215 lines
26 KiB
XML
215 lines
26 KiB
XML
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<mods:title id="6D201A82ACD3FE91E818DCE3E41F5B78">The genus Symmela Erichson, 1835 (Coleoptera, Scarabaeidae, Sericini): taxonomy and phylogenetic analysis</mods:title>
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<mods:namePart id="6BDD2CE3E84C4AB45B2ED67D4975C807">Pacheco, Thaynara L.</mods:namePart>
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<mods:namePart id="382DE53A57C9C8FADEDC03C9C2D63EF5">Wipfler, Benedict</mods:namePart>
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<mods:namePart id="27632482062664720AAE7E1DFE22CE0C">Monné, Marcela L.</mods:namePart>
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<mods:namePart id="F8047F39A10847B3852E5EF1327553A5">Ahrens, Dirk</mods:namePart>
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<mods:title id="790AC07AA6EBA68A9D451A95DD57F8E8">Journal of Natural History</mods:title>
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<mods:part id="17D8CA1939B098667AE07527C134A664">
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<mods:date id="2552A2465EE3955E26DFA113FD55197D">2022</mods:date>
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<mods:number id="FA2602180637A4A43C9EBFF000F39920">2022-07-11</mods:number>
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<mods:number id="ED9E77FCDE748C05FEE355566C2FA405">56</mods:number>
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<mods:number id="0503D9B5AD54D936B3069D35634395FF">9 - 12</mods:number>
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<mods:identifier id="5710AE2D83F4E54816371431512EC13A" type="DOI">10.1080/00222933.2022.2084649</mods:identifier>
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<treatment id="5D554F4F212BFFFBFF47FE1C3A63F969" ID-DOI="http://doi.org/10.5281/zenodo.7019039" ID-GBIF-Taxon="199728127" ID-Zenodo-Dep="7019039" LSID="urn:lsid:plazi:treatment:5D554F4F212BFFFBFF47FE1C3A63F969" httpUri="http://treatment.plazi.org/id/5D554F4F212BFFFBFF47FE1C3A63F969" lastPageId="31" lastPageNumber="638" pageId="29" pageNumber="636">
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<subSubSection id="9DE6ADD2212BFFF9FF47FE1C3D15FDA2" box="[253,1062,497,523]" pageId="29" pageNumber="636" type="nomenclature">
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<paragraph id="D543FE59212BFFF9FF47FE1C3D15FDA2" blockId="29.[253,1062,497,523]" box="[253,1062,497,523]" pageId="29" pageNumber="636">
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<heading id="8E0B4935212BFFF9FF47FE1C3D15FDA2" bold="true" box="[253,1062,497,523]" centered="true" fontSize="11" level="2" pageId="29" pageNumber="636" reason="3">
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<emphasis id="E788224B212BFFF9FF47FE1C3D15FDA2" bold="true" box="[253,1062,497,523]" pageId="29" pageNumber="636">
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<taxonomicName id="12FC85DA212BFFF9FF47FE1C3A8FFDA2" authority="Pacheco, Wipfler, Monne and Ahrens" authorityName="Pacheco, Wipfler, Monne and Ahrens" authorityYear="2022" box="[253,956,497,523]" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="636" phylum="Arthropoda" rank="species" species="erichsoni" status="sp. nov.">
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<emphasis id="E788224B212BFFF9FF47FE1C38D0FDA2" bold="true" box="[253,483,497,523]" italics="true" pageId="29" pageNumber="636">Symmela erichsoni</emphasis>
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Pacheco, Wipfler, Monné and Ahrens
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</taxonomicName>
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||
<taxonomicNameLabel id="FCBB9F30212BFFF9FC79FE1C3D15FDA2" box="[963,1062,497,523]" pageId="29" pageNumber="636" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="9DE6ADD2212BFFF9FDB0FDC83A2AFD96" box="[522,793,549,575]" pageId="29" pageNumber="636" type="description">
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<paragraph id="D543FE59212BFFF9FDB0FDC83A2AFD96" blockId="29.[522,793,549,575]" box="[522,793,549,575]" pageId="29" pageNumber="636">
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(
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<figureCitation id="4DC7E2DC212BFFF9FDA8FDC83B4EFD96" box="[530,637,549,575]" captionStart="Figure 8" captionStartId="30.[160,225,1710,1733]" captionTargetBox="[166,1151,144,1690]" captionTargetId="figure-3@30.[163,1153,142,1693]" captionTargetPageId="30" captionText="Figure 8. Symmela erichsoni sp. nov., holotype ♂. (a) Habitus, dorsal view; (b) habitus, lateral view; (c) aedeagus, lateral view; (d) aedeagus, dorsal view. Scale bars: a, b = 1 mm; c, d = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7012540" httpUri="https://zenodo.org/record/7012540/files/figure.png" pageId="29" pageNumber="636">Figures 8</figureCitation>
