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<document id="D1B50F65F5D18BA3EDFCAB4DB3039D63" ID-CLB-Dataset="86210" ID-DOI="10.5852/ejt.2022.826.1839" ID-GBIF-Dataset="ed37ce13-a67c-4146-b99e-db8ede0a145b" ID-ISSN="2118-9773" ID-Zenodo-Dep="6784127" ID-ZooBank="34BA1AB7-6107-4636-9645-B1C0216DCE5E" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1656607314270" checkinUser="felipe" docAuthor="Michelsen, Verner" docDate="2022" docId="03E0DC63FFEFFFA2FE057C17E96199D3" docLanguage="en" docName="ejt-826_94-134.pdf" docOrigin="European Journal of Taxonomy 826" docSource="http://dx.doi.org/10.5852/ejt.2022.826.1839" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Reinwardtiinae Brauer &amp; Bergenstamm 1889" docType="treatment" docVersion="7" lastPageNumber="117" masterDocId="FFD9A41BFFF9FFB5FFC17D1EEC209B5E" masterDocTitle="Costal vein chaetotaxy, a neglected character source in Fanniidae and Muscidae (Diptera: Calyptratae)" masterLastPageNumber="134" masterPageNumber="94" pageNumber="116" updateTime="1699353813789" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="259055649BFAF0CB6434566D149AC9D1">Costal vein chaetotaxy, a neglected character source in Fanniidae and Muscidae (Diptera: Calyptratae)</mods:title>
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<mods:namePart id="93DA1C66B4FD49C632EEDFF6EAD3FC01">Michelsen, Verner</mods:namePart>
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<mods:affiliation id="FF18220283832A6A579C2040CF722689">Natural History Museum of Denmark, University of Copenhagen, Denmark.</mods:affiliation>
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<mods:title id="E562250037572B23C395F1E0BC8C9E7B">European Journal of Taxonomy</mods:title>
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<treatment id="03E0DC63FFEFFFA2FE057C17E96199D3" ID-DOI="http://doi.org/10.5281/zenodo.6788827" ID-GBIF-Taxon="196374605" ID-Zenodo-Dep="6788827" LSID="urn:lsid:plazi:treatment:03E0DC63FFEFFFA2FE057C17E96199D3" httpUri="http://treatment.plazi.org/id/03E0DC63FFEFFFA2FE057C17E96199D3" lastPageId="23" lastPageNumber="117" pageId="22" pageNumber="116">
<subSubSection id="C3533EFEFFEFFFA3FE057C17E8509A7D" box="[452,1136,265,291]" pageId="22" pageNumber="116" type="nomenclature">
<paragraph id="8BF66D75FFEFFFA3FE057C17E8509A7D" blockId="22.[452,1136,265,291]" box="[452,1136,265,291]" pageId="22" pageNumber="116">
Subfamily
<taxonomicName id="4C4916F6FFEFFFA3FD847C17E8509A7D" authority="Brauer &amp; Bergenstamm, 1889" authorityName="Brauer &amp; Bergenstamm" authorityYear="1889" box="[581,1136,265,291]" family="Muscidae" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="116" rank="subFamily" subFamily="Reinwardtiinae">
<emphasis id="B93DB167FFEFFFA3FD847C17EF279A7D" bold="true" box="[581,775,265,291]" pageId="22" pageNumber="116">Reinwardtiinae</emphasis>
Brauer &amp; Bergenstamm, 1889
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3533EFEFFEFFFA2FF7C7C54E96199D3" lastPageId="23" lastPageNumber="117" pageId="22" pageNumber="116" type="discussion">
<paragraph id="8BF66D75FFEFFFA3FF7C7C54ED179AF5" blockId="22.[189,1399,330,427]" pageId="22" pageNumber="116">
This taxon is a heterogeneous, para- or polyphyletic assemblage of genera which is also reflected in its diverse costal chaetotaxy. Species from seven among the 15 currently recognized genera (
<tableCitation id="C6CB58CEFFEFFFA3FB157C70E90B9AD6" box="[1236,1323,366,392]" captionStart="Table 1" captionStartId="5.[189,258,265,291]" captionTargetBox="[238,1379,394,1980]" captionText="Table 1 (continued on next 4 pages). Summary of vein C chaetotaxy with respect to multistate characterA for genus-group taxa of Fanniidae Schnabel &amp; Dziedcicki, 1911 and Muscidae Latreille, 1802. Examined genera highlighted in yellow. See ‘Material and methods’ for explanation of character states A0–8." httpUri="http://table.plazi.org/id/DF363DFDFFFCFFB0FF7C7C17E91E9A37" pageId="22" pageNumber="116" tableUuid="DF363DFDFFFCFFB0FF7C7C17E91E9A37">Table 1</tableCitation>
) were examined.
</paragraph>
<paragraph id="8BF66D75FFEFFFA3FF7C7CC5EDA49949" blockId="22.[189,1398,474,535]" pageId="22" pageNumber="116">
<emphasis id="B93DB167FFEFFFA3FF7C7CC5ECE79AAA" box="[189,199,475,500]" italics="true" pageId="22" pageNumber="116">•</emphasis>
<taxonomicName id="4C4916F6FFEFFFA3FF1C7CC5EE5D9AAA" authority="Robineau-Desvoidy, 1830" authorityName="Robineau-Desvoidy" authorityYear="1830" box="[221,637,474,500]" class="Insecta" family="Muscidae" genus="Muscina" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B93DB167FFEFFFA3FF1C7CC5ED629AAA" box="[221,322,475,500]" italics="true" pageId="22" pageNumber="116">Muscina</emphasis>
Robineau-Desvoidy, 1830
</taxonomicName>
(
<figureCitation id="137271F0FFEFFFA3FD4C7CC4EEDA9AAA" box="[653,762,474,500]" captionStart="Fig" captionStartId="24.[189,232,1876,1902]" captionTargetBox="[190,1398,677,1852]" captionTargetId="figure-150@24.[189,1399,670,1856]" captionTargetPageId="24" captionText="Fig. 12. ♀♀ of Muscidae Latreille, 1802. A. Muscina stabulans (Fallén, 1823), detail of wing showing CS1‒3, dorsal view. B. Palpibracus trivittatus (Malloch, 1934), detail of wing showing CS1‒3, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6784155" httpUri="https://zenodo.org/record/6784155/files/figure.png" pageId="22" pageNumber="116">Fig. 12A</figureCitation>
),
<taxonomicName id="4C4916F6FFEFFFA3FCD17CC5E8DD9AAB" authority="Rodhain &amp; Villeneuve, 1915" authorityName="Rodhain &amp; Villeneuve" authorityYear="1915" box="[784,1277,474,501]" class="Insecta" family="Muscidae" genus="Passeromyia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B93DB167FFEFFFA3FCD17CC5EF869AAA" box="[784,934,475,500]" italics="true" pageId="22" pageNumber="116">Passeromyia</emphasis>
Rodhain &amp; Villeneuve, 1915
</taxonomicName>
,
<taxonomicName id="4C4916F6FFEFFFA3FAC87CC4EDA49949" authority="Meinert, 1890" authorityName="Meinert" authorityYear="1890" class="Insecta" family="Muscidae" genus="Philornis" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B93DB167FFEFFFA3FAC87CC4E9569AAA" box="[1289,1398,474,500]" italics="true" pageId="22" pageNumber="116">Philornis</emphasis>
Meinert, 1890
</taxonomicName>
</paragraph>
<paragraph id="8BF66D75FFEFFFA3FF7C7F35E83D99D2" blockId="22.[188,1399,555,653]" pageId="22" pageNumber="116">
The examined species belonging to the above genera of
<taxonomicName id="4C4916F6FFEFFFA3FC887F35EFDC991B" authorityName="Brauer &amp; Bergenstamm" authorityYear="1889" box="[841,1020,555,581]" pageId="22" pageNumber="116" rank="subFamily" subFamily="Reinwardtiinae">Reinwardtiinae</taxonomicName>
share the prevalent muscid state
<emphasis id="B93DB167FFEFFFA3FF7D7F50ECC09936" bold="true" box="[188,224,590,616]" pageId="22" pageNumber="116">
A
<quantity id="4CB1C090FFEFFFA3FF137F50ECC09936" box="[210,224,590,616]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="22" pageNumber="116" unit="in" value="4.0">4</quantity>
</emphasis>
in which vein C is bare dorsally and extensively setulose ventrally:
<taxonomicName id="4C4916F6FFEFFFA3FBD17F4EE8559937" authorityName="Robineau-Desvoidy" authorityYear="1830" box="[1040,1141,592,617]" class="Insecta" family="Muscidae" genus="Muscina" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B93DB167FFEFFFA3FBD17F4EE8559937" box="[1040,1141,592,617]" italics="true" pageId="22" pageNumber="116">Muscina</emphasis>
</taxonomicName>
(5 spp),
<taxonomicName id="4C4916F6FFEFFFA3FB207F4EEE0599D3" authority="Villeneuve, 1915" authorityName="Villeneuve" authorityYear="1915" class="Insecta" family="Muscidae" genus="Passeromyia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="116" phylum="Arthropoda" rank="species" species="heterochaeta">
<emphasis id="B93DB167FFEFFFA3FB207F4EED7599D2" italics="true" pageId="22" pageNumber="116">Passeromyia heterochaeta</emphasis>
Villeneuve, 1915
</taxonomicName>
and
<taxonomicName id="4C4916F6FFEFFFA3FD9F7F6DEF6399D2" authority="Pont, 1970" authorityName="Pont" authorityYear="1970" box="[606,835,626,652]" class="Insecta" family="Muscidae" genus="Passeromyia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="116" phylum="Arthropoda" rank="species" species="steini">
<emphasis id="B93DB167FFEFFFA3FD9F7F6DEE9A99D2" box="[606,698,627,652]" italics="true" pageId="22" pageNumber="116">P. steini</emphasis>
<bibRefCitation id="EFD81084FFEFFFA3FD007F6CEF6399D2" author="Pont A. C. &amp; Shinonaga S." box="[705,835,626,652]" pageId="22" pageNumber="116" pagination="193 - 199" refId="ref15633" refString="Pont A. C. &amp; Shinonaga S. 1970. A new genus of Muscidae from Palaearctic East Asia (Diptera, Muscidae). Japanese Journal of Sanitary Zoology 21: 193 - 199." type="journal article" year="1970">Pont, 1970</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C4916F6FFEFFFA3FC8E7F6CEF9C99D2" authorityName="Meinert" authorityYear="1890" box="[847,956,626,652]" class="Insecta" family="Muscidae" genus="Philornis" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B93DB167FFEFFFA3FC8E7F6CEF9C99D2" box="[847,956,626,652]" italics="true" pageId="22" pageNumber="116">Philornis</emphasis>
</taxonomicName>
(5 spp).
</paragraph>
<caption id="DF363DFDFFEFFFA3FF7C7A94ED939C99" ID-DOI="http://doi.org/10.5281/zenodo.6784153" ID-Zenodo-Dep="6784153" httpUri="https://zenodo.org/record/6784153/files/figure.png" pageId="22" pageNumber="116" startId="22.[189,232,1930,1956]" targetBox="[189,1398,704,1908]" targetPageId="22">
<paragraph id="8BF66D75FFEFFFA3FF7C7A94ED939C99" blockId="22.[189,1398,1930,1991]" pageId="22" pageNumber="116">
<emphasis id="B93DB167FFEFFFA3FF7C7A94ED3C9CFA" bold="true" box="[189,284,1930,1956]" pageId="22" pageNumber="116">Fig. 11.</emphasis>
<taxonomicName id="4C4916F6FFEFFFA3FEE77A94EEF99CFA" authority="Schiner, 1868" authorityName="Schiner" authorityYear="1868" box="[294,729,1930,1956]" class="Insecta" family="Muscidae" genus="Atherigona" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="116" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B93DB167FFEFFFA3FEE77A94EE059CFA" box="[294,549,1930,1956]" italics="true" pageId="22" pageNumber="116">Atherigona orientalis</emphasis>
Schiner, 1868
</taxonomicName>
, ♀.
<emphasis id="B93DB167FFEFFFA3FCD07A94EF069CFA" bold="true" box="[785,806,1930,1956]" pageId="22" pageNumber="116">A</emphasis>
. Wing, ventral view.
<emphasis id="B93DB167FFEFFFA3FBF07A94E8659CFA" bold="true" box="[1073,1093,1930,1956]" pageId="22" pageNumber="116">B</emphasis>
. Detail of wing showing CS1‒3, ventral view.
</paragraph>
</caption>
<paragraph id="8BF66D75FFEEFFA2FF7C7C14EEC29A7D" blockId="23.[189,738,265,292]" box="[189,738,265,292]" pageId="23" pageNumber="117">
<emphasis id="B93DB167FFEEFFA2FF7C7C14ECE79A7D" box="[189,199,266,291]" italics="true" pageId="23" pageNumber="117">•</emphasis>
<taxonomicName id="4C4916F6FFEEFFA2FF177C17EEC29A7D" authority="Brauer &amp; Bergenstamm, 1893" authorityName="Brauer &amp; Bergenstamm" authorityYear="1893" box="[214,738,265,292]" class="Insecta" family="Muscidae" genus="Synthesiomyia" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B93DB167FFEEFFA2FF177C17ED5F9A7D" box="[214,383,265,291]" italics="true" pageId="23" pageNumber="117">Synthesiomyia</emphasis>
Brauer &amp; Bergenstamm, 1893
</taxonomicName>
</paragraph>
<paragraph id="8BF66D75FFEEFFA2FF7C7C29ED369AC6" blockId="23.[188,1399,311,408]" pageId="23" pageNumber="117">
The only included species,
<taxonomicName id="4C4916F6FFEEFFA2FE367C29EF229A0F" box="[503,770,311,337]" class="Insecta" family="Muscidae" genus="Synthesiomyia" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="117" phylum="Arthropoda" rank="species" species="nudipes">
<emphasis id="B93DB167FFEEFFA2FE367C29EF229A0F" box="[503,770,311,337]" italics="true" pageId="23" pageNumber="117">Synthesiomyia nudipes</emphasis>
</taxonomicName>
, has vein C only fully setulose ventrally on CS2 (state
<emphasis id="B93DB167FFEEFFA2FF7D7C44ECFF9A2A" bold="true" box="[188,223,346,372]" pageId="23" pageNumber="117">A2</emphasis>
), although some irregular ventral costal setulae extent onto CS2 or (females only) even the basal part of CS3.
</paragraph>
<paragraph id="8BF66D75FFEEFFA2FF7C7CA7E9749A8C" blockId="23.[189,1364,440,467]" box="[189,1364,440,467]" pageId="23" pageNumber="117">
<emphasis id="B93DB167FFEEFFA2FF7C7CA7ECE79A8C" box="[189,199,441,466]" italics="true" pageId="23" pageNumber="117">•</emphasis>
<taxonomicName id="4C4916F6FFEEFFA2FF177CA6EE7D9A8D" authority="Macquart, 1851" authorityName="Macquart" authorityYear="1851" box="[214,605,440,467]" class="Insecta" family="Muscidae" genus="Brachygasterina" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B93DB167FFEEFFA2FF177CA6EDBA9A8C" box="[214,410,440,466]" italics="true" pageId="23" pageNumber="117">Brachygasterina</emphasis>
Macquart, 1851
</taxonomicName>
,
<taxonomicName id="4C4916F6FFEEFFA2FDA87CA6EF8D9A8C" authority="Rondani, 1863" authorityName="Rondani" authorityYear="1863" box="[617,941,440,466]" class="Insecta" family="Muscidae" genus="Palpibracus" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B93DB167FFEEFFA2FDA87CA6EED89A8C" box="[617,760,440,466]" italics="true" pageId="23" pageNumber="117">Palpibracus</emphasis>
Rondani, 1863
</taxonomicName>
(
<figureCitation id="137271F0FFEEFFA2FC7C7CA6E8059A8C" box="[957,1061,440,466]" captionStart="Fig" captionStartId="24.[189,232,1876,1902]" captionTargetBox="[190,1398,677,1852]" captionTargetId="figure-150@24.[189,1399,670,1856]" captionTargetPageId="24" captionText="Fig. 12. ♀♀ of Muscidae Latreille, 1802. A. Muscina stabulans (Fallén, 1823), detail of wing showing CS1‒3, dorsal view. B. Palpibracus trivittatus (Malloch, 1934), detail of wing showing CS1‒3, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6784155" httpUri="https://zenodo.org/record/6784155/files/figure.png" pageId="23" pageNumber="117">Fig. 12B</figureCitation>
),
<taxonomicName id="4C4916F6FFEEFFA2FBFC7CA6E9749A8C" authority="Stein, 1911" authorityName="Stein" authorityYear="1911" box="[1085,1364,440,466]" class="Insecta" family="Muscidae" genus="Psilochaeta" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B93DB167FFEEFFA2FBFC7CA6E8E89A8C" box="[1085,1224,440,466]" italics="true" pageId="23" pageNumber="117">Psilochaeta</emphasis>
Stein, 1911
</taxonomicName>
</paragraph>
<paragraph id="8BF66D75FFEEFFA2FF7C7CF8E96199D3" blockId="23.[189,1399,486,654]" pageId="23" pageNumber="117">
The examined species of these three exclusively South American genera differ substantially from the preceding reinwardtiine genera by the dorsally extensively setulose vein C (state
<emphasis id="B93DB167FFEEFFA2FB047F17E8C9997D" bold="true" box="[1221,1257,521,547]" pageId="23" pageNumber="117">A8</emphasis>
,
<figureCitation id="137271F0FFEEFFA2FB3A7F17E949997A" box="[1275,1385,521,548]" captionStart="Fig" captionStartId="24.[189,232,1876,1902]" captionTargetBox="[190,1398,677,1852]" captionTargetId="figure-150@24.[189,1399,670,1856]" captionTargetPageId="24" captionText="Fig. 12. ♀♀ of Muscidae Latreille, 1802. A. Muscina stabulans (Fallén, 1823), detail of wing showing CS1‒3, dorsal view. B. Palpibracus trivittatus (Malloch, 1934), detail of wing showing CS1‒3, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6784155" httpUri="https://zenodo.org/record/6784155/files/figure.png" pageId="23" pageNumber="117">Fig. 12B</figureCitation>
):
<taxonomicName id="4C4916F6FFEEFFA2FF7C7F33EF389919" authority="(Malloch, 1922)" baseAuthorityName="Malloch" baseAuthorityYear="1922" box="[189,792,557,583]" class="Insecta" family="Muscidae" genus="Brachygasterina" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="117" phylum="Arthropoda" rank="species" species="fulvohumeralis">
<emphasis id="B93DB167FFEEFFA2FF7C7F33EE609919" box="[189,576,557,583]" italics="true" pageId="23" pageNumber="117">Brachygasterina fulvohumeralis</emphasis>
(Malloch, 1922)
</taxonomicName>
,
<taxonomicName id="4C4916F6FFEEFFA2FCEA7F30E8F39916" authority="Macquart, 1851" authorityName="Macquart" authorityYear="1851" box="[811,1235,557,584]" class="Insecta" family="Muscidae" genus="Brachygasterina" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="117" phylum="Arthropoda" rank="species" species="violaceiventris">
<emphasis id="B93DB167FFEEFFA2FCEA7F30E8219919" box="[811,1025,557,583]" italics="true" pageId="23" pageNumber="117">B. violaceiventris</emphasis>
Macquart, 1851
</taxonomicName>
,
<taxonomicName id="4C4916F6FFEEFFA2FB267F33E9569919" authorityName="Rondani" authorityYear="1863" box="[1255,1398,557,583]" class="Insecta" family="Muscidae" genus="Palpibracus" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B93DB167FFEEFFA2FB267F33E9569919" box="[1255,1398,557,583]" italics="true" pageId="23" pageNumber="117">Palpibracus</emphasis>
</taxonomicName>
(12 spp) and
<taxonomicName id="4C4916F6FFEEFFA2FE9E7F4EEF789934" authority="(Wiedemann, 1830)" baseAuthorityName="Wiedemann" baseAuthorityYear="1830" box="[351,856,592,618]" class="Insecta" family="Muscidae" genus="Psilochaeta" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="117" phylum="Arthropoda" rank="species" species="chalybea">
<emphasis id="B93DB167FFEEFFA2FE9E7F4EEE7E9934" box="[351,606,592,618]" italics="true" pageId="23" pageNumber="117">Psilochaeta chalybea</emphasis>
(Wiedemann, 1830)
</taxonomicName>
. Only
<taxonomicName id="4C4916F6FFEEFFA2FC6A7F4EE9579934" authority="Malloch, 1934" authorityName="Malloch" authorityYear="1934" box="[939,1399,592,618]" class="Insecta" family="Muscidae" genus="Brachygasterina" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="117" phylum="Arthropoda" rank="species" species="major">
<emphasis id="B93DB167FFEEFFA2FC6A7F4EE89E9934" box="[939,1214,592,618]" italics="true" pageId="23" pageNumber="117">Brachygasterina major</emphasis>
Malloch, 1934
</taxonomicName>
differs by having the dorsal setulae restricted to CS2 and CS3 (females) or CS3 (males) (state
<emphasis id="B93DB167FFEEFFA2FACD7F6DE90F99D3" bold="true" box="[1292,1327,627,653]" pageId="23" pageNumber="117">A7</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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