293 lines
34 KiB
XML
293 lines
34 KiB
XML
<document id="3B2CC1DA8EF7A2933DA0CC79A9BF8662" ID="10.11646/zootaxa.4147.5.7" ID-DOI="10.11646/zootaxa.4147.5.7" ID-GBIF-Dataset="5e3f6857-bc78-4a29-8047-72ebe5b3f54c" ID-ISSN="1175-5326" ID-Zenodo-Dep="266236" ID-ZooBank="8C7CB847-76EE-4738-BF59-DAB89BCBA6EF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1470637718685" checkinUser="plazi" docAuthor="Prokop, Jakub, Garrouste, Romain & Nel, André" docDate="2016" docId="03E0821EFFACFFD5FF705C12840A23CE" docLanguage="en" docName="zootaxa.4147.5.7.pdf" docOrigin="Zootaxa 4147 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hypopsylla belmontensis Prokop, Garrouste & Nel, 2016, sp. nov." docType="treatment" docVersion="8" lastPageNumber="592" masterDocId="FFD9FA66FFADFFD6FFE75F7D812A2132" masterDocTitle="A new permopsocidan genus and species from the Late Permian of Australia (Insecta: Acercaria: Psocidiidae)" masterLastPageNumber="592" masterPageNumber="589" pageNumber="590" updateTime="1698367179025" updateUser="plazi">
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<mods:mods id="5607D239AF258B05CE51689EEE2CA06A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="28DD61022AA3F556DD674AA4B0FAEF53">
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<mods:title id="F5406551BF05858A94FA972F1681B372">A new permopsocidan genus and species from the Late Permian of Australia (Insecta: Acercaria: Psocidiidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="547B880DAF60B3BC6204732F2DED4FEB" type="personal">
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<mods:role id="45D2FAA592A2B3FD3C7E3CAA5ABDE9A9">
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<mods:roleTerm id="62D050DF3824D233C80CE067986D3AF9">Author</mods:roleTerm>
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<mods:namePart id="30D9BD844D2575E33DF6B56AE5B4CDFF">Prokop, Jakub</mods:namePart>
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</mods:name>
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<mods:name id="748195C23D55C03D92A3CF36A7FC3250" type="personal">
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<mods:roleTerm id="361B234B9FC6EC0BFE741DF3F3F64FEC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="96D145DB1CB743787B48189FF9CCEF77">Garrouste, Romain</mods:namePart>
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</mods:name>
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<mods:name id="4AB4734AA261A0BC247FE07939B4D009" type="personal">
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<mods:role id="00458809BD9E8F18099752F957529707">
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<mods:roleTerm id="DE1D39B7E787303BA9BB627EADA27D10">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="08DCED4F2D198E0FED44CB582FC469B7">Nel, André</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="6BA84CA6C72499CBD886B12DFC463E71">text</mods:typeOfResource>
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<mods:title id="451B9A15B0FB2D620935F14CE4B5149A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B0C1CFB59A9B726B3CE9A51B7232C261">
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<mods:date id="B933459DF1BEE3B357069DDB5FA99C4D">2016</mods:date>
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<mods:detail id="8DA62C510E998257F5081C4297BAA6F5" type="volume">
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<mods:number id="F5975C8F2B6E8A686675F7719980B962">4147</mods:number>
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<mods:detail id="FCF50B69066CF390C37BD9F9363685D7" type="issue">
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<mods:number id="41607A729E7F1E8E39F76E2BFBA97892">5</mods:number>
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<mods:end id="E680CFF6D03150A10D0718B76735C698">592</mods:end>
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<mods:classification id="F6554A497D9BF83A3DF8C02A34D3A8A1">journal article</mods:classification>
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<mods:identifier id="BE3C98CD966AE1A4D2922750E85DAC80" type="DOI">10.11646/zootaxa.4147.5.7</mods:identifier>
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<mods:identifier id="B6D18FB6EB500E024E467DBDDA1C3FDB" type="GBIF-Dataset">5e3f6857-bc78-4a29-8047-72ebe5b3f54c</mods:identifier>
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<mods:identifier id="C5A11007E7BDF8F8E9ACD7D45121FDCF" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="87647D6BDA67ED4BA1B102C7E6D83A10" type="Zenodo-Dep">266236</mods:identifier>
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<treatment id="03E0821EFFACFFD5FF705C12840A23CE" ID-DOI="http://doi.org/10.5281/zenodo.6089814" ID-GBIF-Taxon="123514753" ID-Zenodo-Dep="6089814" LSID="urn:lsid:plazi:treatment:03E0821EFFACFFD5FF705C12840A23CE" httpUri="http://treatment.plazi.org/id/03E0821EFFACFFD5FF705C12840A23CE" lastPageId="3" lastPageNumber="592" pageId="1" pageNumber="590">
