treatments-xml/data/5D/55/4F/5D554F4F2139FFF5FF46FF633B55FADE.xml
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<document id="E54B0AD41E75A6820275ED1283B461D6" ID-CLB-Dataset="128003" ID-DOI="10.1080/00222933.2022.2084649" ID-GBIF-Dataset="006719bf-f416-4310-9626-29381fd9c26d" ID-ISSN="1464-5262" ID-Zenodo-Dep="7012522" ID-ZooBank="5147CC1B-6144-4CA4-BA63-E39ED0A20E1C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1660992512912" checkinUser="plazi" docAuthor="Pacheco, Thaynara L., Wipfler, Benedict, Monné, Marcela L. &amp; Ahrens, Dirk" docDate="2022" docId="5D554F4F2139FFF5FF46FF633B55FADE" docLanguage="en" docName="JNATHIST.56.9-12.607-705.pdf" docOrigin="Journal of Natural History 56 (9 - 12)" docSource="http://dx.doi.org/10.1080/00222933.2022.2084649" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Symmela capixaba Pacheco, Wipfler, Monne and Ahrens 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="624" masterDocId="A16C37372136FFE4FFBAFFED3933FFA9" masterDocTitle="The genus Symmela Erichson, 1835 (Coleoptera, Scarabaeidae, Sericini): taxonomy and phylogenetic analysis" masterLastPageNumber="705" masterPageNumber="607" pageNumber="622" updateTime="1699372031290" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0ABB6023B61EF2D9C575AA8AC65932ED">The genus Symmela Erichson, 1835 (Coleoptera, Scarabaeidae, Sericini): taxonomy and phylogenetic analysis</mods:title>
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<mods:namePart id="02C426620E589B908E43959636F409A8">Pacheco, Thaynara L.</mods:namePart>
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<mods:namePart id="C312BDB830EC1874ADCC3583A3BED882">Wipfler, Benedict</mods:namePart>
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<mods:namePart id="48C30787630C59A34F20395EC9A22306">Monné, Marcela L.</mods:namePart>
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<mods:namePart id="5C002B2CF9D76C0306EB7FF935F41FF0">Ahrens, Dirk</mods:namePart>
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<mods:title id="3B79685EC6AF4430751375E2E5E808B4">Journal of Natural History</mods:title>
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<mods:date id="42C298063897B2C17A573E133EF59ECF">2022</mods:date>
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<treatment id="5D554F4F2139FFF5FF46FF633B55FADE" ID-DOI="http://doi.org/10.5281/zenodo.7019029" ID-GBIF-Taxon="199728135" ID-Zenodo-Dep="7019029" LSID="urn:lsid:plazi:treatment:5D554F4F2139FFF5FF46FF633B55FADE" httpUri="http://treatment.plazi.org/id/5D554F4F2139FFF5FF46FF633B55FADE" lastPageId="17" lastPageNumber="624" pageId="15" pageNumber="622">
<subSubSection id="9DE6ADD22139FFEBFF46FF633D15FF01" box="[252,1062,142,168]" pageId="15" pageNumber="622" type="nomenclature">
<paragraph id="D543FE592139FFEBFF46FF633D15FF01" blockId="15.[252,1062,142,168]" box="[252,1062,142,168]" pageId="15" pageNumber="622">
<heading id="8E0B49352139FFEBFF46FF633D15FF01" bold="true" box="[252,1062,142,168]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="622" reason="3">
<emphasis id="E788224B2139FFEBFF46FF633D15FF01" bold="true" box="[252,1062,142,168]" pageId="15" pageNumber="622">
<taxonomicName id="12FC85DA2139FFEBFF46FF633A8FFF01" authority="Pacheco, Wipfler, Monne and Ahrens" authorityName="Pacheco, Wipfler, Monne and Ahrens" authorityYear="2022" box="[252,956,142,168]" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="622" phylum="Arthropoda" rank="species" species="capixaba" status="sp. nov.">
