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<document id="7AD01C842A2F446FD2E4EC429AD936FC" ID-CLB-Dataset="32249" ID-DOI="10.11646/zootaxa.4313.1.1" ID-GBIF-Dataset="d8cc2cd8-8410-49aa-a75d-7a41d9f52b26" ID-ISSN="1175-5326" ID-Zenodo-Dep="883161" ID-ZooBank="A5Fdfba5-F992-44A8-84C2-1756C943C19B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1504182989959" checkinUser="plazi" docAuthor="Bush, Sarah E." docDate="2017" docId="832187E9FF2AFF61FF746664FC9EFDC3" docLanguage="en" docName="zootaxa.4313.1.1.pdf" docOrigin="Zootaxa 4313 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Harpactrox pontifrons Bush 2017, new species" docType="treatment" docVersion="16" lastPageNumber="163" masterDocId="7F18FF91FF8AFFC3FFE36200FFC9FFA3" masterDocTitle="Morphological revision of the hyperdiverse Brueelia - complex (Insecta: Phthiraptera: Ischnocera: Philopteridae) with new taxa, checklists and generic key" masterLastPageNumber="443" masterPageNumber="1" pageNumber="161" updateTime="1698467717597" updateUser="ExternalLinkService">
<mods:mods id="E3FA300C941340A88F0240B20F48A819" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3D02BB1AF0E010E3914B70889E8BD413">
<mods:title id="ED3BA689F33757A78EBEBCC2CEF943AD">Morphological revision of the hyperdiverse Brueelia - complex (Insecta: Phthiraptera: Ischnocera: Philopteridae) with new taxa, checklists and generic key</mods:title>
</mods:titleInfo>
<mods:name id="361D0C48E9BA2B88B6B3B8AC60EF162A" type="personal">
<mods:role id="D4FCF624A1BC31A393146CB9637EB32A">
<mods:roleTerm id="07E4466AD93E455314C2B7CFAD33A020">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="859C806ABE8FBB412923BBB499830F2A">Bush, Sarah E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="44FE1E22C1FE371F063C4C7CD55DFA3A">text</mods:typeOfResource>
<mods:relatedItem id="8411BFA14C6EDCABF2DDAE31F7DC42CC" type="host">
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<mods:title id="5033EA52A48E0933D674900DC3FA775E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="BDB3C2D05E0384E55810D8901C4C1390">
<mods:date id="AF0BD2850CD353DB341B657C4038FE1A">2017</mods:date>
<mods:detail id="AF912184260B9C01BCA6CF03A4119EBF" type="pubDate">
<mods:number id="682855BFA2BFE5A43AF9C3ED651564F6">2017-08-31</mods:number>
</mods:detail>
<mods:detail id="6633467542E0B54DBD58442A2C6DB6DE" type="volume">
<mods:number id="3EF6F2A10BF8F2BBB672F7EF964FA38F">4313</mods:number>
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<mods:classification id="625A60E3EB4FFEA7BF7C47566E692F86">journal article</mods:classification>
<mods:identifier id="24C94073EB557A37125BCAD91A08FE19" type="CLB-Dataset">32249</mods:identifier>
<mods:identifier id="2B4986962FCEA907C67343EE47FAE5C5" type="DOI">10.11646/zootaxa.4313.1.1</mods:identifier>
<mods:identifier id="3B649352A68A830C426BEF7B4BC6B580" type="GBIF-Dataset">d8cc2cd8-8410-49aa-a75d-7a41d9f52b26</mods:identifier>
<mods:identifier id="930A4B30C9B173A3A2C1BD740A55594C" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
<treatment id="832187E9FF2AFF61FF746664FC9EFDC3" ID-DOI="http://doi.org/10.5281/zenodo.5296967" ID-GBIF-Taxon="133100192" ID-Zenodo-Dep="5296967" LSID="urn:lsid:plazi:treatment:832187E9FF2AFF61FF746664FC9EFDC3" httpUri="http://treatment.plazi.org/id/832187E9FF2AFF61FF746664FC9EFDC3" lastPageId="162" lastPageNumber="163" pageId="160" pageNumber="161">
<subSubSection id="43926574FF2AFF63FF746664FE8AFB3C" pageId="160" pageNumber="161" type="nomenclature">
<paragraph id="0B3736FFFF2AFF63FF746664FCF7FBDD" blockId="160.[151,830,1124,1183]" box="[151,830,1124,1150]" pageId="160" pageNumber="161">
<heading id="507F8193FF2AFF63FF746664FCF7FBDD" bold="true" box="[151,830,1124,1150]" fontSize="11" level="1" pageId="160" pageNumber="161" reason="1">
<emphasis id="39FCEAEDFF2AFF63FF746664FCF7FBDD" bold="true" box="[151,830,1124,1150]" pageId="160" pageNumber="161">
<taxonomicName id="CC884D7CFF2AFF63FF746664FD68FBDD" authority="Gustafsson &amp; Bush" authorityName="Bush" authorityYear="2017" box="[151,673,1124,1150]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="160" pageNumber="161" phylum="Arthropoda" rank="species" species="pontifrons" status="sp. nov.">
<emphasis id="39FCEAEDFF2AFF63FF746664FE6EFBDD" bold="true" box="[151,423,1124,1150]" italics="true" pageId="160" pageNumber="161">Harpactrox pontifrons</emphasis>
Gustafsson &amp; Bush
</taxonomicName>
,
<taxonomicNameLabel id="22CF5796FF2AFF63FD4C6664FCF7FBDD" box="[687,830,1124,1150]" pageId="160" pageNumber="161" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="0B3736FFFF2AFF63FF746686FE8AFB3C" blockId="160.[151,830,1124,1183]" box="[151,323,1158,1183]" pageId="160" pageNumber="161">
(
