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<document id="0FF659F04D7EDE01F46B62E617A8DB02" ID-CLB-Dataset="128003" ID-DOI="10.1080/00222933.2022.2084649" ID-GBIF-Dataset="006719bf-f416-4310-9626-29381fd9c26d" ID-ISSN="1464-5262" ID-Zenodo-Dep="7012522" ID-ZooBank="5147CC1B-6144-4CA4-BA63-E39ED0A20E1C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1660992512912" checkinUser="plazi" docAuthor="Pacheco, Thaynara L., Wipfler, Benedict, Monné, Marcela L. &amp; Ahrens, Dirk" docDate="2022" docId="5D554F4F2114FFC1FF4EFA9639ECFD5A" docLanguage="en" docName="JNATHIST.56.9-12.607-705.pdf" docOrigin="Journal of Natural History 56 (9 - 12)" docSource="http://dx.doi.org/10.1080/00222933.2022.2084649" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Symmela fuhrmanni Pacheco, Wipfler, Monne and Ahrens 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="644" masterDocId="A16C37372136FFE4FFBAFFED3933FFA9" masterDocTitle="The genus Symmela Erichson, 1835 (Coleoptera, Scarabaeidae, Sericini): taxonomy and phylogenetic analysis" masterLastPageNumber="705" masterPageNumber="607" pageNumber="641" updateTime="1699372031290" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="EE24142E59C2B6D79A9E7C7CBEF19B30">The genus Symmela Erichson, 1835 (Coleoptera, Scarabaeidae, Sericini): taxonomy and phylogenetic analysis</mods:title>
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<mods:namePart id="A4B219C5B733F0CC0F64FF9BECE57C94">Pacheco, Thaynara L.</mods:namePart>
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<mods:namePart id="CEEB125E8DEBF1E751C562495CE82319">Wipfler, Benedict</mods:namePart>
</mods:name>
<mods:name id="3B67A9D6E99322E969A7873C0CDC750E" type="personal">
<mods:role id="AE7256C308A2E4BC7E2BBE4688033D9C">
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</mods:role>
<mods:namePart id="54CBE6842F22F1F9EB92693D4F9A416F">Monné, Marcela L.</mods:namePart>
</mods:name>
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<mods:role id="AD2347634C051E0BBECC24854B30FCA5">
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</mods:role>
<mods:namePart id="E5EF3B1E95F564268CE9D98929CF8CF6">Ahrens, Dirk</mods:namePart>
</mods:name>
<mods:typeOfResource id="C1D33E4D28D45CE989DF5C50925DFDB3">text</mods:typeOfResource>
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<mods:title id="6FD717837C791B8B6494264C88CC4307">Journal of Natural History</mods:title>
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<mods:part id="8A005923620404AADA3FC49DBEF454AE">
<mods:date id="75BBBB67C53907CC5FF0966677CACC72">2022</mods:date>
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<mods:number id="DD28A09B61CD0CEC516E837057711B42">2022-07-11</mods:number>
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<mods:number id="D4FC229F650B0F6F5EBFADCBCA83DA46">56</mods:number>
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<mods:number id="9DED68F2012A2D5C6CC1DD63646E26D3">9 - 12</mods:number>
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<mods:start id="26C83CC11EA9E3CB05ACB8F0FE1081B0">607</mods:start>
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<mods:classification id="45711BF540E9225746D2ED783DA61A50">journal article</mods:classification>
<mods:identifier id="60B44827003256CA186C3BA4B3068310" type="CLB-Dataset">128003</mods:identifier>
<mods:identifier id="258E197268DFD437657EE563F36D9330" type="DOI">10.1080/00222933.2022.2084649</mods:identifier>
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</mods:mods>
