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<document id="3D611B4BF36D3554EAEC7A53BB954053" ID-DOI="10.5281/zenodo.276621" ID-GBIF-Dataset="a6eb9c5a-9b0b-47df-a207-6c5ee54acebd" ID-ISSN="1175-5326" ID-Zenodo-Dep="276621" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460225034079" checkinUser="plazi" docAuthor="Latha, Mathews, Cuccodoro, Giulio &amp; Sabu, Thomas K." docDate="2011" docId="03FF87EFFFD3B850FF7230E4E3B8FE43" docLanguage="en" docName="zt02745p029.pdf" docOrigin="Zootaxa 2745" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ochicanthon ceylonicus Cuccodoro, sp. nov." docType="treatment" docVersion="8" lastPageNumber="6" masterDocId="FFC6FF97FFD1B855FFE53643E06BFFEF" masterDocTitle="Taxonomy of the dung beetle genus Ochicanthon Vaz-de-Mello (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian subcontinent, with notes on distribution patterns and flightlessness" masterLastPageNumber="29" masterPageNumber="1" pageNumber="3" updateTime="1698238191473" updateUser="plazi">
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<mods:title id="732B693B28E758E5B44D4B2FD68CB8F4">Taxonomy of the dung beetle genus Ochicanthon Vaz-de-Mello (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian subcontinent, with notes on distribution patterns and flightlessness</mods:title>
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<mods:namePart id="32CC5C59EC7B9DFF43FF73ED3BAD00B4">Latha, Mathews</mods:namePart>
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<mods:namePart id="4A12ECADBBF6C7FE05ABC6F5873ECF26">Cuccodoro, Giulio</mods:namePart>
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<mods:namePart id="5FB667B2B264171D5D151EE3382B3588">Sabu, Thomas K.</mods:namePart>
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<mods:title id="09D06F1F4A643AC91C3642DB86E8260D">Zootaxa</mods:title>
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<mods:date id="446E6C33A0C6A3C8E364DA81F2036D81">2011</mods:date>
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<mods:number id="E4B18FBB419361BCEDB3D87CD57F2420">2745</mods:number>
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<treatment id="03FF87EFFFD3B850FF7230E4E3B8FE43" ID-DOI="http://doi.org/10.5281/zenodo.6182038" ID-GBIF-Taxon="119386108" ID-Zenodo-Dep="6182038" LSID="urn:lsid:plazi:treatment:03FF87EFFFD3B850FF7230E4E3B8FE43" httpUri="http://treatment.plazi.org/id/03FF87EFFFD3B850FF7230E4E3B8FE43" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3">
<subSubSection id="C34C6572FFD3B857FF7230E4E17AF90B" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BE936F9FFD3B857FF7230E4E2F3F92F" blockId="2.[151,664,1703,1764]" box="[151,664,1703,1728]" pageId="2" pageNumber="3">
<heading id="D0A18195FFD3B857FF7230E4E2F3F92F" bold="true" box="[151,664,1703,1728]" fontSize="10" level="2" pageId="2" pageNumber="3" reason="2">
<emphasis id="B922EAEBFFD3B857FF7230E4E2F3F92F" bold="true" box="[151,664,1703,1728]" pageId="2" pageNumber="3">
<taxonomicName id="4C564D7AFFD3B857FF7230E4E25CF92F" ID-CoL="48CV5" authority="Cuccodoro" authorityName="Cuccodoro" box="[151,567,1703,1728]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ceylonicus" status="sp. nov.">
<emphasis id="B922EAEBFFD3B857FF7230E4E1C5F92F" bold="true" box="[151,430,1703,1728]" italics="true" pageId="2" pageNumber="3">Ochicanthon ceylonicus</emphasis>
Cuccodoro
</taxonomicName>
<taxonomicNameLabel id="A2115790FFD3B857FDDB30EBE2F3F92F" box="[574,664,1704,1728]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE936F9FFD3B857FF72308FE17AF90B" blockId="2.[151,664,1703,1764]" box="[151,273,1740,1764]" pageId="2" pageNumber="3">
(
<figureCitation id="136D2A7CFFD3B857FF7A308FE161F90B" box="[159,266,1740,1764]" captionStart="FIGURES 6 8" captionStartId="4.[151,264,1628,1651]" captionTargetBox="[162,1425,184,1605]" captionTargetId="figure@4.[162,1427,184,1606]" captionTargetPageId="4" captionText="FIGURES 6 8. Ochicanthon ceylonicus (holotype). Dorsal habitus (6); base of pronotum and elytra (7); mesosternum and metasternum (8). Scale bars = 1 mm." httpUri="https://zenodo.org/record/276623/files/figure.png" pageId="2" pageNumber="3">Figs. 68</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34C6572FFD3B851FF723150E523F84C" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="description">
<paragraph id="8BE936F9FFD3B851FF723150E084F8B4" blockId="2.[151,1437,1811,2016]" lastBlockId="4.[151,1436,1715,2027]" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
<emphasis id="B922EAEBFFD3B857FF723150E142F8C3" bold="true" box="[151,297,1811,1836]" pageId="2" pageNumber="3">Description.</emphasis>
Body (
<figureCitation id="136D2A7CFFD3B857FE9A3157E1AEF8C3" box="[383,453,1812,1836]" captionStart="FIGURES 6 8" captionStartId="4.[151,264,1628,1651]" captionTargetBox="[162,1425,184,1605]" captionTargetId="figure@4.[162,1427,184,1606]" captionTargetPageId="4" captionText="FIGURES 6 8. Ochicanthon ceylonicus (holotype). Dorsal habitus (6); base of pronotum and elytra (7); mesosternum and metasternum (8). Scale bars = 1 mm." httpUri="https://zenodo.org/record/276623/files/figure.png" pageId="2" pageNumber="3">Fig. 6</figureCitation>
) predominantly black. Dorsal surface of head with annular setiferous punctures separated by intervals almost equal to their diameter. Pronotum in lateral view weakly convex, outline at base almost in line with that of elytral suture; laterobasal paramarginal ridge absent. Pronotal punctation (
<figureCitation id="136D2A7CFFD3B857FB69311FE4BBF89B" box="[1164,1232,1884,1908]" captionStart="FIGURES 6 8" captionStartId="4.[151,264,1628,1651]" captionTargetBox="[162,1425,184,1605]" captionTargetId="figure@4.[162,1427,184,1606]" captionTargetPageId="4" captionText="FIGURES 6 8. Ochicanthon ceylonicus (holotype). Dorsal habitus (6); base of pronotum and elytra (7); mesosternum and metasternum (8). Scale bars = 1 mm." httpUri="https://zenodo.org/record/276623/files/figure.png" pageId="2" pageNumber="3">Fig. 7</figureCitation>
) consists of annular setiferous punctures separated by interval equal to their diameter, becomes finer anteromedially; setae indistinct, straight and recumbent. Elytra moderately convex in lateral view; suture not raised; striae shallow, narrow, consisting of chains of oval depressions joined by straight sulci (
<figureCitation id="136D2A7CFFD3B857FC8E318BE3C4F80F" box="[875,943,1992,2016]" captionStart="FIGURES 6 8" captionStartId="4.[151,264,1628,1651]" captionTargetBox="[162,1425,184,1605]" captionTargetId="figure@4.[162,1427,184,1606]" captionTargetPageId="4" captionText="FIGURES 6 8. Ochicanthon ceylonicus (holotype). Dorsal habitus (6); base of pronotum and elytra (7); mesosternum and metasternum (8). Scale bars = 1 mm." httpUri="https://zenodo.org/record/276623/files/figure.png" pageId="2" pageNumber="3">Fig. 7</figureCitation>
); interstriae slightly convex, uneven, finely punctate, irregular; background microsculpture smooth; setae fine, straight and recumbent; lateral ridge posteriorly extended to level of apex of second stria. Mesosternum impunctate, meso-metasternal line arcuate, metasternal shield coarsely punctate along the sides and finely punctate in the centre (
<figureCitation id="136D2A7CFFD5B851FC3F30B8E474F8FC" box="[986,1055,1787,1811]" captionStart="FIGURES 6 8" captionStartId="4.[151,264,1628,1651]" captionTargetBox="[162,1425,184,1605]" captionTargetId="figure@4.[162,1427,184,1606]" captionTargetPageId="4" captionText="FIGURES 6 8. Ochicanthon ceylonicus (holotype). Dorsal habitus (6); base of pronotum and elytra (7); mesosternum and metasternum (8). Scale bars = 1 mm." httpUri="https://zenodo.org/record/276623/files/figure.png" pageId="4" pageNumber="5">Fig. 8</figureCitation>
). Pygidium covered with shallow setiferous punctures becoming finer and sparser ventrally. Protrochanter with outer margin arcuate.
<collectingCountry id="F3417669FFD5B851FAF2315CE524F8D8" box="[1303,1359,1823,1847]" name="India" pageId="4" pageNumber="5">Hind</collectingCountry>
wings present.
</paragraph>
<caption id="DF296671FFD2B856FF7231E2E40BF838" httpUri="https://zenodo.org/record/276622/files/figure.png" pageId="3" pageNumber="4" targetBox="[215,1365,195,1896]" targetPageId="3">
<paragraph id="8BE936F9FFD2B856FF7231E2E40BF838" blockId="3.[151,1436,1953,2007]" pageId="3" pageNumber="4">
<emphasis id="B922EAEBFFD2B856FF7231E2E156F858" bold="true" box="[151,317,1953,1976]" pageId="3" pageNumber="4">FIGURES 15.</emphasis>
<taxonomicName id="4C564D7AFFD2B856FEA231E1E258F857" box="[327,563,1954,1976]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="besucheti">
<emphasis id="B922EAEBFFD2B856FEA231E1E258F857" box="[327,563,1954,1976]" italics="true" pageId="3" pageNumber="4">Ochicanthon besucheti</emphasis>
</taxonomicName>
(holotype). Dorsal habitus (1); base of pronotum and elytra (2); mesosternum and metasternum (3); aedeagus, ventral view (4); aedeagus, lateral left view (5). Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="DF296671FFD5B851FF72301FE265F97E" httpUri="https://zenodo.org/record/276623/files/figure.png" pageId="4" pageNumber="5" targetBox="[162,1425,184,1605]" targetPageId="4">
<paragraph id="8BE936F9FFD5B851FF72301FE265F97E" blockId="4.[151,1435,1628,1682]" pageId="4" pageNumber="5">
<emphasis id="B922EAEBFFD5B851FF72301FE156F99D" bold="true" box="[151,317,1628,1651]" pageId="4" pageNumber="5">FIGURES 68.</emphasis>
<taxonomicName id="4C564D7AFFD5B851FEA0301EE257F99C" box="[325,572,1629,1651]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis id="B922EAEBFFD5B851FEA0301EE257F99C" box="[325,572,1629,1651]" italics="true" pageId="4" pageNumber="5">Ochicanthon ceylonicus</emphasis>
</taxonomicName>
(holotype). Dorsal habitus (6); base of pronotum and elytra (7); mesosternum and metasternum (8). Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="8BE936F9FFD5B851FF203124E11EF890" blockId="4.[151,1436,1715,2027]" box="[197,373,1895,1919]" pageId="4" pageNumber="5">Male unknown.</paragraph>
<paragraph id="8BE936F9FFD5B851FF2031CFE523F84C" blockId="4.[151,1436,1715,2027]" box="[197,1352,1931,1956]" pageId="4" pageNumber="5">
<emphasis id="B922EAEBFFD5B851FF2031CFE11FF84B" bold="true" box="[197,372,1932,1956]" pageId="4" pageNumber="5">Measurements</emphasis>
(mm; n = 3): TL = 4.24.4; BW = 2.42.5; PL = 1.11.2; PW = 2.02.1; EL = 2.12.2.
</paragraph>
</subSubSection>
<subSubSection id="C34C6572FFD5B850FF2031F3E343FF40" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="8BE936F9FFD5B851FF2031F3E414F804" blockId="4.[151,1436,1715,2027]" pageId="4" pageNumber="5">
<emphasis id="B922EAEBFFD5B851FF2031F3E11EF827" bold="true" box="[197,373,1967,1992]" pageId="4" pageNumber="5">
<typeStatus id="54ED885BFFD5B851FF2031F3E16BF827" box="[197,256,1968,1992]" pageId="4" pageNumber="5">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54ED885BFFD5B851FE9831ECE18EF828" box="[381,485,1967,1991]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
(female,
<collectionCode id="ED47AE3CFFD5B851FDB131ECE2DBF828" box="[596,688,1967,1991]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="4" pageNumber="5">MHNG</collectionCode>
): “
<collectingCountry id="F3417669FFD5B851FD2B31ECE34BF828" box="[718,800,1967,1991]" name="Sri Lanka" pageId="4" pageNumber="5">Ceylan</collectingCountry>
[
<collectingCountry id="F3417669FFD5B851FCCA31ECE3CCF828" box="[815,935,1967,1991]" name="Sri Lanka" pageId="4" pageNumber="5">Sri Lanka</collectingCountry>
], Central, Peradeniya,
<quantity id="4CAE9B1CFFD5B851FB4C31ECE487F828" box="[1193,1260,1967,1991]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="4" pageNumber="5" unit="m" value="550.0">550m</quantity>
,
<date id="FFE81039FFD5B851FB1C31ECE50EF828" box="[1273,1381,1967,1991]" pageId="4" pageNumber="5" value="1970-01-19">19.i.1970</date>
, leg. Besuchet, Löbl &amp; Mussard, #10” [sifting in forest near experimental agricultural station].
</paragraph>
<paragraph id="8BE936F9FFD4B850FF2036DBE343FF40" blockId="5.[151,1436,151,428]" box="[197,808,151,176]" pageId="5" pageNumber="6">
<typeStatus id="54ED885BFFD4B850FF2036DBE159FF5F" box="[197,306,152,176]" pageId="5" pageNumber="6" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D50FD70FFD4B850FEA436D4E1DAFF40" box="[321,433,151,175]" pageId="5" pageNumber="6" type="female">2 females</specimenCount>
,
<collectionCode id="ED47AE3CFFD4B850FE5936D4E27CFF40" box="[444,535,151,175]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="5" pageNumber="6">MHNG</collectionCode>
): same data as
<typeStatus id="54ED885BFFD4B850FD5B36D4E34FFF40" box="[702,804,151,175]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C6572FFD4B850FF2036F8E503FF3B" box="[197,1384,187,212]" pageId="5" pageNumber="6" type="distribution">
<paragraph id="8BE936F9FFD4B850FF2036F8E503FF3B" blockId="5.[151,1436,151,428]" box="[197,1384,187,212]" pageId="5" pageNumber="6">
<emphasis id="B922EAEBFFD4B850FF2036F8E220FF3B" bold="true" box="[197,587,187,212]" pageId="5" pageNumber="6">Distribution and natural history.</emphasis>
<collectingCountry id="F3417669FFD4B850FDB736FFE2A8FF3B" box="[594,707,188,212]" name="Sri Lanka" pageId="5" pageNumber="6">Sri Lanka</collectingCountry>
. (Peradeniya in the
<collectingCountry id="F3417669FFD4B850FC4536FFE465FF3B" box="[928,1038,188,212]" name="Sri Lanka" pageId="5" pageNumber="6">Sri Lanka</collectingCountry>
lowland rainforest ecoregion).
</paragraph>
</subSubSection>
<subSubSection id="C34C6572FFD4B850FF2036A3E3DBFF18" box="[197,944,223,248]" pageId="5" pageNumber="6" type="etymology">
<paragraph id="8BE936F9FFD4B850FF2036A3E3DBFF18" blockId="5.[151,1436,151,428]" box="[197,944,223,248]" pageId="5" pageNumber="6">
<emphasis id="B922EAEBFFD4B850FF2036A3E12FFF17" bold="true" box="[197,324,224,248]" pageId="5" pageNumber="6">Etymology</emphasis>
. Named after “
<collectingCountry id="F3417669FFD4B850FE0A369CE22DFF18" box="[495,582,223,247]" name="Sri Lanka" pageId="5" pageNumber="6">Ceylon</collectingCountry>
”, the former name of
<collectingCountry id="F3417669FFD4B850FCDF369CE3C7FF18" box="[826,940,223,247]" name="Sri Lanka" pageId="5" pageNumber="6">Sri Lanka</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C6572FFD4B850FF203740E3B8FE43" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8BE936F9FFD4B850FF203740E3B8FE43" blockId="5.[151,1436,151,428]" pageId="5" pageNumber="6">
<emphasis id="B922EAEBFFD4B850FF203740E154FEF3" bold="true" box="[197,319,259,284]" pageId="5" pageNumber="6">Diagnosis.</emphasis>
<taxonomicName id="4C564D7AFFD4B850FEA33747E23FFEF3" box="[326,596,260,284]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis id="B922EAEBFFD4B850FEA33747E23FFEF3" box="[326,596,260,284]" italics="true" pageId="5" pageNumber="6">Ochicanthon ceylonicus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C564D7AFFD4B850FD693746E347FEF3" box="[652,812,260,284]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="cingalensis">
<emphasis id="B922EAEBFFD4B850FD693746E347FEF3" box="[652,812,260,284]" italics="true" pageId="5" pageNumber="6">O. cingalensis</emphasis>
</taxonomicName>
are the only
<taxonomicName id="4C564D7AFFD4B850FC253747E43AFEF3" box="[960,1105,260,284]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAEBFFD4B850FC253747E43AFEF3" box="[960,1105,260,284]" italics="true" pageId="5" pageNumber="6">Ochicanthon</emphasis>
</taxonomicName>
known from
<collectingCountry id="F3417669FFD4B850FB0C3747E530FEF3" box="[1257,1371,260,284]" name="Sri Lanka" pageId="5" pageNumber="6">Sri Lanka</collectingCountry>
. Both species can be easily distinguished by the pygidium, which is markedly punctate in
<taxonomicName id="4C564D7AFFD4B850FBBC376BE498FEAF" box="[1113,1267,296,320]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis id="B922EAEBFFD4B850FBBC376BE498FEAF" box="[1113,1267,296,320]" italics="true" pageId="5" pageNumber="6">O. ceylonicus</emphasis>
</taxonomicName>
and appearing impunctate in
<taxonomicName id="4C564D7AFFD4B850FEDD370EE1B3FE8B" box="[312,472,332,356]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="cingalensis">
<emphasis id="B922EAEBFFD4B850FEDD370EE1B3FE8B" box="[312,472,332,356]" italics="true" pageId="5" pageNumber="6">O. cingalensis</emphasis>
</taxonomicName>
. They uniquely share unicolored elytra and striae consisting of chains of oval depressions joined by straight sulci with the South Indian species
<taxonomicName id="4C564D7AFFD4B850FCDD3733E3A2FE67" box="[824,969,368,392]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="gauricola">
<emphasis id="B922EAEBFFD4B850FCDD3733E3A2FE67" box="[824,969,368,392]" italics="true" pageId="5" pageNumber="6">O. gauricola</emphasis>
</taxonomicName>
and
<emphasis id="B922EAEBFFD4B850FBE53733E401FE67" box="[1024,1130,368,392]" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="4C564D7AFFD4B850FBE53733E40DFE67" box="[1024,1126,368,392]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="tristis">O. tristis</taxonomicName>
.
</emphasis>
However the South Indian species have the middle portion of the meso-metasternal line subangulate.
</paragraph>
</subSubSection>
</treatment>
</document>