297 lines
37 KiB
XML
297 lines
37 KiB
XML
<document id="9BEB6875DD23A70C2167DC8E6278E695" ID-CLB-Dataset="9412" ID-DOI="10.11646/zootaxa.4885.3.5" ID-GBIF-Dataset="fd568ba9-4fd4-4f51-9a5f-42061140c137" ID-ISSN="1175-5326" ID-Zenodo-Dep="4296773" ID-ZooBank="AFE72E8B-A590-4B89-96A1-880C2D2AE14B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1606681170164" checkinUser="plazi" docAuthor="Salazar-Silva, Patricia, López-Sánchez, Daniel A. & Salazar-Vallejo, Sergio I." docDate="2020" docId="03EC4719FFC7E55466D0F9F6FCD36283" docLanguage="en" docName="zootaxa.4885.3.5.pdf" docOrigin="Zootaxa 4885 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chaetacanthus harrisae Salazar-Silva & López-Sánchez & Salazar-Vallejo 2020, n. sp." docType="treatment" docVersion="7" lastPageNumber="407" masterDocId="FFD53F61FFCCE5586647FF96FFDD6766" masterDocTitle="Revision of Chaetacanthus Seidler, 1922 (Annelida, Phyllodocida, Polynoidae)" masterLastPageNumber="422" masterPageNumber="395" pageNumber="406" updateTime="1698846816447" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="38CD08B599141AE50A65A75CF6CAC49B">Revision of Chaetacanthus Seidler, 1922 (Annelida, Phyllodocida, Polynoidae)</mods:title>
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<mods:namePart id="0DD33F3E51D868121210327C9E8C7485">Salazar-Silva, Patricia</mods:namePart>
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<mods:namePart id="F2F55582E27DC692CB300DCDE97AAF87">López-Sánchez, Daniel A.</mods:namePart>
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<mods:namePart id="38320228757D1F67514D091DB66D45C5">Salazar-Vallejo, Sergio I.</mods:namePart>
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<mods:title id="A0282F1BDE6B79B5BA6F2EBA2A12B632">Zootaxa</mods:title>
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<mods:part id="D5DFB5B431FC9C0D4CB642DA7F69324E">
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<mods:date id="36C6A0424DABA8FD602A8E5A76263E36">2020</mods:date>
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<mods:number id="5FDA5E8867D57C970986369C405EBB85">2020-11-26</mods:number>
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<treatment id="03EC4719FFC7E55466D0F9F6FCD36283" ID-DOI="http://doi.org/10.5281/zenodo.4329446" ID-GBIF-Taxon="170712087" ID-Zenodo-Dep="4329446" LSID="urn:lsid:plazi:treatment:03EC4719FFC7E55466D0F9F6FCD36283" httpUri="http://treatment.plazi.org/id/03EC4719FFC7E55466D0F9F6FCD36283" lastPageId="12" lastPageNumber="407" pageId="11" pageNumber="406">
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<subSubSection id="C35FA584FFC7E55366D0F9F6FC6261F9" pageId="11" pageNumber="406" type="nomenclature">
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<paragraph id="8BFAF60FFFC7E55366D0F9F6FDD9611D" blockId="11.[151,959,1632,1731]" box="[151,516,1632,1659]" pageId="11" pageNumber="406">
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<heading id="D0B24163FFC7E55366D0F9F6FDD9611D" bold="true" box="[151,516,1632,1659]" fontSize="11" level="1" pageId="11" pageNumber="406" reason="1">
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<emphasis id="B9312A1DFFC7E55366D0F9F6FDD9611D" bold="true" box="[151,516,1632,1659]" pageId="11" pageNumber="406">
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<taxonomicName id="4C458D8CFFC7E55366D0F9F6FE67611D" authority="Salazar-Silva & López-Sánchez & Salazar-Vallejo, 2020" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" box="[151,442,1632,1659]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="11" pageNumber="406" phylum="Annelida" rank="species" species="harrisae" status="sp. nov.">
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<emphasis id="B9312A1DFFC7E55366D0F9F6FE67611D" bold="true" box="[151,442,1632,1659]" italics="true" pageId="11" pageNumber="406">Chaetacanthus harrisae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2029766FFC7E5536786F9F7FDD9611D" box="[449,516,1633,1659]" pageId="11" pageNumber="406" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BFAF60FFFC7E55366D0F913FC6261F9" blockId="11.[151,959,1632,1731]" box="[151,959,1669,1695]" pageId="11" pageNumber="406">
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<uri id="FFD4FA0DFFC7E55366D0F913FC6261F9" box="[151,959,1669,1695]" pageId="11" pageNumber="406">
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urn:lsid:zoobank.org:act:
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<uuid id="FFE3CCDAFFC7E55367EEF913FC6261F9" box="[425,959,1669,1695]" pageId="11" pageNumber="406">E304CEF1-D40F-404C-89D3-710D8087C618</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35FA584FFC7E55366D0F93FFF2861A5" box="[151,245,1705,1731]" pageId="11" pageNumber="406" type="description">
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<paragraph id="8BFAF60FFFC7E55366D0F93FFF2861A5" blockId="11.[151,959,1632,1731]" box="[151,245,1705,1731]" pageId="11" pageNumber="406">
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<figureCitation id="137EEA8AFFC7E55366D0F93FFF2861A5" box="[151,245,1705,1731]" captionStart="FIGURE 9" captionStartId="11.[151,250,1454,1478]" captionTargetBox="[158,1429,181,1430]" captionTargetId="figure-21@11.[158,1429,181,1430]" captionTargetPageId="11" captionText="FIGURE 9. Chaetacanthus harrisae n. sp., holotype (LACM 245). A. First elytron, dorsal view. B. Same, spiny microtubercles. C–E. Same, close up of spiny microtubercles. F. Elytron of middle segment, dorsal view. G. Globular central macrotubercles from first elytron. Scale bars: A, F: 1.1 mm; B, G: 0.1 mm; C: 0.3 mm; D, E: 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4296791" httpUri="https://zenodo.org/record/4296791/files/figure.png" pageId="11" pageNumber="406">Figure 9</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35FA584FFC7E55366D0F97BFCC56042" pageId="11" pageNumber="406" type="reference_group">
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<paragraph id="8BFAF60FFFC7E55366D0F97BFC2A6063" blockId="11.[151,1015,1773,1828]" box="[151,1015,1773,1797]" pageId="11" pageNumber="406">
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<treatmentCitationGroup id="AB55D121FFC7E55366D0F97BFC2A6063" box="[151,1015,1773,1797]" pageId="11" pageNumber="406">
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<treatmentCitation id="0AE4D01EFFC7E55366D0F97BFD386063" author="Hartman, O." box="[151,741,1773,1797]" page="46" pageId="11" pageNumber="406" year="1939">
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<taxonomicName id="4C458D8CFFC7E55366D0F97BFD386063" ID-CoL="854ZC" authority="Hartman 1939 a: 44 - 46" authorityName="Hartman" authorityPageNumber="44 - 46" authorityYear="1939" box="[151,741,1773,1797]" class="Polychaeta" family="Polynoidae" genus="Lepidonotus" kingdom="Animalia" order="Phyllodocida" pageId="11" pageNumber="406" phylum="Annelida" rank="subSpecies" species="pomareae" subSpecies="panamensis">
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||
<emphasis id="B9312A1DFFC7E55366D0F97BFE2A6063" box="[151,503,1773,1797]" italics="true" pageId="11" pageNumber="406">Lepidonotus pomareae panamensis</emphasis>
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<bibRefCitation id="EFD48BFEFFC7E55367BAF978FD386063" author="Hartman, O." box="[509,741,1773,1797]" pageId="11" pageNumber="406" pagination="1 - 156" refId="ref13886" refString="Hartman, O. (1939 a) Polychaetous annelids, 1. Aphroditidae to Pisionidae. Allan Hancock Pacific Expeditions, 7, 1 - 156, pls. 1 - 28." type="journal article" year="1939">Hartman 1939a: 44–46</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