<figureCitation id="93B32A7AFF2AFF63FF7C6686FEF2FB3C" box="[159,315,1158,1183]" captionStart="FIGURES 260 – 262" captionStartId="161.[151,264,1970,1992]" captionTargetBox="[194,1356,212,1926]" captionTargetId="figure@161.[163,1404,193,1927]" captionTargetPageId="161" captionText="FIGURES 260 – 262. Harpactrox pontifrons n. gen. &amp; n. sp. ex Harpactes ardens ardens: 260, female head, dorsal and ventral views. 261, female habitus, dorsal and ventral views. 262, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883298/files/figure.png" pageId="160" pageNumber="161">Figs 260–262</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="43926574FF2AFF63FF7466CEFD62FAA0" pageId="160" pageNumber="161" type="materials_examined">
<paragraph id="0B3736FFFF2AFF63FF7466CEFD62FAA0" blockId="160.[151,925,1230,1283]" pageId="160" pageNumber="161">
<materialsCitation id="BBE03CA2FF2AFF63FF7466CEFD62FAA0" ID-GBIF-Occurrence="1587279399" country="Philippines" location="Tucay" municipality="Mindanao" pageId="160" pageNumber="161" specimenCount="2" typeStatus="holotype">
<emphasis id="39FCEAEDFF2AFF63FF7466CEFECCFB47" bold="true" box="[151,261,1230,1252]" pageId="160" pageNumber="161">
<typeStatus id="D433885DFF2AFF63FF7466CEFF05FB47" box="[151,204,1230,1252]" pageId="160" pageNumber="161">Type</typeStatus>
host.
</emphasis>
<taxonomicName id="CC884D7CFF2AFF63FEEF66D0FD00FB47" authority="ardens (Temminck, 1826)" authorityName="ardens (Temminck" authorityYear="1826" box="[268,713,1230,1253]" class="Aves" family="Trogonidae" genus="Harpactes" kingdom="Animalia" order="Trogoniformes" pageId="160" pageNumber="161" phylum="Chordata" rank="subSpecies" species="ardens" subSpecies="ardens">
<emphasis id="39FCEAEDFF2AFF63FEEF66D0FDC3FB47" box="[268,522,1230,1253]" italics="true" pageId="160" pageNumber="161">Harpactes ardens ardens</emphasis>
(Temminck, 1826)
</taxonomicName>
—Philippine trogon.
<emphasis id="39FCEAEDFF2AFF63FF7466EDFEECFAA0" bold="true" box="[151,293,1261,1283]" pageId="160" pageNumber="161">
<typeStatus id="D433885DFF2AFF63FF7466EDFF05FAA0" box="[151,204,1261,1283]" pageId="160" pageNumber="161">Type</typeStatus>
locality.
</emphasis>
“
<location id="0E576024FF2AFF63FED566EDFEBFFAA0" LSID="urn:lsid:plazi:treatment:832187E9FF2AFF61FF746664FC9EFDC3:0E576024FF2AFF63FED566EDFEBFFAA0" box="[310,374,1261,1283]" country="Philippines" municipality="Mindanao" name="Tucay" pageId="160" pageNumber="161">Tucay</location>
E-el”,
<collectingMunicipality id="EB53AC85FF2AFF63FE5D66EDFDEEFAA0" box="[446,551,1261,1283]" pageId="160" pageNumber="161">Mindanao</collectingMunicipality>
,
<collectingCountry id="739F766FFF2AFF63FDD266EDFD6FFAA0" box="[561,678,1261,1283]" name="Philippines" pageId="160" pageNumber="161">Philippines</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="43926574FF2AFF63FF74672FFC55FA5E" pageId="160" pageNumber="161" type="diagnosis">
<paragraph id="0B3736FFFF2AFF63FF74672FFC55FA5E" blockId="160.[151,1437,1327,2037]" pageId="160" pageNumber="161">
<emphasis id="39FCEAEDFF2AFF63FF74672FFED8FAEB" bold="true" box="[151,273,1327,1352]" pageId="160" pageNumber="161">Diagnosis.</emphasis>
Head shape of
<taxonomicName id="CC884D7CFF2AFF63FE5F6731FD7EFAEB" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[444,695,1328,1352]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="160" pageNumber="161" phylum="Arthropoda" rank="species" species="pontifrons" status="sp. nov.">
<emphasis id="39FCEAEDFF2AFF63FE5F6731FD7EFAEB" box="[444,695,1328,1352]" italics="true" pageId="160" pageNumber="161">Harpactrox pontifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF2AFF63FD5E672FFD32FAEB" box="[701,763,1327,1352]" pageId="160" pageNumber="161" rank="species">
<emphasis id="39FCEAEDFF2AFF63FD5E672FFD32FAEB" bold="true" box="[701,763,1327,1352]" pageId="160" pageNumber="161">n. sp.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="93B32A7AFF2AFF63FCE9672FFCA0FAEB" box="[778,873,1327,1352]" captionStart="FIGURES 260 – 262" captionStartId="161.[151,264,1970,1992]" captionTargetBox="[194,1356,212,1926]" captionTargetId="figure@161.[163,1404,193,1927]" captionTargetPageId="161" captionText="FIGURES 260 – 262. Harpactrox pontifrons n. gen. &amp; n. sp. ex Harpactes ardens ardens: 260, female head, dorsal and ventral views. 261, female habitus, dorsal and ventral views. 262, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883298/files/figure.png" pageId="160" pageNumber="161">Fig. 260</figureCitation>
) intermediate between the other two species in the genus, but as in
<taxonomicName id="CC884D7CFF2AFF63FEAD6755FE28FACE" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[334,481,1364,1389]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="160" pageNumber="161" phylum="Arthropoda" rank="species" species="loeiensis" status="sp. nov.">
<emphasis id="39FCEAEDFF2AFF63FEAD6755FEB9FACF" box="[334,368,1365,1388]" italics="true" pageId="160" pageNumber="161">Ha</emphasis>
.
<emphasis id="39FCEAEDFF2AFF63FE9C6754FE28FACE" box="[383,481,1364,1389]" italics="true" pageId="160" pageNumber="161">loeiensis</emphasis>
</taxonomicName>
<emphasis id="39FCEAEDFF2AFF63FE0B6754FDE1FACE" bold="true" box="[488,552,1364,1389]" pageId="160" pageNumber="161">
<taxonomicNameLabel id="22CF5796FF2AFF63FE0B6754FDE1FACE" box="[488,552,1364,1389]" pageId="160" pageNumber="161" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="93B32A7AFF2AFF63FDD46754FD51FACF" box="[567,664,1364,1389]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="160" pageNumber="161">Fig. 248</figureCitation>
) the head is slightly wider than long. As in
<taxonomicName id="CC884D7CFF2AFF63FB656755FAD0FACE" authorityName="Bush" authorityYear="2017" box="[1158,1305,1364,1389]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="160" pageNumber="161" phylum="Arthropoda" rank="species" species="loeiensis">
<emphasis id="39FCEAEDFF2AFF63FB656755FB61FACF" box="[1158,1192,1365,1388]" italics="true" pageId="160" pageNumber="161">Ha</emphasis>
.
<emphasis id="39FCEAEDFF2AFF63FB546754FAD0FACE" box="[1207,1305,1364,1389]" italics="true" pageId="160" pageNumber="161">loeiensis</emphasis>
</taxonomicName>
, all
<taxonomicName id="CC884D7CFF2AFF63FAA86755FABBFACF" box="[1355,1394,1365,1388]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="160" pageNumber="161" phylum="Arthropoda" rank="species" species="mts">
<emphasis id="39FCEAEDFF2AFF63FAA86755FABBFACF" box="[1355,1394,1365,1388]" italics="true" pageId="160" pageNumber="161">mts</emphasis>
</taxonomicName>
are of roughly equal length, and the dorsal preantennal suture does not reach
<emphasis id="39FCEAEDFF2AFF63FC246777FC36FA33" box="[967,1023,1399,1424]" italics="true" pageId="160" pageNumber="161">dsms</emphasis>
. Dorsal sinuous ridge at the osculum found in
<taxonomicName id="CC884D7CFF2AFF63FEE3679DFE6CFA17" authorityName="Bush" authorityYear="2017" box="[256,421,1436,1460]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="160" pageNumber="161" phylum="Arthropoda" rank="species" species="pontifrons">
<emphasis id="39FCEAEDFF2AFF63FEE3679DFEEBFA17" box="[256,290,1437,1460]" italics="true" pageId="160" pageNumber="161">Ha</emphasis>
.
<emphasis id="39FCEAEDFF2AFF63FED0679CFE6CFA17" box="[307,421,1436,1460]" italics="true" pageId="160" pageNumber="161">pontifrons</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF2AFF63FE55679CFDD0FA16" box="[438,537,1436,1461]" captionStart="FIGURES 260 – 262" captionStartId="161.[151,264,1970,1992]" captionTargetBox="[194,1356,212,1926]" captionTargetId="figure@161.[163,1404,193,1927]" captionTargetPageId="161" captionText="FIGURES 260 – 262. Harpactrox pontifrons n. gen. &amp; n. sp. ex Harpactes ardens ardens: 260, female head, dorsal and ventral views. 261, female habitus, dorsal and ventral views. 262, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883298/files/figure.png" pageId="160" pageNumber="161">Fig. 260</figureCitation>
) is not present in
<taxonomicName id="CC884D7CFF2AFF63FD08679DFC49FA16" authorityName="Bush" authorityYear="2017" box="[747,896,1436,1461]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="160" pageNumber="161" phylum="Arthropoda" rank="species" species="loeiensis">
<emphasis id="39FCEAEDFF2AFF63FD08679DFCC4FA17" box="[747,781,1437,1460]" italics="true" pageId="160" pageNumber="161">Ha</emphasis>
.
<emphasis id="39FCEAEDFF2AFF63FCFD679CFC49FA16" box="[798,896,1436,1461]" italics="true" pageId="160" pageNumber="161">loeiensis</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF2AFF63FC72679CFC3FFA17" box="[913,1014,1436,1461]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="160" pageNumber="161">Fig. 248</figureCitation>
). Like in
<taxonomicName id="CC884D7CFF2AFF63FB84679DFADBFA16" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[1127,1298,1436,1461]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="160" pageNumber="161" phylum="Arthropoda" rank="species" species="geminodus" status="sp. nov.">
<emphasis id="39FCEAEDFF2AFF63FB84679DFB40FA17" box="[1127,1161,1437,1460]" italics="true" pageId="160" pageNumber="161">Ha</emphasis>
.
<emphasis id="39FCEAEDFF2AFF63FB7A679CFADBFA16" box="[1177,1298,1436,1461]" italics="true" pageId="160" pageNumber="161">geminodus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF2AFF63FAFF679CFA94FA16" box="[1308,1373,1436,1461]" pageId="160" pageNumber="161" rank="species">
<emphasis id="39FCEAEDFF2AFF63FAFF679CFA94FA16" bold="true" box="[1308,1373,1436,1461]" pageId="160" pageNumber="161">n. sp.</emphasis>
</taxonomicNameLabel>
(Fig. 255), the lateral margins of the clypeo-labral suture are clearly delimited anterior to the pulvinus in
<taxonomicName id="CC884D7CFF2AFF63FB1067C1FA5CFA7B" authorityName="Bush" authorityYear="2017" box="[1267,1429,1472,1496]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="160" pageNumber="161" phylum="Arthropoda" rank="species" species="pontifrons">
<emphasis id="39FCEAEDFF2AFF63FB1067C1FADCFA7B" box="[1267,1301,1473,1496]" italics="true" pageId="160" pageNumber="161">Ha</emphasis>
.
<emphasis id="39FCEAEDFF2AFF63FAC067C0FA5CFA7B" box="[1315,1429,1472,1496]" italics="true" pageId="160" pageNumber="161">pontifrons</emphasis>
</taxonomicName>
, and the ventral anterior plate is much wider than the marginal carina.
</paragraph>
</subSubSection>
<subSubSection id="43926574FF2AFF63FF246407FA48F82B" pageId="160" pageNumber="161" type="description">
<paragraph id="0B3736FFFF2AFF63FF246407FD22F976" blockId="160.[151,1437,1327,2037]" pageId="160" pageNumber="161">
<emphasis id="39FCEAEDFF2AFF63FF246407FE90F983" bold="true" box="[199,345,1543,1568]" pageId="160" pageNumber="161">Description.</emphasis>
Head shape, structure, and chaetotaxy as in genus description and
<figureCitation id="93B32A7AFF2AFF63FBA66407FB6CF983" box="[1093,1189,1543,1568]" captionStart="FIGURES 260 – 262" captionStartId="161.[151,264,1970,1992]" captionTargetBox="[194,1356,212,1926]" captionTargetId="figure@161.[163,1404,193,1927]" captionTargetPageId="161" captionText="FIGURES 260 – 262. Harpactrox pontifrons n. gen. &amp; n. sp. ex Harpactes ardens ardens: 260, female head, dorsal and ventral views. 261, female habitus, dorsal and ventral views. 262, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883298/files/figure.png" pageId="160" pageNumber="161">Fig. 260</figureCitation>
. Ventral anterior plate much wider than marginal carina. Ventral carinae clearly defined anterior to pulvinus. Dorsal preantennal suture reaches
<emphasis id="39FCEAEDFF2AFF63FF17644FFED2F9CB" box="[244,283,1615,1640]" italics="true" pageId="160" pageNumber="161">ads</emphasis>
but not
<emphasis id="39FCEAEDFF2AFF63FE9F644FFE7DF9CB" box="[380,436,1615,1640]" italics="true" pageId="160" pageNumber="161">dsms</emphasis>
. Clear dorsal thickening present anterior to suture. Preantennal nodi wide and rounded, distorting the lateral head margin just anterior to antennae. All
<taxonomicName id="CC884D7CFF2AFF63FC9E6475FC6DF92F" box="[893,932,1653,1676]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="160" pageNumber="161" phylum="Arthropoda" rank="species" species="mts">
<emphasis id="39FCEAEDFF2AFF63FC9E6475FC6DF92F" box="[893,932,1653,1676]" italics="true" pageId="160" pageNumber="161">mts</emphasis>
</taxonomicName>
of roughly equal length. Gular plate short, indistinct. Thoracic and abdominal segments as in genus description and
<figureCitation id="93B32A7AFF2AFF63FC2D6497FBE6F913" box="[974,1071,1687,1712]" captionStart="FIGURES 260 – 262" captionStartId="161.[151,264,1970,1992]" captionTargetBox="[194,1356,212,1926]" captionTargetId="figure@161.[163,1404,193,1927]" captionTargetPageId="161" captionText="FIGURES 260 – 262. Harpactrox pontifrons n. gen. &amp; n. sp. ex Harpactes ardens ardens: 260, female head, dorsal and ventral views. 261, female habitus, dorsal and ventral views. 262, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883298/files/figure.png" pageId="160" pageNumber="161">Fig. 261</figureCitation>
. Posterior margin of pterothorax strikingly more convergent than in other two species.
</paragraph>
<paragraph id="0B3736FFFF2AFF63FF2464DFFE4FF95B" blockId="160.[151,1437,1327,2037]" box="[199,390,1759,1784]" pageId="160" pageNumber="161">
<materialsCitation id="BBE03CA2FF2AFF63FF2464DFFE4FF95B" ID-GBIF-Occurrence="1587279417" box="[199,390,1759,1784]" location="Male" pageId="160" pageNumber="161" specimenCount="1" stateProvince="Male">
<collectingRegion id="C94CF81DFF2AFF63FF2464DFFECBF95B" box="[199,258,1759,1784]" country="Maldives" name="Maale" pageId="160" pageNumber="161">
<emphasis id="39FCEAEDFF2AFF63FF2464DFFECBF95B" bold="true" box="[199,258,1759,1784]" italics="true" pageId="160" pageNumber="161">Male</emphasis>
</collectingRegion>
. Unknown.
</materialsCitation>
</paragraph>
<paragraph id="0B3736FFFF2AFF63FF246504FA48F82B" blockId="160.[151,1437,1327,2037]" pageId="160" pageNumber="161">
<emphasis id="39FCEAEDFF2AFF63FF246504FED7F8BE" bold="true" box="[199,286,1796,1821]" italics="true" pageId="160" pageNumber="161">Female</emphasis>
. Abdominal chaetotaxy as in
<tableCitation id="460A0344FF2AFF63FD916504FD0FF8BF" box="[626,710,1796,1821]" captionStart="TABLE 2" captionStartId="30.[151,239,796,819]" captionText="TABLE 2. Summary of abdominal chaetotaxy of segments II – VIII for the Brueelia - complex, including both genera and subgenera. In genera where chaetotaxy differs between species group (marked with *), only that of the type species is given here, and more detailed accounts are given separately under the respective genus entries and in Tables 3 – 12. Trichoid setae of segment VIII are present in all genera, and are not listed. Abbreviations: ps = paratergal seta; aps = accessory post-spiracular seta; psps = principal post-spiracular seta; tps = tergal posterior seta; ss = sutural seta; sts = sternal seta." pageId="160" pageNumber="161">Table 2</tableCitation>
and
<figureCitation id="93B32A7AFF2AFF63FD1C6504FCA8F8BE" box="[767,865,1796,1821]" captionStart="FIGURES 260 – 262" captionStartId="161.[151,264,1970,1992]" captionTargetBox="[194,1356,212,1926]" captionTargetId="figure@161.[163,1404,193,1927]" captionTargetPageId="161" captionText="FIGURES 260 – 262. Harpactrox pontifrons n. gen. &amp; n. sp. ex Harpactes ardens ardens: 260, female head, dorsal and ventral views. 261, female habitus, dorsal and ventral views. 262, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883298/files/figure.png" pageId="160" pageNumber="161">Fig. 261</figureCitation>
. Subgenital plate sinuously trapezoidal (
<figureCitation id="93B32A7AFF2AFF63FACF6504FA59F8BE" box="[1324,1424,1796,1821]" captionStart="FIGURES 260 – 262" captionStartId="161.[151,264,1970,1992]" captionTargetBox="[194,1356,212,1926]" captionTargetId="figure@161.[163,1404,193,1927]" captionTargetPageId="161" captionText="FIGURES 260 – 262. Harpactrox pontifrons n. gen. &amp; n. sp. ex Harpactes ardens ardens: 260, female head, dorsal and ventral views. 261, female habitus, dorsal and ventral views. 262, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883298/files/figure.png" pageId="160" pageNumber="161">Fig. 262</figureCitation>
). Vulval margin (
<figureCitation id="93B32A7AFF2AFF63FEA56527FE6CF8E3" box="[326,421,1831,1856]" captionStart="FIGURES 260 – 262" captionStartId="161.[151,264,1970,1992]" captionTargetBox="[194,1356,212,1926]" captionTargetId="figure@161.[163,1404,193,1927]" captionTargetPageId="161" captionText="FIGURES 260 – 262. Harpactrox pontifrons n. gen. &amp; n. sp. ex Harpactes ardens ardens: 260, female head, dorsal and ventral views. 261, female habitus, dorsal and ventral views. 262, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883298/files/figure.png" pageId="160" pageNumber="161">Fig. 262</figureCitation>
) gently rounded, with 2 short, slender
<emphasis id="39FCEAEDFF2AFF63FCB76529FC49F8E3" box="[852,896,1833,1856]" italics="true" pageId="160" pageNumber="161">vms</emphasis>
on each side, and 3 short, thorn-like
<emphasis id="39FCEAEDFF2AFF63FAFE6529FAF6F8E3" box="[1309,1343,1833,1856]" italics="true" pageId="160" pageNumber="161">vss</emphasis>
on each side; 7–8 long, slender
<emphasis id="39FCEAEDFF2AFF63FE7A654DFE77F8C7" box="[409,446,1869,1892]" italics="true" pageId="160" pageNumber="161">vos</emphasis>
on each side; 2 distal
<emphasis id="39FCEAEDFF2AFF63FD57654DFD10F8C7" box="[692,729,1869,1892]" italics="true" pageId="160" pageNumber="161">vos</emphasis>
median to
<emphasis id="39FCEAEDFF2AFF63FCB6654DFCBEF8C7" box="[853,887,1869,1892]" italics="true" pageId="160" pageNumber="161">vss</emphasis>
. Measurements ex
<taxonomicName id="CC884D7CFF2AFF63FBB3654DFAA1F8C6" authority="ardens" authorityName="ardens (Temminck" authorityYear="1824" box="[1104,1384,1868,1893]" class="Aves" family="Trogonidae" genus="Harpactes" kingdom="Animalia" order="Trogoniformes" pageId="160" pageNumber="161" phylum="Chordata" rank="subSpecies" species="ardens" subSpecies="ardens">
<emphasis id="39FCEAEDFF2AFF63FBB3654DFAA1F8C6" box="[1104,1384,1868,1893]" italics="true" pageId="160" pageNumber="161">Harpactes ardens ardens</emphasis>
</taxonomicName>
(n = 3): TL = 1.79–2.01; HL = 0.38–0.42; HW = 0.44–0.49; PRW = 0.26–0.28; PTW = 0.44–0.48; AW = 0.56 –0.67.
</paragraph>
</subSubSection>
<subSubSection id="43926574FF2AFF63FF246594FD85F856" pageId="160" pageNumber="161" type="etymology">
<paragraph id="0B3736FFFF2AFF63FF246594FD85F856" blockId="160.[151,1437,1327,2037]" pageId="160" pageNumber="161">
<emphasis id="39FCEAEDFF2AFF63FF246594FE87F80E" bold="true" box="[199,334,1940,1965]" pageId="160" pageNumber="161">Etymology.</emphasis>
The species epithet is derived from Latin “
<emphasis id="39FCEAEDFF2AFF63FCDC6595FC4AF80F" box="[831,899,1941,1964]" italics="true" pageId="160" pageNumber="161">pontis</emphasis>
” for “bridge” and “
<emphasis id="39FCEAEDFF2AFF63FB8A6594FB68F80F" box="[1129,1185,1940,1964]" italics="true" pageId="160" pageNumber="161">frons</emphasis>
” for “forehead”. This refers to the thickening of the dorsal head plate across the clypeo-labral suture, which forms a “bridge” from one side of the head to the other (
<figureCitation id="93B32A7AFF2AFF63FE3E65DCFD89F856" box="[477,576,2012,2037]" captionStart="FIGURES 260 – 262" captionStartId="161.[151,264,1970,1992]" captionTargetBox="[194,1356,212,1926]" captionTargetId="figure@161.[163,1404,193,1927]" captionTargetPageId="161" captionText="FIGURES 260 – 262. Harpactrox pontifrons n. gen. &amp; n. sp. ex Harpactes ardens ardens: 260, female head, dorsal and ventral views. 261, female habitus, dorsal and ventral views. 262, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883298/files/figure.png" pageId="160" pageNumber="161">Fig. 260</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="5FF76677FF2BFF62FF7465B2FAE8F844" httpUri="https://zenodo.org/record/883298/files/figure.png" pageId="161" pageNumber="162" targetBox="[194,1356,212,1926]" targetPageId="161">
<paragraph id="0B3736FFFF2BFF62FF7465B2FAE8F844" blockId="161.[151,1435,1970,2023]" pageId="161" pageNumber="162">
<emphasis id="39FCEAEDFF2BFF62FF7465B2FEA5F86B" bold="true" box="[151,364,1970,1992]" pageId="161" pageNumber="162">FIGURES 260–262.</emphasis>
<taxonomicName id="CC884D7CFF2BFF62FE9065B3FD9CF86B" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[371,597,1970,1992]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="161" pageNumber="162" phylum="Arthropoda" rank="species" species="pontifrons" status="gen. nov.">
<emphasis id="39FCEAEDFF2BFF62FE9065B3FD9CF86B" box="[371,597,1970,1992]" italics="true" pageId="161" pageNumber="162">Harpactrox pontifrons</emphasis>
</taxonomicName>
<emphasis id="39FCEAEDFF2BFF62FDBF65B2FD34F86B" bold="true" box="[604,765,1970,1992]" pageId="161" pageNumber="162">
<taxonomicNameLabel id="22CF5796FF2BFF62FDBF65B2FD6AF86B" box="[604,675,1970,1992]" pageId="161" pageNumber="162" rank="genus">n. gen.</taxonomicNameLabel>
&amp; n. sp.
</emphasis>
ex
<taxonomicName id="CC884D7CFF2BFF62FCC265B3FBD6F86B" authority="ardens" authorityName="ardens (Temminck" authorityYear="1824" box="[801,1055,1970,1992]" class="Aves" family="Trogonidae" genus="Harpactes" kingdom="Animalia" order="Trogoniformes" pageId="161" pageNumber="162" phylum="Chordata" rank="subSpecies" species="ardens" subSpecies="ardens">
<emphasis id="39FCEAEDFF2BFF62FCC265B3FBD6F86B" box="[801,1055,1970,1992]" italics="true" pageId="161" pageNumber="162">Harpactes ardens ardens</emphasis>
</taxonomicName>
:
<emphasis id="39FCEAEDFF2BFF62FBCF65B2FB90F86B" bold="true" box="[1068,1113,1970,1992]" pageId="161" pageNumber="162">260,</emphasis>
female head, dorsal and ventral views.
<emphasis id="39FCEAEDFF2BFF62FF3D65D1FEC2F844" bold="true" box="[222,267,2001,2023]" pageId="161" pageNumber="162">261,</emphasis>
female habitus, dorsal and ventral views.
<emphasis id="39FCEAEDFF2BFF62FD5665D1FD2BF844" bold="true" box="[693,738,2001,2023]" pageId="161" pageNumber="162">262,</emphasis>
female subgenital plate and vulval margin, ventral view.
</paragraph>
</caption>
<subSubSection id="43926574FF28FF61FF246297FA50FF76" pageId="162" pageNumber="163" type="materials_examined">
<paragraph id="0B3736FFFF28FF61FF246297FA50FF76" blockId="162.[151,1436,151,608]" pageId="162" pageNumber="163">
<emphasis id="39FCEAEDFF28FF61FF246297FEBEFF13" bold="true" box="[199,375,151,176]" pageId="162" pageNumber="163">Type material.</emphasis>
Ex
<emphasis id="39FCEAEDFF28FF61FE4B6298FC85FF13" bold="true" box="[424,844,151,176]" pageId="162" pageNumber="163">
<taxonomicName id="CC884D7CFF28FF61FE4B6298FD04FF13" authority="ardens" authorityName="ardens (Temminck" authorityYear="1824" box="[424,717,151,176]" class="Aves" family="Trogonidae" genus="Harpactes" kingdom="Animalia" order="Trogoniformes" pageId="162" pageNumber="163" phylum="Chordata" rank="subSpecies" species="ardens" subSpecies="ardens">
<emphasis id="39FCEAEDFF28FF61FE4B6298FD04FF13" bold="true" box="[424,717,151,176]" italics="true" pageId="162" pageNumber="163">Harpactes ardens ardens</emphasis>
</taxonomicName>
: Holotype
</emphasis>
♀, “Tucay E-el”, Mindanao, Philippines, SUBBM- 1507 (BPBM).
<emphasis id="39FCEAEDFF28FF61FEA762BCFE0CFF76" bold="true" box="[324,453,188,213]" pageId="162" pageNumber="163">Paratypes:</emphasis>
2♀, same data as holotype (PIPeR) [one slide contains unidentified
<taxonomicName id="CC884D7CFF28FF61FB2062BCFA8CFF76" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[1219,1349,188,213]" class="Insecta" family="Philopteridae" genus="Philopterus" kingdom="Animalia" order="Phthiraptera" pageId="162" pageNumber="163" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="39FCEAEDFF28FF61FB2062BCFA8CFF76" box="[1219,1349,188,213]" italics="true" pageId="162" pageNumber="163">Philopterus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF28FF61FAAE62BDFA44FF76" box="[1357,1421,188,213]" pageId="162" pageNumber="163" sensu="lato">s. lat.</taxonomicNameLabel>
].
</paragraph>
</subSubSection>
<subSubSection id="43926574FF28FF61FF2462DFFC9EFDC3" pageId="162" pageNumber="163" type="discussion">
<paragraph id="0B3736FFFF28FF61FF2462DFFC9EFDC3" blockId="162.[151,1436,151,608]" pageId="162" pageNumber="163">
<emphasis id="39FCEAEDFF28FF61FF2462DFFEF2FF5B" bold="true" box="[199,315,223,248]" pageId="162" pageNumber="163">Remarks.</emphasis>
Despite not having any male, we describe
<taxonomicName id="CC884D7CFF28FF61FCF462E1FBD8FF5B" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[791,1041,224,248]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="162" pageNumber="163" phylum="Arthropoda" rank="species" species="pontifrons" status="sp. nov.">
<emphasis id="39FCEAEDFF28FF61FCF462E1FBD8FF5B" box="[791,1041,224,248]" italics="true" pageId="162" pageNumber="163">Harpactrox pontifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF28FF61FBF462DFFB9CFF5B" box="[1047,1109,223,248]" pageId="162" pageNumber="163" rank="species">
<emphasis id="39FCEAEDFF28FF61FBF462DFFB9CFF5B" bold="true" box="[1047,1109,223,248]" pageId="162" pageNumber="163">n. sp.</emphasis>
</taxonomicNameLabel>
as a new species on the basis of the unique preantennal area of the females. Several genera of the
<taxonomicName id="CC884D7CFF28FF61FC776304FC3DFEBE" authorityName="Bush" authorityYear="2017" box="[916,1012,260,285]" class="Insecta" family="Philopteridae" genus="Brueelia" kingdom="Animalia" order="Phthiraptera" pageId="162" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF28FF61FC776304FC3DFEBE" box="[916,1012,260,285]" italics="true" pageId="162" pageNumber="163">Brueelia</emphasis>
</taxonomicName>
-complex have species with relatively well-defined dorsal thickenings near the frons (e.g.
<taxonomicName id="CC884D7CFF28FF61FD186327FB90FEE3" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[763,1113,295,320]" class="Insecta" family="Philopteridae" genus="Indoceoplanetes" kingdom="Animalia" order="Phthiraptera" pageId="162" pageNumber="163" phylum="Arthropoda" rank="species" species="laurocorythes" status="sp. nov.">
<emphasis id="39FCEAEDFF28FF61FD186327FB90FEE3" box="[763,1113,295,320]" italics="true" pageId="162" pageNumber="163">Indoceoplanetes laurocorythes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF28FF61FB866327FB60FEE3" box="[1125,1193,295,320]" pageId="162" pageNumber="163" rank="species">
<emphasis id="39FCEAEDFF28FF61FB866327FB60FEE3" bold="true" box="[1125,1193,295,320]" pageId="162" pageNumber="163">n. sp.</emphasis>
</taxonomicNameLabel>
;
<figureCitation id="93B32A7AFF28FF61FB5F6327FAEAFEE3" box="[1212,1315,295,320]" captionStart="FIGURES 226 – 230" captionStartId="144.[151,264,1898,1920]" captionTargetBox="[204,1361,204,1841]" captionTargetId="figure@144.[191,1396,193,1877]" captionTargetPageId="144" captionText="FIGURES 226 – 230. Indoceoplanetes (Capnodella) laurocorythes n. gen., n. subgen. &amp; n. sp. ex Edolisoma holopolium holopolium: 226, male head, dorsal and ventral views. 227, male genitalia, dorsal view. 228, male mesosome, ventral view. 229, male paramere, dorsal view. 230, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883282/files/figure.png" pageId="162" pageNumber="163">Fig. 226</figureCitation>
). We here interpret these thickenings as part of the marginal carina displaced posteriorly, as suggested by the shape of this feature in e.g.
<taxonomicName id="CC884D7CFF28FF61FEA26370FDEBFE2B" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[321,546,367,392]" class="Insecta" family="Philopteridae" genus="Aratricerca" kingdom="Animalia" order="Phthiraptera" pageId="162" pageNumber="163" phylum="Arthropoda" rank="species" species="cirithra" status="sp. nov.">
<emphasis id="39FCEAEDFF28FF61FEA26370FDEBFE2B" box="[321,546,367,392]" italics="true" pageId="162" pageNumber="163">Aratricerca cirithra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF28FF61FDCF636FFDA4FE2B" box="[556,621,367,392]" pageId="162" pageNumber="163" rank="species">
<emphasis id="39FCEAEDFF28FF61FDCF636FFDA4FE2B" bold="true" box="[556,621,367,392]" pageId="162" pageNumber="163">n. sp.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="93B32A7AFF28FF61FD63636FFD2CFE2B" box="[640,741,367,392]" captionStart="FIGURES 170 – 174" captionStartId="110.[151,264,1915,1937]" captionTargetBox="[210,1343,209,1873]" captionTargetId="figure@110.[192,1395,193,1894]" captionTargetPageId="110" captionText="FIGURES 170 – 174. Aratricerca cirithrus n. gen. &amp; n. sp. ex Ptiloprora guisei guisei: 170, male head, dorsal and ventral views. 171, male genitalia, dorsal view. 172, male mesosome, ventral view. 173, male paramere, dorsal view. 174, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883252/files/figure.png" pageId="162" pageNumber="163">Fig. 170</figureCitation>
). This is not the case in
<taxonomicName id="CC884D7CFF28FF61FBEB6371FB64FE2B" authorityName="Bush" authorityYear="2017" box="[1032,1197,368,392]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="162" pageNumber="163" phylum="Arthropoda" rank="species" species="pontifrons">
<emphasis id="39FCEAEDFF28FF61FBEB6371FB64FE2B" box="[1032,1197,368,392]" italics="true" pageId="162" pageNumber="163">Ha. pontifrons</emphasis>
</taxonomicName>
, where the marginal carina is clearly continuous across the osculum (
<figureCitation id="93B32A7AFF28FF61FD5F6394FCE9FE0E" box="[700,800,404,429]" captionStart="FIGURES 260 – 262" captionStartId="161.[151,264,1970,1992]" captionTargetBox="[194,1356,212,1926]" captionTargetId="figure@161.[163,1404,193,1927]" captionTargetPageId="161" captionText="FIGURES 260 – 262. Harpactrox pontifrons n. gen. &amp; n. sp. ex Harpactes ardens ardens: 260, female head, dorsal and ventral views. 261, female habitus, dorsal and ventral views. 262, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883298/files/figure.png" pageId="162" pageNumber="163">Fig. 260</figureCitation>
).
<collectingRegion id="C94CF81DFF28FF61FCD76395FC9FFE0F" box="[820,854,405,428]" country="Maldives" name="Uthuru" pageId="162" pageNumber="163">No</collectingRegion>
similar structure is found in either of the other two species of
<taxonomicName id="CC884D7CFF28FF61FEF563B9FE51FE73" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[278,408,441,464]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="162" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF28FF61FEF563B9FE51FE73" box="[278,408,441,464]" italics="true" pageId="162" pageNumber="163">Harpactrox</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF28FF61FE4863B7FDDFFE73" box="[427,534,439,464]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="162" pageNumber="163">Figs 248</figureCitation>
, 255) nor, to our knowledge, in any other ischnoceran louse. Females of
<taxonomicName id="CC884D7CFF28FF61FA6263B9FEC0FE57" authorityName="Bush" authorityYear="2017" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="162" pageNumber="163" phylum="Arthropoda" rank="species" species="pontifrons">
<emphasis id="39FCEAEDFF28FF61FA6263B9FEC0FE57" italics="true" pageId="162" pageNumber="163">H. pontifrons</emphasis>
</taxonomicName>
are well separated from their congenerics, as described above. The
<typeStatus id="D433885DFF28FF61FBF863DDFB82FE56" box="[1051,1099,477,501]" pageId="162" pageNumber="163">type</typeStatus>
host of
<taxonomicName id="CC884D7CFF28FF61FB4D63DDFA9BFE57" authorityName="Bush" authorityYear="2017" box="[1198,1362,476,500]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="162" pageNumber="163" phylum="Arthropoda" rank="species" species="pontifrons">
<emphasis id="39FCEAEDFF28FF61FB4D63DDFA9BFE57" box="[1198,1362,476,500]" italics="true" pageId="162" pageNumber="163">Ha. pontifrons</emphasis>
</taxonomicName>
is not uncommon, but decreasing (
<bibRefCitation id="6F194B0EFF28FF61FE3563FFFCDDFDBB" author="BirdLife" box="[470,788,511,536]" pageId="162" pageNumber="163" refString="BirdLife International (2016) Harpactes ardens. In: The IUCN Red List of Threatened Species 2016: e. T 22682839 A 92963343. http: // dx. doi. org / 10.2305 / IUCN. UK. 2016 - 3. RLTS. T 22682839 A 92963343. en" type="journal article" year="2016">BirdLife International 2016</bibRefCitation>
), and collecting in the
<collectingCountry id="739F766FFF28FF61FBF463FFFB5CFDBB" box="[1047,1173,511,536]" name="Philippines" pageId="162" pageNumber="163">Philippines</collectingCountry>
may not be possible in the foreseeable future; hence additional fresh material to find, describe and compare the male with the other two species in the genus may take a long time to become available.
</paragraph>
</subSubSection>
</treatment>
</document>
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