lines
40 KiB
XML
301 lines
40 KiB
XML
<document id="E77046F5B9D97D78EACE75C71052E529" ID-CLB-Dataset="93655" ID-DOI="10.11646/zootaxa.5162.3.2" ID-GBIF-Dataset="85e6bc60-db9a-4d3a-8a6c-1cd2ee4d9870" ID-ISSN="1175-5326" ID-Zenodo-Dep="6810108" ID-ZooBank="8DBF28CE-4AED-4223-B26A-1B9A37A3947F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1657269105949" checkinUser="plazi" docAuthor="Minkina, Łukasz" docDate="2022" docId="03E71465FFB7B134FF766AE5FCFCFEDD" docLanguage="en" docName="zootaxa.5162.3.2.pdf" docOrigin="Zootaxa 5162 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gilletianus miurai Minkina 2022, new species" docType="treatment" docVersion="6" lastPageNumber="234" masterDocId="FFDE6C1DFFBFB13FFFE16A72FFD3FFC0" masterDocTitle="Studies on subfamily Aphodiinae (Coleoptera: Scarabaeoidea: Scarabaeidae) with description of three new species from the genera Agrilinus Mulsant & Rey, 1869 Gilletianus Balthasar, 1933 and Trichiorhyssemus Clouët, 1901" masterLastPageNumber="242" masterPageNumber="223" pageNumber="231" updateTime="1699356733416" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="D66A46B3DC2EB9DC59DDCBEC5E38DC9A">
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<mods:title id="CD31D6E0636C41EF2D218FC77668F611">Studies on subfamily Aphodiinae (Coleoptera: Scarabaeoidea: Scarabaeidae) with description of three new species from the genera Agrilinus Mulsant & Rey, 1869 Gilletianus Balthasar, 1933 and Trichiorhyssemus Clouët, 1901</mods:title>
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<mods:name id="C6730C44892429BF47A2B80CBFEE6A79" type="personal">
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<mods:roleTerm id="9619C0881F1A334F6751A68F1D5E32FB">Author</mods:roleTerm>
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<mods:namePart id="73B2A87F6F59A24D97DB4A35C0DA68E8">Minkina, Łukasz</mods:namePart>
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<mods:typeOfResource id="BBCCADD36D3CC396B41919817024011D">text</mods:typeOfResource>
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<mods:title id="7A813E183985C95B18182858B34DA60D">Zootaxa</mods:title>
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<mods:part id="FF33994C92BD88EC8CE3670952C1B9AF">
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<mods:date id="329000D6EE011C2F950CABF3E13F73D2">2022</mods:date>
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<mods:number id="9896030EB917FC7FA480506B8B8B7A52">2022-07-07</mods:number>
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<mods:number id="62780A1C51F487459C6E4774EB5F582B">5162</mods:number>
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<mods:number id="0C65117F01FEE5D237E7A1626C771495">3</mods:number>
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<mods:identifier id="24AF6AC67600CDCF55A2D63181C0AE21" type="DOI">10.11646/zootaxa.5162.3.2</mods:identifier>
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<mods:identifier id="3C1CB09CF432D32E035A2243B4C352A6" type="GBIF-Dataset">85e6bc60-db9a-4d3a-8a6c-1cd2ee4d9870</mods:identifier>
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<mods:identifier id="4321F678DEC013D1FF1F6CAC678EEDAB" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E71465FFB7B134FF766AE5FCFCFEDD" ID-DOI="http://doi.org/10.5281/zenodo.6810091" ID-GBIF-Taxon="197216209" ID-Zenodo-Dep="6810091" LSID="urn:lsid:plazi:treatment:03E71465FFB7B134FF766AE5FCFCFEDD" httpUri="http://treatment.plazi.org/id/03E71465FFB7B134FF766AE5FCFCFEDD" lastPageId="11" lastPageNumber="234" pageId="8" pageNumber="231">
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<subSubSection id="C354F6F8FFB7B137FF766AE5FD54FF71" box="[151,647,151,178]" pageId="8" pageNumber="231" type="nomenclature">
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<paragraph id="8BF1A573FFB7B137FF766AE5FD54FF71" blockId="8.[151,647,151,213]" box="[151,647,151,178]" pageId="8" pageNumber="231">
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<heading id="D0B9121FFFB7B137FF766AE5FD54FF71" bold="true" box="[151,647,151,178]" fontSize="11" level="1" pageId="8" pageNumber="231" reason="1">
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<emphasis id="B93A7961FFB7B137FF766AE5FD54FF71" bold="true" box="[151,647,151,178]" pageId="8" pageNumber="231">
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<taxonomicName id="4C4EDEF0FFB7B137FF766AE5FE39FF71" authority="Minkina" authorityName="Minkina" authorityYear="2022" box="[151,490,151,178]" class="Insecta" family="Aphodiidae" genus="Gilletianus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="231" phylum="Arthropoda" rank="species" species="miurai" status="sp. nov.">
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<emphasis id="B93A7961FFB7B137FF766AE5FEA5FF71" bold="true" box="[151,374,151,178]" italics="true" pageId="8" pageNumber="231">Gilletianus miurai</emphasis>
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Minkina
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</taxonomicName>
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,
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<taxonomicNameLabel id="A209C41AFFB7B137FE166AE5FD54FF71" box="[503,647,151,177]" pageId="8" pageNumber="231" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C354F6F8FFB7B137FF766AC9FE76FF15" box="[151,421,187,213]" pageId="8" pageNumber="231" type="description">
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<paragraph id="8BF1A573FFB7B137FF766AC9FE76FF15" blockId="8.[151,647,151,213]" box="[151,421,187,213]" pageId="8" pageNumber="231">
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(
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<figureCitation id="1375B9F6FFB7B137FF7E6AC9FF25FF15" box="[159,246,187,213]" captionStart="FIGURES 13–20" captionStartId="7.[151,264,1726,1750]" captionTargetBox="[151,1434,184,1692]" captionTargetId="figure-15@7.[151,1436,181,1702]" captionTargetPageId="7" captionText="FIGURES 13–20. Heads and pronotum. 13, Trichiorhyssemus immaturus Minkina, new species, holotype, female, head; 14, T. immaturus Minkina, new species, holotype, female, pronotum; 15, Agrilinus matusiaki Minkina, new species, holotype, male, head; 16, Agrilinus inexspectatus (Balthasar, 1935), male, head; 17, Agrilinus striatus (Schmidt, 1910), male, head; 18, Carinaulus meghalayensis (Červenka, 2000), male, head; 19, Gilletianus miurai Minkina, new species, holotype, male, head; 20, Gilletianus sundainsulanus Ochi, Kon & Kawahara, 2014, male, head." figureDoi="http://doi.org/10.5281/zenodo.6810124" httpUri="https://zenodo.org/record/6810124/files/figure.png" pageId="8" pageNumber="231">Figs 19</figureCitation>
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,