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<subSubSection id="C3536083FFACFFD7FF705C12802F229B" pageId="1" pageNumber="590" type="nomenclature">
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<paragraph id="8BF63308FFACFFD7FF705C12830222B8" blockId="1.[151,1435,879,1076]" box="[151,552,879,906]" pageId="1" pageNumber="590">
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<heading id="D0BE8464FFACFFD7FF705C12830222B8" bold="true" box="[151,552,879,906]" fontSize="11" level="1" pageId="1" pageNumber="590" reason="1">
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<emphasis id="B93DEF1AFFACFFD7FF705C12830222B8" bold="true" box="[151,552,879,906]" pageId="1" pageNumber="590">
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<taxonomicName id="4C49488BFFACFFD7FF705C1280E922BB" box="[151,451,879,905]" class="Insecta" family="Psocidiidae" genus="Hypopsylla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Permopsocida" pageId="1" pageNumber="590" phylum="Arthropoda" rank="species" species="belmontensis" status="sp. nov.">
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<emphasis id="B93DEF1AFFACFFD7FF705C1280E922BB" bold="true" box="[151,451,879,905]" italics="true" pageId="1" pageNumber="590">Hypopsylla belmontensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20E5261FFACFFD7FE2C5C0D830222B8" box="[459,552,880,906]" pageId="1" pageNumber="590" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="8BF63308FFACFFD7FF705CEF802F229B" blockId="1.[151,1435,879,1076]" box="[151,261,914,937]" pageId="1" pageNumber="590">
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(
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<figureCitation id="13722F8DFFACFFD7FF795CEF81C8229B" box="[158,226,914,937]" captionStart="FIGURE 1" captionStartId="1.[151,244,1705,1727]" captionTargetBox="[151,1431,1138,1681]" captionTargetId="figure@1.[151,1436,1127,1683]" captionTargetPageId="1" captionText="FIGURE 1. Hypopsylla belmontensis, line drawing of forewing venation. Scale bar represents 1 mm." httpUri="https://zenodo.org/record/266237/files/figure.png" pageId="1" pageNumber="590">Figs 1</figureCitation>
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,
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<figureCitation id="13722F8DFFACFFD7FF085CEF81D7229B" box="[239,253,914,937]" captionStart="FIGURE 2" captionStartId="2.[151,244,1624,1646]" captionTargetBox="[150,1435,434,1602]" captionTargetId="figure@2.[150,1436,434,1603]" captionTargetPageId="2" captionText="FIGURE 2. Hypopsylla belmontensis, holotype In 45397 / In 46044. A, photograph print under dry condition; B, photograph imprint under alcohol (copyright Natural History Museum, London). Scale bars represent 1 mm." httpUri="https://zenodo.org/record/266238/files/figure.png" pageId="1" pageNumber="590">2</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3536083FFACFFD7FF705CC883C42506" pageId="1" pageNumber="590" type="materials_examined">
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||
<paragraph id="8BF63308FFACFFD7FF705CC884B122DD" blockId="1.[151,1435,879,1076]" pageId="1" pageNumber="590">
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<materialsCitation id="3B213955FFACFFD7FF705CC884B122DD" ID-GBIF-Occurrence="1291043942" county="In" location="Age" municipality="In" pageId="1" pageNumber="590" specimenCount="1" stateProvince="Newcastle" typeStatus="holotype">
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<emphasis id="B93DEF1AFFACFFD7FF705CC8802B22FE" bold="true" box="[151,257,949,972]" pageId="1" pageNumber="590">Material.</emphasis>
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<typeStatus id="54F28DAAFFACFFD7FEE05CC8804022FE" box="[263,362,949,972]" pageId="1" pageNumber="590" type="holotype">Holotype</typeStatus>
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<collectingCounty id="62974B84FFACFFD7FE965CC880AD22FE" box="[369,391,949,972]" pageId="1" pageNumber="590">In</collectingCounty>
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45397/
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<collectingMunicipality id="6B92A972FFACFFD7FE3E5CC880C522FE" box="[473,495,949,972]" pageId="1" pageNumber="590">In</collectingMunicipality>
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46044 (print and counterprint) (The Natural Museum, London).
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<emphasis id="B93DEF1AFFACFFD7FF205CA580A622DD" bold="true" box="[199,396,984,1007]" pageId="1" pageNumber="590">
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<location id="8E9665D3FFACFFD7FF205CA581D922DD" LSID="urn:lsid:plazi:treatment:03E0821EFFACFFD5FF705C12840A23CE:8E9665D3FFACFFD7FF205CA581D922DD" box="[199,243,984,1007]" county="In" municipality="In" name="Age" pageId="1" pageNumber="590" stateProvince="Newcastle">Age</location>
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and outcrop.
