treatments-xml/data/83/21/87/832187E9FF0EFF46FF74634CFE12F9D7.xml

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<mods:title id="7C1AC72D49B731308316E9D7FF8E8A7A">Morphological revision of the hyperdiverse Brueelia - complex (Insecta: Phthiraptera: Ischnocera: Philopteridae) with new taxa, checklists and generic key</mods:title>
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<subSubSection id="43926574FF0EFF47FF74634CFD0BFEC5" box="[151,706,332,358]" pageId="132" pageNumber="133" type="nomenclature">
<paragraph id="0B3736FFFF0EFF47FF74634CFD0BFEC5" blockId="132.[151,706,332,358]" box="[151,706,332,358]" pageId="132" pageNumber="133">
<heading id="507F8193FF0EFF47FF74634CFD0BFEC5" bold="true" box="[151,706,332,358]" fontSize="11" level="1" pageId="132" pageNumber="133" reason="1">
<emphasis id="39FCEAEDFF0EFF47FF74634CFD0BFEC5" bold="true" box="[151,706,332,358]" pageId="132" pageNumber="133">
<taxonomicName id="CC884D7CFF0EFF47FF74634CFDFDFEC5" authority="Gustafsson &amp; Bush" authorityName="Bush" authorityYear="2017" box="[151,564,332,358]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FF74634CFEF0FEC5" bold="true" box="[151,313,332,358]" italics="true" pageId="132" pageNumber="133">Titanomessor</emphasis>
Gustafsson &amp; Bush
</taxonomicName>
, 
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</emphasis>
</heading>
</paragraph>
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<subSubSection id="43926574FF0EFF47FF746392FD41FE0B" box="[151,648,402,424]" pageId="132" pageNumber="133" type="materials_examined">
<paragraph id="0B3736FFFF0EFF47FF746392FD41FE0B" blockId="132.[151,648,402,424]" box="[151,648,402,424]" pageId="132" pageNumber="133">
<emphasis id="39FCEAEDFF0EFF47FF746392FEEBFE0B" bold="true" box="[151,290,402,424]" pageId="132" pageNumber="133">Type species.</emphasis>
<taxonomicName id="CC884D7CFF0EFF47FECA6393FDCCFE0B" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[297,517,402,424]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="sexloba" status="sp. nov.">
<emphasis id="39FCEAEDFF0EFF47FECA6393FDCCFE0B" box="[297,517,402,424]" italics="true" pageId="132" pageNumber="133">Titanomessor sexloba</emphasis>
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<taxonomicNameLabel id="22CF5796FF0EFF47FDEF6392FD41FE0B" box="[524,648,402,424]" pageId="132" pageNumber="133" rank="species">
<emphasis id="39FCEAEDFF0EFF47FDEF6392FD41FE0B" bold="true" box="[524,648,402,424]" pageId="132" pageNumber="133">new species</emphasis>
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</paragraph>
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<subSubSection id="43926574FF0EFF47FF7463D5FAECF915" pageId="132" pageNumber="133" type="diagnosis">
<paragraph id="0B3736FFFF0EFF47FF7463D5FD73FDDD" blockId="132.[151,1437,469,2006]" pageId="132" pageNumber="133">
<emphasis id="39FCEAEDFF0EFF47FF7463D5FED8FE4D" bold="true" box="[151,273,469,494]" pageId="132" pageNumber="133">Diagnosis.</emphasis>
<taxonomicName id="CC884D7CFF0EFF47FEF963D6FE78FE4E" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[282,433,470,493]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FEF963D6FE78FE4E" box="[282,433,470,493]" italics="true" pageId="132" pageNumber="133">Titanomessor</emphasis>
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<emphasis id="39FCEAEDFF0EFF47FE5A63D5FDC0FE4D" bold="true" box="[441,521,469,494]" pageId="132" pageNumber="133">n. gen.</emphasis>
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is separated from all other genera treated here by the unique male genitalia and the fact that 
<emphasis id="39FCEAEDFF0EFF47FEC763F9FEBCFDB2" box="[292,373,505,529]" italics="true" pageId="132" pageNumber="133">
<taxonomicName id="CC884D7CFF0EFF47FEC763F9FE83FDB2" box="[292,330,505,529]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
<quantity id="CC709B1AFF0EFF47FEA963F9FEBCFDB2" box="[330,373,505,529]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="132" pageNumber="133" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">3–4</quantity>
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in both sexes, as well as 
<emphasis id="39FCEAEDFF0EFF47FD7563FAFD02FDB2" box="[662,715,506,529]" italics="true" pageId="132" pageNumber="133">
<taxonomicName id="CC884D7CFF0EFF47FD7563FAFD74FDB2" box="[662,701,506,529]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
<quantity id="CC709B1AFF0EFF47FD5E63FAFD02FDB2" box="[701,715,506,529]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="132" pageNumber="133" unit="in" value="5.0">5</quantity>
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in males, are all of equal length, and much longer than 
<emphasis id="39FCEAEDFF0EFF47FAA763F9FA5CFDB2" box="[1348,1429,505,529]" italics="true" pageId="132" pageNumber="133">
<taxonomicName id="CC884D7CFF0EFF47FAA763F9FAA5FDB2" box="[1348,1388,505,529]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
1–2
</emphasis>
. Within the 
<taxonomicName id="CC884D7CFF0EFF47FEF0601DFEBAFD95" authorityName="Keler" authorityYear="1936" box="[275,371,541,566]" class="Insecta" family="Philopteridae" genus="Brueelia" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF0EFF47FEF0601DFEBAFD95" box="[275,371,541,566]" italics="true" pageId="132" pageNumber="133">Brueelia</emphasis>
</taxonomicName>
-complex, 
<taxonomicName id="CC884D7CFF0EFF47FE08601EFDA4FD96" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[491,621,542,565]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FE08601EFDA4FD96" box="[491,621,542,565]" italics="true" pageId="132" pageNumber="133">Harpactrox</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF0EFF47FD90601DFD08FD95" box="[627,705,541,566]" pageId="132" pageNumber="133" rank="genus">
<emphasis id="39FCEAEDFF0EFF47FD90601DFD08FD95" bold="true" box="[627,705,541,566]" pageId="132" pageNumber="133">n. gen.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="93B32A7AFF0EFF47FD2C601DFCFCFD95" box="[719,821,541,566]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="132" pageNumber="133">Figs 248</figureCitation>
, 255, 260) is the only other genus that has a transversal dorsal preantennal suture, but these two genera are otherwise not similar in head chaetotaxy or structure, abdominal chaetotaxy, or the structure of the male genitalia.
</paragraph>
<paragraph id="0B3736FFFF0EFF47FF246089FD21FBD5" blockId="132.[151,1437,469,2006]" pageId="132" pageNumber="133">
Overall, the genus most similar to 
<taxonomicName id="CC884D7CFF0EFF47FDAE608AFD2DFD02" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[589,740,650,673]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FDAE608AFD2DFD02" box="[589,740,650,673]" italics="true" pageId="132" pageNumber="133">Titanomessor</emphasis>
</taxonomicName>
is 
<taxonomicName id="CC884D7CFF0EFF47FCE56089FC74FD01" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" baseAuthorityName="Capnodella" box="[774,957,649,674]" class="Insecta" family="Philopteridae" genus="Indoceoplanetes" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FCE56089FC74FD01" box="[774,957,649,674]" italics="true" pageId="132" pageNumber="133">Indoceoplanetes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF0EFF47FC266089FBDDFD01" box="[965,1044,649,674]" pageId="132" pageNumber="133" rank="genus">
<emphasis id="39FCEAEDFF0EFF47FC266089FBDDFD01" bold="true" box="[965,1044,649,674]" pageId="132" pageNumber="133">n. gen.</emphasis>
</taxonomicNameLabel>
, particularly 
<emphasis id="39FCEAEDFF0EFF47FB4F608AFB02FD02" box="[1196,1227,650,673]" italics="true" pageId="132" pageNumber="133">In.</emphasis>
(
<taxonomicName id="CC884D7CFF0EFF47FB386089FA5BFD01" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" baseAuthorityName="Capnodella" box="[1243,1426,649,674]" class="Insecta" family="Philopteridae" genus="Indoceoplanetes" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="137" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FB386089FA5BFD01" box="[1243,1426,649,674]" italics="true" pageId="132" pageNumber="133">Indoceoplanetes</emphasis>
</taxonomicName>
) 
<emphasis id="39FCEAEDFF0EFF47FF7460ADFEDAFD65" bold="true" box="[151,275,685,710]" pageId="132" pageNumber="133">n. subgen.</emphasis>
These two genera have similar abdominal chaetotaxy (
<tableCitation id="460A0344FF0EFF47FC7560ADFC39FD65" box="[918,1008,685,710]" captionStart="TABLE 2" captionStartId="30.[151,239,796,819]" captionText="TABLE 2. Summary of abdominal chaetotaxy of segments II – VIII for the Brueelia - complex, including both genera and subgenera. In genera where chaetotaxy differs between species group (marked with *), only that of the type species is given here, and more detailed accounts are given separately under the respective genus entries and in Tables 3 – 12. Trichoid setae of segment VIII are present in all genera, and are not listed. Abbreviations: ps = paratergal seta; aps = accessory post-spiracular seta; psps = principal post-spiracular seta; tps = tergal posterior seta; ss = sutural seta; sts = sternal seta." pageId="132" pageNumber="133">Table 2</tableCitation>
), except that 
<taxonomicName id="CC884D7CFF0EFF47FB7360AEFB78FD66" box="[1168,1201,686,709]" class="Insecta" family="Philopteridae" genus="Brueelia" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="subSpecies" species="brachythorax" subSpecies="tps">
<emphasis id="39FCEAEDFF0EFF47FB7360AEFB78FD66" box="[1168,1201,686,709]" italics="true" pageId="132" pageNumber="133">tps</emphasis>
</taxonomicName>
are present in more anterior segments in males of 
<emphasis id="39FCEAEDFF0EFF47FE0560D2FDCCFD4A" box="[486,517,722,745]" italics="true" pageId="132" pageNumber="133">In.</emphasis>
(
<taxonomicName id="CC884D7CFF0EFF47FDF760D1FD02FD49" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" baseAuthorityName="Capnodella" box="[532,715,721,746]" class="Insecta" family="Philopteridae" genus="Indoceoplanetes" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="137" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FDF760D1FD02FD49" box="[532,715,721,746]" italics="true" pageId="132" pageNumber="133">Indoceoplanetes</emphasis>
</taxonomicName>
) (
<figureCitation id="93B32A7AFF0EFF47FD0760D1FC8DFD49" box="[740,836,721,746]" captionStart="FIGURES 217 – 218" captionStartId="140.[151,264,1869,1891]" captionTargetBox="[220,1338,235,1844]" captionTargetId="figure@140.[151,1436,193,1847]" captionTargetPageId="140" captionText="FIGURES 217 – 218. Indoceoplanetes (Indoceoplanetes) indonesiana (Eichler, 1947) n. comb. ex Coracina striata difficilis: 217, male habitus, dorsal and ventral views. 218, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883276/files/figure.png" pageId="132" pageNumber="133">Fig. 217</figureCitation>
) than in males of 
<taxonomicName id="CC884D7CFF0EFF47FBEF60D2FB6AFD4A" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[1036,1187,722,745]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FBEF60D2FB6AFD4A" box="[1036,1187,722,745]" italics="true" pageId="132" pageNumber="133">Titanomessor</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF0EFF47FB5160D1FADDFD49" box="[1202,1300,721,746]" captionStart="FIGURES 210 – 211" captionStartId="134.[151,264,1837,1859]" captionTargetBox="[193,1373,229,1811]" captionTargetId="figure@134.[151,1436,193,1815]" captionTargetPageId="134" captionText="FIGURES 210 – 211. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 210, male habitus, dorsal and ventral views. 211, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883272/files/figure.png" pageId="132" pageNumber="133">Fig. 210</figureCitation>
). In both 
<emphasis id="39FCEAEDFF0EFF47FA9E60D2FA55FD4A" box="[1405,1436,722,745]" italics="true" pageId="132" pageNumber="133">In.</emphasis>
(
<taxonomicName id="CC884D7CFF0EFF47FF4360F5FE91FCAD" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" baseAuthorityName="Capnodella" box="[160,344,757,782]" class="Insecta" family="Philopteridae" genus="Indoceoplanetes" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="137" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FF4360F5FE91FCAD" box="[160,344,757,782]" italics="true" pageId="132" pageNumber="133">Indoceoplanetes</emphasis>
</taxonomicName>
) (
<figureCitation id="93B32A7AFF0EFF47FE9060F5FE1FFCAD" box="[371,470,757,782]" captionStart="FIGURES 219 – 223" captionStartId="141.[151,264,1915,1937]" captionTargetBox="[228,1354,243,1866]" captionTargetId="figure@141.[200,1387,193,1894]" captionTargetPageId="141" captionText="FIGURES 219 – 223. Indoceoplanetes (Indoceoplanetes) indonesiana (Eichler, 1947) n. comb. ex Coracina striata difficilis: 219, male head, dorsal and ventral views. 220, male genitalia, dorsal view. 221, male mesosome, ventral view. 222, male paramere, dorsal view. 223, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883278/files/figure.png" pageId="132" pageNumber="133">Fig. 219</figureCitation>
) and 
<taxonomicName id="CC884D7CFF0EFF47FDF860F6FD7BFCAE" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[539,690,758,781]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FDF860F6FD7BFCAE" box="[539,690,758,781]" italics="true" pageId="132" pageNumber="133">Titanomessor</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF0EFF47FD2760F5FCE3FCAD" box="[708,810,757,782]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="132" pageNumber="133">Fig. 212</figureCitation>
), the marginal carina is uninterrupted but displaced at osculum, the female submarginal plate is broad distally, and approaches but does not reach the vulval margin (
<figureCitation id="93B32A7AFF0EFF47FA896119FF0DFCF5" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="132" pageNumber="133">Figs 216</figureCitation>
, 
<figureCitation id="93B32A7AFF0EFF47FF32613EFF36FCF5" box="[209,255,830,854]" captionStart="FIGURES 219 – 223" captionStartId="141.[151,264,1915,1937]" captionTargetBox="[228,1354,243,1866]" captionTargetId="figure@141.[200,1387,193,1894]" captionTargetPageId="141" captionText="FIGURES 219 – 223. Indoceoplanetes (Indoceoplanetes) indonesiana (Eichler, 1947) n. comb. ex Coracina striata difficilis: 219, male head, dorsal and ventral views. 220, male genitalia, dorsal view. 221, male mesosome, ventral view. 222, male paramere, dorsal view. 223, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883278/files/figure.png" pageId="132" pageNumber="133">223</figureCitation>
). In both genera, 
<emphasis id="39FCEAEDFF0EFF47FE29613EFE36FCF6" box="[458,511,830,853]" italics="true" pageId="132" pageNumber="133">
<taxonomicName id="CC884D7CFF0EFF47FE29613EFE38FCF6" box="[458,497,830,853]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
5
</emphasis>
and sometimes 
<emphasis id="39FCEAEDFF0EFF47FD58613EFD26FCF5" box="[699,751,830,854]" italics="true" pageId="132" pageNumber="133">
<taxonomicName id="CC884D7CFF0EFF47FD58613EFD2BFCF5" box="[699,738,830,854]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
4
</emphasis>
are typically clearly dorsal, rather than marginal. However, there are large differences between the two groups. There is no dorsal preantennal suture in 
<emphasis id="39FCEAEDFF0EFF47FB486162FB03FCDA" box="[1195,1226,866,889]" italics="true" pageId="132" pageNumber="133">In.</emphasis>
(
<taxonomicName id="CC884D7CFF0EFF47FB386161FA5BFCD9" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" baseAuthorityName="Capnodella" box="[1243,1426,865,890]" class="Insecta" family="Philopteridae" genus="Indoceoplanetes" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="137" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FB386161FA5BFCD9" box="[1243,1426,865,890]" italics="true" pageId="132" pageNumber="133">Indoceoplanetes</emphasis>
</taxonomicName>
) (
<figureCitation id="93B32A7AFF0EFF47FF7C6185FECBFC3D" box="[159,258,901,926]" captionStart="FIGURES 219 – 223" captionStartId="141.[151,264,1915,1937]" captionTargetBox="[228,1354,243,1866]" captionTargetId="figure@141.[200,1387,193,1894]" captionTargetPageId="141" captionText="FIGURES 219 – 223. Indoceoplanetes (Indoceoplanetes) indonesiana (Eichler, 1947) n. comb. ex Coracina striata difficilis: 219, male head, dorsal and ventral views. 220, male genitalia, dorsal view. 221, male mesosome, ventral view. 222, male paramere, dorsal view. 223, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883278/files/figure.png" pageId="132" pageNumber="133">Fig. 219</figureCitation>
), but there is in 
<taxonomicName id="CC884D7CFF0EFF47FE596186FD98FC3E" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[442,593,902,925]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FE596186FD98FC3E" box="[442,593,902,925]" italics="true" pageId="132" pageNumber="133">Titanomessor</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF0EFF47FD826185FD0DFC3D" box="[609,708,901,926]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="132" pageNumber="133">Fig. 212</figureCitation>
). In 
<taxonomicName id="CC884D7CFF0EFF47FD146186FC47FC3E" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[759,910,902,925]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FD146186FC47FC3E" box="[759,910,902,925]" italics="true" pageId="132" pageNumber="133">Titanomessor</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF0EFF47FC7E6185FBF3FC3D" box="[925,1082,901,926]" captionStart="FIGURES 210 – 211" captionStartId="134.[151,264,1837,1859]" captionTargetBox="[193,1373,229,1811]" captionTargetId="figure@134.[151,1436,193,1815]" captionTargetPageId="134" captionText="FIGURES 210 – 211. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 210, male habitus, dorsal and ventral views. 211, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883272/files/figure.png" pageId="132" pageNumber="133">Figs 210–211</figureCitation>
) 
<emphasis id="39FCEAEDFF0EFF47FBAA6186FB53FC3D" box="[1097,1178,902,926]" italics="true" pageId="132" pageNumber="133">
<taxonomicName id="CC884D7CFF0EFF47FBAA6186FBB9FC3D" box="[1097,1136,902,926]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
<quantity id="CC709B1AFF0EFF47FB8C6186FB53FC3D" box="[1135,1178,902,926]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.27" metricValueMin="0.762" pageId="132" pageNumber="133" unit="in" value="4.0" valueMax="5.0" valueMin="3.0">3–5</quantity>
</emphasis>
in male and 
<emphasis id="39FCEAEDFF0EFF47FACE6186FAB7FC3D" box="[1325,1406,902,926]" italics="true" pageId="132" pageNumber="133">
<taxonomicName id="CC884D7CFF0EFF47FACE6186FA9AFC3D" box="[1325,1363,902,926]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
<quantity id="CC709B1AFF0EFF47FAB76186FAB7FC3D" box="[1364,1406,902,926]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="132" pageNumber="133" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">3–4</quantity>
</emphasis>
in female are mesosetae, whereas 
<emphasis id="39FCEAEDFF0EFF47FDC561AAFD95FC61" box="[550,604,938,962]" italics="true" pageId="132" pageNumber="133">
<taxonomicName id="CC884D7CFF0EFF47FDC561AAFD87FC61" box="[550,590,938,962]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
3
</emphasis>
is a macroseta and 
<emphasis id="39FCEAEDFF0EFF47FC9061AAFC0DFC61" box="[883,964,938,962]" italics="true" pageId="132" pageNumber="133">
<taxonomicName id="CC884D7CFF0EFF47FC9061AAFC50FC61" box="[883,921,938,962]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
<quantity id="CC709B1AFF0EFF47FC7A61AAFC0DFC61" box="[921,964,938,962]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.27" metricValueMin="1.016" pageId="132" pageNumber="133" unit="in" value="4.5" valueMax="5.0" valueMin="4.0">4–5</quantity>
</emphasis>
in both sexes are microsetae in 
<emphasis id="39FCEAEDFF0EFF47FA9E61AAFA55FC62" box="[1405,1436,938,961]" italics="true" pageId="132" pageNumber="133">In.</emphasis>
(
<taxonomicName id="CC884D7CFF0EFF47FF4361CDFE91FC45" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" baseAuthorityName="Capnodella" box="[160,344,973,998]" class="Insecta" family="Philopteridae" genus="Indoceoplanetes" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="137" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FF4361CDFE91FC45" box="[160,344,973,998]" italics="true" pageId="132" pageNumber="133">Indoceoplanetes</emphasis>
</taxonomicName>
) (
<figureCitation id="93B32A7AFF0EFF47FE9261CDFDDAFC45" box="[369,531,973,998]" captionStart="FIGURES 217 – 218" captionStartId="140.[151,264,1869,1891]" captionTargetBox="[220,1338,235,1844]" captionTargetId="figure@140.[151,1436,193,1847]" captionTargetPageId="140" captionText="FIGURES 217 – 218. Indoceoplanetes (Indoceoplanetes) indonesiana (Eichler, 1947) n. comb. ex Coracina striata difficilis: 217, male habitus, dorsal and ventral views. 218, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883276/files/figure.png" pageId="132" pageNumber="133">Figs 217–218</figureCitation>
). In females of 
<emphasis id="39FCEAEDFF0EFF47FD2461CEFD2CFC46" box="[711,741,974,997]" italics="true" pageId="132" pageNumber="133">In.</emphasis>
(
<taxonomicName id="CC884D7CFF0EFF47FD1461CDFC67FC45" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" baseAuthorityName="Capnodella" box="[759,942,973,998]" class="Insecta" family="Philopteridae" genus="Indoceoplanetes" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="137" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FD1461CDFC67FC45" box="[759,942,973,998]" italics="true" pageId="132" pageNumber="133">Indoceoplanetes</emphasis>
</taxonomicName>
) (
<figureCitation id="93B32A7AFF0EFF47FC2A61CDFBE3FC45" box="[969,1066,973,998]" captionStart="FIGURES 217 – 218" captionStartId="140.[151,264,1869,1891]" captionTargetBox="[220,1338,235,1844]" captionTargetId="figure@140.[151,1436,193,1847]" captionTargetPageId="140" captionText="FIGURES 217 – 218. Indoceoplanetes (Indoceoplanetes) indonesiana (Eichler, 1947) n. comb. ex Coracina striata difficilis: 217, male habitus, dorsal and ventral views. 218, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883276/files/figure.png" pageId="132" pageNumber="133">Fig. 218</figureCitation>
) tergopleurite XI is not fused to tergopleurite IX+X, but these plates are fused in 
<taxonomicName id="CC884D7CFF0EFF47FD3861F2FCBBFBAA" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[731,882,1010,1033]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FD3861F2FCBBFBAA" box="[731,882,1010,1033]" italics="true" pageId="132" pageNumber="133">Titanomessor</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF0EFF47FC6661F1FC22FBA9" box="[901,1003,1009,1034]" captionStart="FIGURES 210 – 211" captionStartId="134.[151,264,1837,1859]" captionTargetBox="[193,1373,229,1811]" captionTargetId="figure@134.[151,1436,193,1815]" captionTargetPageId="134" captionText="FIGURES 210 – 211. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 210, male habitus, dorsal and ventral views. 211, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883272/files/figure.png" pageId="132" pageNumber="133">Fig. 211</figureCitation>
). Finally, the male genitalia are very different between these two groups. The parameres in 
<emphasis id="39FCEAEDFF0EFF47FCFE6616FCF5FB8E" box="[797,828,1046,1069]" italics="true" pageId="132" pageNumber="133">In.</emphasis>
(
<taxonomicName id="CC884D7CFF0EFF47FCB26615FBC1FB8D" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" baseAuthorityName="Capnodella" box="[849,1032,1045,1070]" class="Insecta" family="Philopteridae" genus="Indoceoplanetes" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="137" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FCB26615FBC1FB8D" box="[849,1032,1045,1070]" italics="true" pageId="132" pageNumber="133">Indoceoplanetes</emphasis>
</taxonomicName>
) (
<figureCitation id="93B32A7AFF0EFF47FBC56615FB43FB8D" box="[1062,1162,1045,1070]" captionStart="FIGURES 219 – 223" captionStartId="141.[151,264,1915,1937]" captionTargetBox="[228,1354,243,1866]" captionTargetId="figure@141.[200,1387,193,1894]" captionTargetPageId="141" captionText="FIGURES 219 – 223. Indoceoplanetes (Indoceoplanetes) indonesiana (Eichler, 1947) n. comb. ex Coracina striata difficilis: 219, male head, dorsal and ventral views. 220, male genitalia, dorsal view. 221, male mesosome, ventral view. 222, male paramere, dorsal view. 223, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883278/files/figure.png" pageId="132" pageNumber="133">Fig. 222</figureCitation>
) are curved, wheras in 
<taxonomicName id="CC884D7CFF0EFF47FF74663AFEE7FBF2" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[151,302,1082,1105]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FF74663AFEE7FBF2" box="[151,302,1082,1105]" italics="true" pageId="132" pageNumber="133">Titanomessor</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF0EFF47FEA26639FE6DFBF1" box="[321,420,1081,1106]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="132" pageNumber="133">Fig. 215</figureCitation>
) the parameres are straight and convergent, and the structure of the mesosome of both genera are very different (
<emphasis id="39FCEAEDFF0EFF47FE58665EFE06FBD5" box="[443,463,1118,1142]" italics="true" pageId="132" pageNumber="133">cf</emphasis>
. 
<figureCitation id="93B32A7AFF0EFF47FE3E665DFDF3FBD5" box="[477,570,1117,1142]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="132" pageNumber="133">Fig. 214</figureCitation>
with 
<figureCitation id="93B32A7AFF0EFF47FD99665DFD12FBD5" box="[634,731,1117,1142]" captionStart="FIGURES 219 – 223" captionStartId="141.[151,264,1915,1937]" captionTargetBox="[228,1354,243,1866]" captionTargetId="figure@141.[200,1387,193,1894]" captionTargetPageId="141" captionText="FIGURES 219 – 223. Indoceoplanetes (Indoceoplanetes) indonesiana (Eichler, 1947) n. comb. ex Coracina striata difficilis: 219, male head, dorsal and ventral views. 220, male genitalia, dorsal view. 221, male mesosome, ventral view. 222, male paramere, dorsal view. 223, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883278/files/figure.png" pageId="132" pageNumber="133">Fig. 221</figureCitation>
).
</paragraph>
<paragraph id="0B3736FFFF0EFF47FF246682FD97FAED" blockId="132.[151,1437,469,2006]" pageId="132" pageNumber="133">
In 
<taxonomicName id="CC884D7CFF0EFF47FF056681FDF3FB39" authority="Capnodella" authorityName="Bush" authorityYear="2017" box="[230,570,1153,1178]" class="Insecta" family="Philopteridae" genus="Indoceoplanetes" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="subGenus" status="n. gen. et n. subgen." subGenus="Capnodella">
<emphasis id="39FCEAEDFF0EFF47FF056681FE57FB39" box="[230,414,1153,1178]" italics="true" pageId="132" pageNumber="133">Indoceoplanetes</emphasis>
(
<emphasis id="39FCEAEDFF0EFF47FE4D6681FDF8FB39" box="[430,561,1153,1178]" italics="true" pageId="132" pageNumber="133">Capnodella</emphasis>
)
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF0EFF47FDA16681FCF8FB39" box="[578,817,1153,1178]" pageId="132" pageNumber="133" rank="subGenus">
<emphasis id="39FCEAEDFF0EFF47FDA16681FCF8FB39" bold="true" box="[578,817,1153,1178]" pageId="132" pageNumber="133">n. gen. et n. subgen.</emphasis>
</taxonomicNameLabel>
female tergopleurite IX+X is fused to tergopleurite XI (
<figureCitation id="93B32A7AFF0EFF47FF7C66A5FECCFB1D" box="[159,261,1189,1214]" captionStart="FIGURES 224 – 225" captionStartId="143.[151,264,1968,1990]" captionTargetBox="[204,1393,235,1893]" captionTargetId="figure@143.[163,1407,193,1929]" captionTargetPageId="143" captionText="FIGURES 224 – 225. Indoceoplanetes (Capnodella) laurocorythes n. gen., n. subgen. &amp; n. sp. ex Edolisoma holopolium holopolium: 224, male habitus, dorsal and ventral views. 225, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883280/files/figure.png" pageId="132" pageNumber="133">Figs 225</figureCitation>
, 
<figureCitation id="93B32A7AFF0EFF47FEF366A6FEF5FB1D" box="[272,316,1190,1214]" captionStart="FIGURES 231 – 232" captionStartId="147.[151,264,1952,1974]" captionTargetBox="[264,1397,228,1919]" captionTargetId="figure@147.[173,1417,195,1939]" captionTargetPageId="147" captionText="FIGURES 231 – 232. Indoceoplanetes (Capnodella) loboccupatrix n. gen., n. subgen. &amp; n. sp. ex Lobotos oriolinus: 231, male habitus, dorsal and ventral views. 232, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883284/files/figure.png" pageId="132" pageNumber="133">232</figureCitation>
) as in 
<taxonomicName id="CC884D7CFF0EFF47FE6466A6FDD7FB1E" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[391,542,1190,1213]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FE6466A6FDD7FB1E" box="[391,542,1190,1213]" italics="true" pageId="132" pageNumber="133">Titanomessor</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF0EFF47FDCD66A5FD46FB1D" box="[558,655,1189,1214]" captionStart="FIGURES 210 – 211" captionStartId="134.[151,264,1837,1859]" captionTargetBox="[193,1373,229,1811]" captionTargetId="figure@134.[151,1436,193,1815]" captionTargetPageId="134" captionText="FIGURES 210 – 211. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 210, male habitus, dorsal and ventral views. 211, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883272/files/figure.png" pageId="132" pageNumber="133">Fig. 211</figureCitation>
), but unlike in 
<taxonomicName id="CC884D7CFF0EFF47FCD966A6FC18FB1E" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[826,977,1190,1213]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FCD966A6FC18FB1E" box="[826,977,1190,1213]" italics="true" pageId="132" pageNumber="133">Titanomessor</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF0EFF47FC0266A5FB8DFB1D" box="[993,1092,1189,1214]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="132" pageNumber="133">Fig. 212</figureCitation>
), the dorsal preantennal suture of 
<emphasis id="39FCEAEDFF0EFF47FF5466CAFF1FFB42" box="[183,214,1226,1249]" italics="true" pageId="132" pageNumber="133">In.</emphasis>
(
<emphasis id="39FCEAEDFF0EFF47FF0B66C9FEA2FB41" box="[232,363,1225,1250]" italics="true" pageId="132" pageNumber="133">Capnodella</emphasis>
) (
<figureCitation id="93B32A7AFF0EFF47FE6666C9FE24FB41" box="[389,493,1225,1250]" captionStart="FIGURES 226 – 230" captionStartId="144.[151,264,1898,1920]" captionTargetBox="[204,1361,204,1841]" captionTargetId="figure@144.[191,1396,193,1877]" captionTargetPageId="144" captionText="FIGURES 226 – 230. Indoceoplanetes (Capnodella) laurocorythes n. gen., n. subgen. &amp; n. sp. ex Edolisoma holopolium holopolium: 226, male head, dorsal and ventral views. 227, male genitalia, dorsal view. 228, male mesosome, ventral view. 229, male paramere, dorsal view. 230, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883282/files/figure.png" pageId="132" pageNumber="133">Figs 226</figureCitation>
, 
<figureCitation id="93B32A7AFF0EFF47FE1966CAFDEFFB41" box="[506,550,1226,1250]" captionStart="FIGURES 233 – 237" captionStartId="148.[151,264,1915,1937]" captionTargetBox="[228,1343,228,1862]" captionTargetId="figure@148.[208,1379,193,1894]" captionTargetPageId="148" captionText="FIGURES 233 – 237. Indoceoplanetes (Capnodella) loboccupatrix n. gen., n. subgen. &amp; n. sp. ex Lobotos oriolinus: 233, male head, dorsal and ventral views. 234, male genitalia, dorsal view. 235, male mesosome, ventral view. 236, male paramere, dorsal view. 237, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883286/files/figure.png" pageId="132" pageNumber="133">233</figureCitation>
) is longitudinal rather than latitudinal, and does interrupt the marginal carina submedianly. 
<collectingRegion id="C94CF81DFF0EFF47FED766EDFEA7FAA5" box="[308,366,1261,1286]" country="Maldives" name="Maale" pageId="132" pageNumber="133">Male</collectingRegion>
<emphasis id="39FCEAEDFF0EFF47FE9466EEFE5CFAA6" box="[375,405,1262,1285]" italics="true" pageId="132" pageNumber="133">In.</emphasis>
(
<emphasis id="39FCEAEDFF0EFF47FE4B66EDFDE2FAA5" box="[424,555,1261,1286]" italics="true" pageId="132" pageNumber="133">Capnodella</emphasis>
) also lack 
<taxonomicName id="CC884D7CFF0EFF47FD4866EEFD05FAA6" box="[683,716,1262,1285]" class="Insecta" family="Philopteridae" genus="Brueelia" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="subSpecies" species="brachythorax" subSpecies="tps">
<emphasis id="39FCEAEDFF0EFF47FD4866EEFD05FAA6" box="[683,716,1262,1285]" italics="true" pageId="132" pageNumber="133">tps</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF0EFF47FD3F66EDFC8DFAA5" box="[732,836,1261,1286]" captionStart="FIGURES 224 – 225" captionStartId="143.[151,264,1968,1990]" captionTargetBox="[204,1393,235,1893]" captionTargetId="figure@143.[163,1407,193,1929]" captionTargetPageId="143" captionText="FIGURES 224 – 225. Indoceoplanetes (Capnodella) laurocorythes n. gen., n. subgen. &amp; n. sp. ex Edolisoma holopolium holopolium: 224, male habitus, dorsal and ventral views. 225, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883280/files/figure.png" pageId="132" pageNumber="133">Figs 224</figureCitation>
, 
<figureCitation id="93B32A7AFF0EFF47FCB266EEFCB7FAA5" box="[849,894,1262,1286]" captionStart="FIGURES 231 – 232" captionStartId="147.[151,264,1952,1974]" captionTargetBox="[264,1397,228,1919]" captionTargetId="figure@147.[173,1417,195,1939]" captionTargetPageId="147" captionText="FIGURES 231 – 232. Indoceoplanetes (Capnodella) loboccupatrix n. gen., n. subgen. &amp; n. sp. ex Lobotos oriolinus: 231, male habitus, dorsal and ventral views. 232, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883284/files/figure.png" pageId="132" pageNumber="133">231</figureCitation>
), which are present in male 
<taxonomicName id="CC884D7CFF0EFF47FB2566EEFA94FAA6" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[1222,1373,1262,1285]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FB2566EEFA94FAA6" box="[1222,1373,1262,1285]" italics="true" pageId="132" pageNumber="133">Titanomessor</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF0EFF47FA8D66EDFF0CFA89" captionStart="FIGURES 210 – 211" captionStartId="134.[151,264,1837,1859]" captionTargetBox="[193,1373,229,1811]" captionTargetId="figure@134.[151,1436,193,1815]" captionTargetPageId="134" captionText="FIGURES 210 – 211. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 210, male habitus, dorsal and ventral views. 211, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883272/files/figure.png" pageId="132" pageNumber="133">Fig. 210</figureCitation>
), and while both 
<emphasis id="39FCEAEDFF0EFF47FE676712FE6AFA8A" box="[388,419,1298,1321]" italics="true" pageId="132" pageNumber="133">In.</emphasis>
(
<taxonomicName id="CC884D7CFF0EFF47FE506711FDA3FA89" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" baseAuthorityName="Capnodella" box="[435,618,1297,1322]" class="Insecta" family="Philopteridae" genus="Indoceoplanetes" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="137" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FE506711FDA3FA89" box="[435,618,1297,1322]" italics="true" pageId="132" pageNumber="133">Indoceoplanetes</emphasis>
</taxonomicName>
) (
<figureCitation id="93B32A7AFF0EFF47FD606711FD2BFA89" box="[643,738,1297,1322]" captionStart="FIGURES 219 – 223" captionStartId="141.[151,264,1915,1937]" captionTargetBox="[228,1354,243,1866]" captionTargetId="figure@141.[200,1387,193,1894]" captionTargetPageId="141" captionText="FIGURES 219 – 223. Indoceoplanetes (Indoceoplanetes) indonesiana (Eichler, 1947) n. comb. ex Coracina striata difficilis: 219, male head, dorsal and ventral views. 220, male genitalia, dorsal view. 221, male mesosome, ventral view. 222, male paramere, dorsal view. 223, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883278/files/figure.png" pageId="132" pageNumber="133">Fig. 219</figureCitation>
) and 
<taxonomicName id="CC884D7CFF0EFF47FCC36712FC7EFA8A" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[800,951,1298,1321]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FCC36712FC7EFA8A" box="[800,951,1298,1321]" italics="true" pageId="132" pageNumber="133">Titanomessor</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF0EFF47FC256711FBECFA89" box="[966,1061,1297,1322]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="132" pageNumber="133">Fig. 212</figureCitation>
) lack a ventral anterior plate, this is present in 
<emphasis id="39FCEAEDFF0EFF47FEC66736FE8DFAEE" box="[293,324,1334,1357]" italics="true" pageId="132" pageNumber="133">In.</emphasis>
(
<emphasis id="39FCEAEDFF0EFF47FEB76735FE1EFAED" box="[340,471,1333,1358]" italics="true" pageId="132" pageNumber="133">Capnodella</emphasis>
) (
<figureCitation id="93B32A7AFF0EFF47FE0C6735FD9BFAED" box="[495,594,1333,1358]" captionStart="FIGURES 226 – 230" captionStartId="144.[151,264,1898,1920]" captionTargetBox="[204,1361,204,1841]" captionTargetId="figure@144.[191,1396,193,1877]" captionTargetPageId="144" captionText="FIGURES 226 – 230. Indoceoplanetes (Capnodella) laurocorythes n. gen., n. subgen. &amp; n. sp. ex Edolisoma holopolium holopolium: 226, male head, dorsal and ventral views. 227, male genitalia, dorsal view. 228, male mesosome, ventral view. 229, male paramere, dorsal view. 230, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883282/files/figure.png" pageId="132" pageNumber="133">Fig. 226</figureCitation>
).
</paragraph>
<paragraph id="0B3736FFFF0EFF47FF246759FC8BF931" blockId="132.[151,1437,469,2006]" pageId="132" pageNumber="133">
The large size (see Measurements below) and the shape of the mesomere (
<figureCitation id="93B32A7AFF0EFF47FBE26759FBA9FAD1" box="[1025,1120,1369,1394]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="132" pageNumber="133">Fig. 214</figureCitation>
) suggests that 
<taxonomicName id="CC884D7CFF0EFF47FAE6675AFA55FAD2" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[1285,1436,1370,1393]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FAE6675AFA55FAD2" box="[1285,1436,1370,1393]" italics="true" pageId="132" pageNumber="133">Titanomessor</emphasis>
</taxonomicName>
could be closely related to 
<taxonomicName id="CC884D7CFF0EFF47FE34677DFDA8FA35" authorityName="Eichler" authorityYear="1952" box="[471,609,1405,1430]" class="Insecta" family="Philopteridae" genus="Turdinirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF0EFF47FE34677DFDA8FA35" box="[471,609,1405,1430]" italics="true" pageId="132" pageNumber="133">Turdinirmus</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF0EFF47FD97677DFD13FA35" box="[628,730,1405,1430]" captionStart="FIGURES 184 – 188" captionStartId="121.[151,264,1915,1937]" captionTargetBox="[198,1349,204,1875]" captionTargetId="figure@121.[192,1395,193,1894]" captionTargetPageId="121" captionText="FIGURES 184 – 188. Turdinirmus merulensis (Denny, 1842) ex Turdus merula: 184, male head, dorsal and ventral views. 185, male genitalia, dorsal view. 186, male mesosome, ventral view. 187, male paramere, dorsal view. 188, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883260/files/figure.png" pageId="132" pageNumber="133">Fig. 186</figureCitation>
). Abdominal chaetotaxy is identical between the two genera (
<tableCitation id="460A0344FF0EFF47FF7D67A1FF31FA19" box="[158,248,1441,1466]" captionStart="TABLE 2" captionStartId="30.[151,239,796,819]" captionText="TABLE 2. Summary of abdominal chaetotaxy of segments II – VIII for the Brueelia - complex, including both genera and subgenera. In genera where chaetotaxy differs between species group (marked with *), only that of the type species is given here, and more detailed accounts are given separately under the respective genus entries and in Tables 3 – 12. Trichoid setae of segment VIII are present in all genera, and are not listed. Abbreviations: ps = paratergal seta; aps = accessory post-spiracular seta; psps = principal post-spiracular seta; tps = tergal posterior seta; ss = sutural seta; sts = sternal seta." pageId="132" pageNumber="133">Table 2</tableCitation>
), except for the presence of 
<taxonomicName id="CC884D7CFF0EFF47FDA667A2FDAFFA1A" box="[581,614,1442,1465]" class="Insecta" family="Philopteridae" genus="Brueelia" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="subSpecies" species="brachythorax" subSpecies="tps">
<emphasis id="39FCEAEDFF0EFF47FDA667A2FDAFFA1A" box="[581,614,1442,1465]" italics="true" pageId="132" pageNumber="133">tps</emphasis>
</taxonomicName>
on male tergopleurites VII–VIII in 
<taxonomicName id="CC884D7CFF0EFF47FBE467A2FB57FA1A" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[1031,1182,1442,1465]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FBE467A2FB57FA1A" box="[1031,1182,1442,1465]" italics="true" pageId="132" pageNumber="133">Titanomessor</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF0EFF47FB5367A1FADCFA19" box="[1200,1301,1441,1466]" captionStart="FIGURES 210 – 211" captionStartId="134.[151,264,1837,1859]" captionTargetBox="[193,1373,229,1811]" captionTargetId="figure@134.[151,1436,193,1815]" captionTargetPageId="134" captionText="FIGURES 210 – 211. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 210, male habitus, dorsal and ventral views. 211, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883272/files/figure.png" pageId="132" pageNumber="133">Fig. 210</figureCitation>
), which are absent in all 
<taxonomicName id="CC884D7CFF0EFF47FEC567C5FE66FA7D" authorityName="Eichler" authorityYear="1952" box="[294,431,1477,1502]" class="Insecta" family="Philopteridae" genus="Turdinirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF0EFF47FEC567C5FE66FA7D" box="[294,431,1477,1502]" italics="true" pageId="132" pageNumber="133">Turdinirmus</emphasis>
</taxonomicName>
. The marginal thickening of the mesosomal lobes of both genera do not deviate from the margin (
<figureCitation id="93B32A7AFF0EFF47FF1B67E9FEA8F9A1" box="[248,353,1513,1538]" captionStart="FIGURES 184 – 188" captionStartId="121.[151,264,1915,1937]" captionTargetBox="[198,1349,204,1875]" captionTargetId="figure@121.[192,1395,193,1894]" captionTargetPageId="121" captionText="FIGURES 184 – 188. Turdinirmus merulensis (Denny, 1842) ex Turdus merula: 184, male head, dorsal and ventral views. 185, male genitalia, dorsal view. 186, male mesosome, ventral view. 187, male paramere, dorsal view. 188, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883260/files/figure.png" pageId="132" pageNumber="133">Figs 186</figureCitation>
, 
<figureCitation id="93B32A7AFF0EFF47FE9267EAFE56F9A1" box="[369,415,1514,1538]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="132" pageNumber="133">214</figureCitation>
), and in both genera the gonopore consists of a pair of convergent ventral thickenings. However, parameral shape differs between these two genera, and while 
<emphasis id="39FCEAEDFF0EFF47FC59640EFC20F985" box="[954,1001,1550,1574]" italics="true" pageId="132" pageNumber="133">pst1</emphasis>
is present in 
<taxonomicName id="CC884D7CFF0EFF47FB9D640DFACEF985" authorityName="Eichler" authorityYear="1952" box="[1150,1287,1549,1574]" class="Insecta" family="Philopteridae" genus="Turdinirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF0EFF47FB9D640DFACEF985" box="[1150,1287,1549,1574]" italics="true" pageId="132" pageNumber="133">Turdinirmus</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF0EFF47FAF4640DFAB1F985" box="[1303,1400,1549,1574]" captionStart="FIGURES 184 – 188" captionStartId="121.[151,264,1915,1937]" captionTargetBox="[198,1349,204,1875]" captionTargetId="figure@121.[192,1395,193,1894]" captionTargetPageId="121" captionText="FIGURES 184 – 188. Turdinirmus merulensis (Denny, 1842) ex Turdus merula: 184, male head, dorsal and ventral views. 185, male genitalia, dorsal view. 186, male mesosome, ventral view. 187, male paramere, dorsal view. 188, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883260/files/figure.png" pageId="132" pageNumber="133">Fig. 186</figureCitation>
), it is absent in 
<taxonomicName id="CC884D7CFF0EFF47FEC36432FE7EF9EA" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[288,439,1586,1609]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FEC36432FE7EF9EA" box="[288,439,1586,1609]" italics="true" pageId="132" pageNumber="133">Titanomessor</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF0EFF47FE246431FDE2F9E9" box="[455,555,1585,1610]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="132" pageNumber="133">Fig. 214</figureCitation>
). Preantennal structure also differs between the two, as the marginal carina is interrupted at least submedianly in 
<taxonomicName id="CC884D7CFF0EFF47FDD66455FD77F9CD" authorityName="Eichler" authorityYear="1952" box="[565,702,1621,1646]" class="Insecta" family="Philopteridae" genus="Turdinirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF0EFF47FDD66455FD77F9CD" box="[565,702,1621,1646]" italics="true" pageId="132" pageNumber="133">Turdinirmus</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF0EFF47FD316455FCF1F9CD" box="[722,824,1621,1646]" captionStart="FIGURES 184 – 188" captionStartId="121.[151,264,1915,1937]" captionTargetBox="[198,1349,204,1875]" captionTargetId="figure@121.[192,1395,193,1894]" captionTargetPageId="121" captionText="FIGURES 184 – 188. Turdinirmus merulensis (Denny, 1842) ex Turdus merula: 184, male head, dorsal and ventral views. 185, male genitalia, dorsal view. 186, male mesosome, ventral view. 187, male paramere, dorsal view. 188, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883260/files/figure.png" pageId="132" pageNumber="133">Fig. 184</figureCitation>
), but not in 
<taxonomicName id="CC884D7CFF0EFF47FC2D6456FBACF9CE" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[974,1125,1622,1645]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FC2D6456FBACF9CE" box="[974,1125,1622,1645]" italics="true" pageId="132" pageNumber="133">Titanomessor</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF0EFF47FB9A6455FB16F9CD" box="[1145,1247,1621,1646]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="132" pageNumber="133">Fig. 212</figureCitation>
), and the dorsal preantennal suture is structured differently in the two genera.
</paragraph>
<paragraph id="0B3736FFFF0EFF47FF24649EFAECF915" blockId="132.[151,1437,469,2006]" box="[199,1317,1693,1718]" pageId="132" pageNumber="133">
For differences between 
<taxonomicName id="CC884D7CFF0EFF47FE3F649EFDBAF916" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[476,627,1694,1717]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0EFF47FE3F649EFDBAF916" box="[476,627,1694,1717]" italics="true" pageId="132" pageNumber="133">Titanomessor</emphasis>
</taxonomicName>
and 
<taxonomicName id="CC884D7CFF0EFF47FD4A649DFC8FF915" authorityName="Zlotorzycka" authorityYear="1964" box="[681,838,1693,1718]" class="Insecta" family="Philopteridae" genus="Maculinirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF0EFF47FD4A649DFC8FF915" box="[681,838,1693,1718]" italics="true" pageId="132" pageNumber="133">Maculinirmus</emphasis>
</taxonomicName>
, see the diagnosis of 
<taxonomicName id="CC884D7CFF0EFF47FBD4649DFB1DF915" authorityName="Zlotorzycka" authorityYear="1964" box="[1079,1236,1693,1718]" class="Insecta" family="Philopteridae" genus="Maculinirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF0EFF47FBD4649DFB1DF915" box="[1079,1236,1693,1718]" italics="true" pageId="132" pageNumber="133">Maculinirmus</emphasis>
</taxonomicName>
above.
</paragraph>
</subSubSection>
<subSubSection id="43926574FF0EFF46FF2464C1FB01FCDC" lastPageId="133" lastPageNumber="134" pageId="132" pageNumber="133" type="description">
<paragraph id="0B3736FFFF0EFF46FF2464C1FF25FF5B" blockId="132.[151,1437,469,2006]" lastBlockId="133.[151,1437,151,1544]" lastPageId="133" lastPageNumber="134" pageId="132" pageNumber="133">
<emphasis id="39FCEAEDFF0EFF47FF2464C1FE14F97A" bold="true" box="[199,477,1729,1754]" pageId="132" pageNumber="133">
Description. 
<emphasis id="39FCEAEDFF0EFF47FE8364C1FE14F97A" bold="true" box="[352,477,1729,1754]" italics="true" pageId="132" pageNumber="133">Both sexes</emphasis>
</emphasis>
. Head rounded pentagonal, very broad (
<figureCitation id="93B32A7AFF0EFF47FC4464C1FBC2F979" box="[935,1035,1729,1754]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="132" pageNumber="133">Fig. 212</figureCitation>
). Marginal carina broad, narrowing anteriorly, not interrupted, but displaced posteriorly and dorsally at osculum. Frons hyaline. Ventral carinae broad, diffuse anterior to pulvinus and not clearly continuous with marginal carina. Dorsal preantennal suture transversal, seemingly reaching 
<emphasis id="39FCEAEDFF0EFF47FE99652DFE7BF8E5" box="[378,434,1837,1862]" italics="true" pageId="132" pageNumber="133">dsms</emphasis>
and maybe 
<emphasis id="39FCEAEDFF0EFF47FDDF652DFDAAF8E5" box="[572,611,1837,1862]" italics="true" pageId="132" pageNumber="133">ads</emphasis>
, but this cannot be established with certainty due to the state of examine dmaterial. Suture does not reach lateral margins of head. Dorsal anterior plate connected to main head plate laterally. Ventral anterior plate absent. Head chaetotaxy as in 
<figureCitation id="93B32A7AFF0EFF47FCA06575FC6DF82D" box="[835,932,1909,1934]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="132" pageNumber="133">Fig. 212</figureCitation>
; 
<emphasis id="39FCEAEDFF0EFF47FC4C6576FC1CF82D" box="[943,981,1910,1934]" italics="true" pageId="132" pageNumber="133">as3</emphasis>
present; 
<emphasis id="39FCEAEDFF0EFF47FBDF6576FB51F82E" box="[1084,1176,1910,1933]" italics="true" pageId="132" pageNumber="133">pns, pts,</emphasis>
and 
<emphasis id="39FCEAEDFF0EFF47FB2D6576FACAF82D" box="[1230,1283,1910,1934]" italics="true" pageId="132" pageNumber="133">s1–4</emphasis>
not visible in the material examined; 
<emphasis id="39FCEAEDFF0EFF47FE4A659AFE19F812" box="[425,464,1946,1969]" italics="true" pageId="132" pageNumber="133">pos</emphasis>
posterior to eye, ventral. Ventral anterior plate absent. Coni short, blunt. Antennae monomorphic. Marginal and occipital carinae not visible. Marginal temporal carina very narrow. Uniquely in complex, 
<taxonomicName id="CC884D7CFF0FFF46FEE46299FEE7FF13" box="[263,302,153,176]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="mts">
<emphasis id="39FCEAEDFF0FFF46FEE46299FEE7FF13" box="[263,302,153,176]" italics="true" pageId="133" pageNumber="134">mts</emphasis>
</taxonomicName>
are sexually dimorphic: in males 
<emphasis id="39FCEAEDFF0FFF46FD576298FCCCFF13" box="[692,773,152,176]" italics="true" pageId="133" pageNumber="134">
<taxonomicName id="CC884D7CFF0FFF46FD576298FD13FF13" box="[692,730,152,176]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
3–5
</emphasis>
long, of roughly equal length, in females 
<emphasis id="39FCEAEDFF0FFF46FB0A6298FAF3FF13" box="[1257,1338,152,176]" italics="true" pageId="133" pageNumber="134">
<taxonomicName id="CC884D7CFF0FFF46FB0A6298FAC6FF13" box="[1257,1295,152,176]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
3–4
</emphasis>
long, of roughly equal length, and 
<emphasis id="39FCEAEDFF0FFF46FE2262BDFE3CFF77" box="[449,501,189,212]" italics="true" pageId="133" pageNumber="134">
<taxonomicName id="CC884D7CFF0FFF46FE2262BDFE21FF77" box="[449,488,189,212]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
5
</emphasis>
thorn-like; in both sexes 
<emphasis id="39FCEAEDFF0FFF46FCFA62BCFCA3FF77" box="[793,874,188,212]" italics="true" pageId="133" pageNumber="134">
<taxonomicName id="CC884D7CFF0FFF46FCFA62BCFCF6FF77" box="[793,831,188,212]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
4–5
</emphasis>
often clearly dorsal. Gular plate broad, diamondshaped.
</paragraph>
<paragraph id="0B3736FFFF0FFF46FF246304FBFEFE2B" blockId="133.[151,1437,151,1544]" pageId="133" pageNumber="134">
Prothorax (
<figureCitation id="93B32A7AFF0FFF46FEA66304FE28FEBF" box="[325,481,260,285]" captionStart="FIGURES 210 – 211" captionStartId="134.[151,264,1837,1859]" captionTargetBox="[193,1373,229,1811]" captionTargetId="figure@134.[151,1436,193,1815]" captionTargetPageId="134" captionText="FIGURES 210 – 211. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 210, male habitus, dorsal and ventral views. 211, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883272/files/figure.png" pageId="133" pageNumber="134">Figs 210–211</figureCitation>
) rectangular; 
<emphasis id="39FCEAEDFF0FFF46FD9D6305FD79FEBF" box="[638,688,261,284]" italics="true" pageId="133" pageNumber="134">ppss</emphasis>
on postero-lateral corner. Proepimera and metepisterna with blunt median ends. Pterothorax roughly trapezoidal; lateral margins divergent, posterior margin gently rounded; 
<emphasis id="39FCEAEDFF0FFF46FA8B6329FA52FEE3" box="[1384,1435,297,320]" italics="true" pageId="133" pageNumber="134">mms</emphasis>
widely separated medianly. Meso- and metasterna not fused, 1 seta on postero-lateral corner on each side of each plate. Leg chaetotaxy as in 
<figureCitation id="93B32A7AFF0FFF46FE2A636FFDD2FE2B" box="[457,539,367,392]" captionStart="FIGURES 25" captionStartId="14.[151,264,1012,1034]" captionTargetBox="[173,1428,205,974]" captionTargetId="figure@14.[151,1436,193,991]" captionTargetPageId="14" captionText="FIGURES 25. Dorso-ventral views of coxae I – III, trochanters I – III, and femora I – III, showing terminology of the leg setae used here. Other parts of the thorax removed for clarity. Setae situated marginally are illustrated and named on both dorsal (lefthand) and ventral (right-hand) sides. Terminology for individual seta as used in the text obtained by combining the leg segment (larger font) and the setal name (smaller font). Abbreviations: a 1 – 5, anterior setae 1 – 5; cI – III, coxae I – III; d 1 – 3, dorsal setae 1 – 3; dm 1 – 4, distal marginal setae 1 – 4; fI – III, femur I – III; p 1 – 3, posterior setae 1 – 3; s 1 – 6, sensilla 1 – 6; tI – III, trochanter I – III; v 1 – 4, ventral setae 1 – 4." httpUri="https://zenodo.org/record/883171/files/figure.png" pageId="133" pageNumber="134">Fig. 25</figureCitation>
, except 
<emphasis id="39FCEAEDFF0FFF46FD956370FD39FE2B" box="[630,752,368,392]" italics="true" pageId="133" pageNumber="134">fI-p2, fI-v4</emphasis>
absent; 
<emphasis id="39FCEAEDFF0FFF46FCAE6370FC2DFE2B" box="[845,996,368,392]" italics="true" pageId="133" pageNumber="134">fII-a2, fIII-a2</emphasis>
dorsal.
</paragraph>
<paragraph id="0B3736FFFF0FFF46FF246394FACDFD04" blockId="133.[151,1437,151,1544]" pageId="133" pageNumber="134">
Abdomen (
<figureCitation id="93B32A7AFF0FFF46FEAC6394FE3CFE0F" box="[335,501,404,429]" captionStart="FIGURES 210 – 211" captionStartId="134.[151,264,1837,1859]" captionTargetBox="[193,1373,229,1811]" captionTargetId="figure@134.[151,1436,193,1815]" captionTargetPageId="134" captionText="FIGURES 210 – 211. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 210, male habitus, dorsal and ventral views. 211, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883272/files/figure.png" pageId="133" pageNumber="134">Figs 210–211</figureCitation>
) oval. Distal end of abdomen bulging in male. Tergopleurites rectangular; tergopleurites II–IX+X in male and tergopleurites II–VIII in female narrowly separated medianly. Female tergopleurites IX+X and XI fused medianly. Sternal plates narrow, lateral margins rounded. Sternal plates II–IV translucent and illustrated approximately. Pleural incrassations moderate to large. Re-entrant heads large, blunt, translucent. Abdominal chaetotaxy as in 
<tableCitation id="460A0344FF0FFF46FD886024FD0AFD9F" box="[619,707,548,573]" captionStart="TABLE 2" captionStartId="30.[151,239,796,819]" captionText="TABLE 2. Summary of abdominal chaetotaxy of segments II – VIII for the Brueelia - complex, including both genera and subgenera. In genera where chaetotaxy differs between species group (marked with *), only that of the type species is given here, and more detailed accounts are given separately under the respective genus entries and in Tables 3 – 12. Trichoid setae of segment VIII are present in all genera, and are not listed. Abbreviations: ps = paratergal seta; aps = accessory post-spiracular seta; psps = principal post-spiracular seta; tps = tergal posterior seta; ss = sutural seta; sts = sternal seta." pageId="133" pageNumber="134">Table 2</tableCitation>
. 
<collectingRegion id="C94CF81DFF0FFF46FD326024FCC3FD9E" box="[721,778,548,573]" country="Maldives" name="Maale" pageId="133" pageNumber="134">Male</collectingRegion>
subgenital plate oblong, narrowing distally, and reaching distal part of abdomen. Female subgenital plate slender, roughly rectangular, narrowing only slightly posteriorly, approaching vulval margin. Vulval margin (
<figureCitation id="93B32A7AFF0FFF46FD67606CFD2DFD26" box="[644,740,620,645]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="133" pageNumber="134">Fig. 216</figureCitation>
) gently rounded, with slender 
<emphasis id="39FCEAEDFF0FFF46FBDE606DFBA0FD27" box="[1085,1129,621,644]" italics="true" pageId="133" pageNumber="134">vms</emphasis>
, thorn-like 
<emphasis id="39FCEAEDFF0FFF46FB0E606DFAC6FD27" box="[1261,1295,621,644]" italics="true" pageId="133" pageNumber="134">vss</emphasis>
; slender 
<emphasis id="39FCEAEDFF0FFF46FA95606DFA52FD27" box="[1398,1435,621,644]" italics="true" pageId="133" pageNumber="134">vos</emphasis>
follow lateral margins of subgenital plate, 
<emphasis id="39FCEAEDFF0FFF46FD926091FD5FFD0B" box="[625,662,657,680]" italics="true" pageId="133" pageNumber="134">vos</emphasis>
typically in two parallel rows; distal 
<emphasis id="39FCEAEDFF0FFF46FBDB6091FB94FD0B" box="[1080,1117,657,680]" italics="true" pageId="133" pageNumber="134">vos</emphasis>
median to 
<emphasis id="39FCEAEDFF0FFF46FB386091FB34FD0B" box="[1243,1277,657,680]" italics="true" pageId="133" pageNumber="134">vss</emphasis>
.
</paragraph>
<paragraph id="0B3736FFFF0FFF46FF2460B4FB01FCDC" blockId="133.[151,1437,151,1544]" pageId="133" pageNumber="134">
<collectingRegion id="C94CF81DFF0FFF46FF2460B4FEC8FD6E" box="[199,257,692,717]" country="Maldives" name="Maale" pageId="133" pageNumber="134">Male</collectingRegion>
genitalia (
<figureCitation id="93B32A7AFF0FFF46FE9D60B4FDD0FD6E" box="[382,537,692,717]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="133" pageNumber="134">Fig. 213–215</figureCitation>
) unique within 
<taxonomicName id="CC884D7CFF0FFF46FD3060B4FCFAFD6E" authorityName="Keler" authorityYear="1936" box="[723,819,692,717]" class="Insecta" family="Philopteridae" genus="Brueelia" kingdom="Animalia" order="Phthiraptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF0FFF46FD3060B4FCFAFD6E" box="[723,819,692,717]" italics="true" pageId="133" pageNumber="134">Brueelia</emphasis>
</taxonomicName>
-complex. Basal apodeme (
<figureCitation id="93B32A7AFF0FFF46FB8C60B4FB1BFD6F" box="[1135,1234,692,717]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="133" pageNumber="134">Fig. 213</figureCitation>
) slender, roughly rectangular, anterior margin rounded. Proximal mesosome (
<figureCitation id="93B32A7AFF0FFF46FCCC60D7FC47FD53" box="[815,910,727,752]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="133" pageNumber="134">Fig. 214</figureCitation>
) broadly flattened. Gonopore open distally and proximally, as broad convergent thickenings; protruding distally to lobes. Mesosomal lobes angular, in some specimens slightly rugose distally; 2 
<emphasis id="39FCEAEDFF0FFF46FDB26121FD42FC9B" box="[593,651,801,824]" italics="true" pageId="133" pageNumber="134">pmes</emphasis>
sensilla visible on mesosome lateral to gonopore and near lateral margins. Parameral heads (
<figureCitation id="93B32A7AFF0FFF46FE366144FDF0FCFE" box="[469,569,836,861]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="133" pageNumber="134">Fig. 215</figureCitation>
) blunt. Parameral blades as fleshy lobes, converging distal to mesosome. Parameral blades rounded distally; 
<emphasis id="39FCEAEDFF0FFF46FDC16168FD99FC23" box="[546,592,872,896]" italics="true" pageId="133" pageNumber="134">pst1</emphasis>
not visible; 
<emphasis id="39FCEAEDFF0FFF46FD3F6168FCC2FC23" box="[732,779,872,896]" italics="true" pageId="133" pageNumber="134">pst2</emphasis>
as microseta on distal end of paramere.
</paragraph>
</subSubSection>
<subSubSection id="43926574FF0FFF46FF24618CFDA6FA93" pageId="133" pageNumber="134" type="biology_ecology">
<paragraph id="0B3736FFFF0FFF46FF24618CFF04FB4B" blockId="133.[151,1437,151,1544]" pageId="133" pageNumber="134">
<emphasis id="39FCEAEDFF0FFF46FF24618CFE6BFC06" bold="true" box="[199,418,908,933]" pageId="133" pageNumber="134">Host distribution.</emphasis>
Presently know only from the type host, 
<taxonomicName id="CC884D7CFF0FFF46FC4C618DFB08FC06" authorityName="Bush" authorityYear="2017" baseAuthorityName="Cretzschmar" baseAuthorityYear="1829" box="[943,1217,908,933]" class="Aves" family="Malaconotidae" genus="Laniarius" kingdom="Animalia" order="Passeriformes" pageId="133" pageNumber="134" phylum="Chordata" rank="species" species="erythrogaster">
<emphasis id="39FCEAEDFF0FFF46FC4C618DFB08FC06" box="[943,1217,908,933]" italics="true" pageId="133" pageNumber="134">Laniarius erythrogaster</emphasis>
</taxonomicName>
, belonging to the 
<taxonomicName id="CC884D7CFF0FFF46FF7461AFFE88FC6B" authorityName="Swainson" authorityYear="1824" box="[151,321,943,968]" class="Aves" family="Malaconotidae" kingdom="Animalia" order="Passeriformes" pageId="133" pageNumber="134" phylum="Chordata" rank="family">Malaconotidae</taxonomicName>
. We have examined lice from the following malaconotid hosts: 
<taxonomicName id="CC884D7CFF0FFF46FBB861B1FE8AFC4E" authority="Hartlaub, 1863" authorityName="Hartlaub" authorityYear="1863" class="Aves" family="Malaconotidae" genus="Dryoscopus" kingdom="Animalia" order="Passeriformes" pageId="133" pageNumber="134" phylum="Chordata" rank="subSpecies" species="cubla" subSpecies="hamatus">
<emphasis id="39FCEAEDFF0FFF46FBB861B1FA55FC6B" box="[1115,1436,943,968]" italics="true" pageId="133" pageNumber="134">Dryoscopus cubla hamatus</emphasis>
Hartlaub, 1863
</taxonomicName>
, 
<taxonomicName id="CC884D7CFF0FFF46FEAC61D5FC08FC4F" authority="gambensis (Lichtenstein, 1832)" authorityName="Lichtenstein" authorityYear="1832" box="[335,961,980,1005]" class="Aves" family="Malaconotidae" genus="Dryoscopus" kingdom="Animalia" order="Passeriformes" pageId="133" pageNumber="134" phylum="Chordata" rank="subSpecies" species="gambensis" subSpecies="gambensis">
<emphasis id="39FCEAEDFF0FFF46FEAC61D5FD18FC4E" box="[335,721,980,1005]" italics="true" pageId="133" pageNumber="134">Dryoscopus gambensis gambensis</emphasis>
(Lichtenstein, 1832)
</taxonomicName>
, 
<taxonomicName id="CC884D7CFF0FFF46FC2E61D5FA5EFC4F" authority="major (Hartlaub, 1848)" authorityName="Hartlaub" authorityYear="1848" box="[973,1431,980,1005]" class="Aves" family="Malaconotidae" genus="Laniarius" kingdom="Animalia" order="Passeriformes" pageId="133" pageNumber="134" phylum="Chordata" rank="subSpecies" species="major" subSpecies="major">
<emphasis id="39FCEAEDFF0FFF46FC2E61D5FB19FC4F" box="[973,1232,981,1004]" italics="true" pageId="133" pageNumber="134">Laniarius major major</emphasis>
(Hartlaub, 1848)
</taxonomicName>
, 
<taxonomicName id="CC884D7CFF0FFF46FF7461F9FCBAFBB3" authority="(Fischer and Reichenow, 1880)" baseAuthorityName="Fischer and Reichenow" baseAuthorityYear="1880" box="[151,883,1015,1040]" class="Aves" family="Malaconotidae" genus="Laniarius" kingdom="Animalia" order="Passeriformes" pageId="133" pageNumber="134" phylum="Chordata" rank="subSpecies" species="major" subSpecies="mossambicus">
<emphasis id="39FCEAEDFF0FFF46FF7461F9FE3CFBB3" box="[151,501,1015,1040]" italics="true" pageId="133" pageNumber="134">Laniarius major mossambicus</emphasis>
(Fischer and Reichenow, 1880)
</taxonomicName>
, 
<taxonomicName id="CC884D7CFF0FFF46FC6761F7FF1BFB97" authority="Hartlaub, 1844" authorityName="Hartlaub" authorityYear="1844" class="Aves" family="Malaconotidae" genus="Malaconotus" kingdom="Animalia" order="Passeriformes" pageId="133" pageNumber="134" phylum="Chordata" rank="subSpecies" species="blanchoti" subSpecies="hypopyrrhus">
<emphasis id="39FCEAEDFF0FFF46FC6761F7FAEFFBB3" box="[900,1318,1015,1040]" italics="true" pageId="133" pageNumber="134">Malaconotus blanchoti hypopyrrhus</emphasis>
Hartlaub, 1844
</taxonomicName>
, 
<taxonomicName id="CC884D7CFF0FFF46FF3E661CFD43FB97" authority="(Lesson, 1831)" baseAuthorityName="Lesson" baseAuthorityYear="1831" box="[221,650,1052,1077]" class="Aves" family="Malaconotidae" genus="Malaconotus" kingdom="Animalia" order="Passeriformes" pageId="133" pageNumber="134" phylum="Chordata" rank="species" species="cruentus">
<emphasis id="39FCEAEDFF0FFF46FF3E661CFE1EFB97" box="[221,471,1052,1077]" italics="true" pageId="133" pageNumber="134">Malaconotus cruentus</emphasis>
(Lesson, 1831)
</taxonomicName>
, 
<taxonomicName id="CC884D7CFF0FFF46FD76661CFB76FB96" authority="blanchoti Stephens, 1826" authorityName="Stephens" authorityYear="1826" box="[661,1215,1052,1077]" class="Aves" family="Malaconotidae" genus="Malaconotus" kingdom="Animalia" order="Passeriformes" pageId="133" pageNumber="134" phylum="Chordata" rank="subSpecies" species="blanchoti" subSpecies="blanchoti">
<emphasis id="39FCEAEDFF0FFF46FD76661CFBC0FB96" box="[661,1033,1052,1077]" italics="true" pageId="133" pageNumber="134">Malaconotus blanchoti blanchoti</emphasis>
Stephens, 1826
</taxonomicName>
, 
<taxonomicName id="CC884D7CFF0FFF46FB29661CFE1CFBFB" authority="(Oberholser, 1906)" baseAuthorityName="Oberholser" baseAuthorityYear="1906" class="Aves" family="Malaconotidae" genus="Tchagra" kingdom="Animalia" order="Passeriformes" pageId="133" pageNumber="134" phylum="Chordata" rank="subSpecies" species="senegalus" subSpecies="armenus">
<emphasis id="39FCEAEDFF0FFF46FB29661CFF3EFBFB" italics="true" pageId="133" pageNumber="134">Tchagra senegalus armenus</emphasis>
(Oberholser, 1906)
</taxonomicName>
, 
<taxonomicName id="CC884D7CFF0FFF46FE02663FFBC0FBFB" authority="(Temminck, 1840)" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[481,1033,1087,1112]" class="Aves" family="Malaconotidae" genus="Tchagra" kingdom="Animalia" order="Passeriformes" pageId="133" pageNumber="134" phylum="Chordata" rank="subSpecies" species="senegalus" subSpecies="cucullatus">
<emphasis id="39FCEAEDFF0FFF46FE02663FFCE7FBFB" box="[481,814,1087,1112]" italics="true" pageId="133" pageNumber="134">Tchagra senegalus cucullatus</emphasis>
(Temminck, 1840)
</taxonomicName>
, 
<taxonomicName id="CC884D7CFF0FFF46FBF7663FFE88FBDF" authority="(Bangs, 1931)" baseAuthorityName="Bangs" baseAuthorityYear="1931" class="Aves" family="Malaconotidae" genus="Telophorus" kingdom="Animalia" order="Passeriformes" pageId="133" pageNumber="134" phylum="Chordata" rank="subSpecies" species="nigrifrons" subSpecies="sandgroundi">
<emphasis id="39FCEAEDFF0FFF46FBF7663FFA52FBFB" box="[1044,1435,1087,1112]" italics="true" pageId="133" pageNumber="134">Telophorus nigrifrons sandgroundi</emphasis>
(Bangs, 1931)
</taxonomicName>
, 
<taxonomicName id="CC884D7CFF0FFF46FEB76664FC0AFBDE" authority="(Smith, A., 1936)" authorityName="Smith, A." authorityYear="1936" box="[340,963,1124,1149]" class="Aves" family="Malaconotidae" genus="Telophorus" kingdom="Animalia" order="Passeriformes" pageId="133" pageNumber="134" phylum="Chordata" rank="subSpecies" species="sulfureopectus" subSpecies="similis">
<emphasis id="39FCEAEDFF0FFF46FEB76664FD12FBDE" box="[340,731,1124,1149]" italics="true" pageId="133" pageNumber="134">Telophorus sulfureopectus similis</emphasis>
(Smith, A., 1936)
</taxonomicName>
. However, all this material belongs to 
<emphasis id="39FCEAEDFF0FFF46FF746687FE8CFB03" box="[151,325,1159,1184]" italics="true" pageId="133" pageNumber="134">Guimaraesiella</emphasis>
rather than 
<taxonomicName id="CC884D7CFF0FFF46FE2E6689FDAAFB03" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[461,611,1161,1184]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="133" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0FFF46FE2E6689FDAAFB03" box="[461,611,1161,1184]" italics="true" pageId="133" pageNumber="134">Titanomessor</emphasis>
</taxonomicName>
, making the occurrence of 
<taxonomicName id="CC884D7CFF0FFF46FC746689FBD8FB03" authorityName="Bush" authorityYear="2017" box="[919,1041,1159,1184]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="sexloba">
<emphasis id="39FCEAEDFF0FFF46FC746689FC62FB03" box="[919,939,1161,1184]" italics="true" pageId="133" pageNumber="134">Ti</emphasis>
. 
<emphasis id="39FCEAEDFF0FFF46FC586687FBD8FB03" box="[955,1041,1159,1184]" italics="true" pageId="133" pageNumber="134">sexloba</emphasis>
</taxonomicName>
unique within the host family. The material we have examined is from two different collection events, suggesting that 
<taxonomicName id="CC884D7CFF0FFF46FBA966ADFAC9FB66" authorityName="Bush" authorityYear="2017" baseAuthorityName="Cretzschmar" baseAuthorityYear="1829" box="[1098,1280,1196,1221]" class="Aves" family="Malaconotidae" genus="Laniarius" kingdom="Animalia" order="Passeriformes" pageId="133" pageNumber="134" phylum="Chordata" rank="species" species="erythrogaster">
<emphasis id="39FCEAEDFF0FFF46FBA966ADFB90FB67" box="[1098,1113,1197,1220]" italics="true" pageId="133" pageNumber="134">L</emphasis>
. 
<emphasis id="39FCEAEDFF0FFF46FB8A66ACFAC9FB66" box="[1129,1280,1196,1221]" italics="true" pageId="133" pageNumber="134">erythrogaster</emphasis>
</taxonomicName>
is the correct host.
</paragraph>
<paragraph id="0B3736FFFF0FFF46FF2466F4FDA6FA93" blockId="133.[151,1437,151,1544]" pageId="133" pageNumber="134">
<emphasis id="39FCEAEDFF0FFF46FF2466F4FE77FAAE" bold="true" box="[199,446,1268,1293]" pageId="133" pageNumber="134">Geographical range.</emphasis>
Known only from Uganda, but the host genus 
<taxonomicName id="CC884D7CFF0FFF46FC0F66F4FB9FFAAE" authorityName="Vieillot" authorityYear="1816" box="[1004,1110,1268,1293]" class="Aves" family="Malaconotidae" genus="Laniarius" kingdom="Animalia" order="Passeriformes" pageId="133" pageNumber="134" phylum="Chordata" rank="genus">Laniarius</taxonomicName>
occurs across most of Sub- Saharan Africa (
<bibRefCitation id="6F194B0EFF0FFF46FEAD6717FDAAFA93" author="Harris" box="[334,611,1303,1328]" pageId="133" pageNumber="134" refString="Harris, T. &amp; Franklin, K. (2000) Shrikes and Bush-shrikes. Christopher Helm (Publishers) Ltd, London. 392 pp." type="book" year="2000">Harris &amp; Franklin, 2000</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="43926574FF0FFF46FF24673CFD5FFADB" pageId="133" pageNumber="134" type="etymology">
<paragraph id="0B3736FFFF0FFF46FF24673CFD5FFADB" blockId="133.[151,1437,151,1544]" pageId="133" pageNumber="134">
<emphasis id="39FCEAEDFF0FFF46FF24673CFE87FAF6" bold="true" box="[199,334,1340,1365]" pageId="133" pageNumber="134">Etymology.</emphasis>
The genus name derived from Greek “ 
<emphasis id="39FCEAEDFF0FFF46FCE9673DFCF4FAF7" box="[778,829,1341,1364]" italics="true" pageId="133" pageNumber="134">titan</emphasis>
” for the large size of these lice, combined with Latin “ 
<emphasis id="39FCEAEDFF0FFF46FF406761FF3BFADB" box="[163,242,1377,1400]" italics="true" pageId="133" pageNumber="134">messor</emphasis>
” for “harvester”. Gender: masculine.
</paragraph>
</subSubSection>
<subSubSection id="43926574FF0FFF46FF246784FE12F9D7" pageId="133" pageNumber="134" type="discussion">
<paragraph id="0B3736FFFF0FFF46FF246784FB3CF9A4" blockId="133.[151,1437,151,1544]" pageId="133" pageNumber="134">
<emphasis id="39FCEAEDFF0FFF46FF246784FEF2FA3E" bold="true" box="[199,315,1412,1437]" pageId="133" pageNumber="134">Remarks.</emphasis>
<taxonomicName id="CC884D7CFF0FFF46FEA56785FDF7FA3E" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[326,574,1412,1437]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="sexloba" status="sp. nov.">
<emphasis id="39FCEAEDFF0FFF46FEA56785FDF7FA3E" box="[326,574,1412,1437]" italics="true" pageId="133" pageNumber="134">Titanomessor sexloba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF0FFF46FDA96784FD47FA3E" box="[586,654,1412,1437]" pageId="133" pageNumber="134" rank="species">
<emphasis id="39FCEAEDFF0FFF46FDA96784FD47FA3E" bold="true" box="[586,654,1412,1437]" pageId="133" pageNumber="134">n. sp.</emphasis>
</taxonomicNameLabel>
was not included in the phylogeny of Bush 
<emphasis id="39FCEAEDFF0FFF46FB446785FB14FA3E" box="[1191,1245,1412,1437]" italics="true" pageId="133" pageNumber="134">et al</emphasis>
. (2016), and its affinities within the 
<taxonomicName id="CC884D7CFF0FFF46FE7767A7FE3DFA63" authorityName="Keler" authorityYear="1936" box="[404,500,1447,1472]" class="Insecta" family="Philopteridae" genus="Brueelia" kingdom="Animalia" order="Phthiraptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF0FFF46FE7767A7FE3DFA63" box="[404,500,1447,1472]" italics="true" pageId="133" pageNumber="134">Brueelia</emphasis>
</taxonomicName>
-complex are unknown. The great differences in the male genitalia between 
<emphasis id="39FCEAEDFF0FFF46FF7467CDFEE1FA47" box="[151,296,1485,1508]" italics="true" pageId="133" pageNumber="134">Titanimessor</emphasis>
and all other genera treated here makes the genus hard to place, but abdominal chaetotaxy and overall pigmentation patterns are very similar to those of 
<taxonomicName id="CC884D7CFF0FFF46FD2667EFFCABF9AB" authorityName="Zlotorzycka" authorityYear="1964" box="[709,866,1519,1544]" class="Insecta" family="Philopteridae" genus="Maculinirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF0FFF46FD2667EFFCABF9AB" box="[709,866,1519,1544]" italics="true" pageId="133" pageNumber="134">Maculinirmus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CC884D7CFF0FFF46FC9367EFFC30F9AB" authorityName="Eichler" authorityYear="1952" box="[880,1017,1519,1544]" class="Insecta" family="Philopteridae" genus="Turdinirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF0FFF46FC9367EFFC30F9AB" box="[880,1017,1519,1544]" italics="true" pageId="133" pageNumber="134">Turdinirmus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="CC884D7CFF0FFF46FBD467EFFB27F9AB" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" baseAuthorityName="Capnodella" box="[1079,1262,1519,1544]" class="Insecta" family="Philopteridae" genus="Indoceoplanetes" kingdom="Animalia" order="Phthiraptera" pageId="133" pageNumber="137" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF0FFF46FBD467EFFB27F9AB" box="[1079,1262,1519,1544]" italics="true" pageId="133" pageNumber="134">Indoceoplanetes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="0B3736FFFF0FFF46FF746437FE98F9F3" blockId="133.[151,475,1591,1653]" box="[151,337,1591,1616]" pageId="133" pageNumber="134">Included species</paragraph>
<paragraph id="0B3736FFFF0FFF46FF74645DFE12F9D7" blockId="133.[151,475,1591,1653]" box="[151,475,1628,1653]" pageId="133" pageNumber="134">
<emphasis id="39FCEAEDFF0FFF46FF74645DFE50F9D6" box="[151,409,1628,1653]" italics="true" pageId="133" pageNumber="134">
* 
<taxonomicName id="CC884D7CFF0FFF46FF41645DFE50F9D6" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[162,409,1628,1653]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="sexloba" status="sp. nov.">Titanomessor sexloba</taxonomicName>
</emphasis>
<taxonomicNameLabel id="22CF5796FF0FFF46FE7C645DFE12F9D7" box="[415,475,1629,1652]" pageId="133" pageNumber="134" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>