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<figureCitation id="1375B9F6FFB7B137FEE36AC9FE99FF15" box="[258,330,187,213]" captionStart="FIGURES 21–23" captionStartId="9.[151,264,1764,1788]" captionTargetBox="[163,1428,203,1723]" captionTargetId="figure-15@9.[151,1436,181,1741]" captionTargetPageId="9" captionText="FIGURES 21–23. Gilletianus miurai Minkina, new species. 21, habitus, holotype, male, dorsal view; 22, habitus, holotype, male, ventral view; 23, habitus, holotype, male, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6810128" httpUri="https://zenodo.org/record/6810128/files/figure.png" pageId="8" pageNumber="231">21–23</figureCitation>
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,
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<figureCitation id="1375B9F6FFB7B137FEB76AC9FE4DFF15" box="[342,414,187,213]" captionStart="FIGURES 36–41" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[238,1348,197,1905]" captionTargetId="figure-15@15.[220,1367,181,1920]" captionTargetPageId="15" captionText="FIGURES 36–41. Aedeagi. 36–38, Gilletianus miurai Minkina, new species, holotype: 36, aedeagus, dorsal view; 37, aedeagus, lateral view; 38, apex of parameres, lateral view. 39–41, Gilletianus sundainsulanus Ochi, Kon & Kawahara, 2014: 39, aedeagus, lateral view; 40, aedeagus, dorsal view; 41, apex of parameres, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6810136" httpUri="https://zenodo.org/record/6810136/files/figure.png" pageId="8" pageNumber="231">36–38</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C354F6F8FFB7B137FF766B71FE3BFEA5" pageId="8" pageNumber="231" type="materials_examined">
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<paragraph id="8BF1A573FFB7B137FF766B71FC8FFEDD" blockId="8.[151,1437,259,2014]" box="[151,860,259,286]" pageId="8" pageNumber="231">
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<materialsCitation id="3B26AF2EFFB7B137FF766B71FC8BFEDD" ID-GBIF-Occurrence="3861601305" box="[151,856,259,286]" collectorName="N Keningau" country="Malaysia" location="Sabah" pageId="8" pageNumber="231" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
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<emphasis id="B93A7961FFB7B137FF766B71FEE5FEDD" bold="true" box="[151,310,259,285]" pageId="8" pageNumber="231">
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<typeStatus id="54F51BD1FFB7B137FF766B71FF01FEDD" box="[151,210,259,285]" pageId="8" pageNumber="231">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F359E5E3FFB7B137FEDD6B71FE76FEDD" box="[316,421,259,285]" name="Malaysia" pageId="8" pageNumber="231">Malaysia</collectingCountry>
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, Borneo,
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<collectingRegion id="498A6B91FFB7B137FDF06B71FD8BFEDD" box="[529,600,259,285]" country="Malaysia" name="Sabah" pageId="8" pageNumber="231">Sabah</collectingRegion>
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,
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<collectorName id="26BBC0A5FFB7B137FD856B76FD3CFEDD" box="[612,751,259,286]" pageId="8" pageNumber="231">N Keningau</collectorName>
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, Bingkor
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BF1A573FFB7B137FF266B55FE3BFEA5" blockId="8.[151,1437,259,2014]" pageId="8" pageNumber="231">
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<emphasis id="B93A7961FFB7B137FF266B55FEA9FE81" bold="true" box="[199,378,295,321]" pageId="8" pageNumber="231">Type material.</emphasis>
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<materialsCitation id="3B26AF2EFFB7B137FE676B55FE3BFEA5" ID-GBIF-Occurrence="3861601310" collectingDate="1996-11-20" collectorName="D. Grimm" country="Malaysia" elevation="450" location="Bingkor" municipality="Borneo" pageId="8" pageNumber="231" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
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<typeStatus id="54F51BD1FFB7B137FE676B55FE3DFE81" box="[390,494,295,321]" pageId="8" pageNumber="231" type="holotype">Holotype</typeStatus>
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(
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<specimenCount id="9D486EFAFFB7B137FDE56B55FDCFFE81" box="[516,540,295,321]" pageId="8" pageNumber="231" type="male">♂</specimenCount>
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): “[
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<collectingCountry id="F359E5E3FFB7B137FDA76B55FD67FE81" box="[582,692,295,321]" name="Malaysia" pageId="8" pageNumber="231">Malaysia</collectingCountry>
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]
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||
<collectingMunicipality id="6B953F09FFB7B137FD266B5AFCCDFE82" box="[711,798,296,322]" pageId="8" pageNumber="231">Borneo</collectingMunicipality>
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:
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<collectingRegion id="498A6B91FFB7B137FCCF6B55FCA6FE81" box="[814,885,295,321]" country="Malaysia" name="Sabah" pageId="8" pageNumber="231">Sabah</collectingRegion>
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,
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||
<location id="8E91F3A8FFB7B137FC646B55FC32FE81" LSID="urn:lsid:plazi:treatment:03E71465FFB7B134FF766AE5FCFCFEDD:8E91F3A8FFB7B137FC646B55FC32FE81" box="[901,993,295,321]" country="Malaysia" municipality="Borneo" name="Bingkor" pageId="8" pageNumber="231" stateProvince="Sabah">Bingkor</location>
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/ N Keningau,
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<quantity id="4CB60896FFB7B137FB406B55FAC8FE82" box="[1185,1307,295,322]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="8" pageNumber="231" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">
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<elevation id="00634240FFB7B137FB406B55FAC8FE82" box="[1185,1307,295,322]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="8" pageNumber="231" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">400-500m</elevation>
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</quantity>
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., at / light,
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<date id="FFF083B3FFB7B137FF766B39FEC4FEA5" box="[151,279,331,357]" pageId="8" pageNumber="231" value="1996-11-20">
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<collectingDate id="EFB47A5BFFB7B137FF766B39FEC4FEA5" box="[151,279,331,357]" pageId="8" pageNumber="231" value="1996-11-20">20.11.1996</collectingDate>
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</date>
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||
, leg. /
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||
<collectorName id="26BBC0A5FFB7B137FE806B3EFE09FEA5" box="[353,474,331,358]" pageId="8" pageNumber="231">D. Grimm</collectorName>
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”.