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</emphasis>
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<location id="8E9665D3FFACFFD7FE735CA5830C22DD" LSID="urn:lsid:plazi:treatment:03E0821EFFACFFD5FF705C12840A23CE:8E9665D3FFACFFD7FE735CA5830C22DD" box="[404,550,984,1007]" county="In" municipality="In" name="Late Permian" pageId="1" pageNumber="590" stateProvince="Newcastle">Late Permian</location>
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, mid
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<location id="8E9665D3FFACFFD7FD855CA583F822DD" LSID="urn:lsid:plazi:treatment:03E0821EFFACFFD5FF705C12840A23CE:8E9665D3FFACFFD7FD855CA583F822DD" box="[610,722,984,1007]" county="In" municipality="In" name="Lopingian" pageId="1" pageNumber="590" stateProvince="Newcastle">Lopingian</location>
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, cca 255
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<location id="8E9665D3FFACFFD7FCDE5CA5824722DD" LSID="urn:lsid:plazi:treatment:03E0821EFFACFFD5FF705C12840A23CE:8E9665D3FFACFFD7FCDE5CA5824722DD" box="[825,877,984,1007]" county="In" municipality="In" name="Mya" pageId="1" pageNumber="590" stateProvince="Newcastle">Mya</location>
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,
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<collectingRegion id="498DFDEAFFACFFD7FC9F5CA582CD22DD" box="[888,999,984,1007]" country="United Kingdom" name="Newcastle upon Tyne" pageId="1" pageNumber="590">Newcastle</collectingRegion>
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Coal Measures,
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<location id="8E9665D3FFACFFD7FB7A5CA5840222DD" LSID="urn:lsid:plazi:treatment:03E0821EFFACFFD5FF705C12840A23CE:8E9665D3FFACFFD7FB7A5CA5840222DD" box="[1181,1320,984,1007]" county="In" municipality="In" name="Warners Bay" pageId="1" pageNumber="590" stateProvince="Newcastle">Warners Bay</location>
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- Belmont
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</materialsCitation>
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</paragraph>
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<paragraph id="8BF63308FFACFFD7FF705C8783C42506" blockId="1.[151,1435,879,1076]" pageId="1" pageNumber="590">
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<materialsCitation id="3B213955FFACFFD7FF705C8783C42506" ID-GBIF-Occurrence="1291043943" country="Australia" location="Newcastle" pageId="1" pageNumber="590" specimenCount="1" stateProvince="New South Wales" typeStatus="holotype">
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area near
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<location id="8E9665D3FFACFFD7FF195C87805A2523" LSID="urn:lsid:plazi:treatment:03E0821EFFACFFD5FF705C12840A23CE:8E9665D3FFACFFD7FF195C87805A2523" box="[254,368,1018,1041]" country="Australia" name="Newcastle" pageId="1" pageNumber="590" stateProvince="New South Wales">Newcastle</location>
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,
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<collectingRegion id="498DFDEAFFACFFD7FE9D5C8783162523" box="[378,572,1018,1041]" country="Australia" name="New South Wales" pageId="1" pageNumber="590">
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New
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<collectingRegion id="498DFDEAFFACFFD7FE535C8783162523" box="[436,572,1018,1041]" inferredBySuffix="true" pageId="1" pageNumber="590">South Wales</collectingRegion>
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</collectingRegion>
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||
,
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<collectingCountry id="F35E7398FFACFFD7FDA15C8783822523" box="[582,680,1018,1041]" name="Australia" pageId="1" pageNumber="590">Australia</collectingCountry>
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||
(
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<bibRefCitation id="EFD84EF9FFACFFD7FD515C8782722523" author="Shi" box="[694,856,1017,1041]" pageId="1" pageNumber="590" refString="Shi, G. R., Waterhouse, J. B. & McLoughlin, S. (2010) The Lopingian of Australasia: a review of biostratigraphy, correlations, palaeogeography and palaeobiogeography. Geological Journal, 45, 230 - 263. http: // dx. doi. org / 10.1002 / gj. 1213" type="journal article" year="2010">
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Shi
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<emphasis id="B93DEF1AFFACFFD7FD055C8782322523" box="[738,792,1017,1041]" italics="true" pageId="1" pageNumber="590">et al.</emphasis>
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||
2010
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||
</bibRefCitation>
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||
).
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||
<emphasis id="B93DEF1AFFACFFD7FF205B60806D2506" bold="true" box="[199,327,1053,1076]" pageId="1" pageNumber="590">Etymology.</emphasis>
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Named after the
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<typeStatus id="54F28DAAFFACFFD7FDE75B6083042506" box="[512,558,1053,1076]" pageId="1" pageNumber="590">type</typeStatus>
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locality Belmont.
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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<caption id="DF366380FFACFFD7FF7059D4854F278C" box="[151,1125,1705,1727]" httpUri="https://zenodo.org/record/266237/files/figure.png" pageId="1" pageNumber="590" targetBox="[151,1431,1138,1681]" targetPageId="1">
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<paragraph id="8BF63308FFACFFD7FF7059D4854F278C" blockId="1.[151,1125,1705,1727]" box="[151,1125,1705,1727]" pageId="1" pageNumber="590">
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<emphasis id="B93DEF1AFFACFFD7FF7059D48027278D" bold="true" box="[151,269,1705,1727]" pageId="1" pageNumber="590">FIGURE 1.</emphasis>
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<taxonomicName id="4C49488BFFACFFD7FEF559D48328278C" box="[274,514,1705,1726]" class="Insecta" family="Psocidiidae" genus="Hypopsylla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Permopsocida" pageId="1" pageNumber="590" phylum="Arthropoda" rank="species" species="belmontensis">
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<emphasis id="B93DEF1AFFACFFD7FEF559D48328278C" box="[274,514,1705,1726]" italics="true" pageId="1" pageNumber="590">Hypopsylla belmontensis</emphasis>
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</taxonomicName>
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, line drawing of forewing venation. Scale bar represents 1 mm.
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</paragraph>
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</caption>
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<subSubSection id="C3536083FFACFFD4FF2059998468204D" lastPageId="2" lastPageNumber="591" pageId="1" pageNumber="590" type="description">
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<paragraph id="8BF63308FFACFFD4FF2059998468204D" blockId="1.[151,1436,1764,2030]" lastBlockId="2.[151,1435,152,383]" lastPageId="2" lastPageNumber="591" pageId="1" pageNumber="590">
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<emphasis id="B93DEF1AFFACFFD7FF205999806127C9" bold="true" box="[199,331,1764,1787]" pageId="1" pageNumber="590">Description</emphasis>
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. Print and counterprint of two wings partly overlapping and deformed, these wings being the fore- and the hind wings due to their relative positions. Their venations are nearly identical. Forewing nearly complete, fragment 6.0 mm long,
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||
<quantity id="4CB19EEDFFACFFD7FED1585480A62672" box="[310,396,1833,1856]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="590" unit="mm" value="2.5">2.5 mm</quantity>
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||
wide in widest part; ScP ending on RA about
|
||
<quantity id="4CB19EEDFFACFFD7FC79585482D82672" box="[926,1010,1833,1856]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="590" unit="mm" value="2.7">2.7 mm</quantity>
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||
from wing base, but with a crossvein between it and costa; area between radius and costa broad,
|
||
<quantity id="4CB19EEDFFACFFD7FCF3583182592651" box="[788,883,1868,1891]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="590" unit="mm" value="0.25">0.25 mm</quantity>
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||
wide; R, M and CuA fused into a common stem at wing base, making a weak posterior curve for about
|
||
<quantity id="4CB19EEDFFACFFD7FD2E5812823426B4" box="[713,798,1903,1926]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="590" unit="mm" value="1.1">1.1 mm</quantity>
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||
; then M+CuA and radius separating, with R and basal stem R+M+CuA making a pronounced angle at this point; RP and RA separating
|
||
<quantity id="4CB19EEDFFACFFD7FC3158EC8502269A" box="[982,1064,1937,1960]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="590" unit="mm" value="1.2">1.2 mm</quantity>
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||
distal of base of M+CuA; convex RA with a deep posterior curve limiting a dark pigmented pterostigma, at least
|
||
<quantity id="4CB19EEDFFACFFD7FBCB58C985A926F9" box="[1068,1155,1972,1995]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="590" unit="mm" value="1.2">1.2 mm</quantity>
|
||
long and
|
||
<quantity id="4CB19EEDFFACFFD7FB1958C9847F26F9" box="[1278,1365,1972,1995]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="590" unit="mm" value="0.5">0.5 mm</quantity>
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||