<emphasis id="E788224B2139FFEBFF46FF6338D0FF01" bold="true" box="[252,483,142,168]" italics="true" pageId="15" pageNumber="622">Symmela capixaba</emphasis>
Pacheco, Wipfler, Monné and Ahrens
</taxonomicName>
<taxonomicNameLabel id="FCBB9F302139FFEBFC7EFF633D15FF01" box="[964,1062,142,168]" pageId="15" pageNumber="622" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="9DE6ADD22139FFEBFDB6FF283A24FF76" box="[524,791,197,223]" pageId="15" pageNumber="622" type="description">
<paragraph id="D543FE592139FFEBFDB6FF283A24FF76" blockId="15.[524,791,197,223]" box="[524,791,197,223]" pageId="15" pageNumber="622">
(
<figureCitation id="4DC7E2DC2139FFEBFDAEFF283BB3FF76" box="[532,640,197,223]" captionStart="Figure 3" captionStartId="16.[160,225,1662,1685]" captionTargetBox="[273,1043,145,1623]" captionTargetId="figure-3@16.[270,1047,142,1645]" captionTargetPageId="16" captionText="Figure 3. Symmela capixaba sp. nov., holotype ♂. (a) Habitus, dorsal view; (b) habitus, lateral view; (c) aedeagus, lateral view; (d, e) parameres, dorsal view. Scale bars: a, b = 1 mm; ce = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7012530" httpUri="https://zenodo.org/record/7012530/files/figure.png" pageId="15" pageNumber="622">Figures 3</figureCitation>
,
<figureCitation id="4DC7E2DC2139FFEBFD34FF283B9DFF76" box="[654,686,197,223]" captionStart="Figure 29" captionStartId="86.[160,225,1430,1453]" captionTargetBox="[163,1152,142,1413]" captionTargetId="figure-75@86.[163,1153,142,1413]" captionTargetPageId="86" captionText="Figure 29. Distribution map of the Symmela species." figureDoi="http://doi.org/10.5281/zenodo.7012584" httpUri="https://zenodo.org/record/7012584/files/figure.png" pageId="15" pageNumber="622">29</figureCitation>
(a), 30(c))
</paragraph>
</subSubSection>
<subSubSection id="9DE6ADD22139FFEBFF1AFEC13A14FE04" pageId="15" pageNumber="622" type="materials_examined">
<paragraph id="D543FE592139FFEBFF1AFEC1388CFEEF" blockId="15.[160,1156,300,429]" box="[160,447,300,326]" pageId="15" pageNumber="622">
<emphasis id="E788224B2139FFEBFF1AFEC1388CFEEF" bold="true" box="[160,447,300,326]" italics="true" pageId="15" pageNumber="622">Type material examined</emphasis>
</paragraph>
<paragraph id="D543FE592139FFEBFF1AFEA33A14FE04" blockId="15.[160,1156,300,429]" pageId="15" pageNumber="622">
<materialsCitation id="6594F4042139FFEBFF1AFEA33AB2FEC1" ID-GBIF-Occurrence="3892866468" box="[160,897,334,360]" collectionCode="ZMHB" pageId="15" pageNumber="622" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="E788224B2139FFEBFF1AFEA33821FEC1" bold="true" box="[160,274,334,360]" italics="true" pageId="15" pageNumber="622">
<typeStatus id="0A4740FB2139FFEBFF1AFEA3383EFEC1" box="[160,269,334,360]" pageId="15" pageNumber="622" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="C3FA35D02139FFEBFE9BFEA23806FECF" box="[289,309,335,358]" pageId="15" pageNumber="622" type="male"></specimenCount>
Espir. Santo/81 Neotropical Sericini spec. (
<collectionCode id="B3ED669C2139FFEBFC8BFEA23A48FEC1" box="[817,891,335,360]" pageId="15" pageNumber="622">ZMHB</collectionCode>
)
</materialsCitation>
.
<typeStatus id="0A4740FB2139FFEBFC34FEA23ACDFEC1" box="[910,1022,335,360]" pageId="15" pageNumber="622" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="C3FA35D02139FFEBFBB6FEA23D0BFECF" box="[1036,1080,335,360]" pageId="15" pageNumber="622" type="male">1 ♂</specimenCount>
Espir.
<materialsCitation id="6594F4042139FFEBFF1AFE9F3A09FE22" ID-GBIF-Occurrence="3892866486" box="[160,826,370,395]" collectionCode="ZMHB" collectorName="La Plata &amp; Berlin" location="Santo'" pageId="15" pageNumber="622" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<location id="D023A8822139FFEBFF1AFE9F39D5FE22" LSID="urn:lsid:plazi:treatment:5D554F4F2139FFF5FF46FF633B55FADE:D023A8822139FFEBFF1AFE9F39D5FE22" box="[160,230,370,395]" name="Santo'" pageId="15" pageNumber="622">Santo</location>
(
<collectionCode id="B3ED669C2139FFEBFF4DFE9F3872FE22" box="[247,321,370,395]" pageId="15" pageNumber="622">ZMHB</collectionCode>
),
<specimenCount id="C3FA35D02139FFEBFEEFFE9F38B2FE20" box="[341,385,370,395]" pageId="15" pageNumber="622" type="male">1 ♂</specimenCount>
<collectorName id="78099B8F2139FFEBFE35FE9F38DDFE22" box="[399,494,370,395]" pageId="15" pageNumber="622">La Plata</collectorName>
/
<collectorName id="78099B8F2139FFEBFDBBFE9F3B71FE22" box="[513,578,370,395]" pageId="15" pageNumber="622">Berlin</collectorName>
/
<taxonomicName id="12FC85DA2139FFEBFDEFFE9F3A34FE22" authorityName="Erichson" authorityYear="1835" box="[597,775,370,395]" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="622" phylum="Arthropoda" rank="species" species="opaca">
<emphasis id="E788224B2139FFEBFDEFFE9F3A34FE22" box="[597,775,370,395]" italics="true" pageId="15" pageNumber="622">Symmela opaca</emphasis>
</taxonomicName>
det
</materialsCitation>
<materialsCitation id="6594F4042139FFEBFC80FE9F3A6FFE22" ID-GBIF-Occurrence="3892866313" box="[826,860,370,395]" collectionCode="ZMHB" collectorName="La Plata &amp; Berlin" isTailEnumeration="true" location="Santo'" pageId="15" pageNumber="622" specimenCount="1" typeStatus="paratype">. G</materialsCitation>
<materialsCitation id="6594F4042139FFEBFCE1FE9F3AC0FE22" ID-GBIF-Occurrence="3892866443" box="[859,1011,370,395]" location="Frey" pageId="15" pageNumber="622" specimenCount="1" typeStatus="paratype">
.
<location id="D023A8822139FFEBFCD0FE9F3AA9FE22" LSID="urn:lsid:plazi:treatment:5D554F4F2139FFF5FF46FF633B55FADE:D023A8822139FFEBFCD0FE9F3AA9FE22" box="[874,922,370,395]" name="Frey" pageId="15" pageNumber="622">Frey</location>
ex coll
</materialsCitation>
<materialsCitation id="6594F4042139FFEBFC48FE9F3A10FE04" ID-GBIF-Occurrence="3892866350" collectionCode="CF, NHMB, MNHN" collectorName="Reische" country="Brazil" location="Moser'" pageId="15" pageNumber="622" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
.
<location id="D023A8822139FFEBFBBBFE9F3D78FE22" LSID="urn:lsid:plazi:treatment:5D554F4F2139FFF5FF46FF633B55FADE:D023A8822139FFEBFBBBFE9F3D78FE22" box="[1025,1099,370,395]" country="Brazil" name="Moser'" pageId="15" pageNumber="622">Moser</location>
(
<collectionCode id="B3ED669C2139FFEBFBE6FE9F3D4FFE22" box="[1116,1148,370,395]" pageId="15" pageNumber="622">CF</collectionCode>
/
<collectionCode id="B3ED669C2139FFEBFF1AFE7939D8FE04" box="[160,235,404,429]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="15" pageNumber="622">NHMB</collectionCode>
),
<specimenCount id="C3FA35D02139FFEBFF41FE793816FE05" box="[251,293,404,429]" pageId="15" pageNumber="622" type="male">1 ♂</specimenCount>
<taxonomicName id="12FC85DA2139FFEBFE8BFE7938DCFE04" authorityName="Erichson" authorityYear="1835" box="[305,495,404,429]" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="622" phylum="Arthropoda" rank="species" species="nitidula">
<emphasis id="E788224B2139FFEBFE8BFE7938DCFE04" box="[305,495,404,429]" italics="true" pageId="15" pageNumber="622">Symmela nitidula</emphasis>
</taxonomicName>
<collectingCountry id="ADEBBEC92139FFEBFE4DFE793B00FE04" box="[503,563,404,429]" name="Brazil" pageId="15" pageNumber="622">Brésil</collectingCountry>
<collectorName id="78099B8F2139FFEBFD80FE793BBCFE04" box="[570,655,404,429]" pageId="15" pageNumber="622">Reische</collectorName>
1/2 (
<collectionCode id="B3ED669C2139FFEBFD77FE793A2EFE04" box="[717,797,404,429]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="622">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9DE6ADD22139FFF5FF1AFE043D5AFC45" lastPageId="17" lastPageNumber="624" pageId="15" pageNumber="622" type="description">
<paragraph id="D543FE592139FFEBFF1AFE043815FDAA" blockId="15.[160,1156,489,1693]" box="[160,294,489,515]" pageId="15" pageNumber="622">
<emphasis id="E788224B2139FFEBFF1AFE043815FDAA" bold="true" box="[160,294,489,515]" italics="true" pageId="15" pageNumber="622">Description</emphasis>
</paragraph>
<paragraph id="D543FE592139FFEBFF1AFDE13D56FC41" blockId="15.[160,1156,489,1693]" pageId="15" pageNumber="622">
Length:
<quantity id="120453BC2139FFEBFF45FDE13869FD8C" box="[255,346,524,549]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="15" pageNumber="622" unit="mm" value="6.5">6.5 mm</quantity>
. Elytral length:
<quantity id="120453BC2139FFEBFDAAFDE13B64FD8C" box="[528,599,524,549]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="622" unit="mm" value="4.0">4 mm</quantity>
. Width:
<quantity id="120453BC2139FFEBFD02FDE13A20FD8C" box="[696,787,524,549]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="15" pageNumber="622" unit="mm" value="3.1">3.1 mm</quantity>
. Head yellow, shiny, with erect, sparse setae. Labroclypeus as wide as frons, subtrapezoidal; surface convex; plain, without an anterior concavity on each side; anterior margin straight, weakly reflexed, with a rim of sparse setae behind anterior margin; lateral margin not incised anteriorly; punctation of surface posteriorly dense; suture between labrum and clypeus invisible. Smooth area in front of eye distinctly separated from frons by a suture. Frons anteriorly flat, punctation irregular, dense. Ocular canthus with terminal setae, ratio of length of ocular canthus/eye diameter: 1/3. Eyes evenly spherical, ratio of diameter/interocular width: 0.5. Antenna with eight antennomeres. Antennal club with three antennomeres, blackish, straight; width of antennal lamella equal to width of antennomeres of funiculus; apical lamella subequal to basal and median ones; length ratio of antennal club vs remaining antennomeres combined: 1/1.1; length ratio of third vs fourth antennomere: 1/2; fourth antennomere without lateral projection; length ratio of fifth vs fourth antennomere: 1/3. Labrum anteriorly bluntly angled on each side, concave at middle. Setae on mentum moderately dense.
</paragraph>
<paragraph id="D543FE592139FFEBFF7AFC1F3824FAEA" blockId="15.[160,1156,489,1693]" pageId="15" pageNumber="622">
Head and pronotum without copper iridescent shine. Pronotum unicoloured, shiny, yellowish; widest at base, width subequal to width of elytral base at humerus; anterior marginal line not interrupted medially, posterior marginal line absent; lateral margin not serrated, generally weakly convex, behind anterior angles convex, posteriorly straight; setae of lateral margin sparse; basal margin evenly curved; transverse impression beside basal margin absent; anterior angles produced, sharply pointed; antero-lateral and postero-lateral angles not flattened dorso-ventrally; posterior angles blunt; surface without median longitudinal impression, transversal furrows or setae; punctation on disc moderately dense, simple and fine, without well-defined impunctate areas.
<taxonomicName id="12FC85DA2139FFEBFC09FAEA3D11FA89" box="[947,1058,1287,1312]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="15" pageNumber="613" phylum="Arthropoda" rank="genus">Scutellum</taxonomicName>
without punctures.
</paragraph>
<paragraph id="D543FE592139FFEBFF7AFAA13B1AFA64" blockId="15.[160,1156,489,1693]" pageId="15" pageNumber="622">Elytra unicoloured, yellowish brown; surface iridescent, without setae and microsculpture; striae and intervals distinct; punctation between striae sparse, diameter of punctures compared to those of striae smaller; intervals flat; posterior edge convex; apical margin chitinous, without microtrichomes.</paragraph>
<paragraph id="D543FE592139FFEBFF7AFA3A3A46F934" blockId="15.[160,1156,489,1693]" pageId="15" pageNumber="622">
Abdomen: sternites flat in sagittal plane; lateral carina absent; first visible ventrite with row of setae in posterior edge on an elevated carina; second visible ventrite with row of setae in medial region; basal ventrites medially with half length as laterally; ultimate and penultimate apical ventrites simple and flat; apical ventrite short (as long as preceding ventrite); penultimate apical ventrite with row of setae like preceding one; beside the robust row of setae, almost glabrous.
<taxonomicName id="12FC85DA2139FFEBFDF0F9693B87F934" box="[586,692,1668,1693]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="613" phylum="Chordata" rank="genus">Pygidium</taxonomicName>
strongly convex.
</paragraph>
<caption id="8183AED12126FFF4FF1AF9933D08F91A" ID-DOI="http://doi.org/10.5281/zenodo.7012530" ID-Zenodo-Dep="7012530" httpUri="https://zenodo.org/record/7012530/files/figure.png" pageId="16" pageNumber="623" startId="16.[160,225,1662,1685]" targetBox="[273,1043,145,1623]" targetPageId="16">
<paragraph id="D543FE592126FFF4FF1AF9933D08F91A" blockId="16.[160,1156,1662,1715]" pageId="16" pageNumber="623">
<emphasis id="E788224B2126FFF4FF1AF99339CAF93C" bold="true" box="[160,249,1662,1685]" pageId="16" pageNumber="623">Figure 3.</emphasis>
<taxonomicName id="12FC85DA2126FFF4FEBAF9933880F93C" authority="Pacheco &amp; Wipfler &amp; Monné &amp; Ahrens, 2022" authorityName="Pacheco &amp; Wipfler &amp; Monné &amp; Ahrens" authorityYear="2022" box="[256,435,1662,1685]" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="623" phylum="Arthropoda" rank="species" species="capixaba" status="sp. nov.">
<emphasis id="E788224B2126FFF4FEBAF9933880F93C" box="[256,435,1662,1685]" italics="true" pageId="16" pageNumber="623">Symmela capixaba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FCBB9F302126FFF4FE03F9933B3BF93C" box="[441,520,1662,1685]" pageId="16" pageNumber="623" rank="species">sp. nov.</taxonomicNameLabel>
, holotype ♂. (a) Habitus, dorsal view; (b) habitus, lateral view; (c) aedeagus, lateral view; (d, e) parameres, dorsal view. Scale bars: a, b = 1 mm; ce = 0.5 mm.
</paragraph>
</caption>
<paragraph id="D543FE592127FFF5FF7AFF633B92FEA6" blockId="17.[160,1156,142,341]" pageId="17" pageNumber="624">Legs yellow. Ratio of length of metepisternum/metacoxa: 1/1.24. Metacoxa entirely plain, without elevated lateral longitudinal carina; without longer lateral bristles and setae on basal surface, posterior margin glabrous. Metafemur with posterior margin straight. Protibia with two distinct teeth; spur straight.</paragraph>
<paragraph id="D543FE592127FFF5FF7AFEF438E0FE9B" blockId="17.[160,1156,142,341]" box="[192,467,281,306]" pageId="17" pageNumber="624">
Aedeagus:
<figureCitation id="4DC7E2DC2127FFF5FE86FEF438ABFE9B" box="[316,408,281,306]" captionStart="Figure 3" captionStartId="16.[160,225,1662,1685]" captionTargetBox="[273,1043,145,1623]" captionTargetId="figure-3@16.[270,1047,142,1645]" captionTargetPageId="16" captionText="Figure 3. Symmela capixaba sp. nov., holotype ♂. (a) Habitus, dorsal view; (b) habitus, lateral view; (c) aedeagus, lateral view; (d, e) parameres, dorsal view. Scale bars: a, b = 1 mm; ce = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7012530" httpUri="https://zenodo.org/record/7012530/files/figure.png" pageId="17" pageNumber="624">Figure 3</figureCitation>
(ce).
</paragraph>
<paragraph id="D543FE592127FFF5FF7AFED138BAFEFC" blockId="17.[160,1156,142,341]" box="[192,393,316,341]" pageId="17" pageNumber="624">Female unknown.</paragraph>
<paragraph id="D543FE592127FFF5FF1AFE68383DFE36" blockId="17.[160,1156,389,1004]" box="[160,270,389,415]" pageId="17" pageNumber="624">
<emphasis id="E788224B2127FFF5FF1AFE68383DFE36" bold="true" box="[160,270,389,415]" italics="true" pageId="17" pageNumber="624">Variation</emphasis>
</paragraph>
<paragraph id="D543FE592127FFF5FF1AFE453AB6FE68" blockId="17.[160,1156,389,1004]" box="[160,901,424,449]" pageId="17" pageNumber="624">
Length:
<quantity id="120453BC2127FFF5FF46FE4538B1FE68" box="[252,386,424,449]" metricMagnitude="-3" metricUnit="m" metricValue="6.55" metricValueMax="6.6" metricValueMin="6.5" pageId="17" pageNumber="624" unit="mm" value="6.55" valueMax="6.6" valueMin="6.5">6.56.6 mm</quantity>
. Elytral length:
<quantity id="120453BC2127FFF5FD94FE453B92FE68" box="[558,673,424,449]" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.1" metricValueMin="4.0" pageId="17" pageNumber="624" unit="mm" value="4.05" valueMax="4.1" valueMin="4.0">44.1 mm</quantity>
. Width:
<quantity id="120453BC2127FFF5FD46FE453AB2FE68" box="[764,897,424,449]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="3.5" metricValueMin="3.1" pageId="17" pageNumber="624" unit="mm" value="3.3" valueMax="3.5" valueMin="3.1">3.13.5 mm</quantity>
.
</paragraph>
<paragraph id="D543FE592127FFF5FF7AFE263D5AFC45" blockId="17.[160,1156,389,1004]" pageId="17" pageNumber="624">Metatibia widest at middle; dorsal margin not carinated, smooth; interior surface more or less glabrous; median longitudinal punctation present on interior surface; interior margin of apex sharply and deeply truncate towards ventral margin; punctation on entire lateral face absent; beside dorsal margin without a row of robust setae along entire length; basal group of spines comprising at least two robust spines at same level of metatibial length; distal groups of spines disposed at two-thirds of metatibia length; ventral margin not serrated. Dorsal surface of tarsi glabrous, with dense punctures. Length ratio of dorsal tibial spur/first metatarsomere: 1/2. Metatarsi at apex weakly widened; first tarsomere straight, its length subequal to second and third tarsomeres combined; laterally not carinate; ventral margin serrated; ventral margin glabrous; medial face glabrous. Protibia with two distinct teeth; spur straight. First protarsomere straight, a little longer than second; width ratio of first protarsomere vs second protarsomere: 1.5/ 1; second protarsomere narrow and straight; third and fourth protarsomere elongate and narrow (distinctly longer than wide); fifth protarsomere circular in cross section, a little shorter than third and fourth protarsomere combined. Protarsal claws asymmetrical, shape of inner protarsal claw bifid, apex of basal tooth of inner protarsal claw convex.</paragraph>
</subSubSection>
<subSubSection id="9DE6ADD22127FFF5FF1AFBF13AA3FBD2" pageId="17" pageNumber="624" type="diagnosis">
<paragraph id="D543FE592127FFF5FF1AFBF13827FB9F" blockId="17.[160,1156,1052,1147]" box="[160,276,1052,1078]" pageId="17" pageNumber="624">
<emphasis id="E788224B2127FFF5FF1AFBF13827FB9F" bold="true" box="[160,276,1052,1078]" italics="true" pageId="17" pageNumber="624">Diagnosis</emphasis>
</paragraph>
<paragraph id="D543FE592127FFF5FF1AFBD23AA3FBD2" blockId="17.[160,1156,1052,1147]" pageId="17" pageNumber="624">
<taxonomicName id="12FC85DA2127FFF5FF1AFBD2385DFBF1" authority="Pacheco &amp; Wipfler &amp; Monné &amp; Ahrens, 2022" authorityName="Pacheco &amp; Wipfler &amp; Monné &amp; Ahrens" authorityYear="2022" box="[160,366,1087,1112]" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="624" phylum="Arthropoda" rank="species" species="capixaba" status="sp. nov.">
<emphasis id="E788224B2127FFF5FF1AFBD2385DFBF1" box="[160,366,1087,1112]" italics="true" pageId="17" pageNumber="624">Symmela capixaba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FCBB9F302127FFF5FECCFBD238FDFBF1" box="[374,462,1087,1112]" pageId="17" pageNumber="624" rank="species">sp. nov.</taxonomicNameLabel>
is in external appearance similar to
<taxonomicName id="12FC85DA2127FFF5FCDDFBD23D4DFBF1" authority="Erichson, 1835" authorityName="Erichson" authorityYear="1835" box="[871,1150,1087,1112]" class="Insecta" family="Scarabaeidae" genus="Symela" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="624" phylum="Arthropoda" rank="species" species="longula">
<emphasis id="E788224B2127FFF5FCDDFBD23AE0FBF1" box="[871,979,1087,1112]" italics="true" pageId="17" pageNumber="624">S. longula</emphasis>
<bibRefCitation id="B16D83A82127FFF5FC61FBD23D4DFBF1" author="Erichson WF" box="[987,1150,1087,1112]" pageId="17" pageNumber="624" pagination="256 - 270" refId="ref42712" refString="Erichson WF. 1835. Neue Sudamerikanische Kafergattungen aus der Familie der Blatterhorner. Arch Naturgeschichte. 1: 256 - 270." type="journal article" year="1835">Erichson, 1835</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="12FC85DA2127FFF5FF1AFB8F385DFBD2" authority="Pacheco &amp; Wipfler &amp; Monné &amp; Ahrens, 2022" authorityName="Pacheco &amp; Wipfler &amp; Monné &amp; Ahrens" authorityYear="2022" box="[160,366,1122,1147]" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="624" phylum="Arthropoda" rank="species" species="capixaba" status="sp. nov.">
<emphasis id="E788224B2127FFF5FF1AFB8F385DFBD2" box="[160,366,1122,1147]" italics="true" pageId="17" pageNumber="624">Symmela capixaba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FCBB9F302127FFF5FECFFB8F38FEFBD2" box="[373,461,1122,1147]" pageId="17" pageNumber="624" rank="species">sp. nov.</taxonomicNameLabel>
differs by its stout and trifid parameres.
</paragraph>
</subSubSection>
<subSubSection id="9DE6ADD22127FFF5FF1AFB463871FAA3" pageId="17" pageNumber="624" type="etymology">
<paragraph id="D543FE592127FFF5FF1AFB46382EFB6C" blockId="17.[160,1155,1195,1290]" box="[160,285,1195,1221]" pageId="17" pageNumber="624">
<emphasis id="E788224B2127FFF5FF1AFB46382EFB6C" bold="true" box="[160,285,1195,1221]" italics="true" pageId="17" pageNumber="624">Etymology</emphasis>
</paragraph>
<paragraph id="D543FE592127FFF5FF1AFB233871FAA3" blockId="17.[160,1155,1195,1290]" pageId="17" pageNumber="624">
The new species name is derived from the demonym of
<collectingRegion id="173830BB2127FFF5FCADFB233AC7FB4E" box="[791,1012,1230,1255]" country="Brazil" name="Espirito Santo" pageId="17" pageNumber="624">Espírito Santo State</collectingRegion>
,
<collectingCountry id="ADEBBEC92127FFF5FC44FB233D09FB4E" box="[1022,1082,1230,1255]" name="Brazil" pageId="17" pageNumber="624">Brazil</collectingCountry>
(noun in apposition).
</paragraph>
</subSubSection>
<subSubSection id="9DE6ADD22127FFF5FF1AFAD73B55FADE" box="[160,614,1338,1399]" pageId="17" pageNumber="624" type="discussion">
<paragraph id="D543FE592127FFF5FF1AFAD73835FAFD" blockId="17.[160,614,1338,1399]" box="[160,262,1338,1364]" pageId="17" pageNumber="624">
<emphasis id="E788224B2127FFF5FF1AFAD73835FAFD" bold="true" box="[160,262,1338,1364]" italics="true" pageId="17" pageNumber="624">Remarks</emphasis>
</paragraph>
<paragraph id="D543FE592127FFF5FF1AFAB33B55FADE" blockId="17.[160,614,1338,1399]" box="[160,614,1374,1399]" pageId="17" pageNumber="624">
<typeStatus id="0A4740FB2127FFF5FF1AFAB33834FADE" box="[160,263,1374,1399]" pageId="17" pageNumber="624" type="holotype">Holotype</typeStatus>
lacking metatibia and protarsi.
</paragraph>
</subSubSection>
</treatment>
</document>