<treatment id="5D554F4F2114FFC1FF4EFA9639ECFD5A" ID-DOI="http://doi.org/10.5281/zenodo.7018991" ID-GBIF-Taxon="199728130" ID-Zenodo-Dep="7018991" LSID="urn:lsid:plazi:treatment:5D554F4F2114FFC1FF4EFA9639ECFD5A" httpUri="http://treatment.plazi.org/id/5D554F4F2114FFC1FF4EFA9639ECFD5A" lastPageId="37" lastPageNumber="644" pageId="34" pageNumber="641">
<subSubSection id="9DE6ADD22114FFC6FF4EFA963D1CFA3C" box="[244,1071,1403,1429]" pageId="34" pageNumber="641" type="nomenclature">
<paragraph id="D543FE592114FFC6FF4EFA963D1CFA3C" blockId="34.[244,1071,1403,1429]" box="[244,1071,1403,1429]" pageId="34" pageNumber="641">
<heading id="8E0B49352114FFC6FF4EFA963D1CFA3C" bold="true" box="[244,1071,1403,1429]" centered="true" fontSize="11" level="2" pageId="34" pageNumber="641" reason="3">
<emphasis id="E788224B2114FFC6FF4EFA963D1CFA3C" bold="true" box="[244,1071,1403,1429]" pageId="34" pageNumber="641">
<taxonomicName id="12FC85DA2114FFC6FF4EFA963AF6FA3C" authority="Pacheco, Wipfler, Monne and Ahrens" authorityName="Pacheco, Wipfler, Monne and Ahrens" authorityYear="2022" box="[244,965,1403,1429]" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="641" phylum="Arthropoda" rank="species" species="fuhrmanni" status="sp. nov.">
<emphasis id="E788224B2114FFC6FF4EFA9638DFFA3C" bold="true" box="[244,492,1403,1429]" italics="true" pageId="34" pageNumber="641">Symmela fuhrmanni</emphasis>
Pacheco, Wipfler, Monné and Ahrens
</taxonomicName>
<taxonomicNameLabel id="FCBB9F302114FFC6FC77FA963D1CFA3C" box="[973,1071,1403,1429]" pageId="34" pageNumber="641" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9DE6ADD22114FFC6FDBDFA4F3A2EFA15" box="[519,797,1442,1468]" pageId="34" pageNumber="641" type="description">
<paragraph id="D543FE592114FFC6FDBDFA4F3A2EFA15" blockId="34.[519,797,1442,1468]" box="[519,797,1442,1468]" pageId="34" pageNumber="641">
(
<figureCitation id="4DC7E2DC2114FFC6FDB5FA4F3BBAFA15" box="[527,649,1442,1468]" captionStart="Figure 10" captionStartId="36.[160,225,1667,1690]" captionTargetBox="[335,980,145,1647]" captionTargetId="figure-3@36.[333,983,142,1650]" captionTargetPageId="36" captionText="Figure 10. Symmela fuhrmanni sp. nov., holotype ♂. (a) Habitus, dorsal view; (b) habitus, lateral view; (c) aedeagus, lateral view; (d) parameres, dorsal view. Scale bars: a, b = 1 mm; c, d = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7012544" httpUri="https://zenodo.org/record/7012544/files/figure.png" pageId="34" pageNumber="641">Figures 10</figureCitation>
,
<figureCitation id="4DC7E2DC2114FFC6FD2DFA4F3B8AFA15" box="[663,697,1442,1468]" captionStart="Figure 29" captionStartId="86.[160,225,1430,1453]" captionTargetBox="[163,1152,142,1413]" captionTargetId="figure-75@86.[163,1153,142,1413]" captionTargetPageId="86" captionText="Figure 29. Distribution map of the Symmela species." figureDoi="http://doi.org/10.5281/zenodo.7012584" httpUri="https://zenodo.org/record/7012584/files/figure.png" pageId="34" pageNumber="641">29</figureCitation>
(b), 30(j))
</paragraph>
</subSubSection>
<subSubSection id="9DE6ADD22114FFC6FF1AFA143A6BF969" pageId="34" pageNumber="641" type="materials_examined">
<paragraph id="D543FE592114FFC6FF1AFA14388CF9BA" blockId="34.[160,1157,1529,1728]" box="[160,447,1529,1555]" pageId="34" pageNumber="641">
<emphasis id="E788224B2114FFC6FF1AFA14388CF9BA" bold="true" box="[160,447,1529,1555]" italics="true" pageId="34" pageNumber="641">Type material examined</emphasis>
</paragraph>
<paragraph id="D543FE592114FFC6FF1AF9F13A6BF969" blockId="34.[160,1157,1529,1728]" pageId="34" pageNumber="641">
<materialsCitation id="6594F4042114FFC6FF1AF9F13DB2F9F1" ID-GBIF-Occurrence="3892866490" collectingDate="2010-12-01" collectingDateMax="2010-12-15" collectingDateMin="2010-12-01" collectionCode="CEMT" collectorName="M. R. Frizzas &amp; Neotropical Sericini" country="Brazil" location="Aquas Emendadas" longitude="-47.61361" pageId="34" pageNumber="641" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="E788224B2114FFC6FF1AF9F13821F99F" bold="true" box="[160,274,1564,1590]" italics="true" pageId="34" pageNumber="641">
<typeStatus id="0A4740FB2114FFC6FF1AF9F1383EF99F" box="[160,269,1564,1590]" pageId="34" pageNumber="641" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="C3FA35D02114FFC6FE9BF9F03806F99D" box="[289,309,1565,1588]" pageId="34" pageNumber="641" type="male">♂</specimenCount>
‘
<collectingCountry id="ADEBBEC92114FFC6FE85F9F038A1F99F" box="[319,402,1565,1590]" name="Brazil" pageId="34" pageNumber="641">BRASIL</collectingCountry>
: Distrito Federal. Planatina. ESEC
<location id="D023A8822114FFC6FCB3F9F03AE8F99F" LSID="urn:lsid:plazi:treatment:5D554F4F2114FFC1FF4EFA9639ECFD5A:D023A8822114FFC6FCB3F9F03AE8F99F" box="[777,987,1565,1590]" country="Brazil" longitude="-47.61361" name="Aquas Emendadas" pageId="34" pageNumber="641">Áquas Emendadas</location>
,
<geoCoordinate id="B0C8989E2114FFC6FC5EF9F03D6BF99F" box="[996,1112,1565,1590]" pageId="34" pageNumber="641" value="-15.5419">
15°32
<emphasis id="E788224B2114FFC6FB9EF9F03D19F99C" bold="true" box="[1060,1066,1565,1589]" pageId="34" pageNumber="641">ʹ</emphasis>
31′S
</geoCoordinate>
<geoCoordinate id="B0C8989E2114FFC6FBE5F9F03837F9F1" degrees="47" direction="west" minutes="36" orientation="longitude" pageId="34" pageNumber="641" precision="15" seconds="49" value="-47.61361">
47° 36
<emphasis id="E788224B2114FFC6FF06F9D239F1F9FE" bold="true" box="[188,194,1599,1623]" pageId="34" pageNumber="641">ʹ</emphasis>
49″W
</geoCoordinate>
.
<date id="A142D8992114FFC6FEAFF9D23884F9F1" box="[277,439,1599,1624]" pageId="34" pageNumber="641" value="2010-12-01" valueMax="2010-12-15" valueMin="2010-12-01">
<collectingDate id="B10621712114FFC6FEAFF9D23884F9F1" box="[277,439,1599,1624]" pageId="34" pageNumber="641" value="2010-12-01" valueMax="2010-12-15" valueMin="2010-12-01">1.-15-XII-2010</collectingDate>
</date>
,
<collectingMethod id="0CBD864E2114FFC6FE72F9D23B3FF9F1" box="[456,524,1599,1624]" pageId="34" pageNumber="641">Pitfall</collectingMethod>
,
<collectorName id="78099B8F2114FFC6FDA7F9D23B99F9F1" box="[541,682,1599,1624]" pageId="34" pageNumber="641">M.R. Frizzas</collectorName>
/ 38
<collectorName id="78099B8F2114FFC6FD53F9D23AE3F9F1" box="[745,976,1599,1624]" pageId="34" pageNumber="641">Neotropical Sericini</collectorName>
spec.’ (
<collectionCode id="B3ED669C2114FFC6FB89F9D23D48F9F1" box="[1075,1147,1599,1624]" pageId="34" pageNumber="641">CEMT</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="6594F4042114FFC6FF1AF98F3B54F9D2" ID-GBIF-Occurrence="3892866367" box="[160,615,1634,1659]" country="Brazil" location="Distrito Federal" pageId="34" pageNumber="641" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="0A4740FB2114FFC6FF1AF98F3823F9D2" box="[160,272,1634,1659]" pageId="34" pageNumber="641" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="C3FA35D02114FFC6FEA7F98F387AF9D0" box="[285,329,1634,1659]" pageId="34" pageNumber="641" type="male">1 ♂</specimenCount>
‘
<collectingCountry id="ADEBBEC92114FFC6FEECF98F389AF9D2" box="[342,425,1634,1659]" name="Brazil" pageId="34" pageNumber="641">BRASIL</collectingCountry>
:
<location id="D023A8822114FFC6FE0CF98F3B54F9D2" LSID="urn:lsid:plazi:treatment:5D554F4F2114FFC1FF4EFA9639ECFD5A:D023A8822114FFC6FE0CF98F3B54F9D2" box="[438,615,1634,1659]" country="Brazil" name="Distrito Federal" pageId="34" pageNumber="641">Distrito Federal</location>
</materialsCitation>
. Planatina. ESEC Áquas Emendadas, 15°32
<emphasis id="E788224B2114FFC6FBF5F98F3D66F9D3" bold="true" box="[1103,1109,1634,1658]" pageId="34" pageNumber="641">ʹ</emphasis>
31′S
<geoCoordinate id="B0C8989E2114FFC6FF1AF9683816F937" box="[160,293,1669,1694]" degrees="47" direction="west" minutes="36" orientation="longitude" pageId="34" pageNumber="641" precision="15" seconds="49" value="-47.61361">
47°36
<emphasis id="E788224B2114FFC6FF65F96839D6F934" bold="true" box="[223,229,1669,1693]" pageId="34" pageNumber="641">ʹ</emphasis>
49″W
</geoCoordinate>
.
<date id="A142D8992114FFC6FE97F96838F9F937" box="[301,458,1669,1694]" pageId="34" pageNumber="641" value="2010-12-01" valueMax="2010-12-15" valueMin="2010-12-01">1.-15-XII-2010</date>
, Pitfall, M.R. Frizzas/ 38 Neotropical Sericini spec.’ (
<collectionCode id="B3ED669C2114FFC6FC49F9683D0AF937" box="[1011,1081,1669,1694]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="34" pageNumber="641" type="Museum">ZFMK</collectionCode>
),
<materialsCitation id="6594F4042114FFC6FBF2F968382CF969" ID-GBIF-Occurrence="3892866370" country="Brazil" location="Brasil" pageId="34" pageNumber="641" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="C3FA35D02114FFC6FBF2F9683DB0F935" box="[1096,1155,1669,1694]" pageId="34" pageNumber="641" type="male">3 ♂♂</specimenCount>
‘
<collectingCountry id="ADEBBEC92114FFC6FF18F94A39D3F969" box="[162,224,1703,1728]" name="Brazil" pageId="34" pageNumber="641">Brasil</collectingCountry>
GO J
</materialsCitation>
. &amp;. Bechyné/ Faz.
<materialsCitation id="6594F4042114FFC6FE53F94A3A66F969" ID-GBIF-Occurrence="3892866309" box="[489,853,1703,1728]" collectingDate="1962-02-08" collectionCode="UNSM, ZFMK" country="Brazil" pageId="34" pageNumber="641" specimenCount="1" typeStatus="paratype">
Paraiso
<date id="A142D8992114FFC6FDFBF94A3B95F969" box="[577,678,1703,1728]" pageId="34" pageNumber="641" value="1962-02-08">
<collectingDate id="B10621712114FFC6FDFBF94A3B95F969" box="[577,678,1703,1728]" pageId="34" pageNumber="641" value="1962-02-08">8-2-1962</collectingDate>
</date>
’ (
<collectionCode id="B3ED669C2114FFC6FD0CF94A3A33F969" box="[694,768,1703,1728]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="34" pageNumber="641">UNSM</collectionCode>
,
<collectionCode id="B3ED669C2114FFC6FCB0F94A3A7DF969" box="[778,846,1703,1728]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="34" pageNumber="641" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9DE6ADD22115FFC1FF1AFF633A66FE32" lastPageId="37" lastPageNumber="644" pageId="35" pageNumber="642" type="description">
<paragraph id="D543FE592115FFC7FF1AFF633815FF01" blockId="35.[160,1156,142,1728]" box="[160,294,142,168]" pageId="35" pageNumber="642">
<emphasis id="E788224B2115FFC7FF1AFF633815FF01" bold="true" box="[160,294,142,168]" italics="true" pageId="35" pageNumber="642">Description</emphasis>
</paragraph>
<paragraph id="D543FE592115FFC7FF1AFF5F3842FDAA" blockId="35.[160,1156,142,1728]" pageId="35" pageNumber="642">
Length:
<quantity id="120453BC2115FFC7FF47FF5F3866FF62" box="[253,341,178,203]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="35" pageNumber="642" unit="mm" value="5.5">5.5 mm</quantity>
. Elytral length:
<quantity id="120453BC2115FFC7FDBFFF5F3B6EFF62" box="[517,605,178,203]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="35" pageNumber="642" unit="mm" value="2.9">2.9 mm</quantity>
. Width:
<quantity id="120453BC2115FFC7FD03FF5F3A21FF62" box="[697,786,178,203]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="35" pageNumber="642" unit="mm" value="2.4">2.4 mm</quantity>
. Head blackish brown, dull, with erect dense setae. Labroclypeus as wide as frons, subtrapezoidal; surface flat; plain, without an anterior concavity on each side; anterior margin straight, strongly reflexed, with a rim of dense, short setae behind anterior margin; lateral margin not incised anteriorly; punctation of surface posteriorly dense; suture between labrum and clypeus invisible. Smooth area in front of eye distinctly separated from frons by a suture. Frons anteriorly flat, punctation irregular, sparse. Ocular canthus with terminal setae, ratio of length of ocular canthus/eye diameter: 1/3. Eyes evenly spherical, ratio of diameter/ interocular width: 0.47. Labrum anteriorly without blunt processes. Setae on mentum moderately dense.
</paragraph>
<paragraph id="D543FE592115FFC7FF7AFDE1382BFCF4" blockId="35.[160,1156,142,1728]" pageId="35" pageNumber="642">
Head and pronotum without copper iridescent shine. Pronotum unicoloured, dull, black; widest at middle, width subequal to width of elytral base at humerus; anterior marginal line not interrupted medially, posterior marginal line absent; lateral margin not serrated, generally strongly convex, behind anterior angles convex, posteriorly weakly convex; setae of lateral margin sparse; basal margin evenly curved; transverse impression beside basal margin absent; anterior angles produced, sharply pointed; antero-lateral and postero-lateral angles not flattened dorso-ventrally; posterior angles blunt; surface without median longitudinal impression, transversal furrows or setae; punctation on disc dense, simple and fine, without well-defined impunctate areas.
<taxonomicName id="12FC85DA2115FFC7FC10FCCF3D2EFC92" box="[938,1053,802,827]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="35" pageNumber="613" phylum="Arthropoda" rank="genus">Scutellum</taxonomicName>
without punctures.
</paragraph>
<paragraph id="D543FE592115FFC7FF7AFC8A3BB3FC41" blockId="35.[160,1156,142,1728]" pageId="35" pageNumber="642">Elytra unicoloured, blackish brown; surface opaque and iridescent, without setae and microsculpture; striae and intervals distinct; punctation between striae sparse, diameter of punctures compared to those of striae smaller; intervals flat; posterior edge convex; apical margin chitinous, without microtrichomes.</paragraph>
<paragraph id="D543FE592115FFC7FF7AFC1F3B09FB11" blockId="35.[160,1156,142,1728]" pageId="35" pageNumber="642">
Abdomen: sternites flat in sagittal plane; lateral carina absent; first visible ventrite with row of setae on posterior edge; second visible ventrite with row of setae in medial region; basal ventrites medially of same length as laterally; ultimate and penultimate apical ventrites simple and flat; apical ventrite short (as long as preceding ventrite); penultimate apical ventrite with row of setae like preceding one; beside the robust row of setae, almost glabrous.
<taxonomicName id="12FC85DA2115FFC7FEB5FB72384AFB11" box="[271,377,1183,1208]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="35" pageNumber="613" phylum="Chordata" rank="genus">Pygidium</taxonomicName>
strongly convex.
</paragraph>
<paragraph id="D543FE592115FFC7FF7AFB2F3A26F9D2" blockId="35.[160,1156,142,1728]" pageId="35" pageNumber="642">Legs blackish. Ratio of length of metepisternum/metacoxa: 1/1.44. Metacoxa entirely plain, without elevated lateral longitudinal carina; without longer lateral bristles, with setae on basal surface, posterior margin glabrous. Metafemur with posterior margin straight. Metatibia widest at middle; ratio of width/length: 1/2.04; dorsal margin not carinated, smooth; interior surface more or less glabrous; median longitudinal punctation present on interior surface; interior margin of apex deeply concave; punctation on entire lateral face present; beside dorsal margin without a row of robust setae along entire length; basal group of spines reduced to a single robust spine; distal groups of spines disposed at nearly half of metatibia length; ventral margin not serrated. Dorsal surface of tarsi glabrous, with dense punctures. Length ratio of dorsal tibial spur/first metatarsomere: 1/2. Metatarsi at apex weakly widened; first tarsomere straight, its length subequal to second and third tarsomeres combined; laterally not carinate; ventral margin serrated; ventral margin glabrous; medial face glabrous.</paragraph>
<paragraph id="D543FE592115FFC7FF7AF96938E8F934" blockId="35.[160,1156,142,1728]" box="[192,475,1668,1693]" pageId="35" pageNumber="642">
Aedeagus:
<figureCitation id="4DC7E2DC2115FFC7FE86F9693894F934" box="[316,423,1668,1693]" captionStart="Figure 10" captionStartId="36.[160,225,1667,1690]" captionTargetBox="[335,980,145,1647]" captionTargetId="figure-3@36.[333,983,142,1650]" captionTargetPageId="36" captionText="Figure 10. Symmela fuhrmanni sp. nov., holotype ♂. (a) Habitus, dorsal view; (b) habitus, lateral view; (c) aedeagus, lateral view; (d) parameres, dorsal view. Scale bars: a, b = 1 mm; c, d = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7012544" httpUri="https://zenodo.org/record/7012544/files/figure.png" pageId="35" pageNumber="642">Figure 10</figureCitation>
(c,d).
</paragraph>
<paragraph id="D543FE592115FFC7FF7AF94A38BAF969" blockId="35.[160,1156,142,1728]" box="[192,393,1703,1728]" pageId="35" pageNumber="642">Female unknown.</paragraph>
<caption id="8183AED12112FFC0FF1AF96E3D70F911" ID-DOI="http://doi.org/10.5281/zenodo.7012544" ID-Zenodo-Dep="7012544" httpUri="https://zenodo.org/record/7012544/files/figure.png" pageId="36" pageNumber="643" startId="36.[160,225,1667,1690]" targetBox="[335,980,145,1647]" targetPageId="36">
<paragraph id="D543FE592112FFC0FF1AF96E3D70F911" blockId="36.[160,1158,1667,1720]" pageId="36" pageNumber="643">
<emphasis id="E788224B2112FFC0FF1AF96E3835F933" bold="true" box="[160,262,1667,1690]" pageId="36" pageNumber="643">Figure 10.</emphasis>
<taxonomicName id="12FC85DA2112FFC0FEB7F96E38FDF933" authority="Pacheco &amp; Wipfler &amp; Monné &amp; Ahrens, 2022" authorityName="Pacheco &amp; Wipfler &amp; Monné &amp; Ahrens" authorityYear="2022" box="[269,462,1667,1690]" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="643" phylum="Arthropoda" rank="species" species="fuhrmanni" status="sp. nov.">
<emphasis id="E788224B2112FFC0FEB7F96E38FDF933" box="[269,462,1667,1690]" italics="true" pageId="36" pageNumber="643">Symmela fuhrmanni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FCBB9F302112FFC0FE6FF96E3B17F933" box="[469,548,1667,1690]" pageId="36" pageNumber="643" rank="species">sp. nov.</taxonomicNameLabel>
, holotype ♂. (a) Habitus, dorsal view; (b) habitus, lateral view; (c) aedeagus, lateral view; (d) parameres, dorsal view. Scale bars: a, b = 1 mm; c, d = 0.5 mm.
</paragraph>
</caption>
<paragraph id="D543FE592113FFC1FF1AFF63383DFF01" blockId="37.[160,1156,142,411]" box="[160,270,142,168]" pageId="37" pageNumber="644">
<emphasis id="E788224B2113FFC1FF1AFF63383DFF01" bold="true" box="[160,270,142,168]" italics="true" pageId="37" pageNumber="644">Variation</emphasis>
</paragraph>
<paragraph id="D543FE592113FFC1FF1AFF5F3A66FE32" blockId="37.[160,1156,142,411]" pageId="37" pageNumber="644">
Length:
<quantity id="120453BC2113FFC1FF45FF5F38BBFF62" box="[255,392,178,203]" metricMagnitude="-3" metricUnit="m" metricValue="4.95" metricValueMax="5.5" metricValueMin="4.4" pageId="37" pageNumber="644" unit="mm" value="4.95" valueMax="5.5" valueMin="4.4">4.4–5.5 mm</quantity>
. Elytral length:
<quantity id="120453BC2113FFC1FD87FF5F3BF6FF62" box="[573,709,178,203]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="37" pageNumber="644" unit="mm" value="2.7" valueMax="2.9" valueMin="2.5">2.5–2.9 mm</quantity>
. Width:
<quantity id="120453BC2113FFC1FC9FFF5F3A9DFF62" box="[805,942,178,203]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="37" pageNumber="644" unit="mm" value="2.3" valueMax="2.4" valueMin="2.2">2.2–2.4 mm</quantity>
. Protibia with two teeth; spur straight. First protarsomere straight, its length subequal to second and third combined; width ratio of first protarsomere vs second protarsomere: 1; second protarsomere narrow and straight; third and fourth protarsomere elongate and narrow (distinctly longer than wide); fifth protarsomere circular in cross section, as long as third and fourth protarsomere combined. Protarsal claws asymmetrical, shape of inner protarsal claw bifid, apex of basal tooth of inner protarsal claw convex.
</paragraph>
</subSubSection>
<subSubSection id="9DE6ADD22113FFC1FF1AFE3B3D65FD9C" pageId="37" pageNumber="644" type="diagnosis">
<paragraph id="D543FE592113FFC1FF1AFE3B3827FE59" blockId="37.[160,1155,470,565]" box="[160,276,470,496]" pageId="37" pageNumber="644">
<emphasis id="E788224B2113FFC1FF1AFE3B3827FE59" bold="true" box="[160,276,470,496]" italics="true" pageId="37" pageNumber="644">Diagnosis</emphasis>
</paragraph>
<paragraph id="D543FE592113FFC1FF1AFE173D65FD9C" blockId="37.[160,1155,470,565]" pageId="37" pageNumber="644">
<taxonomicName id="12FC85DA2113FFC1FF1AFE173848FDBA" authority="Pacheco &amp; Wipfler &amp; Monné &amp; Ahrens, 2022" authorityName="Pacheco &amp; Wipfler &amp; Monné &amp; Ahrens" authorityYear="2022" box="[160,379,506,531]" class="Insecta" family="Melolonthidae" genus="Symmela" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="644" phylum="Arthropoda" rank="species" species="fuhrmanni" status="sp. nov.">
<emphasis id="E788224B2113FFC1FF1AFE173848FDBA" box="[160,379,506,531]" italics="true" pageId="37" pageNumber="644">Symmela fuhrmanni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FCBB9F302113FFC1FE3BFE1738E4FDBA" box="[385,471,506,531]" pageId="37" pageNumber="644" rank="species">sp. nov.</taxonomicNameLabel>
differs from all other
<taxonomicName id="12FC85DA2113FFC1FD79FE173A4EFDBA" box="[707,893,506,531]" pageId="37" pageNumber="644">
<emphasis id="E788224B2113FFC1FD79FE173A15FDBA" box="[707,806,506,531]" italics="true" pageId="37" pageNumber="644">Symmela</emphasis>
species
</taxonomicName>
by the metatibia which are widest at middle, and the elongated parameres being subparallel in dorsal view.
</paragraph>
</subSubSection>
<subSubSection id="9DE6ADD22113FFC1FF1AFD9C39ECFD5A" pageId="37" pageNumber="644" type="etymology">
<paragraph id="D543FE592113FFC1FF1AFD9C382EFD22" blockId="37.[160,1156,625,755]" box="[160,285,625,651]" pageId="37" pageNumber="644">
<emphasis id="E788224B2113FFC1FF1AFD9C382EFD22" bold="true" box="[160,285,625,651]" italics="true" pageId="37" pageNumber="644">Etymology</emphasis>
</paragraph>
<paragraph id="D543FE592113FFC1FF1AFD7939ECFD5A" blockId="37.[160,1156,625,755]" pageId="37" pageNumber="644">This species is named after Juares Fuhrmann, a Brazillian entomologist who has been making great contributions to Neotropical Melolonthinae taxonomy (noun in genitive case).</paragraph>
</subSubSection>
</treatment>
</document>
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