wide, pterostigma limited by a vein basally (ScP); a crossvein perpendicular to RA and to RP below middle of pterostigma; relatively concave RP without visible fork in preserved part; M and CuA separating immediately distal of point of reemergence of M+CuA; neutral stem of M very long,
|
||
<quantity id="4CB19EEDFFAFFFD4FD2A5FC7823521E3" box="[717,799,186,209]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="591" unit="mm" value="2.5">2.5 mm</quantity>
|
||
long before its first fork; anterior branch of M distally with a deep fork; posterior branch of M with three branches ending on posterior wing margin; convex CuA very short before crossvein cua-cup ends on its, being aligned with distal part of CuA; distal part of CuA very long,
|
||
<quantity id="4CB19EEDFFAFFFD4FB215E7D84322025" box="[1222,1304,256,279]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="591" unit="mm" value="2.5">2.5 mm</quantity>
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||
long before areola postica; areola postica very long and narrow, parallel to posterior wing margin, with CuA1 curved and CuA2 very short; cua-cup weaker than CuA and M,
|
||
<quantity id="4CB19EEDFFAFFFD4FDBA5E38839A206E" box="[605,688,325,348]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="591" unit="mm" value="0.9">0.9 mm</quantity>
|
||
long from base of CuP and CuA; concave CuP weakly curved and simple; two convex simple anal veins basally curved; anal area about
|
||
<quantity id="4CB19EEDFFAFFFD4FC995E1582FA204D" box="[894,976,360,383]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="591" unit="mm" value="2.2">2.2 mm</quantity>
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||
long along posterior wing margin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF366380FFAFFFD4FF70592582C227B8" httpUri="https://zenodo.org/record/266238/files/figure.png" pageId="2" pageNumber="591" targetBox="[150,1435,434,1602]" targetPageId="2">
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<paragraph id="8BF63308FFAFFFD4FF70592582C227B8" blockId="2.[151,1435,1624,1674]" pageId="2" pageNumber="591">
|
||
<emphasis id="B93DEF1AFFAFFFD4FF7059258027275C" bold="true" box="[151,269,1624,1646]" pageId="2" pageNumber="591">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C49488BFFAFFFD4FEF35925832C275F" box="[276,518,1624,1645]" class="Insecta" family="Psocidiidae" genus="Hypopsylla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Permopsocida" pageId="2" pageNumber="591" phylum="Arthropoda" rank="species" species="belmontensis">
|
||
<emphasis id="B93DEF1AFFAFFFD4FEF35925832C275F" box="[276,518,1624,1645]" italics="true" pageId="2" pageNumber="591">Hypopsylla belmontensis</emphasis>
|
||
</taxonomicName>
|
||
, holotype In 45397/In 46044.
|
||
<emphasis id="B93DEF1AFFAFFFD4FCD75925826B275C" bold="true" box="[816,833,1624,1646]" pageId="2" pageNumber="591">A</emphasis>
|
||
, photograph print under dry condition;
|
||
<emphasis id="B93DEF1AFFAFFFD4FB23592585FE275C" bold="true" box="[1220,1236,1624,1646]" pageId="2" pageNumber="591">B</emphasis>
|
||
, photograph imprint under alcohol (copyright Natural History Museum, London). Scale bars represent 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3536083FFAFFFD4FF2059CB843827DD" pageId="2" pageNumber="591" type="discussion">
|
||
<paragraph id="8BF63308FFAFFFD4FF2059CB843827DD" blockId="2.[151,1436,1717,2018]" pageId="2" pageNumber="591">
|
||
<emphasis id="B93DEF1AFFAFFFD4FF2059CB806C27FF" bold="true" box="[199,326,1718,1741]" pageId="2" pageNumber="591">Discussion.</emphasis>
|
||
The pattern of wing venation of
|
||
<taxonomicName id="4C49488BFFAFFFD4FD4359C8823427FF" box="[676,798,1717,1741]" class="Insecta" family="Psocidiidae" genus="Hypopsylla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Permopsocida" pageId="2" pageNumber="590" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B93DEF1AFFAFFFD4FD4359C8823427FF" box="[676,798,1717,1741]" italics="true" pageId="2" pageNumber="591">Hypopsylla</emphasis>
|
||
</taxonomicName>
|
||
is typical of the Acercaria with a common stem R+M+CuA and a neutral crossvein cua-cup between concave CuP and convex CuA weaker than CuA (
|
||
<bibRefCitation id="EFD84EF9FFAFFFD4FB8659A5842C27DD" author="Nel" box="[1121,1286,1751,1776]" pageId="2" pageNumber="591" refString="Nel, A., Prokop, J., Nel, P., Grandcolas, P., Huang, Di-ying, Roques, P., Guilbert, E., Dostal, O. & Szwedo, J. (2012) Traits and evolution of wing venation pattern in paraneopteran insects. Journal of Morphology, 273, 480 - 506. http: // dx. doi. org / 10.1002 / jmor. 11036" type="journal article" year="2012">
|
||