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,
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||
<figureCitation id="4DC7E2DC212BFFF9FD31FDC83B9EFD96" box="[651,685,549,575]" captionStart="Figure 29" captionStartId="86.[160,225,1430,1453]" captionTargetBox="[163,1152,142,1413]" captionTargetId="figure-75@86.[163,1153,142,1413]" captionTargetPageId="86" captionText="Figure 29. Distribution map of the Symmela species." figureDoi="http://doi.org/10.5281/zenodo.7012584" httpUri="https://zenodo.org/record/7012584/files/figure.png" pageId="29" pageNumber="636">29</figureCitation>
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(b), 30(h))
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</paragraph>
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</subSubSection>
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<subSubSection id="9DE6ADD2212BFFF9FF1AFD643B9EFCA2" pageId="29" pageNumber="636" type="materials_examined">
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<paragraph id="D543FE59212BFFF9FF1AFD64388CFD0A" blockId="29.[160,1156,649,779]" box="[160,447,649,675]" pageId="29" pageNumber="636">
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<emphasis id="E788224B212BFFF9FF1AFD64388CFD0A" bold="true" box="[160,447,649,675]" italics="true" pageId="29" pageNumber="636">Type material examined</emphasis>
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</paragraph>
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<paragraph id="D543FE59212BFFF9FF1AFD413B9EFCA2" blockId="29.[160,1156,649,779]" pageId="29" pageNumber="636">
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<materialsCitation id="6594F404212BFFF9FF1AFD413DB3FD6F" ID-GBIF-Occurrence="3892866357" box="[160,1152,684,710]" collectionCode="ZMHB" county="Neotropical Sericini" location="Rio Jan" municipality="Type" pageId="29" pageNumber="636" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<emphasis id="E788224B212BFFF9FF1AFD413821FD6F" bold="true" box="[160,274,684,710]" italics="true" pageId="29" pageNumber="636">
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<typeStatus id="0A4740FB212BFFF9FF1AFD41383EFD6F" box="[160,269,684,710]" pageId="29" pageNumber="636" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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||
<specimenCount id="C3FA35D0212BFFF9FE9BFD403806FD6D" box="[289,309,685,708]" pageId="29" pageNumber="636" type="male">♂</specimenCount>
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‘
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||
<location id="D023A882212BFFF9FE84FD4038A1FD6F" LSID="urn:lsid:plazi:treatment:5D554F4F212BFFFBFF47FE1C3A63F969:D023A882212BFFF9FE84FD4038A1FD6F" box="[318,402,685,710]" county="Neotropical Sericini" municipality="Type" name="Rio Jan" pageId="29" pageNumber="636">Rio Jan</location>
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/
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<collectingMunicipality id="35276423212BFFF9FE1AFD4038E5FD6F" box="[416,470,685,710]" pageId="29" pageNumber="636">Type</collectingMunicipality>
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mgl./
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<taxonomicName id="12FC85DA212BFFF9FDA7FD403BE4FD6F" authority="Er." authorityName="Er." box="[541,727,685,710]" class="Insecta" family="Scarabaeidae" genus="Symela" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="636" phylum="Arthropoda" rank="species" species="opaca">
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<emphasis id="E788224B212BFFF9FDA7FD403B87FD6F" box="[541,692,685,710]" italics="true" pageId="29" pageNumber="636">Symela opaca</emphasis>
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Er.
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</taxonomicName>
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||
/ 80
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<collectingCounty id="3C2286D5212BFFF9FCBDFD403AD3FD6F" box="[775,992,685,710]" pageId="29" pageNumber="636">Neotropical Sericini</collectingCounty>
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spec.’ (
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<collectionCode id="B3ED669C212BFFF9FB8AFD403D49FD6F" box="[1072,1146,685,710]" pageId="29" pageNumber="636">ZMHB</collectionCode>
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)
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</materialsCitation>
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.
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||
<materialsCitation id="6594F404212BFFF9FF1AFD2238D1FD41" ID-GBIF-Occurrence="3892866340" box="[160,482,719,744]" location="Rio Jan" pageId="29" pageNumber="636" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
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||
<typeStatus id="0A4740FB212BFFF9FF1AFD223823FD41" box="[160,272,719,744]" pageId="29" pageNumber="636" type="paratype">Paratypes</typeStatus>
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||
:
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||
<specimenCount id="C3FA35D0212BFFF9FEA0FD22386BFD4E" box="[282,344,719,744]" pageId="29" pageNumber="636" type="male">2 ♂♂</specimenCount>
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||
‘Fry
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||
<location id="D023A882212BFFF9FE35FD2238D1FD41" LSID="urn:lsid:plazi:treatment:5D554F4F212BFFFBFF47FE1C3A63F969:D023A882212BFFF9FE35FD2238D1FD41" box="[399,482,719,744]" name="Rio Jan" pageId="29" pageNumber="636">Rio Jan</location>
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</materialsCitation>
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./ Fry coll. 1905–100./
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||
<taxonomicName id="12FC85DA212BFFF9FD6DFD223ACCFD41" authority="Er. Det." authorityName="Er. Det." box="[727,1023,719,744]" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="636" phylum="Arthropoda" rank="species" species="mutabilis">
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<emphasis id="E788224B212BFFF9FD6DFD223A95FD41" box="[727,934,719,744]" italics="true" pageId="29" pageNumber="636">Symmela mutabilis</emphasis>
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Er. Det.
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</taxonomicName>
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at B.M. H.F. Howden ‘62
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<emphasis id="E788224B212BFFF9FE9DFD1F3803FCA2" box="[295,304,754,779]" italics="true" pageId="29" pageNumber="636">/</emphasis>
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<taxonomicName id="12FC85DA212BFFF9FE8DFD1F3B9AFCA2" authority="' (CNCI, ZFMK)" baseAuthorityName="CNCI, ZFMK" box="[311,681,754,779]" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="636" phylum="Arthropoda" rank="species" species="mutabilis">
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||
<emphasis id="E788224B212BFFF9FE8DFD1F3B35FCA2" box="[311,518,754,779]" italics="true" pageId="29" pageNumber="636">Symmela mutabilis</emphasis>
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’ (CNCI, ZFMK)
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</taxonomicName>
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.
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||
</paragraph>
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</subSubSection>
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<subSubSection id="9DE6ADD2212BFFFBFF1AFCA93D57FAD9" lastPageId="31" lastPageNumber="638" pageId="29" pageNumber="636" type="description">
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<paragraph id="D543FE59212BFFF9FF1AFCA93815FCF7" blockId="29.[160,1156,836,1728]" box="[160,294,836,862]" pageId="29" pageNumber="636">
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<emphasis id="E788224B212BFFF9FF1AFCA93815FCF7" bold="true" box="[160,294,836,862]" italics="true" pageId="29" pageNumber="636">Description</emphasis>
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</paragraph>
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||
<paragraph id="D543FE59212BFFF9FF1AFC8A3B88FACF" blockId="29.[160,1156,836,1728]" pageId="29" pageNumber="636">
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Length:
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||
<quantity id="120453BC212BFFF9FF46FC8A3867FC29" box="[252,340,871,896]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="29" pageNumber="636" unit="mm" value="5.1">5.1 mm</quantity>
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. Elytral length:
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<quantity id="120453BC212BFFF9FDBBFC8A3B6AFC29" box="[513,601,871,896]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="29" pageNumber="636" unit="mm" value="3.2">3.2 mm</quantity>
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. Width:
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<quantity id="120453BC212BFFF9FD0EFC8A3A38FC29" box="[692,779,871,896]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="636" unit="mm" value="2.6">2.6 mm</quantity>
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. Head blackish brown, iridescent, with erect, sparse setae. Labroclypeus as wide as frons, subtrapezoidal; surface convex; plain, without an anterior concavity on each side; anterior margin straight, weakly reflexed, with a rim of sparse setae behind anterior margin; lateral margin not incised anteriorly; punctation of surface posteriorly dense; suture between labrum and clypeus invisible. Smooth area in front of eye distinctly separated from frons by a suture. Frons anteriorly flat, punctation irregular, moderately dense. Ocular canthus with terminal setae, ratio of length of ocular canthus/eye diameter: 1/3. Eyes evenly spherical, ratio of diameter/interocular width: 0.49. Antenna with eight antennomeres. Antennal club with three antennomeres, blackish, straight; width of antennal lamella equal to width of antennomeres of funiculus; apical lamella subequal to basal and median ones; length ratio of antennal club vs remaining antennomeres combined: 1/1.1; length ratio of third vs fourth antennomere: 1/3; fourth antennomere without lateral projection; length ratio of fifth vs fourth antennomere: 1/5. Labrum anteriorly bluntly angled on each side, concave at middle. Setae on mentum moderately dense.
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||
</paragraph>
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||
<paragraph id="D543FE59212BFFF9FF7AFA823A63F969" blockId="29.[160,1156,836,1728]" pageId="29" pageNumber="636">
|
||
Head and pronotum without copper iridescent shine. Pronotum unicoloured, iridescent, blackish brown; widest at middle and base, width subequal to width of elytral base at humerus; anterior marginal line not interrupted medially, posterior marginal line absent; lateral margin not serrated, generally bent at middle, otherwise straight, behind anterior angles convex, posteriorly straight; setae of lateral margin sparse; basal margin evenly curved; transverse impression beside basal margin absent; anterior angles produced, sharply pointed; antero-lateral and postero-lateral angles not flattened dorsoventrally; posterior angles blunt; surface without median longitudinal impression, transversal furrows or setae; punctation on disc moderately dense, simple and fine, without well-defined impunctate areas.
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<taxonomicName id="12FC85DA212BFFF9FDB8F94A3B42F969" box="[514,625,1703,1728]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="29" pageNumber="613" phylum="Arthropoda" rank="genus">Scutellum</taxonomicName>
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without punctures.
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</paragraph>
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<caption id="8183AED12128FFFAFF1AF9433D28F94B" ID-DOI="http://doi.org/10.5281/zenodo.7012540" ID-Zenodo-Dep="7012540" httpUri="https://zenodo.org/record/7012540/files/figure.png" pageId="30" pageNumber="637" startId="30.[160,225,1710,1733]" targetBox="[166,1151,144,1690]" targetPageId="30">
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<paragraph id="D543FE592128FFFAFF1AF9433D28F94B" blockId="30.[160,1156,1710,1762]" pageId="30" pageNumber="637">
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||
<emphasis id="E788224B2128FFFAFF1AF94339C9F96C" bold="true" box="[160,250,1710,1733]" pageId="30" pageNumber="637">Figure 8.</emphasis>
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||
<taxonomicName id="12FC85DA2128FFFAFEBAF9433882F96C" authority="Pacheco & Wipfler & Monné & Ahrens, 2022" authorityName="Pacheco & Wipfler & Monné & Ahrens" authorityYear="2022" box="[256,433,1710,1733]" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="637" phylum="Arthropoda" rank="species" species="erichsoni" status="sp. nov.">
|
||
<emphasis id="E788224B2128FFFAFEBAF9433882F96C" box="[256,433,1710,1733]" italics="true" pageId="30" pageNumber="637">Symmela erichsoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FCBB9F302128FFFAFE02F9433B34F96C" box="[440,519,1710,1733]" pageId="30" pageNumber="637" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype ♂. (a) Habitus, dorsal view; (b) habitus, lateral view; (c) aedeagus, lateral view; (d) aedeagus, dorsal view. Scale bars: a, b = 1 mm; c, d = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D543FE592129FFFBFF7AFF633B1AFEA6" blockId="31.[160,1156,142,1242]" pageId="31" pageNumber="638">Elytra unicoloured, blackish brown; surface iridescent, without setae and microsculpture; striae and intervals distinct; punctation between striae sparse, diameter of punctures compared to those of striae smaller; intervals flat; posterior edge convex; apical margin chitinous, without microtrichomes.</paragraph>
|
||
<paragraph id="D543FE592129FFFBFF7AFEF43A46FE76" blockId="31.[160,1156,142,1242]" pageId="31" pageNumber="638">
|
||
Abdomen: sternites flat in sagittal plane; lateral carina absent; first visible ventrite with row of setae in posterior edge on an elevated carina; second visible ventrite with row of setae in medial region; basal ventrites medially with half length as laterally; ultimate and penultimate apical ventrites simple and flat; apical ventrite short (as long as preceding ventrite); penultimate apical ventrite with row of setae like preceding one; beside the robust row of setae, almost glabrous.
|
||
<taxonomicName id="12FC85DA2129FFFBFDF0FE2B3B87FE76" box="[586,692,454,479]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="31" pageNumber="613" phylum="Chordata" rank="genus">Pygidium</taxonomicName>
|
||
strongly convex.
|
||
</paragraph>
|
||
<paragraph id="D543FE592129FFFBFF7AFE0439C4FB3C" blockId="31.[160,1156,142,1242]" pageId="31" pageNumber="638">Legs blackish and yellow. Ratio of length of metepisternum/metacoxa: 1/1.22. Metacoxa entirely plain, without elevated lateral longitudinal carina; without longer lateral bristles, with setae on basal surface, posterior margin glabrous. Metafemur with posterior margin straight. Metatibia widest at apex; ratio of width/length: 1/3.33; dorsal margin not carinated, smooth; interior surface more or less glabrous; median longitudinal punctation present on interior surface; interior margin of apex sharply and deeply truncate towards ventral margin; punctation on entire lateral face present; beside dorsal margin without a row of robust setae along entire length; basal group of spines reduced to a single robust spine; distal groups of spines disposed at two-thirds of metatibia length; ventral margin not serrated. Dorsal surface of tarsi glabrous, with dense punctures. Length ratio of dorsal tibial spur/first metatarsomere: 1/2. Metatarsi at apex weakly widened; first tarsomere straight, its length subequal to second and third tarsomeres combined; laterally not carinate; ventral margin serrated; ventral margin glabrous; medial face glabrous. Protibia with two distinct teeth; spur straight. First protarsomere curved, its length longer than second, third and fourth combined; width ratio of first protarsomere vs second protarsomere: ca. 1/2; second protarsomere enlarged; third and fourth protarsomere short and transverse (distinctly wider than long); fifth protarsomere circular in cross section, a little shorter than third and fourth protarsomere combined. Protarsal claws asymmetrical, shape of inner protarsal claw bifid, apex of basal tooth of inner protarsal claw convex.</paragraph>
|
||
<paragraph id="D543FE592129FFFBFF7AFB7338FEFB1E" blockId="31.[160,1156,142,1242]" box="[192,461,1182,1207]" pageId="31" pageNumber="638">
|
||
Aedeagus:
|
||
<figureCitation id="4DC7E2DC2129FFFBFE86FB7338AAFB1E" box="[316,409,1182,1207]" captionStart="Figure 8" captionStartId="30.[160,225,1710,1733]" captionTargetBox="[166,1151,144,1690]" captionTargetId="figure-3@30.[163,1153,142,1693]" captionTargetPageId="30" captionText="Figure 8. Symmela erichsoni sp. nov., holotype ♂. (a) Habitus, dorsal view; (b) habitus, lateral view; (c) aedeagus, lateral view; (d) aedeagus, dorsal view. Scale bars: a, b = 1 mm; c, d = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7012540" httpUri="https://zenodo.org/record/7012540/files/figure.png" pageId="31" pageNumber="638">Figure 8</figureCitation>
|
||
(c,d).
|
||
</paragraph>
|
||
<paragraph id="D543FE592129FFFBFF7AFB2C38BAFB73" blockId="31.[160,1156,142,1242]" box="[192,393,1217,1242]" pageId="31" pageNumber="638">Female unknown.</paragraph>
|
||
<paragraph id="D543FE592129FFFBFF1AFAFC383DFA82" blockId="31.[160,1156,1297,1392]" box="[160,270,1297,1323]" pageId="31" pageNumber="638">
|
||
<emphasis id="E788224B2129FFFBFF1AFAFC383DFA82" bold="true" box="[160,270,1297,1323]" italics="true" pageId="31" pageNumber="638">Variation</emphasis>
|
||
</paragraph>
|
||
<paragraph id="D543FE592129FFFBFF1AFAD83D57FAD9" blockId="31.[160,1156,1297,1392]" pageId="31" pageNumber="638">
|
||
Length: 5.1–6.0 mm. Elytral length:
|
||
<quantity id="120453BC2129FFFBFD9CFAD83B99FAE7" box="[550,682,1333,1358]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="3.8" metricValueMin="3.2" pageId="31" pageNumber="638" unit="mm" value="3.5" valueMax="3.8" valueMin="3.2">3.2–3.8 mm</quantity>
|
||
. Width:
|
||
<quantity id="120453BC2129FFFBFCB8FAD83AB6FAE7" box="[770,901,1333,1358]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.1" metricValueMin="2.6" pageId="31" pageNumber="638" unit="mm" value="2.85" valueMax="3.1" valueMin="2.6">2.6–3.1 mm</quantity>
|
||
. Pronotum bicoloured, basal lateral margins yellowish; elytra bicoloured, yellowish brown with dark margins.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9DE6ADD22129FFFBFF1AFA453AEFF983" pageId="31" pageNumber="638" type="diagnosis">
|
||
<paragraph id="D543FE592129FFFBFF1AFA453827FA6B" blockId="31.[160,1156,1448,1578]" box="[160,276,1448,1474]" pageId="31" pageNumber="638">
|
||
<emphasis id="E788224B2129FFFBFF1AFA453827FA6B" bold="true" box="[160,276,1448,1474]" italics="true" pageId="31" pageNumber="638">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="D543FE592129FFFBFF1AFA263AEFF983" blockId="31.[160,1156,1448,1578]" pageId="31" pageNumber="638">
|
||
<taxonomicName id="12FC85DA2129FFFBFF1AFA263859FA4D" authority="Pacheco & Wipfler & Monné & Ahrens, 2022" authorityName="Pacheco & Wipfler & Monné & Ahrens" authorityYear="2022" box="[160,362,1483,1508]" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="638" phylum="Arthropoda" rank="species" species="erichsoni" status="sp. nov.">
|
||
<emphasis id="E788224B2129FFFBFF1AFA263859FA4D" box="[160,362,1483,1508]" italics="true" pageId="31" pageNumber="638">Symmela erichsoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FCBB9F302129FFFBFECAFA2638F4FA4D" box="[368,455,1483,1508]" pageId="31" pageNumber="638" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="12FC85DA2129FFFBFDEEFA263B9DFA4D" authorityName="Pacheco & Wipfler & Monné & Ahrens" authorityYear="2022" box="[596,686,1483,1508]" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="638" phylum="Arthropoda" rank="species" species="beskei">
|
||
<emphasis id="E788224B2129FFFBFDEEFA263B9DFA4D" box="[596,686,1483,1508]" italics="true" pageId="31" pageNumber="638">S. beskei</emphasis>
|
||
</taxonomicName>
|
||
by the enlarged first protarsomere, which is longer than second, third, and fourth protarsomere combined, and by the shorter and stouter parameres which are flattened laterally instead of dorso-ventrally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9DE6ADD22129FFFBFF1AF98C3A63F969" pageId="31" pageNumber="638" type="etymology">
|
||
<paragraph id="D543FE592129FFFBFF1AF98C382EF9D2" blockId="31.[160,1155,1633,1728]" box="[160,285,1633,1659]" pageId="31" pageNumber="638">
|
||
<emphasis id="E788224B2129FFFBFF1AF98C382EF9D2" bold="true" box="[160,285,1633,1659]" italics="true" pageId="31" pageNumber="638">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="D543FE592129FFFBFF1AF9683A63F969" blockId="31.[160,1155,1633,1728]" pageId="31" pageNumber="638">
|
||
The species is named after W.F. Erichson, a German entomologist who described the genus
|
||
<taxonomicName id="12FC85DA2129FFFBFF56F94A387CF969" authorityName="Erichson" authorityYear="1835" box="[236,335,1703,1728]" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="638" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E788224B2129FFFBFF56F94A387CF969" box="[236,335,1703,1728]" italics="true" pageId="31" pageNumber="638">Symmela</emphasis>
|
||
</taxonomicName>
|
||
and many other taxa (noun in genitive case).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |