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<document id="62BBD80420A00BFA58C5FC843F1D48B8" ID-CLB-Dataset="9025" ID-DOI="10.11646/zootaxa.4861.4.6" ID-GBIF-Dataset="59dc47c8-01b5-4f14-baa6-95b57158ca96" ID-ISSN="1175-5326" ID-Zenodo-Dep="4416789" ID-ZooBank="1A9329DD-74BA-4B44-9008-383D4C88C9D1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609801562384" checkinUser="plazi" docAuthor="Ma, Zijun, Huang, Sunbin &amp; Tian, Mingyi" docDate="2020" docId="1D57B212FFA8FFFC7EB9FE1A1080FC1B" docLanguage="en" docName="zootaxa.4861.4.6.pdf" docOrigin="Zootaxa 4861 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sinaphaenops Ueno &amp; Wang 1991" docType="key" docVersion="8" lastPageNumber="587" masterDocId="E16ECA6AFFAEFFFA7E2EFFAD151DFF41" masterDocTitle="Two new species of cavernicolous trechines from central Guizhou and northwestern Guangxi, China (Coleoptera: Carabidae: Trechinae)" masterLastPageNumber="593" masterPageNumber="581" pageNumber="587" updateTime="1698852245223" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8A5889A2A446B68C0B42F62244255F6B">Two new species of cavernicolous trechines from central Guizhou and northwestern Guangxi, China (Coleoptera: Carabidae: Trechinae)</mods:title>
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<mods:namePart id="E1E1FCBCBE93FFE49A12706185D85D3D">Ma, Zijun</mods:namePart>
<mods:nameIdentifier id="D76A5A2298FC9390C9D4B653E225AB90" type="ORCID">0000-0001-8884-3376</mods:nameIdentifier>
<mods:affiliation id="7A8898C385955828EDBE2F4FA4657130">Department of Entomology, College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou, 510642, CHINA &amp; 3275124967 @ qq. com; https: // orcid. org / 0000 - 0001 - 8884 - 3376</mods:affiliation>
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<mods:namePart id="B8725B09ED6B8EF817A7DF5BEE15763A">Huang, Sunbin</mods:namePart>
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<mods:affiliation id="A1409CBB7F20FF2CF5DD0AC507F8DEB6">Department of Entomology, College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou, 510642, CHINA &amp; Mécanismes adaptatifs et évolution (MECADEV), UMR 7179 CNRS-MNHN, Muséum National d’Histoire Naturelle, CP 50, 57 rue Cuvier, 75005 Paris, France. huangsunbin @ 163. com; https: // orcid. org / 0000 - 0001 - 8357 - 6651</mods:affiliation>
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<mods:namePart id="7D156BBCA3F6BE834F552B08C2E98D65">Tian, Mingyi</mods:namePart>
<mods:affiliation id="65899387E98C422155D9D4E1BE1D34A8">Department of Entomology, College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou, 510642, CHINA</mods:affiliation>
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<mods:title id="12D519D001FDCF472F4B515FB7D84B41">Zootaxa</mods:title>
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<mods:date id="253A53A3DBF602B004BABFF132F3DE5E">2020</mods:date>
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<mods:number id="5BCACE927B20B4B27C2FB522F8DBC820">2020-10-20</mods:number>
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<treatment id="1D57B212FFA8FFFC7EB9FE1A1080FC1B" ID-DOI="http://doi.org/10.5281/zenodo.4427106" ID-GBIF-Taxon="176440756" ID-Zenodo-Dep="4427106" LSID="urn:lsid:plazi:treatment:1D57B212FFA8FFFC7EB9FE1A1080FC1B" httpUri="http://treatment.plazi.org/id/1D57B212FFA8FFFC7EB9FE1A1080FC1B" lastPageNumber="587" pageId="6" pageNumber="587">
<subSubSection id="DDE4508FFFA8FFFC7EB9FE1A1128FE90" box="[151,1077,438,465]" pageId="6" pageNumber="587" type="nomenclature">
<paragraph id="95410304FFA8FFFC7EB9FE1A1128FE90" blockId="6.[151,1077,438,501]" box="[151,1077,438,465]" pageId="6" pageNumber="587">
<heading id="CE09B468FFA8FFFC7EB9FE1A1128FE90" bold="true" box="[151,1077,438,465]" fontSize="11" level="1" pageId="6" pageNumber="587" reason="1">
<emphasis id="A78ADF16FFA8FFFC7EB9FE1A1128FE90" bold="true" box="[151,1077,438,465]" pageId="6" pageNumber="587">
Key to species of the
<emphasis id="A78ADF16FFA8FFFC7FB3FE1B172BFE90" bold="true" box="[413,566,438,465]" italics="true" pageId="6" pageNumber="587">banshanicus</emphasis>
species group of the genus
<taxonomicName id="52FE7887FFA8FFFC7DA4FE1B1128FE90" ID-CoL="8KYX4" authorityName="Ueno &amp; Wang" authorityYear="1991" box="[906,1077,438,465]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="587" phylum="Arthropoda" rank="genus">
<emphasis id="A78ADF16FFA8FFFC7DA4FE1B1128FE90" bold="true" box="[906,1077,438,465]" italics="true" pageId="6" pageNumber="587">Sinaphaenops</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="DDE4508FFFA8FFFC7EB9FE761080FC1B" pageId="6" pageNumber="587" type="key">
<paragraph id="95410304FFA8FFFC7EB9FE76179EFEB4" blockId="6.[151,1077,438,501]" box="[151,643,475,501]" pageId="6" pageNumber="587">
(modified from
<bibRefCitation id="F16F7EF5FFA8FFFC7F64FE761766FEB4" author="Chen, M. Z. &amp; Huang, S. B. &amp; Tian, M. Y." box="[330,635,475,501]" pageId="6" pageNumber="587" pagination="1 - 8" refId="ref4718" refString="Chen, M. Z., Huang, S. B. &amp; Tian, M. Y. (2020) A new species of the aphaenopsian genus Sinaphaenops Ueno and Wang, with notes on male genitalia of S. orthogenys Ueno (Coleoptera: Carabidae: Trechini). The Coleopterists Bulletin, 74 (2), 1 - 8. https: // doi. org / 10.1649 / 0010 - 065 X- 74.2.343" type="journal article" year="2020">Chen, Huang &amp; Tian, 2020</bibRefCitation>
)
</paragraph>
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<paragraph id="95410304FFA8FFFC7EB9FD8D1081FD74" blockId="6.[151,1437,544,859]" box="[151,1436,544,566]" pageId="6" pageNumber="587">
<keyLead id="2E0FA031FFA8FFFC7EB9FD8D1081FD74" box="[151,1436,544,566]" pageId="6" pageNumber="587">
1 Right mandible tridentate, protibia with a longitudinal sulcus (subgenus
<taxonomicName id="52FE7887FFA8FFFC7DB6FD8D1102FD74" ID-CoL="8KYX4" authorityName="Ueno &amp; Wang" authorityYear="1991" box="[920,1055,544,565]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="587" phylum="Arthropoda" rank="genus">
<emphasis id="A78ADF16FFA8FFFC7DB6FD8D1102FD74" box="[920,1055,544,565]" italics="true" pageId="6" pageNumber="587">Sinaphaenops</emphasis>
</taxonomicName>
(
<emphasis id="A78ADF16FFA8FFFC7A03FD8D112BFD74" box="[1069,1078,544,565]" italics="true" pageId="6" pageNumber="587">s</emphasis>
.
<emphasis id="A78ADF16FFA8FFFC7A6CFD8D1146FD74" box="[1090,1115,544,565]" italics="true" pageId="6" pageNumber="587">str</emphasis>
.) Uéno &amp; Wang)........... 2
</keyLead>
</paragraph>
<paragraph id="95410304FFA8FFFC7EB9FD901081FD31" blockId="6.[151,1437,544,859]" pageId="6" pageNumber="587">
<keyLead id="2E0FA031FFA8FFFC7EB9FD901081FD31" pageId="6" pageNumber="587">
- Right mandible bidentate, protibia without a longitudinal sulcus........................................................................... subgenera
<taxonomicName id="52FE7887FFA8FFFC7CA7FDF7115FFD31" authority="Magrini, Vanni &amp; Zanon" authorityName="Magrini, Vanni &amp; Zanon" authorityYear="1997" box="[649,1090,602,624]" class="Insecta" family="Carabidae" genus="Thaumastaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="587" phylum="Arthropoda" rank="genus">
<emphasis id="A78ADF16FFA8FFFC7CA7FDF7164DFD2E" box="[649,848,602,623]" italics="true" pageId="6" pageNumber="587">Thaumastaphaenops</emphasis>
Magrini, Vanni &amp; Zanon
</taxonomicName>
and
<taxonomicName id="52FE7887FFA8FFFC7A5FFDF71081FD31" authority="Deuve &amp; Tian" authorityName="Deuve &amp; Tian" authorityYear="2014" box="[1137,1436,602,624]" class="Insecta" family="Carabidae" genus="Dongaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="587" phylum="Arthropoda" rank="genus">
<emphasis id="A78ADF16FFA8FFFC7A5FFDF71013FD2E" box="[1137,1294,602,623]" italics="true" pageId="6" pageNumber="587">Dongaphaenops</emphasis>
Deuve &amp; Tian
</taxonomicName>
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<paragraph id="95410304FFA8FFFC7EB9FDD51081FDCC" blockId="6.[151,1437,544,859]" box="[151,1436,632,654]" pageId="6" pageNumber="587">
<keyLead id="2E0FA031FFA8FFFC7EB9FDD51081FDCC" box="[151,1436,632,654]" pageId="6" pageNumber="587">2 Body stouter, neck constriction comparatively wide.......................................................... 3</keyLead>
</paragraph>
<paragraph id="95410304FFA8FFFC7EB9FD381081FDEB" blockId="6.[151,1437,544,859]" box="[151,1436,661,683]" pageId="6" pageNumber="587">
<keyLead id="2E0FA031FFA8FFFC7EB9FD381081FDEB" box="[151,1436,661,683]" pageId="6" pageNumber="587">
- Body slenderer, neck constriction distinctly narrow....................................
<emphasis id="A78ADF16FFA8FFFC7AA8FD38100EFDEB" box="[1158,1299,661,682]" italics="true" pageId="6" pageNumber="587">mirabilissimus</emphasis>
species group
</keyLead>
</paragraph>
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<paragraph id="95410304FFA8FFFC7EB9FD1F1081FD86" blockId="6.[151,1437,544,859]" box="[151,1436,690,712]" pageId="6" pageNumber="587">
<keyLead id="2E0FA031FFA8FFFC7EB9FD1F1081FD86" box="[151,1436,690,712]" pageId="6" pageNumber="587">
3 Pronotum with pair of lateral-marginal setae (
<taxonomicName id="52FE7887FFA8FFFC7CB7FD1F160CFD86" ID-CoL="4XJBM" box="[665,785,690,711]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="587" phylum="Arthropoda" rank="species" species="banshanicus">
<emphasis id="A78ADF16FFA8FFFC7CB7FD1F160CFD86" box="[665,785,690,711]" italics="true" pageId="6" pageNumber="587">banshanicus</emphasis>
</taxonomicName>
species group)......................................... 4
</keyLead>
</paragraph>
<paragraph id="95410304FFA8FFFC7EB9FD7D1081FDA4" blockId="6.[151,1437,544,859]" box="[151,1436,720,742]" pageId="6" pageNumber="587">
<keyLead id="2E0FA031FFA8FFFC7EB9FD7D1081FDA4" box="[151,1436,720,742]" pageId="6" pageNumber="587">
- Pronotum without lateral-marginal setae.................................................
<emphasis id="A78ADF16FFA8FFFC7A95FD7D100EFDA4" box="[1211,1299,720,741]" italics="true" pageId="6" pageNumber="587">trisetiger</emphasis>
species group
</keyLead>
</paragraph>
</keyStep>
<keyStep id="2E0A1BA1FFA8FFFC7EB9FD401081FC61" pageId="6" pageNumber="587">
<paragraph id="95410304FFA8FFFC7EB9FD401080FC43" blockId="6.[151,1437,544,859]" box="[151,1437,749,771]" pageId="6" pageNumber="587">
<keyLead id="2E0FA031FFA8FFFC7EB9FD401080FC43" box="[151,1437,749,771]" pageId="6" pageNumber="587">
4. Pronotum shorter (
<figureCitation id="0DC51F81FFA8FFFC7FB8FD4014F9FC42" box="[406,484,749,771]" captionStart="FIGURE 2" captionStartId="2.[151,250,1967,1991]" captionTargetBox="[152,1433,1396,1936]" captionTargetId="figure-39@2.[151,1436,1390,1944]" captionTargetPageId="2" captionText="FIGURE 2. Pronotum of Sinaphaenops species (A. S. (s. str.) chengguangyuani n. sp., holotype, male; B. S. (s. str.) lipoi, holotype, male; C. S. (s. str.) banshanicus, holotype, male)." figureDoi="http://doi.org/10.5281/zenodo.4416791" httpUri="https://zenodo.org/record/4416791/files/figure.png" pageId="6" pageNumber="587">Fig. 2A</figureCitation>
), male genitalia with a distinct sagittal aileron (
<figureCitation id="0DC51F81FFA8FFFC7D85FD4016F8FC42" box="[939,997,749,771]" captionStart="FIGURE 4" captionStartId="4.[151,250,1203,1227]" captionTargetBox="[321,1274,208,1155]" captionTargetId="figure-23@4.[295,1292,181,1178]" captionTargetPageId="4" captionText="FIGURE 4. Male genitalia of Sinaphaenops (s. str.) chengguangyuani n. sp., holotype (A. median lobe and parameres, lateral view; B. apex of median lobe, dorsal view; C. genital ring, ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4416795" httpUri="https://zenodo.org/record/4416795/files/figure.png" pageId="6" pageNumber="587">Fig. 4</figureCitation>
)........
<taxonomicName id="52FE7887FFA8FFFC7A7DFD40107CFC43" ID-CoL="8MGFB" authority="Ma &amp; Huang &amp; Tian, 2020" authorityName="Ma &amp; Huang &amp; Tian" authorityYear="2020" box="[1107,1377,749,771]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="587" phylum="Arthropoda" rank="species" species="chengguangyuani" status="sp. nov.">
<emphasis id="A78ADF16FFA8FFFC7A7DFD401142FC43" box="[1107,1119,749,770]" italics="true" pageId="6" pageNumber="587">S</emphasis>
. (
<emphasis id="A78ADF16FFA8FFFC7A5DFD431161FC42" box="[1139,1148,750,771]" italics="true" pageId="6" pageNumber="587">s</emphasis>
.
<emphasis id="A78ADF16FFA8FFFC7AA6FD4311BCFC42" box="[1160,1185,750,771]" italics="true" pageId="6" pageNumber="587">str</emphasis>
.)
<emphasis id="A78ADF16FFA8FFFC7A98FD40107CFC43" box="[1206,1377,749,770]" italics="true" pageId="6" pageNumber="587">chengguangyuani</emphasis>
</taxonomicName>
<emphasis id="A78ADF16FFA8FFFC7B48FD401080FC43" bold="true" box="[1382,1437,749,770]" pageId="6" pageNumber="587">
<taxonomicNameLabel id="BCB9626DFFA8FFFC7B48FD401080FC43" box="[1382,1437,749,770]" pageId="6" pageNumber="587" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="95410304FFA8FFFC7EB9FCA61081FC61" blockId="6.[151,1437,544,859]" box="[151,1436,778,800]" pageId="6" pageNumber="587">
<keyLead id="2E0FA031FFA8FFFC7EB9FCA61081FC61" box="[151,1436,778,800]" pageId="6" pageNumber="587">
- Pronotum longer (
<figureCitation id="0DC51F81FFA8FFFC7FBFFCA714E5FC61" box="[401,504,778,800]" captionStart="FIGURE 2" captionStartId="2.[151,250,1967,1991]" captionTargetBox="[152,1433,1396,1936]" captionTargetId="figure-39@2.[151,1436,1390,1944]" captionTargetPageId="2" captionText="FIGURE 2. Pronotum of Sinaphaenops species (A. S. (s. str.) chengguangyuani n. sp., holotype, male; B. S. (s. str.) lipoi, holotype, male; C. S. (s. str.) banshanicus, holotype, male)." figureDoi="http://doi.org/10.5281/zenodo.4416791" httpUri="https://zenodo.org/record/4416791/files/figure.png" pageId="6" pageNumber="587">Fig. 2B, C</figureCitation>
), male genitalia without sagittal aileron............................................ 5
</keyLead>
</paragraph>
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<keyStep id="2E0A1BA1FFA8FFFC7EB9FC8A1080FC1B" pageId="6" pageNumber="587">
<paragraph id="95410304FFA8FFFC7EB9FC8A1081FC7F" blockId="6.[151,1437,544,859]" box="[151,1436,807,830]" pageId="6" pageNumber="587">
<keyLead id="2E0FA031FFA8FFFC7EB9FC8A1081FC7F" box="[151,1436,807,830]" pageId="6" pageNumber="587">
5 Hind angles acute.........................................................
<taxonomicName id="52FE7887FFA8FFFC7A6EFC851081FC7F" ID-CoL="8MNLR" authority="Chen, Huang &amp; Tian" authorityName="Chen, Huang &amp; Tian" authorityYear="2020" box="[1088,1436,808,830]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="587" phylum="Arthropoda" rank="species" species="lipoi">
<emphasis id="A78ADF16FFA8FFFC7A6EFC851151FC7C" box="[1088,1100,808,829]" italics="true" pageId="6" pageNumber="587">S</emphasis>
. (
<emphasis id="A78ADF16FFA8FFFC7A4EFC851174FC7C" box="[1120,1129,808,829]" italics="true" pageId="6" pageNumber="587">s</emphasis>
.
<emphasis id="A78ADF16FFA8FFFC7A5BFC851193FC7C" box="[1141,1166,808,829]" italics="true" pageId="6" pageNumber="587">str</emphasis>
.)
<emphasis id="A78ADF16FFA8FFFC7A8CFC8511D3FC7C" box="[1186,1230,808,829]" italics="true" pageId="6" pageNumber="587">lipoi</emphasis>
Chen, Huang &amp; Tian
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="95410304FFA8FFFC7EB9FCE81080FC1B" blockId="6.[151,1437,544,859]" box="[151,1437,837,859]" pageId="6" pageNumber="587">
<keyLead id="2E0FA031FFA8FFFC7EB9FCE81080FC1B" box="[151,1437,837,859]" pageId="6" pageNumber="587">
- Hind angles obtuse...................................................
<taxonomicName id="52FE7887FFA8FFFC7A2AFCE81080FC1B" ID-CoL="4XJBM" authority="Tian, Chen &amp; Tang" authorityName="Tian, Chen &amp; Tang" box="[1028,1437,837,859]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="587" phylum="Arthropoda" rank="species" species="banshanicus">
<emphasis id="A78ADF16FFA8FFFC7A2AFCE8110DFC1B" box="[1028,1040,837,858]" italics="true" pageId="6" pageNumber="587">S</emphasis>
. (
<emphasis id="A78ADF16FFA8FFFC7A0AFCEB1130FC1A" box="[1060,1069,838,859]" italics="true" pageId="6" pageNumber="587">s</emphasis>
.
<emphasis id="A78ADF16FFA8FFFC7A14FCEB114EFC1A" box="[1082,1107,838,859]" italics="true" pageId="6" pageNumber="587">str</emphasis>
.)
<emphasis id="A78ADF16FFA8FFFC7A49FCE811C2FC1B" box="[1127,1247,837,858]" italics="true" pageId="6" pageNumber="587">banshanicus</emphasis>
Tian, Chen &amp; Tang
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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