Nel
|
||
<emphasis id="B93DEF1AFFAFFFD4FB7759A485EC27DD" box="[1168,1222,1751,1776]" italics="true" pageId="2" pageNumber="591">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3536083FFAFFFD5FF205987806E2343" lastPageId="3" lastPageNumber="592" pageId="2" pageNumber="591" type="reference_group">
|
||
<paragraph id="8BF63308FFAFFFD5FF20598782DC20F6" blockId="2.[151,1436,1717,2018]" lastBlockId="3.[151,1436,152,764]" lastPageId="3" lastPageNumber="592" pageId="2" pageNumber="591">
|
||
<taxonomicName id="4C49488BFFAFFFD4FF205987806B2620" box="[199,321,1786,1810]" class="Insecta" family="Psocidiidae" genus="Hypopsylla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Permopsocida" pageId="2" pageNumber="590" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B93DEF1AFFAFFFD4FF205987806B2620" box="[199,321,1786,1810]" italics="true" pageId="2" pageNumber="591">Hypopsylla</emphasis>
|
||
</taxonomicName>
|
||
has the wing venation synapomorphies of the
|
||
<taxonomicName id="4C49488BFFAFFFD4FCD6598685432620" authority="Tillyard, 1926" authorityName="Tillyard" authorityYear="1926" box="[817,1129,1787,1810]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Permopsocida" pageId="2" pageNumber="591" phylum="Arthropoda" rank="order">
|
||
Permopsocida
|
||
<bibRefCitation id="EFD84EF9FFAFFFD4FC28598685432620" author="Tillyard" box="[975,1129,1787,1810]" pageId="2" pageNumber="591" refString="Tillyard, R. J. (1926) Kansas Permian insects. 8. Copeognatha. American Journal of Science, Series 5, 11, 314 - 349." type="journal article" year="1926">Tillyard, 1926</bibRefCitation>
|
||
</taxonomicName>
|
||
, as redefined in Huang
|
||
<emphasis id="B93DEF1AFFAFFFD4FA81598684B62620" box="[1382,1436,1786,1810]" italics="true" pageId="2" pageNumber="591">et al.</emphasis>
|
||
(2016), viz. fore- and hind wings with sclerotized pterostigma posteriorly limited by RA, with RA making a strong angle at the point of contact with crossvein between it and RP (synapomorphy of the
|
||
<taxonomicName id="4C49488BFFAFFFD4FC16583D85A12665" box="[1009,1163,1856,1879]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Permopsocida" pageId="2" pageNumber="591" phylum="Arthropoda" rank="order">Permopsocida</taxonomicName>
|
||
, a structure found also in some Psocodean forewing but never in their hind wing); a distinct angle of radial vein at base of M+CuA (a character absent in
|
||
<taxonomicName id="4C49488BFFAFFFD4FF1A58FB804826AF" ID-CoL="8MP8R" box="[253,354,1926,1949]" class="Insecta" kingdom="Animalia" order="Psocodea" pageId="2" pageNumber="591" phylum="Arthropoda" rank="order">Psocodea</taxonomicName>
|
||
and Thripida, but present to a certain point in the forewings of many
|
||
<taxonomicName id="4C49488BFFAFFFD4FBAF58FB859226AF" ID-CoL="HP" box="[1096,1208,1926,1949]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="591" phylum="Arthropoda" rank="order">Hemiptera</taxonomicName>
|
||
of different lineages, viz. the
|
||
<taxonomicName id="4C49488BFFAFFFD4FF1358D5808E268D" box="[244,420,1960,1983]" class="Insecta" family="Archescytinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="591" phylum="Arthropoda" rank="family">Archescytinidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C49488BFFAFFFD4FE5458D5831E268D" ID-CoL="8HBR9" box="[435,564,1960,1983]" class="Insecta" family="Prosbolidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="591" phylum="Arthropoda" rank="family">Prosbolidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C49488BFFAFFFD4FDA558D583F6268D" box="[578,732,1960,1983]" class="Insecta" family="Pincombeidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="591" phylum="Arthropoda" rank="family">Pincombeidae</taxonomicName>
|
||
, etc.); presence of an areola postica in forewings (probably an acercarian symplesiomorphy as this structure is present in
|
||
<taxonomicName id="4C49488BFFAFFFD4FCE458B6824D26D0" ID-CoL="8MP8R" box="[771,871,1995,2018]" class="Insecta" kingdom="Animalia" order="Psocodea" pageId="2" pageNumber="591" phylum="Arthropoda" rank="order">Psocodea</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C49488BFFAFFFD4FC7D58B6852126D0" ID-CoL="HP" box="[922,1035,1995,2018]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="591" phylum="Arthropoda" rank="order">Hemiptera</taxonomicName>
|
||
but reduced in Thripida); presence of an areola postica (forked CuA) in hindwings, a character present in
|
||
<taxonomicName id="4C49488BFFAEFFD5FC715FE5852C219D" ID-CoL="HP" box="[918,1030,152,175]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="592" phylum="Arthropoda" rank="order">Hemiptera</taxonomicName>
|
||
but very rare in
|
||
<taxonomicName id="4C49488BFFAEFFD5FB2C5FE5841A219D" ID-CoL="8MP8R" box="[1227,1328,152,175]" class="Insecta" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="592" phylum="Arthropoda" rank="order">Psocodea</taxonomicName>
|
||
(in some specimens of
|
||
<taxonomicName id="4C49488BFFAEFFD5FECB5FC783EB21E3" ID-CoL="8N87D" authority="Rambur, 1842" authorityName="Rambur" authorityYear="1842" box="[300,705,185,209]" class="Insecta" family="Epipsocidae" genus="Epipsocus" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="592" phylum="Arthropoda" rank="species" species="lucifugus">
|
||
<emphasis id="B93DEF1AFFAEFFD5FECB5FC7832921E3" box="[300,515,185,209]" italics="true" pageId="3" pageNumber="592">Epipsocus lucifugus</emphasis>
|
||
(Rambur, 1842))
|
||
</taxonomicName>
|
||
, and absent in the Thripida; RP with a simple apical fork (structure also present in
|
||
<taxonomicName id="4C49488BFFAEFFD5FED15FA0832D21C6" ID-CoL="8MP8R" authority="Thripida" authorityName="Thripida" box="[310,519,221,244]" class="Insecta" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="592" phylum="Arthropoda" rank="order">Psocodea, Thripida</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C49488BFFAEFFD5FDDA5FA0839921C6" ID-CoL="HP" box="[573,691,221,244]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="592" phylum="Arthropoda" rank="order">Hemiptera</taxonomicName>
|
||
); M with two main branches, each forked again (a character present in
|
||
<taxonomicName id="4C49488BFFAEFFD5FF705E7D80592025" ID-CoL="HP" authority="Thripida" authorityName="Thripida" box="[151,371,256,279]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="592" phylum="Arthropoda" rank="order">Hemiptera, Thripida</taxonomicName>
|
||
have either a simple or a forked M, while
|
||
<taxonomicName id="4C49488BFFAEFFD5FCB45E7D82922025" ID-CoL="8MP8R" box="[851,952,256,279]" class="Insecta" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="592" phylum="Arthropoda" rank="order">Psocodea</taxonomicName>
|
||
have a three- to simple M). The ScP either parallel to radius, or fused to costal margin, re-emerging from it or not, are character states present in various
|
||
<taxonomicName id="4C49488BFFAEFFD5FAC75E5F84B2200B" ID-CoL="8MP8S" box="[1312,1432,290,313]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocoptera" pageId="3" pageNumber="592" phylum="Arthropoda" rank="order">Psocoptera</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C49488BFFAEFFD5FF705E39803B206E" box="[151,273,324,348]" class="Insecta" family="Psocidiidae" genus="Hypopsylla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Permopsocida" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B93DEF1AFFAEFFD5FF705E39803B206E" box="[151,273,324,348]" italics="true" pageId="3" pageNumber="592">Hypopsylla</emphasis>
|
||
</taxonomicName>
|
||
differs from the other representatives of the clade
|
||
<taxonomicName id="4C49488BFFAEFFD5FCCE5E3882EA206E" box="[809,960,325,348]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Permopsocida" pageId="3" pageNumber="592" phylum="Arthropoda" rank="order">Permopsocida</taxonomicName>
|
||
in the much deeper posterior curve of RA in the pterostigma and the presence of three branches of posterior branch of M. This particular pterostigma reminds the pterostigma of the ‘hypoperlid’
|
||
<taxonomicName id="4C49488BFFAEFFD5FE0C5EF483402093" box="[491,618,393,417]" class="Insecta" family="Hypoperlidae" genus="Idelopsocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hypoperlida" pageId="3" pageNumber="592" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93DEF1AFFAEFFD5FE0C5EF483402093" box="[491,618,393,417]" italics="true" pageId="3" pageNumber="592">Idelopsocus</emphasis>
|
||
</taxonomicName>
|
||
, with the important difference in the fact that the pterostigma is covering the area between C and RA but also part of area between RA and RP in
|
||
<emphasis id="B93DEF1AFFAEFFD5FC975ED182C520F6" box="[880,1007,428,452]" italics="true" pageId="3" pageNumber="592">Ideloposcus</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF63308FFAEFFD5FF205EB2806E2343" blockId="3.[151,1436,152,764]" pageId="3" pageNumber="592">
|
||
<taxonomicName id="4C49488BFFAEFFD5FF205EB2806B20D5" box="[199,321,463,487]" class="Insecta" family="Psocidiidae" genus="Hypopsylla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Permopsocida" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B93DEF1AFFAEFFD5FF205EB2806B20D5" box="[199,321,463,487]" italics="true" pageId="3" pageNumber="592">Hypopsylla</emphasis>
|
||
