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<document ID-ISSN="1982-0224" approvalRequired="20" approvalRequired_for_originalDoi="1" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="16" approvalRequired_for_treatments="2" checkinTime="1630377764028" checkinUser="carolina" docAuthor="Paixão, Andrea de Carvalho &amp; Toledo-Piza, Mônica" docDate="2009" docId="4F73A776FA435178FF7AFDE9FC35677C" docLanguage="en" docName="NeotropIchthyol.2009.54.pdf" docOrigin="Neotropical Ichthyology 7 (4)" docStyle="DocumentStyle:85A40B78B715C4497F837D9D78407CE5.4:NeotropIchthyol.2003-2013.journal_article" docStyleId="85A40B78B715C4497F837D9D78407CE5" docStyleName="NeotropIchthyol.2003-2013.journal_article" docStyleVersion="4" docTitle="Lamontichthys stibaros Isbrucker &amp; Nijssen 1978" docType="treatment" docVersion="1" lastPageNumber="535" masterDocId="B34ADF0EFA535168FFD4FFA6FFEA657B" masterDocTitle="Systematics of Lamontichthys Miranda-Ribeiro (Siluriformes: Loricariidae), with the description of two new species" masterLastPageNumber="568" masterPageNumber="519" pageId="16" pageNumber="535" updateTime="1630386870922" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title>Systematics of Lamontichthys Miranda-Ribeiro (Siluriformes: Loricariidae), with the description of two new species</mods:title>
</mods:titleInfo>
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<mods:namePart>Paixão, Andrea de Carvalho</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Toledo-Piza, Mônica</mods:namePart>
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<mods:title>Neotropical Ichthyology</mods:title>
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<mods:date>2009</mods:date>
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<mods:number>7</mods:number>
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<mods:identifier type="ISSN">1982-0224</mods:identifier>
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<subSubSection pageId="16" pageNumber="535" type="nomenclature">
<paragraph blockId="16.[174,740,591,647]" pageId="16" pageNumber="535">
<emphasis bold="true" pageId="16" pageNumber="535">
<taxonomicName authority="Isbrucker &amp; Nijssen, 1978" authorityName="Isbrucker &amp; Nijssen" authorityYear="1978" box="[174,740,591,616]" class="Actinopterygii" family="Loricariidae" genus="Lamontichthys" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="535" phylum="Chordata" rank="species" species="stibaros">
<emphasis bold="true" box="[174,434,591,615]" italics="true" pageId="16" pageNumber="535">Lamontichthys stibaros</emphasis>
Isbrücker &amp; Nijssen, 1978
</taxonomicName>
<figureCitation box="[419,496,623,647]" captionStart="Fig" captionStartId="17.[129,168,1515,1539]" captionTargetBox="[129,1452,174,1502]" captionTargetId="figure-210@17.[129,1453,174,1502]" captionTargetPageId="17" captionText="Fig. 14. Lamontichthys stibaros, INHS 84111, 100.1 mm SL. Dorsal, lateral and ventral views." pageId="16" pageNumber="535">Fig. 14</figureCitation>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="535" type="reference_group">
<paragraph blockId="16.[136,779,687,1095]" pageId="16" pageNumber="535">
<taxonomicName authority="Isbrucker &amp; Nijssen, 1978 b: 66 - 69" authorityName="Isbrucker &amp; Nijssen" authorityPageNumber="66 - 69" authorityYear="1978" box="[136,726,687,712]" class="Actinopterygii" family="Loricariidae" genus="Lamontichthys" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="535" phylum="Chordata" rank="species" species="stibaros">
<emphasis box="[136,373,687,711]" italics="true" pageId="16" pageNumber="535">Lamontichthys stibaros</emphasis>
Isbrücker &amp; Nijssen, 1978b: 66-69
</taxonomicName>
, figs. 8 and 9, 10d, 14, table Ic, Ii-j, IIIb [original description;
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locality:
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, “Prov.
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, rio Amazon system, rio Bobonaza in Chicherota,
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<geoCoordinate box="[522,619,783,807]" degrees="76" direction="west" minutes="38" orientation="longitude" pageId="16" pageNumber="535" precision="925" value="-76.63333">76°38W</geoCoordinate>
, altitude aprox.
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, upper rio
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”];
<bibRefCitation author="Isbrucker, I. J" box="[496,693,815,840]" pageId="16" pageNumber="535" pagination="1 - 181" refId="ref36843" refString="Isbrucker, I. J. H. 1980. Classification and catalogue of the mailed Loricariidae (Pisces, Siluriformes). Verlagen en Technische Genevens, 22: 1 - 181." type="journal article" year="1980">Isbrücker, 1980: 92</bibRefCitation>
[listed]; Ortega &amp; Vari, 1986: 17 [literature compilation;
<collectingCountry box="[713,768,848,871]" name="Peru" pageId="16" pageNumber="535">Peru</collectingCountry>
];
<bibRefCitation author="Burgess, W" box="[166,413,879,904]" pageId="16" pageNumber="535" refId="ref35815" refString="Burgess, W. E. 1989. An atlas of freshwater and marine catfishes. A preliminary survey of the Siluriformes. New Jersey, T. F. H. Publications, 784 p." type="book" year="1989">Burgess, 1989: 426-427</bibRefCitation>
[listed; description];
<bibRefCitation author="Barriga, R" pageId="16" pageNumber="535" pagination="7 - 88" refId="ref35512" refString="Barriga, R. 1991. Peces de agua dulce (Ecuador). Politecnica - Biologia 3, 14 (3): 7 - 88." type="journal article" year="1991">Barriga, 1991: 66</bibRefCitation>
[literature compilation; Oriental
<collectingCountry box="[519,610,911,935]" name="Ecuador" pageId="16" pageNumber="535">Ecuador</collectingCountry>
]; Ferraris &amp; Vari, 1992: 27 [
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listed];
<bibRefCitation author="Eschmeyer, W" box="[425,685,943,968]" pageId="16" pageNumber="535" pagination="1 - 2905" refId="ref36315" refString="Eschmeyer, W. N. 1998. Catalog of Fishes. Center for Biodiversity Research and Information, Spec. Publ. 1. California Academy of Sciences, 1 - 3: 1 - 2905" type="journal article" year="1998">Eschmeyer, 1998: 1611</bibRefCitation>
[listed];
<bibRefCitation author="Isbrucker, I. J" box="[166,370,975,1000]" pageId="16" pageNumber="535" pagination="25 - 32" refId="ref36912" refString="Isbrucker, I. J. H. 2001. Nomenklator der gattungen und arten der harnischwelse, familie Loricariidae Rafinesque, 1815 (Teleostei, Ostariophysi). Datz-Soderheft Harnischwelse, 2: 25 - 32." type="journal article" year="2001">Isbrücker, 2001: 29</bibRefCitation>
[listed]; Ferraris Jr., 2003: 336 [listed]; Ferraris Jr., 2007:264 [listed];
<bibRefCitation author="Sullivan, J" box="[462,660,1007,1031]" pageId="16" pageNumber="535" pagination="636 - 662" refId="ref39057" refString="Sullivan, J. P., J. G. Lundberg &amp; M. Hardman. 2006. A phylogenetic analysis of the major groups of catfishes (Teleostei: Siluriformes) using rag 1 and rag 2 nuclear gene sequences. Molecular Phylogenetics and Evolution, 41: 636 - 662." type="journal article" year="2006">
Sullivan
<emphasis box="[549,590,1007,1031]" italics="true" pageId="16" pageNumber="535">et al</emphasis>
., 2006
</bibRefCitation>
: [Siluriform higher-level relationships; molecular data, specimen not examined].
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="535" type="diagnosis">
<paragraph blockId="16.[136,779,1135,1863]" pageId="16" pageNumber="535">
<emphasis bold="true" box="[136,256,1135,1159]" pageId="16" pageNumber="535">Diagnosis.</emphasis>
<taxonomicName authorityName="Isbrucker &amp; Nijssen" authorityYear="1978" box="[266,528,1135,1159]" class="Actinopterygii" family="Loricariidae" genus="Lamontichthys" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="535" phylum="Chordata" rank="species" species="stibaros">
<emphasis box="[266,528,1135,1159]" italics="true" pageId="16" pageNumber="535">Lamontichthys stibaros</emphasis>
</taxonomicName>
differs from all other species of the genus, with the exception of
<taxonomicName box="[614,779,1168,1191]" class="Actinopterygii" family="Loricariidae" genus="Lamontichthys" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="535" phylum="Chordata" rank="species" species="avacanoeiro">
<emphasis box="[614,629,1168,1191]" italics="true" pageId="16" pageNumber="535">L</emphasis>
.
<emphasis box="[644,779,1168,1191]" italics="true" pageId="16" pageNumber="535">avacanoeiro</emphasis>
</taxonomicName>
and
<taxonomicName box="[183,315,1199,1223]" class="Actinopterygii" family="Loricariidae" genus="Lamontichthys" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="535" phylum="Chordata" rank="species" species="parakana">
<emphasis box="[183,198,1200,1223]" italics="true" pageId="16" pageNumber="535">L</emphasis>
.
<emphasis box="[212,315,1199,1223]" italics="true" pageId="16" pageNumber="535">parakana</emphasis>
</taxonomicName>
, in the lack of plates on the anterior tip of the snout (
<emphasis box="[251,273,1232,1255]" italics="true" pageId="16" pageNumber="535">vs</emphasis>
. anterior portion of snout totally covered with plates or with an inconspicuous naked area), in the teeth with long cusps (
<emphasis box="[264,286,1296,1319]" italics="true" pageId="16" pageNumber="535">vs</emphasis>
. short) (
<figureCitation box="[376,440,1295,1319]" captionStart="Fig" captionStartId="4.[816,855,1008,1032]" captionTargetBox="[818,1449,483,990]" captionTargetId="figure-417@4.[814,1457,470,1002]" captionTargetPageId="4" captionText="Fig. 3. Lateral view, distal portion to top, of left premaxillary tooth of Lamontichthys avacanoeiro (MZUSP 61974, 113.7 mm SL, paratype) (a), and L. filamentosus (LACM 41741-9, 119.4 mm SL) (b). Detail of cusps in anterior view, distal tip to left. Scale bar = 0.5 mm." pageId="16" pageNumber="535">Fig. 3</figureCitation>
), and in the lower lip semi-oval shaped (
<emphasis box="[224,246,1328,1351]" italics="true" pageId="16" pageNumber="535">vs</emphasis>
. semicircular) (
<figureCitation box="[411,474,1327,1351]" captionStart="Fig" captionStartId="4.[816,855,1008,1032]" captionTargetBox="[818,1449,483,990]" captionTargetId="figure-417@4.[814,1457,470,1002]" captionTargetPageId="4" captionText="Fig. 3. Lateral view, distal portion to top, of left premaxillary tooth of Lamontichthys avacanoeiro (MZUSP 61974, 113.7 mm SL, paratype) (a), and L. filamentosus (LACM 41741-9, 119.4 mm SL) (b). Detail of cusps in anterior view, distal tip to left. Scale bar = 0.5 mm." pageId="16" pageNumber="535">Fig. 3</figureCitation>
).
</paragraph>
<paragraph blockId="16.[136,779,1135,1863]" pageId="16" pageNumber="535">
<taxonomicName authorityName="Isbrucker &amp; Nijssen" authorityYear="1978" box="[174,427,1359,1383]" class="Actinopterygii" family="Loricariidae" genus="Lamontichthys" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="535" phylum="Chordata" rank="species" species="stibaros">
<emphasis box="[174,427,1359,1383]" italics="true" pageId="16" pageNumber="535">Lamontichthys stibaros</emphasis>
</taxonomicName>
differs from
<taxonomicName box="[570,733,1360,1383]" class="Actinopterygii" family="Loricariidae" genus="Lamontichthys" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="535" phylum="Chordata" rank="species" species="avacanoeiro">
<emphasis box="[570,585,1360,1383]" italics="true" pageId="16" pageNumber="535">L</emphasis>
.
<emphasis box="[598,733,1360,1383]" italics="true" pageId="16" pageNumber="535">avacanoeiro</emphasis>
</taxonomicName>
and
<taxonomicName box="[136,266,1391,1415]" class="Actinopterygii" family="Loricariidae" genus="Lamontichthys" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="535" phylum="Chordata" rank="species" species="parakana">
<emphasis box="[136,151,1392,1415]" italics="true" pageId="16" pageNumber="535">L</emphasis>
.
<emphasis box="[163,266,1391,1415]" italics="true" pageId="16" pageNumber="535">parakana</emphasis>
</taxonomicName>
in the longer first branched dorsal-fin ray (29.7- 36.8%,
<emphasis box="[213,234,1424,1447]" italics="true" pageId="16" pageNumber="535">vs</emphasis>
. 22.6-31.0% SL), in specimens larger than
<quantity box="[666,738,1423,1447]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="16" pageNumber="535" unit="mm" value="70.0">70 mm</quantity>
SL. It further differs from
<taxonomicName box="[396,565,1456,1479]" class="Actinopterygii" family="Loricariidae" genus="Lamontichthys" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="535" phylum="Chordata" rank="species" species="avacanoeiro">
<emphasis box="[396,411,1456,1479]" italics="true" pageId="16" pageNumber="535">L</emphasis>
.
<emphasis box="[428,565,1456,1479]" italics="true" pageId="16" pageNumber="535">avacanoeiro</emphasis>
</taxonomicName>
in the longer first branched pectoral-fin ray (21.7-25.1%,
<emphasis box="[539,561,1488,1511]" italics="true" pageId="16" pageNumber="535">vs</emphasis>
. 19.2-22.0% SL) and from
<taxonomicName box="[190,314,1519,1543]" class="Actinopterygii" family="Loricariidae" genus="Lamontichthys" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="535" phylum="Chordata" rank="species" species="parakana">
<emphasis box="[190,204,1520,1543]" italics="true" pageId="16" pageNumber="535">L</emphasis>
.
<emphasis box="[215,314,1519,1543]" italics="true" pageId="16" pageNumber="535">parakana</emphasis>
</taxonomicName>
in the narrower head (89.1-101.4%,
<emphasis box="[680,701,1520,1543]" italics="true" pageId="16" pageNumber="535">vs</emphasis>
. 101.3- 106.1% HL).
</paragraph>
<paragraph blockId="16.[136,779,1135,1863]" pageId="16" pageNumber="535">
The presence of 19 to 20 lateral plates along the mid-ventral series (
<emphasis box="[206,227,1616,1639]" italics="true" pageId="16" pageNumber="535">vs</emphasis>
. 14 to 18) and the longer dentary (18.6-23.2%,
<emphasis box="[694,715,1616,1639]" italics="true" pageId="16" pageNumber="535">vs</emphasis>
. 11.5- 17.3% HL) in specimens larger then
<quantity box="[495,566,1647,1671]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="16" pageNumber="535" unit="mm" value="70.0">70 mm</quantity>
SL also distinguishes
<taxonomicName authorityName="Isbrucker &amp; Nijssen" authorityYear="1978" box="[136,246,1679,1703]" class="Actinopterygii" family="Loricariidae" genus="Lamontichthys" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="535" phylum="Chordata" rank="species" species="stibaros">
<emphasis box="[136,151,1680,1703]" italics="true" pageId="16" pageNumber="535">L</emphasis>
.
<emphasis box="[162,246,1679,1703]" italics="true" pageId="16" pageNumber="535">stibaros</emphasis>
</taxonomicName>
from
<taxonomicName box="[306,464,1679,1703]" class="Actinopterygii" family="Loricariidae" genus="Lamontichthys" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="535" phylum="Chordata" rank="species" species="filamentosus">
<emphasis box="[306,321,1680,1703]" italics="true" pageId="16" pageNumber="535">L</emphasis>
.
<emphasis box="[332,464,1679,1703]" italics="true" pageId="16" pageNumber="535">filamentosus</emphasis>
</taxonomicName>
and
<taxonomicName box="[512,613,1679,1703]" class="Actinopterygii" family="Loricariidae" genus="Lamontichthys" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="535" phylum="Chordata" rank="species" species="llanero">
<emphasis box="[512,527,1680,1703]" italics="true" pageId="16" pageNumber="535">L</emphasis>
.
<emphasis box="[538,613,1679,1703]" italics="true" pageId="16" pageNumber="535">llanero</emphasis>
</taxonomicName>
.
<taxonomicName authorityName="Isbrucker &amp; Nijssen" authorityYear="1978" class="Actinopterygii" family="Loricariidae" genus="Lamontichthys" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="535" phylum="Chordata" rank="species" species="stibaros">
<emphasis italics="true" pageId="16" pageNumber="535">Lamontichthys stibaros</emphasis>
</taxonomicName>
differs from
<taxonomicName box="[357,520,1711,1735]" class="Actinopterygii" family="Loricariidae" genus="Lamontichthys" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="535" phylum="Chordata" rank="species" species="maracaibero">
<emphasis box="[357,372,1712,1735]" italics="true" pageId="16" pageNumber="535">L</emphasis>
.
<emphasis box="[384,520,1711,1735]" italics="true" pageId="16" pageNumber="535">maracaibero</emphasis>
</taxonomicName>
in having more than one plate between the base of the last pelvic-fin ray and the anterior border of the anus (
<emphasis box="[350,372,1776,1799]" italics="true" pageId="16" pageNumber="535">vs</emphasis>
. one plate) (
<figureCitation box="[510,585,1775,1799]" captionStart="Fig" captionStartId="2.[136,175,913,937]" captionTargetBox="[142,776,177,896]" captionTargetId="figure-729@2.[136,779,172,902]" captionTargetPageId="2" captionText="Fig. 1. Ventral view of head and anterior portion of body of Lamontichthys filamentosus (INPA 17927, 79.4 mm SL). Odontods, plates on upper lip and papillae on lower lip not represented. Scale bar = 4 mm." pageId="16" pageNumber="535">Figs. 1</figureCitation>
and
<figureCitation box="[640,654,1775,1799]" captionStart="Fig" captionStartId="5.[129,170,1199,1223]" captionTargetBox="[129,772,618,1189]" captionTargetId="figure-184@5.[129,772,618,1189]" captionTargetPageId="5" captionText="Fig. 5. Ventral view of pelvic region of Lamontichthys maracaibero (MCNG 3592, 154.9 mm SL). Arrow points to single plate from base of last pelvic fin-ray to margin of anus." pageId="16" pageNumber="535">5</figureCitation>
) and in the uniform color pattern of the fins (
<emphasis box="[468,489,1808,1831]" italics="true" pageId="16" pageNumber="535">vs</emphasis>
. with brown and gray spots). Morphometric and meristic data are presented in
<tableCitation box="[647,725,1839,1863]" captionStart="Table 3" captionStartId="15.[129,192,177,201]" captionText="Table 3. Morphometric and meristic data for Lamontichthys parakana (n = 3) and L. stibaros. SD = standard deviation. Range of L. parakana includes holotype and paratypes." pageId="16" pageNumber="535">Table 3</tableCitation>
.
</paragraph>
<paragraph blockId="16.[136,779,1903,1960]" pageId="16" pageNumber="535">
<emphasis bold="true" box="[136,411,1903,1927]" pageId="16" pageNumber="535">Geographic distribution.</emphasis>
Upper rio
<collectingRegion box="[519,630,1903,1927]" country="Venezuela" name="Amazonas" pageId="16" pageNumber="535">Amazonas</collectingRegion>
basin (
<figureCitation box="[703,766,1903,1927]" captionStart="Fig" captionStartId="4.[136,175,1744,1768]" captionTargetBox="[140,782,1263,1728]" captionTargetId="figure-347@4.[140,784,1259,1728]" captionTargetPageId="4" captionText="Fig. 2. Map of central and northern portions of SouthAmerica showing geographic distribution of the species of Lamontichthys (numbers indicate type localities): 1. L. avacanoeiro (lozenge), 2. L. filamentosus (triangle), 3. L. llanero (rudder), 4. L. maracaibero (open squares), 5. L. parakana (number 5), and 6. L. stibaros (solid squares). Some symbols represent more than one locality or lot of specimens." pageId="16" pageNumber="535">Fig. 2</figureCitation>
).
<taxonomicName authorityName="Isbrucker &amp; Nijssen" authorityYear="1978" box="[136,425,1935,1959]" class="Actinopterygii" family="Loricariidae" genus="Lamontichthys" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="535" phylum="Chordata" rank="species" species="stibaros">
<emphasis box="[136,425,1935,1959]" italics="true" pageId="16" pageNumber="535">Lamontichthys stibaros</emphasis>
</taxonomicName>
occurs in sympatry with
<emphasis box="[755,770,1936,1959]" italics="true" pageId="16" pageNumber="535">L</emphasis>
.
</paragraph>
<paragraph blockId="16.[816,1459,177,233]" pageId="16" pageNumber="535">
<emphasis box="[816,942,177,201]" italics="true" pageId="16" pageNumber="535">filamentous</emphasis>
in the drainages of río Santiago, río Marañón and
<collectingRegion box="[860,981,209,233]" country="Peru" name="Ucayali" pageId="16" pageNumber="535">río Ucayali</collectingRegion>
, in
<collectingCountry box="[1016,1069,210,233]" name="Peru" pageId="16" pageNumber="535">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="535" type="materials_examined">
<paragraph blockId="16.[816,1460,266,519]" pageId="16" pageNumber="535">
<emphasis bold="true" box="[816,1017,266,288]" pageId="16" pageNumber="535">Material examined</emphasis>
(
<specimenCount box="[1033,1173,266,288]" pageId="16" pageNumber="535" type="generic">25 specimens</specimenCount>
;
<quantity box="[1186,1339,267,288]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" metricValueMax="1.728" metricValueMin="0.27199999999999996" pageId="16" pageNumber="535" unit="mm" value="100.0" valueMax="172.8" valueMin="27.2">27.2-172.8 mm</quantity>
SL):
<materialsCitation country="Peru" location="Amazonas" pageId="16" pageNumber="535" specimenCount="1">
<emphasis bold="true" box="[1401,1458,267,288]" pageId="16" pageNumber="535">
<collectingCountry box="[1401,1455,267,288]" name="Peru" pageId="16" pageNumber="535">Peru</collectingCountry>
:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:4F73A776FA435178FF7AFDE9FC35677C:C20540BBFA435178FCE4FE81FC716447" box="[816,923,295,316]" country="Peru" name="Amazonas" pageId="16" pageNumber="535">Amazonas</location>
</materialsCitation>
:
<materialsCitation box="[936,1376,295,317]" collectionCode="LACM" country="Peru" location="rio Santiago" pageId="16" pageNumber="535" specimenCode="LACM 39897-1, 3, 42.2, 89.5" specimenCount="1">
<specimenCode box="[936,1099,295,316]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="16" pageNumber="535" type="Museum">LACM 39897-1</specimenCode>
,
<specimenCode box="[1113,1126,295,316]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="16" pageNumber="535" type="Museum">3</specimenCode>
,
<specimenCode box="[1140,1186,295,317]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="16" pageNumber="535" type="Museum">42.2</specimenCode>
-
<specimenCode box="[1193,1237,295,317]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="16" pageNumber="535" type="Museum">89.5</specimenCode>
,
<location LSID="urn:lsid:plazi:treatment:4F73A776FA435178FF7AFDE9FC35677C:C20540BBFA435178FB36FE81FA8A6446" box="[1250,1376,295,317]" country="Peru" name="rio Santiago" pageId="16" pageNumber="535">río Santiago</location>
</materialsCitation>
;
<materialsCitation collectionCode="LACM" country="Peru" location="rio Santiago" municipality="Maranon" pageId="16" pageNumber="535" specimenCode="LACM 39884-3, 2, 100.2, 120.1" specimenCount="1">
<specimenCode collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="16" pageNumber="535" type="Museum">LACM 39884-3</specimenCode>
,
<specimenCode box="[910,924,324,345]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="16" pageNumber="535" type="Museum">2</specimenCode>
,
<specimenCode box="[934,992,324,345]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="16" pageNumber="535" type="Museum">100.2</specimenCode>
-
<specimenCode box="[999,1055,324,345]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="16" pageNumber="535" type="Museum">120.1</specimenCode>
,
<collectingMunicipality box="[1064,1154,324,346]" pageId="16" pageNumber="535">Marañón</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:4F73A776FA435178FF7AFDE9FC35677C:C20540BBFA435178FB58FEE2FAEC6421" box="[1164,1286,324,346]" country="Peru" municipality="Maranon" name="rio Santiago" pageId="16" pageNumber="535">río Santiago</location>
</materialsCitation>
;
<materialsCitation collectionCode="LACM" collectorName="Maranon" country="Peru" location="rio Tujushiku." municipality="rio Cenepa" pageId="16" pageNumber="535" specimenCode="LACM 39647-1, 1, 172.6" specimenCount="1" stateProvince="Ayacucho">
<specimenCode box="[1296,1455,324,346]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="16" pageNumber="535" type="Museum">LACM 39647-1</specimenCode>
,
<specimenCode box="[816,828,353,374]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="16" pageNumber="535" type="Museum">1</specimenCode>
,
<specimenCode box="[841,898,353,375]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="16" pageNumber="535" type="Museum">172.6</specimenCode>
,
<collectorName box="[909,999,353,375]" pageId="16" pageNumber="535">Marañón</collectorName>
,
<collectingMunicipality box="[1010,1120,353,375]" pageId="16" pageNumber="535">río Cenepa</collectingMunicipality>
, close to
<location LSID="urn:lsid:plazi:treatment:4F73A776FA435178FF7AFDE9FC35677C:C20540BBFA435178FB68FEC7FAAD640C" box="[1212,1351,353,375]" country="Peru" municipality="rio Cenepa" name="rio Tujushiku." pageId="16" pageNumber="535" stateProvince="Ayacucho">río Tujushiku.</location>
<collectingRegion box="[1356,1457,353,375]" country="Peru" name="Ayacucho" pageId="16" pageNumber="535">Ayacucho</collectingRegion>
</materialsCitation>
:
<materialsCitation accessionNumber="ROM 47093" collectionCode="ROM" country="Peru" location="between rio Mantaro and Tambo" pageId="16" pageNumber="535" specimenCode="2, 122.6, 150.1" specimenCount="1" stateProvince="Apurimac">
<accessionNumber box="[816,937,382,403]" httpUri="http://www.ncbi.nlm.nih.gov/protein/ROM47093" pageId="16" pageNumber="535">
<collectionCode box="[816,870,382,403]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="16" pageNumber="535" type="Museum">ROM</collectionCode>
47093
</accessionNumber>
,
<specimenCode box="[944,958,382,403]" collectionCode="ROM" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="16" pageNumber="535" type="Museum">2</specimenCode>
,
<specimenCode box="[966,1022,381,403]" collectionCode="ROM" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="16" pageNumber="535" type="Museum">122.6</specimenCode>
-
<specimenCode box="[1028,1082,381,403]" collectionCode="ROM" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="16" pageNumber="535" type="Museum">150.1</specimenCode>
, near to confluence
<location LSID="urn:lsid:plazi:treatment:4F73A776FA435178FF7AFDE9FC35677C:C20540BBFA435178FB25FED8FC7564CB" country="Peru" name="between rio Mantaro and Tambo" pageId="16" pageNumber="535" stateProvince="Apurimac">between río Mantaro and Tambo</location>
<collectingRegion box="[935,1036,410,432]" country="Peru" name="Apurimac" pageId="16" pageNumber="535">Apurimac</collectingRegion>
.
<collectingRegion box="[1048,1128,410,432]" country="Peru" name="Ucayali" pageId="16" pageNumber="535">Ucayali</collectingRegion>
</materialsCitation>
:
<materialsCitation collectionCode="FMNH" country="Peru" location="rio San Alejandro" pageId="16" pageNumber="535" specimenCode="FMNH 84137, 7, 27.2, 58.2" specimenCount="1">
<specimenCode box="[1140,1282,411,432]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="535" type="Museum">FMNH 84137</specimenCode>
,
<specimenCode box="[1295,1309,411,432]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="535" type="Museum">7</specimenCode>
,
<specimenCode box="[1323,1368,410,432]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="535" type="Museum">27.2</specimenCode>
-
<specimenCode box="[1374,1419,410,432]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="535" type="Museum">58.2</specimenCode>
,
<location LSID="urn:lsid:plazi:treatment:4F73A776FA435178FF7AFDE9FC35677C:C20540BBFA435178FA4CFE3CFC5764B6" country="Peru" name="rio San Alejandro" pageId="16" pageNumber="535">río San Alejandro</location>
</materialsCitation>
;
<materialsCitation collectionCode="FMNH" country="Peru" location="rio San Alejandro" pageId="16" pageNumber="535" specimenCode="FMNH 84131, 2, 43.8, 47.3, FMNH 84112, 3, 63.3, 86.6" specimenCount="1">
<specimenCode box="[965,1103,439,461]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="535" type="Museum">FMNH 84131</specimenCode>
,
<specimenCode box="[1112,1126,439,460]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="535" type="Museum">2</specimenCode>
,
<specimenCode box="[1135,1180,439,460]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="535" type="Museum">43.8</specimenCode>
-
<specimenCode box="[1187,1232,439,460]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="535" type="Museum">47.3</specimenCode>
;
<specimenCode box="[1240,1377,439,461]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="535" type="Museum">FMNH 84112</specimenCode>
,
<specimenCode box="[1385,1398,439,460]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="535" type="Museum">3</specimenCode>
,
<specimenCode box="[1408,1452,439,461]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="535" type="Museum">63.3</specimenCode>
-
<specimenCode box="[816,861,468,490]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="535" type="Museum">86.6</specimenCode>
,
<location LSID="urn:lsid:plazi:treatment:4F73A776FA435178FF7AFDE9FC35677C:C20540BBFA435178FCB3FE72FBFD6491" box="[871,1047,468,490]" country="Peru" name="rio San Alejandro" pageId="16" pageNumber="535">río San Alejandro</location>
</materialsCitation>
;
<materialsCitation collectionCode="FMNH" country="Peru" location="rio San Alejandro" pageId="16" pageNumber="535" specimenCode="FMNH 84111, 6, 49.3, 100.1, 1" specimenCount="1">
<specimenCode box="[1057,1194,468,489]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="535" type="Museum">FMNH 84111</specimenCode>
,
<specimenCode box="[1204,1218,468,490]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="535" type="Museum">6</specimenCode>
,
<specimenCode box="[1229,1275,468,489]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="535" type="Museum">49.3</specimenCode>
-
<specimenCode box="[1281,1337,468,489]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="535" type="Museum">100.1</specimenCode>
,
<specimenCode box="[1347,1359,468,489]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="535" type="Museum">1</specimenCode>
cs*, 62.4,
<location LSID="urn:lsid:plazi:treatment:4F73A776FA435178FF7AFDE9FC35677C:C20540BBFA435178FCE4FE57FC36677C" box="[816,988,497,519]" country="Peru" name="rio San Alejandro" pageId="16" pageNumber="535">río San Alejandro</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>