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<document id="23342D8016B11D5C21755E6E7F104C7B" ID-DOI="10.5281/zenodo.4619069" ID-ISSN="2345-7600" ID-Zenodo-Dep="4619069" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616093119406" checkinUser="carolina" docAuthor="Shy, Jhy-Yun" docDate="2005" docId="03E0F22D887E45592DAFA0F82F2DFC80" docLanguage="en" docName="RafflesBZool.53.1.99-102.pdf" docOrigin="Raffles Bulletin of Zoology 53 (1)" docStyle="DocumentStyle:19380AF0FBA5B2F218DD253A34ABE57F.5:RafflesBZool.2003-2005.journal_article" docStyleId="19380AF0FBA5B2F218DD253A34ABE57F" docStyleName="RafflesBZool.2003-2005.journal_article" docStyleVersion="5" docTitle="Geothelphusa leeae Shy 2005, new species" docType="treatment" docVersion="3" lastPageNumber="101" masterDocId="FFD98A55887F455B2D51A1192B00FFA2" masterDocTitle="A New Species Of Freshwater Crab Of The Genus Geothelphusa Stimpson, 1858 (Crustacea, Decapoda, Brachyura, Potamidae) From Taiwan" masterLastPageNumber="102" masterPageNumber="99" pageNumber="100" updateTime="1698890766719" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="2725E015BC0C5AC7E768045EBCFDD14A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="538E6FD6AD140768D591EDBB7DA26CF8">
<mods:title id="8F3FE36981BE52C45F0F15CA6CE021C3">A New Species Of Freshwater Crab Of The Genus Geothelphusa Stimpson, 1858 (Crustacea, Decapoda, Brachyura, Potamidae) From Taiwan</mods:title>
</mods:titleInfo>
<mods:name id="70127C99E32B2A52CC559648EDF20E0A" type="personal">
<mods:role id="7019CA012ACE80989362BE31DD06BA9A">
<mods:roleTerm id="95FC5F6728BC850F3378872882F888DB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="57D78B41B27C5C1B46236625D8373359">Shy, Jhy-Yun</mods:namePart>
</mods:name>
<mods:typeOfResource id="5F1BEE1DACEDBEA80AC3A64419086F49">text</mods:typeOfResource>
<mods:relatedItem id="5B3AEF023D2089426D03B6E1FA9E221B" type="host">
<mods:titleInfo id="4034A5FEAD44AAB969E6185B85CA4A21">
<mods:title id="253BFC675BF112A0C24FDD1EB932ADEC">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="7711708EBD79383E1D3B06E49A4EA1E2">
<mods:date id="58D120EB101A92BA1708624049ABB82B">2005</mods:date>
<mods:detail id="DD438101E517E3A03E7FE4D80424FB01" type="volume">
<mods:number id="39F360800A30850BAB7FFCF9B80F65F2">53</mods:number>
</mods:detail>
<mods:detail id="349EBB5AAFF80502AE81B359FA9AAEB4" type="issue">
<mods:number id="D2C61FE1E2F337DF1355FA3CB6D21284">1</mods:number>
</mods:detail>
<mods:extent id="C26156238E9193D794C30744E505C940" unit="page">
<mods:start id="0A095A024A6017E469FD1FA7F68AF3B7">99</mods:start>
<mods:end id="4E1BA7D8231138BB4A0DD0A2671956C9">102</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="5FAA51A06A224E88DCA528431235F96D">journal article</mods:classification>
<mods:identifier id="D93A50E55858C69252FB25008061C7A8" type="DOI">10.5281/zenodo.4619069</mods:identifier>
<mods:identifier id="44990FFDF84EEC16CE9E3CAC5D17B10A" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="A45933525FC273E44F748DACABC15546" type="Zenodo-Dep">4619069</mods:identifier>
</mods:mods>
<treatment id="03E0F22D887E45592DAFA0F82F2DFC80" LSID="urn:lsid:plazi:treatment:03E0F22D887E45592DAFA0F82F2DFC80" httpUri="http://treatment.plazi.org/id/03E0F22D887E45592DAFA0F82F2DFC80" lastPageId="2" lastPageNumber="101" pageId="1" pageNumber="100">
<subSubSection id="C35310B0887E455A2DAFA0F82963FE5A" box="[254,611,481,505]" pageId="1" pageNumber="100" type="nomenclature">
<paragraph id="8BF6433B887E455A2DAFA0F82963FE5A" blockId="1.[254,611,481,536]" box="[254,611,481,505]" pageId="1" pageNumber="100">
<heading id="D0BEF457887E455A2DAFA0F82963FE5A" box="[254,611,481,505]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="100" reason="3">