</taxonomicName>
|
||
differs from the Permopsociidae in the absence of a crossvein between M and CuA. It differs from the
|
||
<taxonomicName id="4C49488BFFAEFFD5FF705E8F801E233B" box="[151,308,498,521]" class="Insecta" family="Archipsyllidae" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="592" phylum="Arthropoda" rank="family">Archipsyllidae</taxonomicName>
|
||
in the ScP ending in RA instead of going into costal margin (and re-emerging or not distally to reach RA) (Huang
|
||
<emphasis id="B93DEF1AFFAEFFD5FF095D68800F231E" box="[238,293,532,556]" italics="true" pageId="3" pageNumber="592">et al.</emphasis>
|
||
2016 suppl. inf.). The presence of a crossvein between RP and the posterior side of the pterostigma is not discriminant as it is present in the three families (
|
||
<taxonomicName id="4C49488BFFAEFFD5FD4E5D4582DC237D" box="[681,1014,568,591]" class="Insecta" family="Tachinidae" genus="Permopsocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="592" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93DEF1AFFAEFFD5FD4E5D458213237D" box="[681,825,568,591]" italics="true" pageId="3" pageNumber="592">Permopsocus</emphasis>
|
||
(Permopsocidae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C49488BFFAEFFD5FBE55D4A8400237D" box="[1026,1322,567,591]" class="Insecta" family="Tachinidae" genus="Dichentomum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Permopsocida" pageId="3" pageNumber="592" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93DEF1AFFAEFFD5FBE55D4A85BD237D" box="[1026,1175,567,591]" italics="true" pageId="3" pageNumber="592">Dichentomum</emphasis>
|
||
(Psocidiidae)
|
||
</taxonomicName>
|
||
and in the
|
||
<taxonomicName id="4C49488BFFAEFFD5FF705D2780132343" box="[151,313,602,625]" class="Insecta" family="Archipsyllidae" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="592" phylum="Arthropoda" rank="family">Archipsyllidae</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3536083FFAEFFD5FF205D01840A23CE" pageId="3" pageNumber="592" type="description">
|
||
<paragraph id="8BF63308FFAEFFD5FF205D01840A23CE" blockId="3.[151,1436,152,764]" pageId="3" pageNumber="592">
|
||
<taxonomicName id="4C49488BFFAEFFD5FF205D01806B23A6" box="[199,321,636,660]" class="Insecta" family="Psocidiidae" genus="Hypopsylla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Permopsocida" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B93DEF1AFFAEFFD5FF205D01806B23A6" box="[199,321,636,660]" italics="true" pageId="3" pageNumber="592">Hypopsylla</emphasis>
|
||
</taxonomicName>
|
||
shares with the
|
||
<taxonomicName id="4C49488BFFAEFFD5FE0A5D00834223A6" box="[493,616,637,660]" class="Insecta" family="Psocidiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Permopsocida" pageId="3" pageNumber="592" phylum="Arthropoda" rank="family">Psocidiidae</taxonomicName>
|
||
the elongate and narrow areola postica, absence of crossvein between M and CuA, presence of a short crossvein between ScP and costa as in
|
||
<taxonomicName id="4C49488BFFAEFFD5FCA85DE282CE2385" box="[847,996,671,695]" class="Insecta" family="Tachinidae" genus="Dichentomum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Permopsocida" pageId="3" pageNumber="592" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93DEF1AFFAEFFD5FCA85DE282CE2385" box="[847,996,671,695]" italics="true" pageId="3" pageNumber="592">Dichentomum</emphasis>
|
||
</taxonomicName>
|
||
, ScP ending on RA distal of base of RP, but closer to this point than to the pterostigma, unlike to the other taxa of this family. The wing venation of
|
||
<taxonomicName id="4C49488BFFAEFFD5FAC45DBC800823CE" class="Insecta" family="Psocidiidae" genus="Hypopsylla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Permopsocida" pageId="3" pageNumber="592" phylum="Arthropoda" rank="species" species="belmontensis">
|
||
<emphasis id="B93DEF1AFFAEFFD5FAC45DBC800823CE" italics="true" pageId="3" pageNumber="592">Hypopsylla belmontensis</emphasis>
|
||
</taxonomicName>
|
||
allows its assignment to the
|
||
<taxonomicName id="4C49488BFFAEFFD5FDB15D9883FE23CE" box="[598,724,741,764]" class="Insecta" family="Psocidiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Permopsocida" pageId="3" pageNumber="592" phylum="Arthropoda" rank="family">Psocidiidae</taxonomicName>
|
||
, increasing the diversity of the Permian
|
||
<taxonomicName id="4C49488BFFAEFFD5FB665D98843623CE" box="[1153,1308,741,764]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Permopsocida" pageId="3" pageNumber="592" phylum="Arthropoda" rank="order">Permopsocida</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |