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<document id="E298AF7D3AC9BB2CACE37CEF5455C44D" ID="10.11646/zootaxa.4169.3.7" ID-DOI="10.11646/zootaxa.4169.3.7" ID-GBIF-Dataset="faeafa7a-a105-4443-bbc3-2bea598d8cd5" ID-ISSN="1175-5326" ID-Zenodo-Dep="258027" ID-ZooBank="8A1FB454-AC73-4D54-A5CB-D12D6CD82AC6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1474440730591" checkinUser="plazi" docAuthor="Tian, Mingyi &amp; Deuve, Thierry" docDate="2016" docId="8706BE21FFB0FF931DC8FB92FDB4A77A" docLanguage="en" docName="zootaxa.4169.3.7.pdf" docOrigin="Zootaxa 4169 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hexachaetus vietnamensis Tian &amp; Deuve, 2016, n. sp." docType="treatment" docVersion="7" lastPageNumber="551" masterDocId="7B3FC659FFBAFF981D5FFF88FFF0A546" masterDocTitle="Re-definition and review of the Oriental genus Hexachaetus Chaudoir, 1871 (Coleoptera: Carabidae: Orthogoniini)" masterLastPageNumber="554" masterPageNumber="540" pageNumber="550" updateTime="1698386596725" updateUser="plazi">
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<mods:title id="60D32989F1FB3688E59658E33F8C6703">Re-definition and review of the Oriental genus Hexachaetus Chaudoir, 1871 (Coleoptera: Carabidae: Orthogoniini)</mods:title>
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<mods:namePart id="80E6DE96A3F7A638D52C93F0BE8B51E9">Tian, Mingyi</mods:namePart>
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<mods:namePart id="75630B4E255F9EA9CB26BC152BF51422">Deuve, Thierry</mods:namePart>
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<mods:title id="559D63398243B79A4FF330CF7A680F6E">Zootaxa</mods:title>
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<mods:date id="B589F8D12BD24135189CE8B4E04428D3">2016</mods:date>
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<treatment id="8706BE21FFB0FF931DC8FB92FDB4A77A" ID-DOI="http://doi.org/10.5281/zenodo.6075837" ID-GBIF-Taxon="124552523" ID-Zenodo-Dep="6075837" LSID="urn:lsid:plazi:treatment:8706BE21FFB0FF931DC8FB92FDB4A77A" httpUri="http://treatment.plazi.org/id/8706BE21FFB0FF931DC8FB92FDB4A77A" lastPageId="11" lastPageNumber="551" pageId="10" pageNumber="550">
<subSubSection id="47B55CBCFFB0FF921DC8FB92FE96A110" pageId="10" pageNumber="550" type="nomenclature">
<paragraph id="0F100F37FFB0FF921DC8FB92FDD2A173" blockId="10.[151,546,1050,1110]" box="[151,546,1050,1077]" pageId="10" pageNumber="550">
<heading id="5458B85BFFB0FF921DC8FB92FDD2A173" bold="true" box="[151,546,1050,1077]" fontSize="11" level="1" pageId="10" pageNumber="550" reason="1">
<emphasis id="3DDBD325FFB0FF921DC8FB92FDD2A173" bold="true" box="[151,546,1050,1077]" pageId="10" pageNumber="550">
<taxonomicName id="C8AF74B4FFB0FF921DC8FB92FE28A172" ID-CoL="3LG2N" box="[151,472,1050,1076]" class="Insecta" family="Carabidae" genus="Hexachaetus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="550" phylum="Arthropoda" rank="species" species="vietnamensis" status="sp. nov.">
<emphasis id="3DDBD325FFB0FF921DC8FB92FE28A172" bold="true" box="[151,472,1050,1076]" italics="true" pageId="10" pageNumber="550">Hexachaetus vietnamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="26E86E5EFFB0FF921CBFFB93FDD2A173" box="[480,546,1051,1077]" pageId="10" pageNumber="550" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="0F100F37FFB0FF921DC8FBB5FE96A110" blockId="10.[151,546,1050,1110]" box="[151,358,1085,1110]" pageId="10" pageNumber="550">
(
<figureCitation id="979413B2FFB0FF921DC1FBB5FEFBA110" box="[158,267,1085,1110]" captionStart="FIGURES 9 11" captionStartId="6.[151,264,1663,1686]" captionTargetBox="[449,1174,205,1627]" captionTargetId="figure@6.[398,1188,193,1643]" captionTargetPageId="6" captionText="FIGURES 9 11. Habitus of Hexachaetus species (9. H. vietnamensis n. sp.; 10. H. prodigus Tian &amp; Deuve, 2006; 11. H. mulan n. sp. Scale bar: 5.0 mm)" httpUri="https://zenodo.org/record/258030/files/figure.png" pageId="10" pageNumber="550">Figures 9</figureCitation>
,
<figureCitation id="979413B2FFB0FF921C48FBB6FEAEA110" box="[279,350,1086,1110]" captionStart="FIGURES 12 19" captionStartId="7.[151,264,1737,1760]" captionTargetBox="[277,1307,185,1721]" captionTargetId="figure@7.[259,1328,173,1732]" captionTargetPageId="7" captionText="FIGURES 12 19. Male genitalia of Hexachaetus species, right lateral view and dorsal view for apical lamella (12 13. H. maculatus Tian &amp; Deuve; 14 15. H. kirschenhoferi n. sp.; 16 17. H. brunki n. sp.; 18 19. H. vietnamensis n. sp.)" httpUri="https://zenodo.org/record/258031/files/figure.png" pageId="10" pageNumber="550">1819</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="47B55CBCFFB0FF921DC8FB0DFAC3A1A0" pageId="10" pageNumber="550" type="materials_examined">
<paragraph id="0F100F37FFB0FF921DC8FB0DFAC3A1A0" blockId="10.[151,1436,1157,2010]" pageId="10" pageNumber="550">
<materialsCitation id="BFC7056AFFB0FF921DC8FB0DFA68A1DB" ID-GBIF-Occurrence="1313880547" box="[151,1432,1157,1182]" collectionCode="MNHN" collectorName="Material. &amp; R. P. C. Corompt" country="Vietnam" location="Annam" pageId="10" pageNumber="550" specimenCount="1" stateProvince="Kontum" typeStatus="holotype">
<collectorName id="A25A6AE1FFB0FF921DC8FB0DFEF6A1D8" box="[151,262,1157,1182]" pageId="10" pageNumber="550">
<emphasis id="3DDBD325FFB0FF921DC8FB0DFEF6A1D8" bold="true" box="[151,262,1157,1182]" pageId="10" pageNumber="550">Material.</emphasis>
</collectorName>
<typeStatus id="D014B195FFB0FF921C4BFB0DFE8CA1D8" box="[276,380,1157,1182]" pageId="10" pageNumber="550" type="holotype">Holotype</typeStatus>
male, “
<location id="0A7059ECFFB0FF921CBEFB0EFDC8A1D8" LSID="urn:lsid:plazi:treatment:8706BE21FFB0FF931DC8FB92FDB4A77A:0A7059ECFFB0FF921CBEFB0EFDC8A1D8" box="[481,568,1158,1182]" country="Vietnam" name="Annam" pageId="10" pageNumber="550" stateProvince="Kontum">Annam</location>
,
<collectingRegion id="CD6BC1D5FFB0FF921F14FB0EFD5AA1DB" box="[587,682,1158,1181]" country="Vietnam" name="Kon Tum" pageId="10" pageNumber="550">Kontum</collectingRegion>
,
<collectorName id="A25A6AE1FFB0FF921FE3FB0EFC65A1D8" box="[700,917,1158,1182]" pageId="10" pageNumber="550">R. P. C. Corompt</collectorName>
, 1914”, from Central
<collectingCountry id="77B84FA7FFB0FF9219F9FB0DFAF8A1D8" box="[1190,1288,1157,1182]" name="Vietnam" pageId="10" pageNumber="550">Vietnam</collectingCountry>
, in
<collectionCode id="69BE97F2FFB0FF921860FB0EFA68A1DB" box="[1343,1432,1158,1181]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="550">MNHN</collectionCode>
</materialsCitation>
;
<materialsCitation id="BFC7056AFFB0FF921DC8FB22FB4AA184" ID-GBIF-Occurrence="1313880544" box="[151,1210,1193,1218]" collectionCode="SCAU" collectorName="Juni-Juli, H." country="Vietnam" location="Vietnam" pageId="10" pageNumber="550" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="D014B195FFB0FF921DC8FB22FEF7A184" box="[151,263,1194,1218]" pageId="10" pageNumber="550" type="paratype">paratypes</typeStatus>
:
<specimenCount id="19A9C4BEFFB0FF921C4EFB22FEAFA184" box="[273,351,1193,1218]" pageId="10" pageNumber="550" type="male">1 male</specimenCount>
, “Tonkin, Than-Moi,
<collectorName id="A25A6AE1FFB0FF921F0BFB21FD2DA187" box="[596,733,1193,1218]" pageId="10" pageNumber="550">Juni-Juli, H.</collectorName>
Fruhstorfer”, northern
<collectingCountry id="77B84FA7FFB0FF921EBCFB21FBB5A184" box="[995,1093,1193,1218]" name="Vietnam" pageId="10" pageNumber="550">Vietnam</collectingCountry>
, in
<collectionCode id="69BE97F2FFB0FF921933FB22FB4AA184" box="[1132,1210,1194,1218]" pageId="10" pageNumber="550">SCAU</collectionCode>
</materialsCitation>
;
<materialsCitation id="BFC7056AFFB0FF921999FB22FADFA1A0" ID-GBIF-Occurrence="1313880539" collectingDate="2008-09" collectionCode="IRSNB" collectorName="Coll. I. R. &amp; Sc. N. B. &amp; Buon Ma Thuot" country="Vietnam" location="Dak Lak Province" pageId="10" pageNumber="550" specimenCount="1" specimenCount-male="1" stateProvince="Dak Lak Province" typeStatus="paratype">
<specimenCount id="19A9C4BEFFB0FF921999FB22FAE6A184" box="[1222,1302,1193,1218]" pageId="10" pageNumber="550" type="male">1 male</specimenCount>
: “
<collectorName id="A25A6AE1FFB0FF921874FB21FF0BA1A0" pageId="10" pageNumber="550">Coll. I. R. Sc. N. B.</collectorName>
/
<collectingCountry id="77B84FA7FFB0FF921C53FB45FE9FA1A0" box="[268,367,1229,1254]" name="Vietnam" pageId="10" pageNumber="550">Vietnam</collectingCountry>
:
<collectingRegion id="CD6BC1D5FFB0FF921C26FB45FDB7A1A0" box="[377,583,1229,1254]" country="Vietnam" name="Dak Lak" pageId="10" pageNumber="550">
Dak
<collectingRegion id="CD6BC1D5FFB0FF921CF0FB45FDB7A1A0" box="[431,583,1229,1254]" inferredBySuffix="true" pageId="10" pageNumber="550">Lak Province</collectingRegion>
</collectingRegion>
, env.
<collectorName id="A25A6AE1FFB0FF921FD7FB46FCCDA1A0" box="[648,829,1229,1254]" pageId="10" pageNumber="550">Buon Ma Thuot</collectorName>
/
<collectingDate id="6B55D01FFFB0FF921E1AFB45FC55A1A0" box="[837,933,1229,1254]" pageId="10" pageNumber="550" value="2008-09">
<date id="7B1129F7FFB0FF921E1AFB45FC55A1A0" box="[837,933,1229,1254]" pageId="10" pageNumber="550" value="2008-09">IX. 2008</date>
</collectingDate>
/ Leg. Local collectors”, in
<collectionCode id="69BE97F2FFB0FF921986FB46FADFA1A0" box="[1241,1327,1230,1254]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="10" pageNumber="550">IRSNB</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="47B55CBCFFB0FF921D98FB79FC9AA0DC" pageId="10" pageNumber="550" type="diagnosis">
<paragraph id="0F100F37FFB0FF921D98FB79FC7DA030" blockId="10.[151,1436,1157,2010]" pageId="10" pageNumber="550">
<emphasis id="3DDBD325FFB0FF921D98FB79FEB1A04C" bold="true" box="[199,321,1265,1290]" pageId="10" pageNumber="550">Diagnosis.</emphasis>
Large sized, reddish brown; body moderately stout, smooth, polish; head and pronotum impunctate, elytra covered fine and sparse punctures; palps dilated, clypeus bisetose; pronotum transverse, widest at about middle, hind angle broadly round off; middle tibia not dilated medially in male, though stout; fore tarsi slightly stouter than middle ones, both much thicker than hind tarsi.
</paragraph>
<paragraph id="0F100F37FFB0FF921D98FA09FC9AA0DC" blockId="10.[151,1436,1157,2010]" box="[199,874,1409,1434]" pageId="10" pageNumber="550">
Length: 16.017.0 mm; width:
<quantity id="C857A2D2FFB0FF921F7AFA09FD8EA0DF" box="[549,638,1409,1434]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="10" pageNumber="550" unit="mm" value="6.5">6.5 mm</quantity>
. Habitus as in
<figureCitation id="979413B2FFB0FF921E7DFA09FC96A0DC" box="[802,870,1409,1434]" captionStart="FIGURES 9 11" captionStartId="6.[151,264,1663,1686]" captionTargetBox="[449,1174,205,1627]" captionTargetId="figure@6.[398,1188,193,1643]" captionTargetPageId="6" captionText="FIGURES 9 11. Habitus of Hexachaetus species (9. H. vietnamensis n. sp.; 10. H. prodigus Tian &amp; Deuve, 2006; 11. H. mulan n. sp. Scale bar: 5.0 mm)" httpUri="https://zenodo.org/record/258030/files/figure.png" pageId="10" pageNumber="550">Fig. 9</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="47B55CBCFFB0FF931D98FA2DFCDEA4CE" lastPageId="11" lastPageNumber="551" pageId="10" pageNumber="550" type="description">
<paragraph id="0F100F37FFB0FF921D98FA2DFB13A340" blockId="10.[151,1436,1157,2010]" pageId="10" pageNumber="550">
<emphasis id="3DDBD325FFB0FF921D98FA2DFEA9A0F8" bold="true" box="[199,345,1445,1470]" pageId="10" pageNumber="550">Description.</emphasis>
Reddish yellow to brown, tibiae and tarsi of legs, antennae, apex of mandibles, frons, clypeus and middle labrum dark brown, femora and vertex lighter; strongly shiny; surface glabrous, especially on pronotum; microsculptural engraved meshes irregularly striate on head and pronotum, isodiametric on elytra.
</paragraph>
<paragraph id="0F100F37FFB0FF921D98F999FD51A398" blockId="10.[151,1436,1157,2010]" pageId="10" pageNumber="550">Head convex, longer than wide, HL/HW=1.13; glabrous, frontal impressions well-marked, long and deep, extending to level between anterior and posterior eyes; clypeus bisetose, labrum straight at front, eyes large and prominent; palps stout; labial palpomere 2 with three setae on inner margin; maxillary palpomere 4 obliquely truncate at subapex; ligula very broad at apex, 12-setose, median two (normal length) much longer than others, three pairs shorter, and two pairs shortest, closed to median ones and at outer sides, respectively. Antennae short, antennomere 3 shorter than 4, but as long as 5.</paragraph>
<paragraph id="0F100F37FFB0FF921D98F962FD10A20C" blockId="10.[151,1436,1157,2010]" pageId="10" pageNumber="550">Pronotum moderately transverse, PL/PW=1.62; bordered at base and front, widest at about middle, lateral expanded margins broad and distinctly reflexed; both fore and hind angle round off, transversal impressions partly faint, basal foveae narrow, striate; median line clear.</paragraph>
<paragraph id="0F100F37FFB0FF921D98F8DDFD43A29F" blockId="10.[151,1436,1157,2010]" pageId="10" pageNumber="550">Elytra elongate-ovate, EL/EW=1.58; shoulders nearly quadrate, widest at middle, parallel-sided, apex obliquely truncated, inner apical angle pointed, outer one round off; striae deep, intervals slightly convex, intervals 2, 4 and 5 slightly wider than 3 and 6, or subequal in length; intervals 3, 5 and 7 with fine, indistinct punctures; interval 3 with basal and apical setiferous pores.</paragraph>
<paragraph id="0F100F37FFB1FF931D98FF10FD3FA45B" blockId="11.[151,1436,151,573]" pageId="11" pageNumber="551">Legs moderate, fore tibia slightly dilated at apex, with a moderate outer angle nearly rectangular, apical margin sinuate, outer margin distinctly serrate; middle tibia not dilate in male, hind tibia slender; apical tibial spurs of hind leg short and blunt, broadly sword-like, hind tarsomeres 34 subequal, tarsomere 4 bilobed at apex, asymmetrically; tarsal claws moderately pectinate.</paragraph>
<paragraph id="0F100F37FFB1FF931D98FEAFFCDEA4CE" blockId="11.[151,1436,151,573]" pageId="11" pageNumber="551">
Male genitalia (
<figureCitation id="979413B2FFB1FF931CDEFEAFFDE0A406" box="[385,528,295,320]" captionStart="FIGURES 12 19" captionStartId="7.[151,264,1737,1760]" captionTargetBox="[277,1307,185,1721]" captionTargetId="figure@7.[259,1328,173,1732]" captionTargetPageId="7" captionText="FIGURES 12 19. Male genitalia of Hexachaetus species, right lateral view and dorsal view for apical lamella (12 13. H. maculatus Tian &amp; Deuve; 14 15. H. kirschenhoferi n. sp.; 16 17. H. brunki n. sp.; 18 19. H. vietnamensis n. sp.)" httpUri="https://zenodo.org/record/258031/files/figure.png" pageId="11" pageNumber="551">Figs. 1819</figureCitation>
): The median lobe of aedeagus short and stout, distinctly enlarged medially, ventral margin bisinuate, strongly expanded near base, gradually narrowed towards apex; in dorsal view, the apical lamella very short and broad, apical margin nearly straight.
</paragraph>
</subSubSection>
<subSubSection id="47B55CBCFFB1FF931D98FE1CFDE3A4B2" pageId="11" pageNumber="551" type="discussion">
<paragraph id="0F100F37FFB1FF931D98FE1CFDE3A4B2" blockId="11.[151,1436,151,573]" pageId="11" pageNumber="551">
<emphasis id="3DDBD325FFB1FF931D98FE1CFECBA4EB" bold="true" box="[199,315,404,429]" pageId="11" pageNumber="551">Remarks.</emphasis>
This species is probably a representative of a unique lineage within the genus
<taxonomicName id="C8AF74B4FFB1FF931990FE1CFA90A4EA" box="[1231,1376,404,428]" class="Insecta" family="Carabidae" genus="Hexachaetus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="3DDBD325FFB1FF931990FE1CFA90A4EA" box="[1231,1376,404,428]" italics="true" pageId="11" pageNumber="551">Hexachaetus</emphasis>
</taxonomicName>
. The peculiar character states of
<taxonomicName id="C8AF74B4FFB1FF931C86FE31FD60A489" box="[473,656,440,464]" class="Insecta" family="Carabidae" genus="Hexachaetus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="551" phylum="Arthropoda" rank="species" species="vietnamensis" status="sp. nov.">
<emphasis id="3DDBD325FFB1FF931C86FE31FD60A489" box="[473,656,440,464]" italics="true" pageId="11" pageNumber="551">H. vietnamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="26E86E5EFFB1FF931FC4FE3FFD2DA496" box="[667,733,439,464]" pageId="11" pageNumber="551" rank="species">
<emphasis id="3DDBD325FFB1FF931FC4FE3FFD2DA496" bold="true" box="[667,733,439,464]" pageId="11" pageNumber="551">n. sp.</emphasis>
</taxonomicNameLabel>
include dilated palps, plurisetose ligula, and stout aedeagus which is broadly rounded at apex.
</paragraph>
</subSubSection>
<subSubSection id="47B55CBCFFB1FF931D98FE77FD64A75E" box="[199,660,511,536]" pageId="11" pageNumber="551" type="etymology">
<paragraph id="0F100F37FFB1FF931D98FE77FD64A75E" blockId="11.[151,1436,151,573]" box="[199,660,511,536]" pageId="11" pageNumber="551">
<emphasis id="3DDBD325FFB1FF931D98FE77FEBEA75E" bold="true" box="[199,334,511,536]" pageId="11" pageNumber="551">Etymology.</emphasis>
Referred to the
<typeStatus id="D014B195FFB1FF931F5CFD88FDC3A75E" box="[515,563,512,536]" pageId="11" pageNumber="551">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="47B55CBCFFB1FF931D98FDACFDB4A77A" box="[199,580,548,573]" pageId="11" pageNumber="551" type="distribution">
<paragraph id="0F100F37FFB1FF931D98FDACFDB4A77A" blockId="11.[151,1436,151,573]" box="[199,580,548,573]" pageId="11" pageNumber="551">
<emphasis id="3DDBD325FFB1FF931D98FDACFE90A77B" bold="true" box="[199,352,548,573]" pageId="11" pageNumber="551">Distribution.</emphasis>
<collectingCountry id="77B84FA7FFB1FF931C38FDACFE36A77B" box="[359,454,548,573]" name="Vietnam" pageId="11" pageNumber="551">Vietnam</collectingCountry>
(
<figureCitation id="979413B2FFB1FF931C8AFDACFDD9A77A" box="[469,553,548,573]" captionStart="FIGURE 20" captionStartId="12.[151,250,1789,1811]" captionTargetBox="[226,1360,198,1765]" captionTargetId="figure@12.[223,1363,195,1768]" captionTargetPageId="12" captionText="FIGURE 20. Distribution of Hexachaetus species (square: angulatus species group; round: lateralis species group; star: laevissimus species group; heart: vietnamensis species group; triangle: mulan species group; a. H. lateralis lateralis; b. H. lateralis maindroni; c. H. maculatus; d. H. kirschenhoferi n. sp.; e. H. brunki n. sp.; f. H. laevissimus; g. H. vietnamensis n. sp.; h. H. prodigus; i. H. mulan n. sp.)" httpUri="https://zenodo.org/record/258032/files/figure.png" pageId="11" pageNumber="551">
Fig.
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g).
</paragraph>
</subSubSection>
</treatment>
</document>