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C354F6F8FFB7B137FF266B1DFBECF81E" pageId="8" pageNumber="231" type="description">
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<paragraph id="8BF1A573FFB7B137FF266B1DFA57FE11" blockId="8.[151,1437,259,2014]" pageId="8" pageNumber="231">
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<emphasis id="B93A7961FFB7B137FF266B1DFDA3FE49" bold="true" box="[199,624,367,393]" pageId="8" pageNumber="231">
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Description of the
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<typeStatus id="54F51BD1FFB7B137FE546B1DFDCEFE49" box="[437,541,367,393]" pageId="8" pageNumber="231" type="holotype">holotype</typeStatus>
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, male.
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</emphasis>
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Dorsum (
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<figureCitation id="1375B9F6FFB7B137FD0D6B1DFC96FE49" box="[748,837,367,393]" captionStart="FIGURES 21–23" captionStartId="9.[151,264,1764,1788]" captionTargetBox="[163,1428,203,1723]" captionTargetId="figure-15@9.[151,1436,181,1741]" captionTargetPageId="9" captionText="FIGURES 21–23. Gilletianus miurai Minkina, new species. 21, habitus, holotype, male, dorsal view; 22, habitus, holotype, male, ventral view; 23, habitus, holotype, male, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6810128" httpUri="https://zenodo.org/record/6810128/files/figure.png" pageId="8" pageNumber="231">Fig. 21</figureCitation>
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). Body length of
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<typeStatus id="54F51BD1FFB7B137FBFE6B1DFB52FE49" box="[1055,1153,367,393]" pageId="8" pageNumber="231" type="holotype">holotype</typeStatus>
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5.4 mm, elongate-oval, moderately convex, shiny, elytra weakly shiny, brownish with orange-brownish elytra and extermities, on each elytron before apex from third to sixth interval with brownish spot. Elytra glabrous except apical third and sides.
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</paragraph>
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<paragraph id="8BF1A573FFB7B137FF266BA9FABFFD6A" blockId="8.[151,1437,259,2014]" pageId="8" pageNumber="231">
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||
Head (
|
||
<figureCitation id="1375B9F6FFB7B137FEF16BA9FEB2FE35" box="[272,353,475,501]" captionStart="FIGURES 13–20" captionStartId="7.[151,264,1726,1750]" captionTargetBox="[151,1434,184,1692]" captionTargetId="figure-15@7.[151,1436,181,1702]" captionTargetPageId="7" captionText="FIGURES 13–20. Heads and pronotum. 13, Trichiorhyssemus immaturus Minkina, new species, holotype, female, head; 14, T. immaturus Minkina, new species, holotype, female, pronotum; 15, Agrilinus matusiaki Minkina, new species, holotype, male, head; 16, Agrilinus inexspectatus (Balthasar, 1935), male, head; 17, Agrilinus striatus (Schmidt, 1910), male, head; 18, Carinaulus meghalayensis (Červenka, 2000), male, head; 19, Gilletianus miurai Minkina, new species, holotype, male, head; 20, Gilletianus sundainsulanus Ochi, Kon & Kawahara, 2014, male, head." figureDoi="http://doi.org/10.5281/zenodo.6810124" httpUri="https://zenodo.org/record/6810124/files/figure.png" pageId="8" pageNumber="231">Fig. 19</figureCitation>
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) widely trapezoidal, weakly convex, shiny, with trace of microreticulation. Clypeus rather thickly bordered, anteriorly weakly sinuate, laterally widely rounded, not notched before genae, clypeal border with very short and very sparse macrosetae. Genae obtuse, distinctly exceeding eyes, with few relatively long setae. Frontal suture not noticeable, without gibbosities; central epistomal gibbosity low. Clypeus distinctly, doubly punctate: smaller punctures variable in size, densely, somewhat regularly distributed, in basal part coarser, larger punctures with diameter two to four times larger than smaller ones, very sparse, irregularly distributed only in basal part.
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</paragraph>
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<paragraph id="8BF1A573FFB7B137FF2668C1FEC3FC41" blockId="8.[151,1437,259,2014]" pageId="8" pageNumber="231">
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Epipharynx (
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<figureCitation id="1375B9F6FFB7B137FEB868C1FE78FD0D" box="[345,427,691,717]" captionStart="FIGURES 30–35" captionStartId="14.[151,264,1357,1381]" captionTargetBox="[169,1424,194,1323]" captionTargetId="figure-18@14.[151,1436,181,1334]" captionTargetPageId="14" captionText="FIGURES 30–35. Details of the morphology. 30–32, Agrilinus matusiaki Minkina, new species, holotype, male: 30, aedeagus, lateral view; 31, aedeagus, dorsal view; 32, epipharynx.33, Gilletianus miurai Minkina, new species, holotype,male, epipharynx; 34, Trichiorhyssemus immaturus Minkina, new species, holotype, female, epipharynx; 35, T. immaturus Minkina, new species, holotype, female, druseform macrosetae on border of pronotum." figureDoi="http://doi.org/10.5281/zenodo.6810134" httpUri="https://zenodo.org/record/6810134/files/figure.png" pageId="8" pageNumber="231">Fig. 33</figureCitation>
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) transverse, with sides broadly rounded, anterior margin of pedia convex, corypha clearly below of margin of pedia, reduced, with two distinct, large celtes. Mesoepitorma distinctly triangular, regularly widened posteriad. Acanthopariae with dense, short and thick chaetae; acropariae with dense, long and thiner chaetae than on acanthopariae; chaetopariae with dense belt of thin, long chaetae; adelochaetae absent; prophobae with very dense, short and thin setae; chaetopediae with relatively sparse, thick and long chaetae. Tormae relatively thin, short.
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</paragraph>
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<paragraph id="8BF1A573FFB7B137FF2669FEFC01FBBD" blockId="8.[151,1437,259,2014]" pageId="8" pageNumber="231">Pronotum transverse, somewhat wider than base of elytra, widest in the basal part, weakly convex, shiny, with trace of microreticulation, with double punctation: larger punctures irregularly spaced, eight to ten times larger than smaller ones, regular in size, three to six times larger than the smaller ones, relatively sparse, quite denser in basal part and on sides; smaller punctures quite regularly spaced, regular in size, rather dense; sides and base thinly bordered, anteriorly not bordered; sides and base with very short, quite dense setae; anterior angles widely rounded, hind angles obtuse, sides before hind angles very widely rounded, base of pronotum before hind angles very widely sinuate, additionally, somewhat more distinctly sinuate before the middle.</paragraph>
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<paragraph id="8BF1A573FFB7B137FF266EF5FEB3FB05" blockId="8.[151,1437,259,2014]" pageId="8" pageNumber="231">Scutellum small, triangular, with sides widely ogival, with very sparse, very fine punctures, weakly shiny, with microreticulation.</paragraph>
|
||
<paragraph id="8BF1A573FFB7B137FF266EBDFECEFA02" blockId="8.[151,1437,259,2014]" pageId="8" pageNumber="231">Elytra oblong, convex, widest near the middle, weakly shiny, with distinct microreticulation; without humeral denticles; with ten striae and ten intervals. Striae distinctly, densely and rather finely punctate; punctures rather weakly indenting margins of intervals. First and tenth striae joined together before apex; all the rest shortened before apex, sixth and seventh striae slightly shortened before base, eighth striae distinctly shortened before base, ninth and tenth striae joined before base. Intervals quite distinctly convex, punctate, punctures fine, quite dense, create two irregular rows nearby sides of each interval. Punctures on sides and in apical third with thin, moderately long macrosetae.</paragraph>
|
||
<paragraph id="8BF1A573FFB7B137FF266FBEFE0CF95A" blockId="8.[151,1437,259,2014]" pageId="8" pageNumber="231">Legs. Femora shiny, without microreticulation, finely and quite densely punctate. Protibiae distinctly tridentate laterally, proximally with few small teeth; its dorsal side shiny, with few very fine punctures; apical spur long, moderately broad, very distinctly downwardly directed, with obtuse apex. Meso- and metatibiae with two strong transverse carinae, fimbriate apically with row of unequal spinules. Metatibiae superior apical spur distinctly shorter than basimetatarsomere, latter distinctly longer than next three following metatarsomeres combined. Claws rather short, thin, distinctly arcuate.</paragraph>
|
||
<paragraph id="8BF1A573FFB7B137FF266CD6FEB9F97E" blockId="8.[151,1437,259,2014]" box="[199,362,1700,1726]" pageId="8" pageNumber="231">Macropterous.</paragraph>
|
||
<paragraph id="8BF1A573FFB7B137FF266CB5FC17F922" blockId="8.[151,1437,259,2014]" box="[199,964,1735,1762]" pageId="8" pageNumber="231">Pygidium with similar structure to ventrites, except long macrosetae</paragraph>
|
||
<paragraph id="8BF1A573FFB7B137FF266C9EFD2EF8B2" blockId="8.[151,1437,259,2014]" pageId="8" pageNumber="231">
|
||
Venter (
|
||
<figureCitation id="1375B9F6FFB7B137FEC76C99FE52F8C5" box="[294,385,1771,1797]" captionStart="FIGURES 21–23" captionStartId="9.[151,264,1764,1788]" captionTargetBox="[163,1428,203,1723]" captionTargetId="figure-15@9.[151,1436,181,1741]" captionTargetPageId="9" captionText="FIGURES 21–23. Gilletianus miurai Minkina, new species. 21, habitus, holotype, male, dorsal view; 22, habitus, holotype, male, ventral view; 23, habitus, holotype, male, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6810128" httpUri="https://zenodo.org/record/6810128/files/figure.png" pageId="8" pageNumber="231">Fig. 22</figureCitation>
|
||
). Meso-metaventral plate shiny, concave, with distinct longitudinal line at middle; with fine, relatively sparse punctures. Abdominal ventrites rather mat, densely, not coarsely punctate; with distinct microreticulation; punctures in basal third forming row and bearing very long and thin macrosetae; remaining punctures bearing much thiner and shorter macrosetae.
|
||
</paragraph>
|
||
<paragraph id="8BF1A573FFB7B137FF266D09FBECF81E" blockId="8.[151,1437,259,2014]" pageId="8" pageNumber="231">
|
||
Aedeagus (
|
||
<figureCitation id="1375B9F6FFB7B137FEA26D09FE16F855" box="[323,453,1915,1941]" captionStart="FIGURES 36–41" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[238,1348,197,1905]" captionTargetId="figure-15@15.[220,1367,181,1920]" captionTargetPageId="15" captionText="FIGURES 36–41. Aedeagi. 36–38, Gilletianus miurai Minkina, new species, holotype: 36, aedeagus, dorsal view; 37, aedeagus, lateral view; 38, apex of parameres, lateral view. 39–41, Gilletianus sundainsulanus Ochi, Kon & Kawahara, 2014: 39, aedeagus, lateral view; 40, aedeagus, dorsal view; 41, apex of parameres, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6810136" httpUri="https://zenodo.org/record/6810136/files/figure.png" pageId="8" pageNumber="231">Figs 36–38</figureCitation>
|
||
). Length about 3.3 mm, parameres about 0.5 x length of phallobase, regularly constricted till 1/3 from apex, then ventral margin obliquely truncate, in lateral view running as a straight line to apex. In dorsal view with a fringe of quite short, thin macrosetae on inner side of bend; apex acute.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF31F5FBFFB6B136FF766C96FD2FF8E1" ID-DOI="http://doi.org/10.5281/zenodo.6810128" ID-Zenodo-Dep="6810128" httpUri="https://zenodo.org/record/6810128/files/figure.png" pageId="9" pageNumber="232" startId="9.[151,264,1764,1788]" targetBox="[163,1428,203,1723]" targetPageId="9">
|
||
<paragraph id="8BF1A573FFB6B136FF766C96FD2FF8E1" blockId="9.[151,1436,1764,1825]" pageId="9" pageNumber="232">
|
||
<emphasis id="B93A7961FFB6B136FF766C96FE85F93D" bold="true" box="[151,342,1764,1789]" pageId="9" pageNumber="232">FIGURES 21–23.</emphasis>
|
||
<taxonomicName id="4C4EDEF0FFB6B136FEBF6C97FDA8F93D" authority="Minkina" authorityName="Minkina" authorityYear="2022" box="[350,635,1765,1789]" class="Insecta" family="Aphodiidae" genus="Gilletianus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="232" phylum="Arthropoda" rank="species" species="miurai" status="sp. nov.">
|
||
<emphasis id="B93A7961FFB6B136FEBF6C97FDC8F93C" box="[350,539,1765,1789]" italics="true" pageId="9" pageNumber="232">Gilletianus miurai</emphasis>
|
||
Minkina
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A209C41AFFB6B136FD666C97FCD6F93D" box="[647,773,1765,1789]" pageId="9" pageNumber="232" rank="species">new species</taxonomicNameLabel>
|
||
. 21, habitus, holotype, male, dorsal view; 22, habitus, holotype, male, ventral view; 23, habitus, holotype, male, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C354F6F8FFB6B136FF266D3DFF30F84D" pageId="9" pageNumber="232" type="etymology">
|
||
<paragraph id="8BF1A573FFB6B136FF266D3DFF30F84D" blockId="9.[151,1436,1871,2042]" pageId="9" pageNumber="232">
|
||
<emphasis id="B93A7961FFB6B136FF266D3DFE9DF8A9" bold="true" box="[199,334,1871,1897]" pageId="9" pageNumber="232">Etymology.</emphasis>
|
||
The new species is dedicated to the memory of famous Japanese manga cartoonist – Kentaro Miura.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C354F6F8FFB6B134FF266DE5FCFCFEDD" lastPageId="11" lastPageNumber="234" pageId="9" pageNumber="232" type="diagnosis">
|
||
<paragraph id="8BF1A573FFB6B135FF266DE5FD4AFEA5" blockId="9.[151,1436,1871,2042]" lastBlockId="10.[151,1437,151,357]" lastPageId="10" lastPageNumber="233" pageId="9" pageNumber="232">
|
||
<emphasis id="B93A7961FFB6B136FF266DE5FE1FF871" bold="true" box="[199,460,1943,1969]" pageId="9" pageNumber="232">Differential diagnosis.</emphasis>
|
||
In a key to species of
|
||
<bibRefCitation id="EFDFD882FFB6B136FD566DE5FCA9F871" author="Ochi, T. & Kon, M. & Kakizoe, S." box="[695,890,1943,1969]" pageId="9" pageNumber="232" pagination="307 - 320" refId="ref8804" refString="Ochi, T., Kon, M. & Kakizoe, S. (2016) Nine new taxa of the genus Gilletianus from Southeast Asia (Coleoptera: Scarabaeidae). Japanese Journal of Systematic Entomology, 22 (2), 307 - 320." type="journal article" year="2016">
|
||
Ochi
|
||
<emphasis id="B93A7961FFB6B136FD136DEAFCFBF871" box="[754,808,1943,1969]" italics="true" pageId="9" pageNumber="232">et al.</emphasis>
|
||
(2016)
|
||
</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C4EDEF0FFB6B136FC626DE5FB87F871" authority="Minkina" authorityName="Minkina" authorityYear="2022" box="[899,1108,1943,1969]" class="Insecta" family="Aphodiidae" genus="Gilletianus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="232" phylum="Arthropoda" rank="species" species="miurai" status="sp. nov.">
|
||
<emphasis id="B93A7961FFB6B136FC626DE5FC3DF871" box="[899,1006,1943,1969]" italics="true" pageId="9" pageNumber="232">G. miurai</emphasis>
|
||
Minkina
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A209C41AFFB6B136FBBC6DE5FB32F871" box="[1117,1249,1943,1969]" pageId="9" pageNumber="232" rank="species">new species</taxonomicNameLabel>
|
||
can be identified as
|
||
<taxonomicName id="4C4EDEF0FFB6B136FF576DC9FD34F815" authority="Ochi, Kon & Kawahara, 2014" authorityName="Ochi, Kon & Kawahara" authorityYear="2014" box="[182,743,1979,2005]" class="Insecta" family="Aphodiidae" genus="Gilletianus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="232" phylum="Arthropoda" rank="species" species="sundainsulanus">
|
||
<emphasis id="B93A7961FFB6B136FF576DC9FE54F815" box="[182,391,1979,2005]" italics="true" pageId="9" pageNumber="232">G. sundainsulanus</emphasis>
|
||
<bibRefCitation id="EFDFD882FFB6B136FE6E6DC9FD34F815" author="Ochi, T. & Kon, M. & Kawahara, M." box="[399,743,1979,2005]" pageId="9" pageNumber="232" pagination="635 - 648" refId="ref8893" refString="Ochi, T., Kon, M. & Kawahara, M. (2014) Four new species of Aphodiini from Southeast Asia (Coleoptera Scarabaeidae). Giornale Italiano di Entomologia, 13 (59), 635 - 648." type="journal article" year="2014">Ochi, Kon & Kawahara, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
. However aedeagi of both species are distinctly different – in
|
||
<taxonomicName id="4C4EDEF0FFB6B136FF766DADFEBAF839" authorityName="Ochi, Kon & Kawahara" authorityYear="2014" box="[151,361,2015,2041]" class="Insecta" family="Aphodiidae" genus="Gilletianus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="232" phylum="Arthropoda" rank="species" species="sundainsulanus">
|
||
<emphasis id="B93A7961FFB6B136FF766DADFEBAF839" box="[151,361,2015,2041]" italics="true" pageId="9" pageNumber="232">G. sundainsulanus</emphasis>
|
||
</taxonomicName>
|
||
the aedeagus is shorter, length about 2.5 mm and the parameres are relatively much shorter in relation to phallobase (about 0.39 x its length), their apices are less sharply upturned and the oblique truncature is less marked.
|
||
<taxonomicName id="4C4EDEF0FFB5B135FECB6AC9FDB2FF15" authority="Minkina" authorityName="Minkina" authorityYear="2022" box="[298,609,187,213]" class="Insecta" family="Aphodiidae" genus="Gilletianus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="233" phylum="Arthropoda" rank="species" species="miurai" status="sp. nov.">
|
||
<emphasis id="B93A7961FFB5B135FECB6AC9FE2BFF14" box="[298,504,187,213]" italics="true" pageId="10" pageNumber="233">Gilletianus miurai</emphasis>
|
||
Minkina
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A209C41AFFB5B135FD8D6AC9FC97FF15" box="[620,836,187,213]" pageId="10" pageNumber="233" rank="genus">
|
||
new species and
|
||
<taxonomicName id="4C4EDEF0FFB5B135FCC86AC9FC97FF15" authorityName="Ochi, Kon & Kawahara" authorityYear="2014" box="[809,836,187,213]" class="Insecta" family="Aphodiidae" genus="Gilletianus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="233" phylum="Arthropoda" rank="species" species="sundainsulanus">
|
||
<emphasis id="B93A7961FFB5B135FCC86AC9FC97FF15" box="[809,836,187,213]" italics="true" pageId="10" pageNumber="233">G.</emphasis>
|
||
</taxonomicName>
|
||
</taxonomicNameLabel>
|
||
<emphasis id="B93A7961FFB5B135FCAA6AC9FC2AFF15" box="[843,1017,187,213]" italics="true" pageId="10" pageNumber="233">sundainsulanus</emphasis>
|
||
are in my opinion almost identical in external view. The male of
|
||
<taxonomicName id="4C4EDEF0FFB5B135FE246AADFD4AFF39" authority="Minkina" authorityName="Minkina" authorityYear="2022" box="[453,665,223,249]" class="Insecta" family="Aphodiidae" genus="Gilletianus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="233" phylum="Arthropoda" rank="species" species="miurai" status="sp. nov.">
|
||
<emphasis id="B93A7961FFB5B135FE246AADFDE2FF38" box="[453,561,223,249]" italics="true" pageId="10" pageNumber="233">G. miurai</emphasis>
|
||
Minkina
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A209C41AFFB5B135FD426AADFCFAFF39" box="[675,809,223,249]" pageId="10" pageNumber="233" rank="species">new species</taxonomicNameLabel>
|
||
has the basal margin of pronotum near the middle rather more distinctly sinuate, somewhat different hind angles of pronotum and weakly sinuate clypeus anteriorly (when clypeus of
|
||
<taxonomicName id="4C4EDEF0FFB5B135FEF36B55FE31FE81" authorityName="Ochi, Kon & Kawahara" authorityYear="2014" box="[274,482,295,321]" class="Insecta" family="Aphodiidae" genus="Gilletianus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="233" phylum="Arthropoda" rank="species" species="sundainsulanus">
|
||
<emphasis id="B93A7961FFB5B135FEF36B55FE31FE81" box="[274,482,295,321]" italics="true" pageId="10" pageNumber="233">G. sundainsulanus</emphasis>
|
||
</taxonomicName>
|
||
is rather trapezoidal) but in my opinion these are very subjective differences and will probably be lost in larger series of specimens.
|
||
</paragraph>
|
||
<caption id="DF31F5FBFFB5B135FF766CD3FD8AF91D" ID-DOI="http://doi.org/10.5281/zenodo.6810130" ID-Zenodo-Dep="6810130" httpUri="https://zenodo.org/record/6810130/files/figure.png" pageId="10" pageNumber="233" startId="10.[151,264,1697,1721]" targetBox="[166,1423,394,1657]" targetPageId="10">
|
||
<paragraph id="8BF1A573FFB5B135FF766CD3FD8AF91D" blockId="10.[151,1436,1697,1757]" pageId="10" pageNumber="233">
|
||
<emphasis id="B93A7961FFB5B135FF766CD3FE87F979" bold="true" box="[151,340,1697,1721]" pageId="10" pageNumber="233">FIGURES 24–26.</emphasis>
|
||
<taxonomicName id="4C4EDEF0FFB5B135FEB86CD3FC75F979" authority="Ochi, Kon & Kawahara, 2014" authorityName="Ochi, Kon & Kawahara" authorityYear="2014" box="[345,934,1697,1721]" class="Insecta" family="Aphodiidae" genus="Gilletianus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="233" phylum="Arthropoda" rank="species" species="sundainsulanus">
|
||
<emphasis id="B93A7961FFB5B135FEB86CD3FDBDF979" box="[345,622,1697,1721]" italics="true" pageId="10" pageNumber="233">Gilletianus sundainsulanus</emphasis>
|
||
<bibRefCitation id="EFDFD882FFB5B135FD956CD3FC75F979" author="Ochi, T. & Kon, M. & Kawahara, M." box="[628,934,1697,1721]" pageId="10" pageNumber="233" pagination="635 - 648" refId="ref8893" refString="Ochi, T., Kon, M. & Kawahara, M. (2014) Four new species of Aphodiini from Southeast Asia (Coleoptera Scarabaeidae). Giornale Italiano di Entomologia, 13 (59), 635 - 648." type="journal article" year="2014">Ochi, Kon & Kawahara, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
. 24, habitus, male, dorsal view; 25, habitus, male, ventral view; 26, habitus, male, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF1A573FFB5B135FF266D79FEDFF876" blockId="10.[151,1437,1803,2010]" pageId="10" pageNumber="233">
|
||
<taxonomicName id="4C4EDEF0FFB5B135FF266D79FDD2F8E5" authority="Minkina" authorityName="Minkina" authorityYear="2022" box="[199,513,1803,1829]" class="Insecta" family="Aphodiidae" genus="Gilletianus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="233" phylum="Arthropoda" rank="species" species="miurai" status="sp. nov.">
|
||
<emphasis id="B93A7961FFB5B135FF266D79FE44F8E5" box="[199,407,1803,1829]" italics="true" pageId="10" pageNumber="233">Gilletianus miurai</emphasis>
|
||
Minkina
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A209C41AFFB5B135FDEC6D7EFD39F8E6" box="[525,746,1803,1830]" pageId="10" pageNumber="233" rank="genus">
|
||
new species and
|
||
<taxonomicName id="4C4EDEF0FFB5B135FD2F6D7EFD39F8E6" authorityName="Ochi, Kon & Kawahara" authorityYear="2014" box="[718,746,1804,1830]" class="Insecta" family="Aphodiidae" genus="Gilletianus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="233" phylum="Arthropoda" rank="species" species="sundainsulanus">
|
||
<emphasis id="B93A7961FFB5B135FD2F6D7EFD39F8E6" box="[718,746,1804,1830]" italics="true" pageId="10" pageNumber="233">G.</emphasis>
|
||
</taxonomicName>
|
||
</taxonomicNameLabel>
|
||
<emphasis id="B93A7961FFB5B135FD136D79FC73F8E5" box="[754,928,1803,1829]" italics="true" pageId="10" pageNumber="233">sundainsulanus</emphasis>
|
||
are additionally closely related to
|
||
<taxonomicName id="4C4EDEF0FFB5B135FACD6D7EFE92F88A" authority="(Harold, 1859)" baseAuthorityName="Harold" baseAuthorityYear="1859" class="Insecta" family="Aphodiidae" genus="Gilletianus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="233" phylum="Arthropoda" rank="species" species="reichei">
|
||
<emphasis id="B93A7961FFB5B135FACD6D7EFA4EF8E5" box="[1324,1437,1803,1830]" italics="true" pageId="10" pageNumber="233">G. reichei</emphasis>
|
||
(Harold, 1859)
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFDFD882FFB5B135FE606D5DFD97F889" author="Ochi, T. & Kon, M. & Kakizoe, S." box="[385,580,1839,1865]" pageId="10" pageNumber="233" pagination="307 - 320" refId="ref8804" refString="Ochi, T., Kon, M. & Kakizoe, S. (2016) Nine new taxa of the genus Gilletianus from Southeast Asia (Coleoptera: Scarabaeidae). Japanese Journal of Systematic Entomology, 22 (2), 307 - 320." type="journal article" year="2016">
|
||
Ochi
|
||
<emphasis id="B93A7961FFB5B135FE236D42FE2EF889" box="[450,509,1839,1865]" italics="true" pageId="10" pageNumber="233">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C4EDEF0FFB5B135FDB86D5DFC45F889" authority="Minkina" authorityName="Minkina" authorityYear="2022" box="[601,918,1839,1865]" class="Insecta" family="Aphodiidae" genus="Gilletianus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="233" phylum="Arthropoda" rank="species" species="miurai" status="sp. nov.">
|
||
<emphasis id="B93A7961FFB5B135FDB86D5DFCF9F889" box="[601,810,1839,1865]" italics="true" pageId="10" pageNumber="233">Gilletianus miurai</emphasis>
|
||
Minkina
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A209C41AFFB5B135FC456D42FBFEF889" box="[932,1069,1839,1866]" pageId="10" pageNumber="233" rank="species">new species</taxonomicNameLabel>
|
||
has additionally a quite similar aedeagus to
|
||
<taxonomicName id="4C4EDEF0FFB5B135FEC96D26FE49F8AD" baseAuthorityName="Harold" baseAuthorityYear="1859" box="[296,410,1875,1902]" class="Insecta" family="Aphodiidae" genus="Gilletianus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="233" phylum="Arthropoda" rank="species" species="reichei">
|
||
<emphasis id="B93A7961FFB5B135FEC96D26FE49F8AD" box="[296,410,1875,1902]" italics="true" pageId="10" pageNumber="233">G. reichei</emphasis>
|
||
</taxonomicName>
|
||
, but still more distinctly elongate. It is easily distinguished from
|
||
<taxonomicName id="4C4EDEF0FFB5B135FB746D26FAD4F8AD" baseAuthorityName="Harold" baseAuthorityYear="1859" box="[1173,1287,1875,1902]" class="Insecta" family="Aphodiidae" genus="Gilletianus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="233" phylum="Arthropoda" rank="species" species="reichei">
|
||
<emphasis id="B93A7961FFB5B135FB746D26FAD4F8AD" box="[1173,1287,1875,1902]" italics="true" pageId="10" pageNumber="233">G. reichei</emphasis>
|
||
</taxonomicName>
|
||
by the same features as
|
||
<taxonomicName id="4C4EDEF0FFB5B135FEF86D0AFE38F851" authorityName="Ochi, Kon & Kawahara" authorityYear="2014" box="[281,491,1911,1938]" class="Insecta" family="Aphodiidae" genus="Gilletianus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="233" phylum="Arthropoda" rank="species" species="sundainsulanus">
|
||
<emphasis id="B93A7961FFB5B135FEF86D0AFE38F851" box="[281,491,1911,1938]" italics="true" pageId="10" pageNumber="233">G. sundainsulanus</emphasis>
|
||
</taxonomicName>
|
||
i.e.: relatively larger body, elytra proportionaly longer, distinctly different sides of pronotum.
|
||
</paragraph>
|
||
<paragraph id="8BF1A573FFB5B134FF266DCDFEE0FF39" blockId="10.[151,1437,1803,2010]" lastBlockId="11.[151,1436,151,285]" lastPageId="11" lastPageNumber="234" pageId="10" pageNumber="233">
|
||
Additionally the shape of aedeagus of
|
||
<taxonomicName id="4C4EDEF0FFB5B135FD996DB2FC9CF819" authority="Minkina" authorityName="Minkina" authorityYear="2022" box="[632,847,1983,2010]" class="Insecta" family="Aphodiidae" genus="Gilletianus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="233" phylum="Arthropoda" rank="species" species="miurai" status="sp. nov.">
|
||
<emphasis id="B93A7961FFB5B135FD996DB2FD35F819" box="[632,742,1984,2010]" italics="true" pageId="10" pageNumber="233">G. miurai</emphasis>
|
||
Minkina
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A209C41AFFB5B135FCBB6DB2FC31F819" box="[858,994,1983,2010]" pageId="10" pageNumber="233" rank="species">new species</taxonomicNameLabel>
|
||
is similar to
|
||
<taxonomicName id="4C4EDEF0FFB5B134FB946DB2FEBFFF71" authority="Ochi, Kon & Kawahara, 2010" authorityName="Ochi, Kon & Kawahara" authorityYear="2010" class="Insecta" family="Aphodiidae" genus="Gilletianus" kingdom="Animalia" lastPageId="11" lastPageNumber="234" order="Coleoptera" pageId="10" pageNumber="233" phylum="Arthropoda" rank="species" species="mindorensis">
|
||
<emphasis id="B93A7961FFB5B135FB946DB2FAF3F819" box="[1141,1312,1983,2010]" italics="true" pageId="10" pageNumber="233">G. mindorensis</emphasis>
|
||
<bibRefCitation id="EFDFD882FFB5B134FAC66DCDFEBFFF71" author="Ochi, T. & Kon, M. & Kawahara, M." lastPageId="11" lastPageNumber="234" pageId="10" pageNumber="233" pagination="111 - 120" refId="ref8852" refString="Ochi, T., Kon, M. & Kawahara, M. (2010) Four new species of the genus Gilletianus from Southeast Asia (Coleoptera, Scarabaeidae). Kogane, 11, 111 - 120." type="journal article" year="2010">Ochi, Kon & Kawahara, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
known only from Mindoro Island. From this species it can be very easily distinguished (males) by macrosetation of elytra present only on sides and in apical third (in
|
||
<taxonomicName id="4C4EDEF0FFB4B134FC516AC9FB89FF15" authorityName="Ochi, Kon & Kawahara" authorityYear="2010" box="[944,1114,187,213]" class="Insecta" family="Aphodiidae" genus="Gilletianus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="234" phylum="Arthropoda" rank="species" species="mindorensis">
|
||
<emphasis id="B93A7961FFB4B134FC516AC9FB89FF15" box="[944,1114,187,213]" italics="true" pageId="11" pageNumber="234">G. mindorensis</emphasis>
|
||
</taxonomicName>
|
||
macrosetation on almost the whole elytra).
|
||
</paragraph>
|
||
<paragraph id="8BF1A573FFB4B134FF266B71FCFCFEDD" blockId="11.[151,1436,151,285]" box="[199,815,258,285]" pageId="11" pageNumber="234">
|
||
<emphasis id="B93A7961FFB4B134FF266B71FE37FEDC" bold="true" box="[199,484,258,285]" pageId="11" pageNumber="234">Photographed material.</emphasis>
|
||
See material for comparison.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |