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<mods:title id="C28B53CCECB88128F8B1CBA2D8D1FBED">An illustrated guide to the lady beetles (Coleoptera: Coccinellidae) of the Indian Subcontinent. Part II. Tribe Chilocorini</mods:title>
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<treatment id="03C44153FF8DFF89FF77F93CFF7DF8E3" ID-DOI="http://doi.org/10.5281/zenodo.11067907" ID-Zenodo-Dep="11067907" LSID="urn:lsid:plazi:treatment:03C44153FF8DFF89FF77F93CFF7DF8E3" httpUri="http://treatment.plazi.org/id/03C44153FF8DFF89FF77F93CFF7DF8E3" lastPageId="36" lastPageNumber="37" pageId="33" pageNumber="34">
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<paragraph id="8BD2F045FF8DFF8CFF77F93CFE35F92B" blockId="33.[151,443,1707,1770]" box="[151,443,1707,1734]" pageId="33" pageNumber="34">
<heading id="D09A4729FF8DFF8CFF77F93CFE35F92B" bold="true" box="[151,443,1707,1734]" fontSize="11" level="2" pageId="33" pageNumber="34" reason="4">
<taxonomicName id="4C6D8BC6FF8DFF8CFF77F93CFE35F92B" authority="Weise" authorityName="Weise" box="[151,443,1707,1734]" class="Insecta" family="Coccinellidae" genus="Chilocorus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="braeti">
<emphasis id="B9192C57FF8DFF8CFF77F93CFE35F92B" bold="true" box="[151,443,1707,1734]" pageId="33" pageNumber="34">
<emphasis id="B9192C57FF8DFF8CFF77F93CFEE4F928" bold="true" box="[151,362,1707,1734]" italics="true" pageId="33" pageNumber="34">Chilocorus braeti</emphasis>
Weise
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C377A3CEFF8DFF8CFF77F958FF77F904" box="[151,249,1743,1770]" pageId="33" pageNumber="34" type="description">
<paragraph id="8BD2F045FF8DFF8CFF77F958FF77F904" blockId="33.[151,443,1707,1770]" box="[151,249,1743,1770]" pageId="33" pageNumber="34">
(
<figureCitation id="1356ECC0FF8DFF8CFF7FF958FF7FF904" box="[159,241,1743,1770]" captionStart="FIGURE 25" captionStartId="36.[151,250,1456,1480]" captionTargetBox="[154,1400,183,1431]" captionTargetId="figure-20@36.[151,1436,181,1432]" captionTargetPageId="36" captionText="FIGURE 25. Chilocorus braeti Weise: a, b. adult, dorsal view (BMNH); b. adult, dorsal view (ZSIK); c, d. male genitalia: c. tegmen, ventral view; d. penis." figureDoi="http://doi.org/10.5281/zenodo.10208973" httpUri="https://zenodo.org/record/10208973/files/figure.png" pageId="33" pageNumber="34">Fig. 25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C377A3CEFF8DFF8CFF77F899FBF2F8AB" pageId="33" pageNumber="34" type="reference_group">
<paragraph id="8BD2F045FF8DFF8CFF77F899FD9BF8C8" blockId="33.[151,1148,1806,1861]" box="[151,533,1806,1830]" pageId="33" pageNumber="34">
<treatmentCitationGroup id="AB7DD76BFF8DFF8CFF77F899FD9BF8C8" box="[151,533,1806,1830]" pageId="33" pageNumber="34">
<treatmentCitation id="0ACCD654FF8DFF8CFF77F899FD9FF8C8" author="Weise, J." box="[151,529,1806,1830]" page="154" pageId="33" pageNumber="34" year="1895">
<taxonomicName id="4C6D8BC6FF8DFF8CFF77F899FD9FF8C8" authority="Weise, 1895 b: 154" authorityName="Weise" authorityPageNumber="154" authorityYear="1895" box="[151,529,1806,1830]" class="Insecta" family="Coccinellidae" genus="Chilocorus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="braeti">
<emphasis id="B9192C57FF8DFF8CFF77F899FEC5F8C8" box="[151,331,1806,1830]" italics="true" pageId="33" pageNumber="34">Chilocorus braeti</emphasis>
<bibRefCitation id="EFFC8DB4FF8DFF8CFEB1F899FD9FF8C8" author="Weise, J." box="[337,529,1806,1830]" pageId="33" pageNumber="34" pagination="151 - 157" refId="ref41480" refString="Weise, J. (1895 b) Insectes du Bengale. Coccinellidae. Annales de la Societe Entomologique du Belgique, 39, 151 - 157." type="journal article" year="1895">Weise, 1895b: 154</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD2F045FF8DFF8CFF77F8BAFBF2F8AB" blockId="33.[151,1148,1806,1861]" box="[151,1148,1837,1861]" pageId="33" pageNumber="34">
<treatmentCitationGroup id="AB7DD76BFF8DFF8CFF77F8BAFBF2F8AB" box="[151,1148,1837,1861]" pageId="33" pageNumber="34">
<taxonomicName id="4C6D8BC6FF8DFF8CFF77F8BAFEC5F8AB" box="[151,331,1837,1861]" class="Insecta" family="Coccinellidae" genus="Chilocorus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="braeti">
<emphasis id="B9192C57FF8DFF8CFF77F8BAFEC5F8AB" box="[151,331,1837,1861]" italics="true" pageId="33" pageNumber="34">Chilocorus braeti</emphasis>
</taxonomicName>
:
<treatmentCitation id="0ACCD654FF8DFF8CFEB8F8BAFDC9F8AB" author="Korschefsky, R." box="[344,583,1837,1861]" page="240" pageId="33" pageNumber="34" year="1932">
<bibRefCitation id="EFFC8DB4FF8DFF8CFEB8F8BAFDC9F8AB" author="Korschefsky, R." box="[344,583,1837,1861]" pageId="33" pageNumber="34" refId="ref36793" refString="Korschefsky, R. (1932) Coleopterorum Catalogus. Pars 120. Coccinellidae II. W. Junk, Berlin, 435 pp." type="book" year="1932">Korschefsky 1932: 240</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ACCD654FF8DFF8CFDB2F8BAFD7DF8AB" author="Kapur, A. P." box="[594,755,1837,1861]" page="24" pageId="33" pageNumber="34" year="1963">
<bibRefCitation id="EFFC8DB4FF8DFF8CFDB2F8BAFD7DF8AB" author="Kapur, A. P." box="[594,755,1837,1861]" pageId="33" pageNumber="34" pagination="1 - 48" refId="ref36509" refString="Kapur, A. P. (1963) The Coccinellidae of the third Mount Everest expedition, 1924 (Coleoptera). Bulletin of the British Museum (Natural History), Entomology, 14, 1 - 48. https: // doi. org / 10.5962 / bhl. part. 8780" type="journal article" year="1963">Kapur 1963: 24</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ACCD654FF8DFF8CFD1EF8BAFC35F8AB" author="Poorani, J." box="[766,955,1837,1861]" page="311" pageId="33" pageNumber="34" year="2002">
<bibRefCitation id="EFFC8DB4FF8DFF8CFD1EF8BAFC35F8AB" author="Poorani, J." box="[766,955,1837,1861]" pageId="33" pageNumber="34" pagination="307 - 383" refId="ref38985" refString="Poorani, J. (2002) An annotated checklist of the Coccinellidae (Coleoptera) (excluding Epilachninae) of the Indian Subregion. Oriental Insects, 36, 307 - 383. https: // doi. org / 10.1080 / 00305316.2002.10417335" type="journal article" year="2002">Poorani 2002: 311</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ACCD654FF8DFF8CFC26F8BAFBF6F8AB" author="Li, W. J. &amp; Huo, L. Z. &amp; Wang, D. &amp; Ahrens, D. &amp; Wang, X. M." box="[966,1144,1837,1861]" page="26" pageId="33" pageNumber="34" year="2018">
<bibRefCitation id="EFFC8DB4FF8DFF8CFC26F8BAFBF6F8AB" author="Li, W. J. &amp; Huo, L. Z. &amp; Wang, D. &amp; Ahrens, D. &amp; Wang, X. M." box="[966,1144,1837,1861]" pageId="33" pageNumber="34" pagination="1 - 34" refId="ref37468" refString="Li, W. J., Huo, L. Z., Wang, D., Ahrens, D. &amp; Wang, X. M. (2018) Contribution to the genus Chilocorus Leach, 1815 (Coleoptera: Coccinellidae: Chilocorini), with descriptions of two new species from China. European Journal of Taxonomy, 469, 1 - 34. https: // doi. org / 10.5852 / ejt. 2018.469" type="journal article" year="2018">
Li
<emphasis id="B9192C57FF8DFF8CFC03F8B9FB81F8AB" box="[995,1039,1837,1861]" italics="true" pageId="33" pageNumber="34">et al</emphasis>
. 2018: 26
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C377A3CEFF8DFF8CFF77F8E4FE81F817" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="8BD2F045FF8DFF8CFF77F8E4FE81F817" blockId="33.[151,1437,1907,2041]" pageId="33" pageNumber="34">
<emphasis id="B9192C57FF8DFF8CFF77F8E4FE9CF863" bold="true" box="[151,274,1907,1933]" pageId="33" pageNumber="34">Diagnosis.</emphasis>
Length:
<quantity id="4C955DA0FF8DFF8CFE9BF8E4FDADF860" box="[379,547,1907,1934]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.5" metricValueMin="4.2" pageId="33" pageNumber="34" unit="mm" value="4.85" valueMax="5.5" valueMin="4.2">4.20–5.50 mm</quantity>
. It can be distinguished from the other
<taxonomicName id="4C6D8BC6FF8DFF8CFC0BF8E4FBBDF863" box="[1003,1075,1907,1933]" class="Aves" family="Phasianidae" genus="Indian" kingdom="Animalia" order="Galliformes" pageId="33" pageNumber="108" phylum="Chordata" rank="genus">Indian</taxonomicName>
congeners by its characteristic, cordate or heart shaped body outline (
<figureCitation id="1356ECC0FF8DFF8CFDA1F800FD32F85F" box="[577,700,1943,1970]" captionStart="FIGURE 25" captionStartId="36.[151,250,1456,1480]" captionTargetBox="[154,1400,183,1431]" captionTargetId="figure-20@36.[151,1436,181,1432]" captionTargetPageId="36" captionText="FIGURE 25. Chilocorus braeti Weise: a, b. adult, dorsal view (BMNH); b. adult, dorsal view (ZSIK); c, d. male genitalia: c. tegmen, ventral view; d. penis." figureDoi="http://doi.org/10.5281/zenodo.10208973" httpUri="https://zenodo.org/record/10208973/files/figure.png" pageId="33" pageNumber="34">Fig. 25a, b</figureCitation>
) that is distinctly narrowed in the apical one-third, uniform black coloration and elytra with finely punctate disc and coarsely punctate lateral margins.
<collectingRegion id="49A93EA7FF8DFF8CFBADF82CFB09F83B" box="[1101,1159,1979,2005]" country="Maldives" name="Maale" pageId="33" pageNumber="34">Male</collectingRegion>
genitalia (
<figureCitation id="1356ECC0FF8DFF8CFB1CF82CFAF8F83B" box="[1276,1398,1979,2006]" captionStart="FIGURE 25" captionStartId="36.[151,250,1456,1480]" captionTargetBox="[154,1400,183,1431]" captionTargetId="figure-20@36.[151,1436,181,1432]" captionTargetPageId="36" captionText="FIGURE 25. Chilocorus braeti Weise: a, b. adult, dorsal view (BMNH); b. adult, dorsal view (ZSIK); c, d. male genitalia: c. tegmen, ventral view; d. penis." figureDoi="http://doi.org/10.5281/zenodo.10208973" httpUri="https://zenodo.org/record/10208973/files/figure.png" pageId="33" pageNumber="34">Fig. 25c, d</figureCitation>
) as illustrated.
</paragraph>
</subSubSection>
<caption id="DF12A0CDFF8EFF8FFF77F801FB6EF83F" ID-DOI="http://doi.org/10.5281/zenodo.10208969" ID-Zenodo-Dep="10208969" httpUri="https://zenodo.org/record/10208969/files/figure.png" pageId="34" pageNumber="35" startId="34.[151,250,1942,1966]" targetBox="[186,1401,181,1902]" targetPageId="34" targetType="figure">
<paragraph id="8BD2F045FF8EFF8FFF77F801FB6EF83F" blockId="34.[151,1437,1942,2002]" pageId="34" pageNumber="35">
<emphasis id="B9192C57FF8EFF8FFF77F801FEACF840" bold="true" box="[151,290,1942,1966]" pageId="34" pageNumber="35">FIGURE 23.</emphasis>
<taxonomicName id="4C6D8BC6FF8EFF8FFECAF801FD41F840" authority="(Li &amp; Wang)" authorityName="POORANI" authorityYear="2023" baseAuthorityName="Li &amp; Wang" baseAuthorityYear="2023" box="[298,719,1942,1966]" class="Insecta" family="Coccinellidae" genus="Chilocorus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="albomarginalis">
<emphasis id="B9192C57FF8EFF8FFECAF801FDB3F840" box="[298,573,1942,1966]" italics="true" pageId="34" pageNumber="35">Chilocorus albomarginalis</emphasis>
(Li &amp; Wang)
</taxonomicName>
: a. adult, dorsal view; b. adult, lateral view; c. adult, frontal view; d. abdomen, male; e–h. male genitalia: e. tegmen, lateral view; f. tegmen, ventral view; g. penis; h. penis apex.
</paragraph>
</caption>
<caption id="DF12A0CDFF8FFF8EFF77F800FACFF83D" ID-DOI="http://doi.org/10.5281/zenodo.10208971" ID-Zenodo-Dep="10208971" httpUri="https://zenodo.org/record/10208971/files/figure.png" pageId="35" pageNumber="36" startId="35.[151,250,1943,1967]" targetBox="[237,1350,191,1917]" targetPageId="35" targetType="figure">
<paragraph id="8BD2F045FF8FFF8EFF77F800FACFF83D" blockId="35.[151,1436,1943,2004]" pageId="35" pageNumber="36">
<emphasis id="B9192C57FF8FFF8EFF77F800FE91F841" bold="true" box="[151,287,1943,1967]" pageId="35" pageNumber="36">FIGURE 24.</emphasis>
<taxonomicName id="4C6D8BC6FF8FFF8EFEC4F800FD03F841" authority="(Linnaeus)" authorityName="Linnaeus" authorityYear="1758" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[292,653,1943,1967]" class="Insecta" family="Coccinellidae" genus="Chilocorus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="bipustulatus">
<emphasis id="B9192C57FF8FFF8EFEC4F800FD98F841" box="[292,534,1943,1967]" italics="true" pageId="35" pageNumber="36">Chilocorus bipustulatus</emphasis>
(Linnaeus)
</taxonomicName>
: a. adult, dorsal view; b. adult, frontal view; c. adult, lateral view; d. abdomen; e–h. male genitalia: e. tegmen, ventral view; f. tegmen, lateral view; g. penis; h. penis apex; i. spermatheca; j. coxites.
</paragraph>
</caption>
<caption id="DF12A0CDFF88FF89FF77FA27FE5EFA02" ID-DOI="http://doi.org/10.5281/zenodo.10208973" ID-Zenodo-Dep="10208973" httpUri="https://zenodo.org/record/10208973/files/figure.png" pageId="36" pageNumber="37" startId="36.[151,250,1456,1480]" targetBox="[154,1400,183,1431]" targetPageId="36" targetType="figure">
<paragraph id="8BD2F045FF88FF89FF77FA27FE5EFA02" blockId="36.[151,1436,1456,1516]" pageId="36" pageNumber="37">
<emphasis id="B9192C57FF88FF89FF77FA27FEAFFA26" bold="true" box="[151,289,1456,1480]" pageId="36" pageNumber="37">FIGURE 25.</emphasis>
<taxonomicName id="4C6D8BC6FF88FF89FEC8FA27FDAAFA26" authority="Weise" authorityName="Weise" box="[296,548,1456,1480]" class="Insecta" family="Coccinellidae" genus="Chilocorus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="braeti">
<emphasis id="B9192C57FF88FF89FEC8FA27FE53FA26" box="[296,477,1456,1480]" italics="true" pageId="36" pageNumber="37">Chilocorus braeti</emphasis>
Weise
</taxonomicName>
: a, b. adult, dorsal view (BMNH); b. adult, dorsal view (ZSIK); c, d. male genitalia: c. tegmen, ventral view; d. penis.
</paragraph>
</caption>
<subSubSection id="C377A3CEFF88FF89FF27F98DFE72F992" pageId="36" pageNumber="37" type="distribution">
<paragraph id="8BD2F045FF88FF89FF27F98DFACDF9DB" blockId="36.[151,1436,1562,1805]" box="[199,1347,1562,1589]" pageId="36" pageNumber="37">
<emphasis id="B9192C57FF88FF89FF27F98DFEEFF9DA" bold="true" box="[199,353,1562,1588]" pageId="36" pageNumber="37">Distribution.</emphasis>
<collectingCountry id="F37AB0D5FF88FF89FE87F98DFE2FF9DA" box="[359,417,1562,1588]" name="India" pageId="36" pageNumber="37">India</collectingCountry>
(North-eastern region:
<collectingRegion id="49A93EA7FF88FF89FD48F98DFD70F9DA" box="[680,766,1562,1588]" country="India" name="Tripura" pageId="36" pageNumber="37">Tripura</collectingRegion>
,
<collectingRegion id="49A93EA7FF88FF89FCE9F98DFC06F9DA" box="[777,904,1562,1588]" country="India" name="Meghalaya" pageId="36" pageNumber="37">Meghalaya</collectingRegion>
,
<collectingRegion id="49A93EA7FF88FF89FC73F98CFBADF9DA" box="[915,1059,1562,1589]" country="India" name="West Bengal" pageId="36" pageNumber="37">West Bengal</collectingRegion>
,
<collectingRegion id="49A93EA7FF88FF89FBCFF98DFB0BF9DA" box="[1071,1157,1562,1588]" country="India" name="Sikkim" pageId="36" pageNumber="37">Sikkim</collectingRegion>
);
<collectingCountry id="F37AB0D5FF88FF89FB77F98DFB77F9DA" box="[1175,1273,1562,1588]" name="Vietnam" pageId="36" pageNumber="37">Vietnam</collectingCountry>
;
<collectingCountry id="F37AB0D5FF88FF89FAE5F98CFAB1F9DB" box="[1285,1343,1563,1589]" name="Laos" pageId="36" pageNumber="37">Laos</collectingCountry>
.
</paragraph>
<paragraph id="8BD2F045FF88FF89FF27F9A9FE72F992" blockId="36.[151,1436,1562,1805]" pageId="36" pageNumber="37">
<emphasis id="B9192C57FF88FF89FF27F9A9FE63F9B6" bold="true" box="[199,493,1598,1624]" pageId="36" pageNumber="37">Prey / associated habitat.</emphasis>
Found on
<taxonomicName id="4C6D8BC6FF88FF89FD80F9A8FCC8F9B6" box="[608,838,1599,1624]" class="Magnoliopsida" family="Primulaceae" genus="Myrsine" kingdom="Plantae" order="Ericales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="semiserrata">
<emphasis id="B9192C57FF88FF89FD80F9A8FCC8F9B6" box="[608,838,1599,1624]" italics="true" pageId="36" pageNumber="37">Myrsine semiserrata</emphasis>
</taxonomicName>
(label data). Collected at high altitudes (
<quantity id="4C955DA0FF88FF89FAE2F9A9FA1BF9B7" box="[1282,1429,1598,1625]" metricMagnitude="3" metricUnit="m" metricValue="1.8288" metricValueMax="2.1336" metricValueMin="1.524" pageId="36" pageNumber="37" unit="ft" value="6000.0" valueMax="7000.0" valueMin="5000.0">5000–7000ft</quantity>
) of the Himalayas (
<bibRefCitation id="EFFC8DB4FF88FF89FE85F9F5FE7EF992" author="Kapur, A. P." box="[357,496,1634,1660]" pageId="36" pageNumber="37" pagination="1 - 48" refId="ref36509" refString="Kapur, A. P. (1963) The Coccinellidae of the third Mount Everest expedition, 1924 (Coleoptera). Bulletin of the British Museum (Natural History), Entomology, 14, 1 - 48. https: // doi. org / 10.5962 / bhl. part. 8780" type="journal article" year="1963">Kapur 1963</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C377A3CEFF88FF89FF27F911FD67F94E" box="[199,745,1670,1697]" pageId="36" pageNumber="37" type="biology_ecology">
<paragraph id="8BD2F045FF88FF89FF27F911FD67F94E" blockId="36.[151,1436,1562,1805]" box="[199,745,1670,1697]" pageId="36" pageNumber="37">
<emphasis id="B9192C57FF88FF89FF27F911FE4FF94E" bold="true" box="[199,449,1670,1696]" pageId="36" pageNumber="37">Seasonal occurrence.</emphasis>
March, June (Label data).
</paragraph>
</subSubSection>
<subSubSection id="C377A3CEFF88FF89FF27F93DFF7DF8E3" pageId="36" pageNumber="37" type="discussion">
<paragraph id="8BD2F045FF88FF89FF27F93DFF7DF8E3" blockId="36.[151,1436,1562,1805]" pageId="36" pageNumber="37">
<emphasis id="B9192C57FF88FF89FF27F93DFE9FF92A" bold="true" box="[199,273,1706,1732]" pageId="36" pageNumber="37">Notes.</emphasis>
It appears to be a rare species apparently endemic to the eastern Himalayan region and is not commonly represented in collections.
<bibRefCitation id="EFFC8DB4FF88FF89FE20F958FDD9F906" author="Kapur, A. P." box="[448,599,1742,1769]" pageId="36" pageNumber="37" pagination="1 - 48" refId="ref36509" refString="Kapur, A. P. (1963) The Coccinellidae of the third Mount Everest expedition, 1924 (Coleoptera). Bulletin of the British Museum (Natural History), Entomology, 14, 1 - 48. https: // doi. org / 10.5962 / bhl. part. 8780" type="journal article" year="1963">Kapur (1963)</bibRefCitation>
gave a brief account of
<taxonomicName id="4C6D8BC6FF88FF89FCBFF958FC4CF906" box="[863,962,1742,1769]" class="Insecta" family="Coccinellidae" genus="Chilocorus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="braeti">
<emphasis id="B9192C57FF88FF89FCBFF958FC4CF906" box="[863,962,1742,1769]" italics="true" pageId="36" pageNumber="37">C. braeti</emphasis>
</taxonomicName>
in his work on the
<taxonomicName id="4C6D8BC6FF88FF89FB74F959FAA0F906" authorityName="Latreille" authorityYear="1807" box="[1172,1326,1742,1768]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="family">Coccinellidae</taxonomicName>
of Mount Everest.
</paragraph>
</subSubSection>
</treatment>
</document>
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