<taxonomicName id="4C4938B8887E455A2DAFA0F82AD9FE5B" ID-CoL="3FT3X" authority="Shy, 2005" authorityName="Shy" authorityYear="2005" box="[254,473,481,505]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="leeae" status="sp. nov.">
<emphasis id="B93D9F29887E455A2DAFA0F82AD9FE5B" bold="true" box="[254,473,481,505]" italics="true" pageId="1" pageNumber="100">Geothelphusa leeae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E2252887E455A2CB1A0F82963FE5A" box="[480,611,481,504]" pageId="1" pageNumber="100" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35310B0887E455A2C25A3182AEDFDBA" box="[372,493,513,536]" pageId="1" pageNumber="100" type="description">
<paragraph id="8BF6433B887E455A2C25A3182AEDFDBA" blockId="1.[254,611,481,536]" box="[372,493,513,536]" pageId="1" pageNumber="100">
(
<figureCitation id="13725FBE887E455A2C2DA3182AC9FDBA" box="[380,457,513,536]" captionStart="Fig" captionStartId="1.[113,149,2065,2086]" captionTargetBox="[113,752,1176,2025]" captionTargetId="figure-228@1.[113,752,1176,1601]" captionTargetPageId="1" captionText="Fig. 2. The morphology of Geothelphusa leeae. (A: frontal view. B: dorsal view.)" figureDoi="http://doi.org/10.5281/zenodo.4619073" httpUri="https://zenodo.org/record/4619073/files/figure.png" pageId="1" pageNumber="100">Figs. 2</figureCitation>
,
<figureCitation id="13725FBE887E455A2C86A3182AE4FDBA" box="[471,484,513,536]" captionStart="Fig" captionStartId="2.[138,174,2063,2084]" captionTargetBox="[144,1467,1254,2018]" captionTargetId="figure-45@2.[97,1518,1191,2079]" captionTargetPageId="2" captionText="Fig. 3. Gonapods of Gelthelphusa leeae. A: ventral view of right G.1 terminal segment. B: ventral view of right G1. C: dorsal view of right G1. D: ventral view of right G2. (scale = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.4619075" httpUri="https://zenodo.org/record/4619075/files/figure.png" pageId="1" pageNumber="100">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35310B0887E455A2D20A3592BC3FBA7" pageId="1" pageNumber="100" type="materials_examined">
<paragraph id="8BF6433B887E455A2D20A3592A24FD2D" blockId="1.[113,753,576,655]" pageId="1" pageNumber="100">
<emphasis id="B93D9F29887E455A2D20A3592A45FDF7" bold="true" box="[113,325,576,597]" italics="true" pageId="1" pageNumber="100">Material examined. –</emphasis>
<materialsCitation id="3B214966887E455A2C1BA3592A24FD2D" accessionNumber="F10986" collectingDate="1999-10-06" collectionCode="NTOU" collectorName="K. Lee &amp; C. L. Lee" location="Nantow county" municipality="Sun-Moon Lake" pageId="1" pageNumber="100" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54F2FD99887E455A2C1BA3592AA4FDF7" box="[330,420,576,597]" pageId="1" pageNumber="100" type="holotype">Holotype</typeStatus>
-
<specimenCount id="9D4F88B2887E455A2CE6A3592AFAFDF7" box="[439,506,576,597]" pageId="1" pageNumber="100" type="male">1 male</specimenCount>
, 22.7 by
<quantity id="4CB1EEDE887E455A2F1EA35929A5FDF7" box="[591,677,576,597]" metricMagnitude="-2" metricUnit="m" metricValue="1.8199999999999998" pageId="1" pageNumber="100" unit="mm" value="18.2">18.2 mm</quantity>
(
<collectionCode id="ED58DBFE887E455A2FFAA35929EFFDF7" box="[683,751,576,597]" country="Taiwan" httpUri="http://grbio.org/cool/be15-j6k8" name="Institute of Marine Biology, National Taiwan Ocean University" pageId="1" pageNumber="100">NTOU</collectionCode>
F10986),
<location id="8E9615E0887E455A2D81A3442A6AFDD0" LSID="urn:lsid:plazi:treatment:03E0F22D887E45592DAFA0F82F2DFC80:8E9615E0887E455A2D81A3442A6AFDD0" box="[208,362,605,626]" municipality="Sun-Moon Lake" name="Nantow county" pageId="1" pageNumber="100">Nantow county</location>
:
<collectingMunicipality id="6B92D941887E455A2C22A3442915FDD0" box="[371,533,605,626]" pageId="1" pageNumber="100">Sun-Moon Lake</collectingMunicipality>
,, coll.
<collectorName id="26BC26ED887E455A2F0DA344299EFDD0" box="[604,670,605,626]" pageId="1" pageNumber="100">K. Lee</collectorName>
&amp;
<collectorName id="26BC26ED887E455A2FEFA3442B98FD2D" pageId="1" pageNumber="100">C. L. Lee</collectorName>
,
<date id="FFF765FB887E455A2DF2A3632A20FD2D" box="[163,288,634,655]" pageId="1" pageNumber="100" value="1999-10-06">
<collectingDate id="EFB39C13887E455A2DF2A3632A20FD2D" box="[163,288,634,655]" pageId="1" pageNumber="100" value="1999-10-06">Oct. 6. 1999</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BF6433B887E455A2D20A3AD2A40FCFB" blockId="1.[113,753,692,857]" pageId="1" pageNumber="100">
<materialsCitation id="3B214966887E455A2D20A3AD2BBBFCBD" accessionNumber="F10987" collectingDate="1999-10-06" collectionCode="NTOU" collectorName="K. Lee &amp; C. L. Lee" county="Nantow county" location="Nantow county" municipality="Sun-Moon Lake" pageId="1" pageNumber="100" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="54F2FD99887E455A2D20A3AD2BD0FD6B" box="[113,208,692,713]" pageId="1" pageNumber="100" type="paratype">Paratypes</typeStatus>
–
<collectingCounty id="62973BB7887E455A2DB8A3AD2A83FD6B" box="[233,387,692,713]" pageId="1" pageNumber="100">Nantow county</collectingCounty>
:
<collectingMunicipality id="6B92D941887E455A2CDCA3AD292DFD6B" box="[397,557,692,713]" pageId="1" pageNumber="100">Sun-Moon Lake</collectingMunicipality>
-
<specimenCount id="9D4F88B2887E455A2F13A3AD2997FD6B" box="[578,663,692,713]" pageId="1" pageNumber="100" type="female">1 female</specimenCount>
(19.4 by
<quantity id="4CB1EEDE887E455A2D20A3C92BCAFD47" box="[113,202,720,741]" metricMagnitude="-2" metricUnit="m" metricValue="1.58" pageId="1" pageNumber="100" unit="mm" value="15.8">15.8 mm</quantity>
)
<location id="8E9615E0887E455A2D86A3C92A71FD47" LSID="urn:lsid:plazi:treatment:03E0F22D887E45592DAFA0F82F2DFC80:8E9615E0887E455A2D86A3C92A71FD47" box="[215,369,720,741]" county="Nantow county" municipality="Sun-Moon Lake" name="Nantow county" pageId="1" pageNumber="100">Nantow county</location>
:
<location id="8E9615E0887E455A2C2AA3C9291EFD47" LSID="urn:lsid:plazi:treatment:03E0F22D887E45592DAFA0F82F2DFC80:8E9615E0887E455A2C2AA3C9291EFD47" box="[379,542,720,741]" county="Nantow county" municipality="Sun-Moon Lake" name="Sun-Moon Lake" pageId="1" pageNumber="100">Sun-Moon Lake</location>
, coll.
<collectorName id="26BC26ED887E455A2F0BA3C9299DFD47" box="[602,669,720,741]" pageId="1" pageNumber="100">K. Lee</collectorName>
&amp;
<collectorName id="26BC26ED887E455A2FEFA3C92B98FCA0" pageId="1" pageNumber="100">C. L. Lee</collectorName>
,
<date id="FFF765FB887E455A2DF0A3F42A1BFCA0" box="[161,283,749,770]" pageId="1" pageNumber="100" value="1999-10-06">
<collectingDate id="EFB39C13887E455A2DF0A3F42A1BFCA0" box="[161,283,749,770]" pageId="1" pageNumber="100" value="1999-10-06">Oct. 6. 1999</collectingDate>
</date>
. (
<collectionCode id="ED58DBFE887E455A2C7BA3F42A6FFCA0" box="[298,367,749,770]" country="Taiwan" httpUri="http://grbio.org/cool/be15-j6k8" name="Institute of Marine Biology, National Taiwan Ocean University" pageId="1" pageNumber="100">NTOU</collectionCode>
F10987), coll.
<collectorName id="26BC26ED887E455A2F53A3F42943FCA0" box="[514,579,749,770]" pageId="1" pageNumber="100">K. Lee</collectorName>
&amp;
<collectorName id="26BC26ED887E455A2F30A3F429BEFCA0" box="[609,702,749,770]" pageId="1" pageNumber="100">C. L. Lee</collectorName>
,
<date id="FFF765FB887E455A2F96A3F42BBBFCBD" pageId="1" pageNumber="100" value="1999-10-06">
<collectingDate id="EFB39C13887E455A2F96A3F42BBBFCBD" pageId="1" pageNumber="100" value="1999-10-06">Oct. 6. 1999</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B214966887E455A2D92A2132A3DFCFB" collectingDate="1999-10-06" collectionCode="ZRC" collectorName="K. Lee &amp; C. L. Lee" location="Sun-Moon Lake" pageId="1" pageNumber="100" specimenCount="1" specimenCount-male="1" stateProvince="Nantou" typeStatus="paratype">
<specimenCount id="9D4F88B2887E455A2D92A2132A02FCBD" box="[195,258,778,799]" pageId="1" pageNumber="100" type="male">1 male</specimenCount>
(17.2 by
<quantity id="4CB1EEDE887E455A2C0BA2132AAAFCBD" box="[346,426,778,799]" metricMagnitude="-2" metricUnit="m" metricValue="1.41" pageId="1" pageNumber="100" unit="mm" value="14.1">14.1mm</quantity>
)
<collectingRegion id="498D8DD9887E455A2CE7A2132947FCBD" box="[438,583,778,799]" country="Taiwan" name="Nantou" pageId="1" pageNumber="100">Nantou county</collectingRegion>
:
<location id="8E9615E0887E455A2F1FA21329ECFCBD" LSID="urn:lsid:plazi:treatment:03E0F22D887E45592DAFA0F82F2DFC80:8E9615E0887E455A2F1FA21329ECFCBD" box="[590,748,778,799]" name="Sun-Moon Lake" pageId="1" pageNumber="100" stateProvince="Nantou">Sun-Moon Lake</location>
, coll.
<collectorName id="26BC26ED887E455A2DF2A23E2BE5FC9E" box="[163,229,807,828]" pageId="1" pageNumber="100">K. Lee</collectorName>
&amp;
<collectorName id="26BC26ED887E455A2C57A23E2A66FC9E" box="[262,358,807,828]" pageId="1" pageNumber="100">C. L. Lee</collectorName>
,
<date id="FFF765FB887E455A2C20A23E2AEEFC9E" box="[369,494,807,828]" pageId="1" pageNumber="100" value="1999-10-06">
<collectingDate id="EFB39C13887E455A2C20A23E2AEEFC9E" box="[369,494,807,828]" pageId="1" pageNumber="100" value="1999-10-06">Oct. 6. 1999</collectingDate>
</date>
. (
<collectionCode id="ED58DBFE887E455A2CAEA23E2933FC9E" box="[511,563,807,828]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="100">ZRC</collectionCode>
), coll.
<collectorName id="26BC26ED887E455A2F27A23E29B9FC9E" box="[630,697,807,828]" pageId="1" pageNumber="100">K. Lee</collectorName>
&amp;
<collectorName id="26BC26ED887E455A2F8BA23E2BB4FCFB" pageId="1" pageNumber="100">C. L. Lee</collectorName>
,
<date id="FFF765FB887E455A2DEEA25D2A3DFCFB" box="[191,317,836,857]" pageId="1" pageNumber="100" value="1999-10-06">
<collectingDate id="EFB39C13887E455A2DEEA25D2A3DFCFB" box="[191,317,836,857]" pageId="1" pageNumber="100" value="1999-10-06">Oct. 6. 1999</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF6433B887E455A2D20A2642BC3FBA7" blockId="1.[113,753,893,1029]" pageId="1" pageNumber="100">
<materialsCitation id="3B214966887E455A2D20A2642BC3FBA7" accessionNumber="F11020" collectingDate="2005-03-11" collectionCode="NTOU" collectorName="J. Y. Shy &amp; K. Lee" county="Sun-Moon Lake" location="Others - Nantow county" municipality="Nantow county" pageId="1" pageNumber="100" specimenCount="6" specimenCount-female="3" specimenCount-male="3">
<location id="8E9615E0887E455A2D20A2642A5EFC30" LSID="urn:lsid:plazi:treatment:03E0F22D887E45592DAFA0F82F2DFC80:8E9615E0887E455A2D20A2642A5EFC30" box="[113,350,893,914]" county="Sun-Moon Lake" municipality="Nantow county" name="Others - Nantow county" pageId="1" pageNumber="100">Others –Nantow county</location>
:
<location id="8E9615E0887E455A2C39A2642907FC30" LSID="urn:lsid:plazi:treatment:03E0F22D887E45592DAFA0F82F2DFC80:8E9615E0887E455A2C39A2642907FC30" box="[360,519,893,914]" county="Sun-Moon Lake" municipality="Nantow county" name="Sun-Moon Lake" pageId="1" pageNumber="100">Sun-Moon Lake</location>
-
<specimenCount id="9D4F88B2887E455A2F4DA2642967FC30" box="[540,615,893,914]" pageId="1" pageNumber="100" type="male">3 males</specimenCount>
(27.8 by
<quantity id="4CB1EEDE887E455A2F94A2642B99FC0D" metricMagnitude="-2" metricUnit="m" metricValue="2.18" pageId="1" pageNumber="100" unit="mm" value="21.8">21.8 mm</quantity>
, 23.3 by
<quantity id="4CB1EEDE887E455A2DA5A2832A4CFC0D" box="[244,332,922,943]" metricMagnitude="-2" metricUnit="m" metricValue="2.07" pageId="1" pageNumber="100" unit="mm" value="20.7">20.7 mm</quantity>
and 22.7 by
<quantity id="4CB1EEDE887E455A2C9DA2832927FC0D" box="[460,551,922,943]" metricMagnitude="-2" metricUnit="m" metricValue="1.86" pageId="1" pageNumber="100" unit="mm" value="18.6">18.6 mm</quantity>
),
<specimenCount id="9D4F88B2887E455A2F68A2832997FC0D" box="[569,663,922,943]" pageId="1" pageNumber="100" type="female">3 females</specimenCount>
(24.3 by 20.0 mm, 23.5 by
<quantity id="4CB1EEDE887E455A2C71A2AE2A76FC6E" box="[288,374,951,972]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="1" pageNumber="100" unit="mm" value="19.1">19.1 mm</quantity>
and 23.0 by
<quantity id="4CB1EEDE887E455A2CA0A2AE2948FC6E" box="[497,584,951,972]" metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" pageId="1" pageNumber="100" unit="mm" value="18.7">18.7 mm</quantity>
)
<collectingMunicipality id="6B92D941887E455A2F05A2AE29EDFC6E" box="[596,749,951,972]" pageId="1" pageNumber="100">Nantow county</collectingMunicipality>
:
<collectingCounty id="62973BB7887E455A2D20A2CD2A12FC4B" box="[113,274,980,1001]" pageId="1" pageNumber="100">Sun-Moon Lake</collectingCounty>
, coll.
<collectorName id="26BC26ED887E455A2C1DA2CD2AA4FC4B" box="[332,420,980,1001]" pageId="1" pageNumber="100">J. Y. Shy</collectorName>
&amp;
<collectorName id="26BC26ED887E455A2C93A2CD2906FC4B" box="[450,518,980,1001]" pageId="1" pageNumber="100">K. Lee</collectorName>
,
<date id="FFF765FB887E455A2F41A2CD299CFC4B" box="[528,668,980,1001]" pageId="1" pageNumber="100" value="2005-03-11">
<collectingDate id="EFB39C13887E455A2F41A2CD299CFC4B" box="[528,668,980,1001]" pageId="1" pageNumber="100" value="2005-03-11">Mar. 11. 2005</collectingDate>
</date>
. (
<collectionCode id="ED58DBFE887E455A2FFAA2CD29F0FC4B" box="[683,752,980,1001]" country="Taiwan" httpUri="http://grbio.org/cool/be15-j6k8" name="Institute of Marine Biology, National Taiwan Ocean University" pageId="1" pageNumber="100">NTOU</collectionCode>
F11020)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35310B0887E455A2D20A5342E4FFA9A" pageId="1" pageNumber="100" type="diagnosis">
<paragraph id="8BF6433B887E455A2D20A53429F0FBC6" blockId="1.[113,752,1069,1124]" pageId="1" pageNumber="100">
<emphasis id="B93D9F29887E455A2D20A5342A0FFBE7" bold="true" box="[113,271,1069,1093]" italics="true" pageId="1" pageNumber="100">Diagnosis. –</emphasis>
Carapace smooth, moderately convex longitudinally and transversely. Carapace length and width
</paragraph>
<paragraph id="8BF6433B887E455A2E7DA1D82FB1FC5A" blockId="1.[812,1452,193,1016]" pageId="1" pageNumber="100">1.6 times and 2.0 times of carapace depth, respectively. Frontal region faintly divided into 2 lobes; supraorbital cristae smooth, without granules; infraorbital cristae smooth to almost smooth, sometimes lined with very low granules. External orbital angle sharp. Anterolateral crista distinct, lined with small granules; epibranchial tooth small, just discernible. Postorbital crista rough, with striae. Frontal, orbital, branchial and anterolateral regions covered with weak striae; gastric, cardiac and intestinal regions smooth. Cervical groove faint; H-shaped median gastro-cardiac depression distinct. Posterior margin of epistome with sharp median projection. Distance between tip of closed male abdomen and anterior margin of thoracic sternite 4 about 1.5 times length of thoracic sternites 1-3. External margin of palm smooth; fingers of right chela forming ovate gape when closed. Ambulatory legs of normal length; total length of second leg about 2.0 times carapace length; length of merus about 3.9 times maximum width. Telson of male abdomen bell-shaped. G1 with subterminal segment moderately curving outwards, outer proximal margin with tooth, inner proximal margin moderately dilated; terminal segment straight, distal one-third slender, with spinules and setae; total length of G1 7.0 times of terminal segment; length of synovial membrane about 3.0 times maximum width. G2 with outer proximal margin of basal segment dilated, faintly divided into 2 lobes; distal segment short, about 0.16 times total length.</paragraph>
<paragraph id="8BF6433B887E455A2E7DA5382F2CFBDA" blockId="1.[812,1451,1057,1144]" pageId="1" pageNumber="100">
<emphasis id="B93D9F29887E455A2E7DA53828D0FB9B" bold="true" box="[812,976,1057,1081]" italics="true" pageId="1" pageNumber="100">Colouration. –</emphasis>
Carapace and ambulatory legs greenish to orange, sometimes ambulatory legs covered with scattered darker coloured stripes.
</paragraph>
<paragraph id="8BF6433B887E455A2E7DA5B828DEFB7A" blockId="1.[812,1451,1185,1240]" pageId="1" pageNumber="100">
<emphasis id="B93D9F29887E455A2E7DA5B8289BFB1B" bold="true" box="[812,923,1185,1209]" italics="true" pageId="1" pageNumber="100">Habitat. –</emphasis>
Probably in burrows along the banks of the lake under the water.
</paragraph>
<paragraph id="8BF6433B887E455A2E7DA4182E4FFA9A" blockId="1.[812,1451,1281,1336]" pageId="1" pageNumber="100">
<emphasis id="B93D9F29887E455A2E7DA4182878FABB" bold="true" box="[812,888,1281,1305]" italics="true" pageId="1" pageNumber="100">Size. –</emphasis>
Largest male 27.1 by
<quantity id="4CB1EEDE887E455A292DA4182FE0FABA" box="[1148,1248,1281,1304]" metricMagnitude="-2" metricUnit="m" metricValue="2.18" pageId="1" pageNumber="100" unit="mm" value="21.8">21.8 mm</quantity>
(NTOU F11020); largest female 24.3 by 20.0 mm (NTOU F11020).
</paragraph>
</subSubSection>
<subSubSection id="C35310B0887E455A2E7DA4782E3BFADA" box="[812,1339,1377,1401]" pageId="1" pageNumber="100" type="distribution">
<paragraph id="8BF6433B887E455A2E7DA4782E3BFADA" blockId="1.[812,1339,1377,1401]" box="[812,1339,1377,1401]" pageId="1" pageNumber="100">
<emphasis id="B93D9F29887E455A2E7DA47828CFFADB" bold="true" box="[812,975,1377,1401]" italics="true" pageId="1" pageNumber="100">Distribution. –</emphasis>
Nantow County, central
<collectingCountry id="F35E03AB887E455A29B3A4782E36FADA" box="[1250,1334,1377,1400]" name="Taiwan" pageId="1" pageNumber="100">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C35310B0887E455A2E7DA4B82881F9BA" pageId="1" pageNumber="100" type="etymology">
<paragraph id="8BF6433B887E455A2E7DA4B82881F9BA" blockId="1.[812,1451,1441,1560]" pageId="1" pageNumber="100">
<emphasis id="B93D9F29887E455A2E7DA4B828BEFA1B" box="[812,958,1441,1465]" italics="true" pageId="1" pageNumber="100">
<emphasis id="B93D9F29887E455A2E7DA4B828A2FA1B" bold="true" box="[812,930,1441,1465]" italics="true" pageId="1" pageNumber="100">Etymology</emphasis>
.
<emphasis id="B93D9F29887E455A2EE0A4B828BEFA1B" bold="true" box="[945,958,1441,1465]" italics="true" pageId="1" pageNumber="100">–</emphasis>
</emphasis>
The species is named after Miss Kari Lee of the National
<collectingCountry id="F35E03AB887E455A2EFEA4D828FDFA7A" box="[943,1021,1473,1496]" name="Taiwan" pageId="1" pageNumber="100">Taiwan</collectingCountry>
Ocean University, Department of Fishery Science, Crustacean Laboratory, all for her help to find this species.
</paragraph>
</subSubSection>
<subSubSection id="C35310B0887E45592E7DA7582F2DFC80" lastPageId="2" lastPageNumber="101" pageId="1" pageNumber="100" type="discussion">
<paragraph id="8BF6433B887E455A2E7DA75828E9F81A" blockId="1.[812,1452,1601,1976]" pageId="1" pageNumber="100">
<emphasis id="B93D9F29887E455A2E7DA75828A6F9FB" box="[812,934,1601,1625]" italics="true" pageId="1" pageNumber="100">
<emphasis id="B93D9F29887E455A2E7DA758288DF9FB" bold="true" box="[812,909,1601,1625]" italics="true" pageId="1" pageNumber="100">Remarks</emphasis>
.
<emphasis id="B93D9F29887E455A2EC8A75828A6F9FB" bold="true" box="[921,934,1601,1625]" italics="true" pageId="1" pageNumber="100">–</emphasis>
</emphasis>
Although Sun-Moon Lake is a natural freshwater lake at an altitude of about
<quantity id="4CB1EEDE887E455A2906A7782F94F9DA" box="[1111,1172,1633,1656]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="1" pageNumber="100" unit="m" value="750.0">750m</quantity>
above sea level, this lake experiences a daily tide. During the day, the water is allowed to flow out of conduits into a lower altitude pond to generate hydroelectric power; and the water level gradually decreases. About at midnight, the company utilizes the excess electric power to pump water up from this pond back into Sun-Moon Lake; and the water level rises back up. Under these special conditions, we have been able to find
<taxonomicName id="4C4938B8887E455A29BEA6582E50F8FA" authorityName="Shy" authorityYear="2005" box="[1263,1360,1857,1880]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="leeae">
<emphasis id="B93D9F29887E455A29BEA6582E50F8FA" box="[1263,1360,1857,1880]" italics="true" pageId="1" pageNumber="100">G. leeae</emphasis>
</taxonomicName>
only at midnight (i.e. at lowest tide). At this time, we can also find many burrows along the bank which we believe have been dug by the crabs.
</paragraph>
<paragraph id="8BF6433B887E45592E7DA6F82972FBA0" blockId="1.[812,1452,2017,2104]" lastBlockId="2.[136,775,683,1026]" lastPageId="2" lastPageNumber="101" pageId="1" pageNumber="100">
Of the known
<taxonomicName id="4C4938B8887E455A2E93A6F82F57F85A" authorityName="STIMPSON" authorityYear="1858" box="[962,1111,2017,2040]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="B93D9F29887E455A2E93A6F82F57F85A" box="[962,1111,2017,2040]" italics="true" pageId="1" pageNumber="100">Geothelphusa</emphasis>
</taxonomicName>
species in
<collectingCountry id="F35E03AB887E455A2998A6F82E1CF85A" box="[1225,1308,2017,2040]" name="Taiwan" pageId="1" pageNumber="100">Taiwan</collectingCountry>
,
<taxonomicName id="4C4938B8887E455A2874A6F82E73F85A" authorityName="Shy, Ng and Yu" authorityYear="1994" box="[1317,1395,2017,2040]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="olea">
<emphasis id="B93D9F29887E455A2874A6F82E73F85A" box="[1317,1395,2017,2040]" italics="true" pageId="1" pageNumber="100">G. olea</emphasis>
</taxonomicName>
(
<figureCitation id="13725FBE887E455A28D1A6F8283AF7BA" captionStart="Fig" captionStartId="3.[119,155,640,661]" captionTargetBox="[116,1398,192,599]" captionTargetPageId="3" captionText="Fig. 4. Dorsal carapace view and gonopods of Geothelphusa olea. A: ventral view of right G.1 terminal segment. B: ventral view of right G1. C: dorsal view of right G1. D: ventral view of right G2. (dorsal view from Shy &amp; Yu, 1999; gonopods from Shy, 1994; scale = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.4619077" httpUri="https://zenodo.org/record/4619077/files/figure.png" pageId="1" pageNumber="100">Fig. 4</figureCitation>
) is most similar to the present new species but there are some distinct morphological differences between them, notably in the carapace and gonopods (Table 1). The most obvious difference is the body size of
<taxonomicName id="4C4938B8887D45592F6BA3D52998FD41" authorityName="Shy" authorityYear="2005" box="[570,664,716,739]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="leeae">
<emphasis id="B93D9F29887D45592F6BA3D52998FD41" box="[570,664,716,739]" italics="true" pageId="2" pageNumber="101">G. leeae</emphasis>
</taxonomicName>
, which is smaller than
<taxonomicName id="4C4938B8887D45592C41A3F52A5EFCA1" authorityName="Shy, Ng and Yu" authorityYear="1994" box="[272,350,748,771]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="olea">
<emphasis id="B93D9F29887D45592C41A3F52A5EFCA1" box="[272,350,748,771]" italics="true" pageId="2" pageNumber="101">G. olea</emphasis>
</taxonomicName>
. Although we have only
<specimenCount id="9D4F88B2887D45592F39A3F229EBFCA0" box="[616,747,747,770]" pageId="2" pageNumber="101" type="generic">9 specimens</specimenCount>
of
<taxonomicName id="4C4938B8887D45592DD9A2152BE9FC81" authorityName="Shy" authorityYear="2005" box="[136,233,780,803]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="leeae">
<emphasis id="B93D9F29887D45592DD9A2152BE9FC81" box="[136,233,780,803]" italics="true" pageId="2" pageNumber="101">G. leeae</emphasis>
</taxonomicName>
, the female specimen is already fully mature. Specimens of
<taxonomicName id="4C4938B8887D45592C70A2352A71FCE1" authorityName="Shy, Ng and Yu" authorityYear="1994" box="[289,369,812,835]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="olea">
<emphasis id="B93D9F29887D45592C70A2352A71FCE1" box="[289,369,812,835]" italics="true" pageId="2" pageNumber="101">G. olea</emphasis>
</taxonomicName>
, the size of this female as
<taxonomicName id="4C4938B8887D45592FC2A23529EDFCE1" authorityName="Shy" authorityYear="2005" box="[659,749,812,835]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="leeae">
<emphasis id="B93D9F29887D45592FC2A23529EDFCE1" box="[659,749,812,835]" italics="true" pageId="2" pageNumber="101">G. leeae</emphasis>
</taxonomicName>
is still immature. Another possible difference is their habitat. Although both species live in burrows, their specific habitats are rather different.
<taxonomicName id="4C4938B8887D45592C07A295291AFC01" authorityName="Shy, Ng and Yu" authorityYear="1994" box="[342,538,908,931]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="olea">
<emphasis id="B93D9F29887D45592C07A295291AFC01" box="[342,538,908,931]" italics="true" pageId="2" pageNumber="101">Geothelphusa olea</emphasis>
</taxonomicName>
lives in burrows along the banks of streams above the water, but
<taxonomicName id="4C4938B8887D45592F1BA2B529A3FC61" authorityName="Shy" authorityYear="2005" box="[586,675,940,963]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="leeae">
<emphasis id="B93D9F29887D45592F1BA2B529A3FC61" box="[586,675,940,963]" italics="true" pageId="2" pageNumber="101">G. leeae</emphasis>
</taxonomicName>
seems to prefer to be underwater. So far, this species is the only lake habitat species of freshwater crab in
<collectingCountry id="F35E03AB887D45592F4BA2F2296EFBA0" box="[538,622,1003,1026]" name="Taiwan" pageId="2" pageNumber="101">Taiwan</collectingCountry>
.
</paragraph>
<caption id="DF3613B3887E455A2D20A9082A0FF7E1" ID-DOI="http://doi.org/10.5281/zenodo.4619073" ID-Zenodo-Dep="4619073" httpUri="https://zenodo.org/record/4619073/files/figure.png" pageId="1" pageNumber="100" startId="1.[113,149,2065,2086]" targetBox="[113,752,1176,2025]" targetPageId="1">
<paragraph id="8BF6433B887E455A2D20A9082A0FF7E1" blockId="1.[113,753,2065,2115]" pageId="1" pageNumber="100">
Fig. 2. The morphology of
<taxonomicName id="4C4938B8887E455A2C28A90B293AF785" authorityName="Shy" authorityYear="2005" box="[377,570,2066,2087]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="leeae">
<emphasis id="B93D9F29887E455A2C28A90B293AF785" box="[377,570,2066,2087]" italics="true" pageId="1" pageNumber="100">Geothelphusa leeae</emphasis>
</taxonomicName>
. (A: frontal view. B: dorsal view.)
</paragraph>
</caption>
<paragraph id="8BF6433B887D45592DD9A5352F2DFC80" blockId="2.[136,775,1067,1187]" lastBlockId="2.[835,1474,683,803]" pageId="2" pageNumber="101">
<taxonomicName id="4C4938B8887D45592DD9A5352AAAFBE1" authorityName="Bott" authorityYear="1967" box="[136,426,1068,1091]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="candidiensis">
<emphasis id="B93D9F29887D45592DD9A5352AAAFBE1" box="[136,426,1068,1091]" italics="true" pageId="2" pageNumber="101">Geothelphusa candidiensis</emphasis>
</taxonomicName>
(
<figureCitation id="13725FBE887D45592CE6A5322AF7FBE0" box="[439,503,1067,1090]" captionStart="Fig" captionStartId="3.[113,149,1444,1465]" captionTargetBox="[232,1328,744,1402]" captionTargetId="figure-11@3.[200,1355,715,1436]" captionTargetPageId="3" captionText="Fig. 5. Gonopods of Geothelphusa candidiensis. A: ventral view of right G.1 terminal segment. B: ventral view of right G1. C: dorsal view of right G1. D: ventral view of right G2. (gonopods from Shy, 1994; scale = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.4619079" httpUri="https://zenodo.org/record/4619079/files/figure.png" pageId="2" pageNumber="101">Fig. 5</figureCitation>
) is also similar to present new species, but we can distinguished by several characters. The major differences are: the epibranchial tooth is small but very clear in
<emphasis id="B93D9F29887D45592C43A5952AC0FB01" box="[274,448,1164,1187]" italics="true" pageId="2" pageNumber="101">
<taxonomicName id="4C4938B8887D45592C43A5952ABDFB01" authorityName="Bott" authorityYear="1967" box="[274,445,1164,1187]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="candidiensis">G. candidiensis</taxonomicName>
,
</emphasis>
but is faint in
<taxonomicName id="4C4938B8887D45592F08A59529B2FB01" authorityName="Shy" authorityYear="2005" box="[601,690,1164,1187]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="leeae">
<emphasis id="B93D9F29887D45592F08A59529B2FB01" box="[601,690,1164,1187]" italics="true" pageId="2" pageNumber="101">G. leeae</emphasis>
</taxonomicName>
; the G1 outer proximal margin is without a tooth in
<taxonomicName id="4C4938B8887D4559284BA3B52EC2FD61" authorityName="Bott" authorityYear="1967" box="[1306,1474,684,707]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="candidiensis">
<emphasis id="B93D9F29887D4559284BA3B52EC2FD61" box="[1306,1474,684,707]" italics="true" pageId="2" pageNumber="101">G. candidiensis</emphasis>
</taxonomicName>
but there is a distinct tooth in
<taxonomicName id="4C4938B8887D4559293EA3D52FC5FD41" authorityName="Shy" authorityYear="2005" box="[1135,1221,716,739]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="leeae">
<emphasis id="B93D9F29887D4559293EA3D52FC5FD41" box="[1135,1221,716,739]" italics="true" pageId="2" pageNumber="101">G. leeae</emphasis>
</taxonomicName>
; and the G1 subterminal segment is almost straight in
<taxonomicName id="4C4938B8887D455929DAA3F52E36FCA1" authorityName="Bott" authorityYear="1967" box="[1163,1334,748,771]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="candidiensis">
<emphasis id="B93D9F29887D455929DAA3F52E36FCA1" box="[1163,1334,748,771]" italics="true" pageId="2" pageNumber="101">G. candidiensis</emphasis>
</taxonomicName>
but directed outwards in
<taxonomicName id="4C4938B8887D45592E9BA2152F26FC81" authorityName="Shy" authorityYear="2005" box="[970,1062,780,803]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="leeae">
<emphasis id="B93D9F29887D45592E9BA2152F26FC81" box="[970,1062,780,803]" italics="true" pageId="2" pageNumber="101">G. leeae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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