, Pl. 6, Figs 70–77 (
|
||
<emphasis id="B9312A1DFFC7E55365E0F978FC346063" box="[935,1001,1774,1797]" italics="true" pageId="11" pageNumber="406">partim</emphasis>
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).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BFAF60FFFC7E55366D0F89AFCC56042" blockId="11.[151,1015,1773,1828]" box="[151,792,1804,1828]" pageId="11" pageNumber="406">
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<treatmentCitationGroup id="AB55D121FFC7E55366D0F89AFCC56042" box="[151,792,1804,1828]" pageId="11" pageNumber="406">
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<treatmentCitation id="0AE4D01EFFC7E55366D0F89AFD856042" author="Hartman, O." box="[151,600,1804,1828]" page="28" pageId="11" pageNumber="406" year="1948">
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<taxonomicName id="4C458D8CFFC7E55366D0F89AFD856042" ID-CoL="854ZC" authority=": Hartman 1948: 28" authorityName=": Hartman" authorityPageNumber="28" authorityYear="1948" box="[151,600,1804,1828]" class="Polychaeta" family="Polynoidae" genus="Lepidonotus" kingdom="Animalia" order="Phyllodocida" pageId="11" pageNumber="406" phylum="Annelida" rank="species" species="panamensis">
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||
<emphasis id="B9312A1DFFC7E55366D0F89AFE4E6042" box="[151,403,1804,1828]" italics="true" pageId="11" pageNumber="406">Lepidonotus panamensis</emphasis>
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||
:
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||
<bibRefCitation id="EFD48BFEFFC7E55367E7F89AFD856042" author="Hartman, O." box="[416,600,1804,1828]" pageId="11" pageNumber="406" pagination="1 - 137" refId="ref14008" refString="Hartman, O. (1948) The marine annelids erected by Kinberg with some notes on some other types in the Swedish State Museum. Arkiv for Zoologi, 42 A (1), 1 - 137." type="journal article" year="1948">Hartman 1948: 28</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
(n. status,
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||
<emphasis id="B9312A1DFFC7E5536480F89BFCD66042" box="[711,779,1805,1828]" italics="true" pageId="11" pageNumber="406">partim</emphasis>
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).
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C35FA584FFC7E55366D0F8C4FA5660F6" pageId="11" pageNumber="406" type="materials_examined">
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<paragraph id="8BFAF60FFFC7E55366D0F8C4FA5660F6" blockId="11.[151,1437,1874,2009]" pageId="11" pageNumber="406">
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||
<emphasis id="B9312A1DFFC7E55366D0F8C4FE43600A" bold="true" box="[151,414,1874,1900]" pageId="11" pageNumber="406">Type material. Pacific</emphasis>
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||
<materialsCitation id="3B2DFC52FFC7E55367E2F8C4FA5660F6" ID-GBIF-Occurrence="2989844311" collectingDate="1934-07-21" collectionCode="LACM" collectorName="V Velero & Sta." country="Panama" elevation="36" latitude="7.738611" location="North West Point Rock" longLatPrecision="21" longitude="-81.58972" municipality="Pacora Island" pageId="11" pageNumber="406" specimenCode="LACM 245-34" specimenCount="1" stateProvince="Panama" typeStatus="holotype">
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<emphasis id="B9312A1DFFC7E55367E2F8C4FDDA600A" bold="true" box="[421,519,1874,1900]" pageId="11" pageNumber="406">
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<collectingCountry id="F352B69FFFC7E55367E2F8C4FDDA600A" box="[421,519,1874,1900]" name="Panama" pageId="11" pageNumber="406">Panama</collectingCountry>
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</emphasis>
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||
.
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<emphasis id="B9312A1DFFC7E5536451F8C4FD5E600A" bold="true" box="[534,643,1874,1900]" pageId="11" pageNumber="406">
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<typeStatus id="54FE48ADFFC7E5536451F8C4FD5E600A" box="[534,643,1874,1900]" pageId="11" pageNumber="406" type="holotype">Holotype</typeStatus>
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</emphasis>
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||
<specimenCode id="DBE35E74FFC7E55364CCF8C4FCE9600B" box="[651,820,1874,1901]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="11" pageNumber="406" type="Museum">LACM 245-34</specimenCode>
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,
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||
<collectingMunicipality id="6B9E6C75FFC7E5536506F8C5FC01600A" box="[833,988,1874,1901]" pageId="11" pageNumber="406">Pacora Island</collectingMunicipality>
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,
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<collectingRegion id="498138EDFFC7E55365AFF8C5FB99600B" box="[1000,1092,1875,1901]" country="Panama" name="Panama" pageId="11" pageNumber="406">Panama</collectingRegion>
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,
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<quantity id="4CBD5BEAFFC7E5536208F8C4FB68600B" box="[1103,1205,1874,1901]" metricMagnitude="1" metricUnit="m" metricValue="3.6" metricValueMax="4.5" metricValueMin="2.7" pageId="11" pageNumber="406" unit="m" value="36.0" valueMax="45.0" valueMin="27.0">
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<elevation id="0068113CFFC7E5536208F8C4FB68600B" box="[1103,1205,1874,1901]" metricMagnitude="1" metricUnit="m" metricValue="3.6" metricValueMax="4.5" metricValueMin="2.7" pageId="11" pageNumber="406" unit="m" value="36.0" valueMax="45.0" valueMin="27.0">27–45 m</elevation>
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</quantity>
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||
, R/
|
||
<collectorName id="26B093D9FFC7E553629BF8C4FAE2600A" box="[1244,1343,1874,1900]" pageId="11" pageNumber="406">V Velero</collectorName>
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III,
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||
<collectorName id="26B093D9FFC7E5536336F8C4FA40600A" box="[1393,1437,1874,1900]" pageId="11" pageNumber="406">Sta.</collectorName>
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||
245-34,
|
||
<geoCoordinate id="EE7190C8FFC7E55366B2F8E0FE5A60F7" box="[245,391,1910,1937]" degrees="07" direction="north" minutes="44" orientation="latitude" pageId="11" pageNumber="406" precision="15" seconds="19" value="7.738611">07°44’19” N</geoCoordinate>
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,
|
||
<geoCoordinate id="EE7190C8FFC7E55367D4F8E0FDFF60F7" box="[403,546,1910,1937]" degrees="81" direction="west" minutes="35" orientation="longitude" pageId="11" pageNumber="406" precision="15" seconds="23" value="-81.58972">81°35’23”W</geoCoordinate>
|
||
, dredged off
|
||
<location id="8E9AA0D4FFC7E55364F1F8E0FC6160F6" LSID="urn:lsid:plazi:treatment:03EC4719FFC7E55466D0F9F6FCD36283:8E9AA0D4FFC7E55364F1F8E0FC6160F6" box="[694,956,1910,1937]" country="Panama" latitude="7.738611" longLatPrecision="21" longitude="-81.58972" municipality="Pacora Island" name="North West Point Rock" pageId="11" pageNumber="406" stateProvince="Panama">North West Point Rock</location>
|
||
, large shells and nullipores,
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||
<date id="FFFBD0CFFFC7E55362BDF8E0FA5A60F6" box="[1274,1415,1910,1936]" pageId="11" pageNumber="406" value="1934-07-21">
|
||
<collectingDate id="EFBF2927FFC7E55362BDF8E0FA5A60F6" box="[1274,1415,1910,1936]" pageId="11" pageNumber="406" value="1934-07-21">21 Jul. 1934</collectingDate>
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||
</date>
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||
.
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||
</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C35FA584FFC7E5546680F80CFAE667D7" lastPageId="12" lastPageNumber="407" pageId="11" pageNumber="406" type="diagnosis">
|
||
<paragraph id="8BFAF60FFFC7E5546680F80CFAE667D7" blockId="11.[151,1437,1874,2009]" lastBlockId="12.[151,1437,150,1509]" lastPageId="12" lastPageNumber="407" pageId="11" pageNumber="406">
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||
<emphasis id="B9312A1DFFC7E5536680F80CFE9F60D2" bold="true" box="[199,322,1946,1972]" pageId="11" pageNumber="406">Diagnosis.</emphasis>
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||
<taxonomicName id="4C458D8CFFC7E553670FF80CFE2D60D2" authorityName="Seidler" authorityYear="1922" box="[328,496,1946,1972]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="11" pageNumber="406" phylum="Annelida" rank="genus">
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||
<emphasis id="B9312A1DFFC7E553670FF80CFE2D60D2" box="[328,496,1946,1972]" italics="true" pageId="11" pageNumber="406">Chaetacanthus</emphasis>
|
||
</taxonomicName>
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||
with branchial filaments single, thick lobes; elytra with pale to brownish round spinous macrotubercles; microtubercles bulbous up to three times longer than wide, spinous surface extended along ½–
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<superScript id="7C305B47FFC0E55466FDFF00FF1F67C2" attach="none" box="[186,194,150,164]" fontSize="6" pageId="12" pageNumber="407">2</superScript>
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/
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<subScript id="17C1F44AFFC0E554668DFF3FFF0F67D1" attach="left" box="[202,210,169,183]" fontSize="6" pageId="12" pageNumber="407">3</subScript>
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basal length; elytra without large, amber-colour macrotubercles arranged in honeycomb like groups.
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</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C35FA584FFC0E5546680FF2CFB19645F" pageId="12" pageNumber="407" type="description">
|
||
<paragraph id="8BFAF60FFFC0E5546680FF2CFD666603" blockId="12.[151,1437,150,1509]" pageId="12" pageNumber="407">
|
||
<emphasis id="B9312A1DFFC0E5546680FF2CFE8467B2" bold="true" box="[199,345,186,212]" pageId="12" pageNumber="407">Description.</emphasis>
|
||
<typeStatus id="54FE48ADFFC0E5546725FF2DFE1767B3" box="[354,458,187,213]" pageId="12" pageNumber="407" type="holotype">Holotype</typeStatus>
|
||
(LACM 245-34) 5.5 cm long, 1.35 cm wide, 26 segments. Prostomium bilobed with facial tubercle prominent, acute. Two pairs of eyes, anterior pair on widest prostomial area, posterior pair hidden under anterior projection of second segment. Median antenna with ceratophore inserted frontally, style long, subdistally swollen. Lateral antennae with ceratophores inserted frontally, styles similar to median antennal one. Palps with rows of thick papillae. Pharynx not everted.
|
||
</paragraph>
|
||
<paragraph id="8BFAF60FFFC0E5546680FEF9FB5166CB" blockId="12.[151,1437,150,1509]" pageId="12" pageNumber="407">Tentacular segment not visible dorsally. Tentaculophores thin, tentacular cirri similar to median antenna. Second segment with anterior border projected over prostomium as a short round nuchal lobe.</paragraph>
|
||
<paragraph id="8BFAF60FFFC0E5546680FE21FDAC65E3" blockId="12.[151,1437,150,1509]" pageId="12" pageNumber="407">
|
||
Elytral margin with fringe of long abundant, filiform papillae (
|
||
<figureCitation id="137EEA8AFFC0E55465D6FE21FC3066B7" box="[913,1005,439,465]" captionStart="FIGURE 9" captionStartId="11.[151,250,1454,1478]" captionTargetBox="[158,1429,181,1430]" captionTargetId="figure-21@11.[158,1429,181,1430]" captionTargetPageId="11" captionText="FIGURE 9. Chaetacanthus harrisae n. sp., holotype (LACM 245). A. First elytron, dorsal view. B. Same, spiny microtubercles. C–E. Same, close up of spiny microtubercles. F. Elytron of middle segment, dorsal view. G. Globular central macrotubercles from first elytron. Scale bars: A, F: 1.1 mm; B, G: 0.1 mm; C: 0.3 mm; D, E: 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4296791" httpUri="https://zenodo.org/record/4296791/files/figure.png" pageId="12" pageNumber="407">Fig. 9A</figureCitation>
|
||
), some with globular tips, including a distinctive bundle of papillae. Elytral surface verrucose with abundant globular, non-sclerotized spinous macrotubercles (
|
||
<figureCitation id="137EEA8AFFC0E55466B0FE69FE8E657F" box="[247,339,511,537]" captionStart="FIGURE 9" captionStartId="11.[151,250,1454,1478]" captionTargetBox="[158,1429,181,1430]" captionTargetId="figure-21@11.[158,1429,181,1430]" captionTargetPageId="11" captionText="FIGURE 9. Chaetacanthus harrisae n. sp., holotype (LACM 245). A. First elytron, dorsal view. B. Same, spiny microtubercles. C–E. Same, close up of spiny microtubercles. F. Elytron of middle segment, dorsal view. G. Globular central macrotubercles from first elytron. Scale bars: A, F: 1.1 mm; B, G: 0.1 mm; C: 0.3 mm; D, E: 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4296791" httpUri="https://zenodo.org/record/4296791/files/figure.png" pageId="12" pageNumber="407">Fig. 9G</figureCitation>
|
||
), and clavate micropapillae (
|
||
<figureCitation id="137EEA8AFFC0E55464DAFE69FD2A657F" box="[669,759,511,537]" captionStart="FIGURE 9" captionStartId="11.[151,250,1454,1478]" captionTargetBox="[158,1429,181,1430]" captionTargetId="figure-21@11.[158,1429,181,1430]" captionTargetPageId="11" captionText="FIGURE 9. Chaetacanthus harrisae n. sp., holotype (LACM 245). A. First elytron, dorsal view. B. Same, spiny microtubercles. C–E. Same, close up of spiny microtubercles. F. Elytron of middle segment, dorsal view. G. Globular central macrotubercles from first elytron. Scale bars: A, F: 1.1 mm; B, G: 0.1 mm; C: 0.3 mm; D, E: 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4296791" httpUri="https://zenodo.org/record/4296791/files/figure.png" pageId="12" pageNumber="407">Fig. 9B</figureCitation>
|
||
). First pair of elytra with scattered microtubercles along its whole surface, globular macrotubercles close to elytrophoral plug area (
|
||
<figureCitation id="137EEA8AFFC0E55465FFFDB5FBEE655B" box="[952,1075,547,573]" captionStart="FIGURE 9" captionStartId="11.[151,250,1454,1478]" captionTargetBox="[158,1429,181,1430]" captionTargetId="figure-21@11.[158,1429,181,1430]" captionTargetPageId="11" captionText="FIGURE 9. Chaetacanthus harrisae n. sp., holotype (LACM 245). A. First elytron, dorsal view. B. Same, spiny microtubercles. C–E. Same, close up of spiny microtubercles. F. Elytron of middle segment, dorsal view. G. Globular central macrotubercles from first elytron. Scale bars: A, F: 1.1 mm; B, G: 0.1 mm; C: 0.3 mm; D, E: 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4296791" httpUri="https://zenodo.org/record/4296791/files/figure.png" pageId="12" pageNumber="407">Fig. 9A, G</figureCitation>
|
||
). On middle and posterior elytra globular macrotubercles scarce (
|
||
<figureCitation id="137EEA8AFFC0E5546446FDD1FD886507" box="[513,597,583,609]" captionStart="FIGURE 9" captionStartId="11.[151,250,1454,1478]" captionTargetBox="[158,1429,181,1430]" captionTargetId="figure-21@11.[158,1429,181,1430]" captionTargetPageId="11" captionText="FIGURE 9. Chaetacanthus harrisae n. sp., holotype (LACM 245). A. First elytron, dorsal view. B. Same, spiny microtubercles. C–E. Same, close up of spiny microtubercles. F. Elytron of middle segment, dorsal view. G. Globular central macrotubercles from first elytron. Scale bars: A, F: 1.1 mm; B, G: 0.1 mm; C: 0.3 mm; D, E: 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4296791" httpUri="https://zenodo.org/record/4296791/files/figure.png" pageId="12" pageNumber="407">Fig. 9F</figureCitation>
|
||
), spinous surface extended along ½–
|
||
<superScript id="7C305B47FFC0E55465A9FDD0FC2B6532" attach="none" box="[1006,1014,582,596]" fontSize="6" pageId="12" pageNumber="407">2</superScript>
|
||
/
|
||
<subScript id="17C1F44AFFC0E55465B9FDCFFBDB6501" attach="left" box="[1022,1030,601,615]" fontSize="6" pageId="12" pageNumber="407">3</subScript>
|
||
basal length, microtubercles numerous, bulbous with long peaks (
|
||
<figureCitation id="137EEA8AFFC0E55467ABFDFDFDF265E3" box="[492,559,619,645]" captionStart="FIGURE 9" captionStartId="11.[151,250,1454,1478]" captionTargetBox="[158,1429,181,1430]" captionTargetId="figure-21@11.[158,1429,181,1430]" captionTargetPageId="11" captionText="FIGURE 9. Chaetacanthus harrisae n. sp., holotype (LACM 245). A. First elytron, dorsal view. B. Same, spiny microtubercles. C–E. Same, close up of spiny microtubercles. F. Elytron of middle segment, dorsal view. G. Globular central macrotubercles from first elytron. Scale bars: A, F: 1.1 mm; B, G: 0.1 mm; C: 0.3 mm; D, E: 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4296791" httpUri="https://zenodo.org/record/4296791/files/figure.png" pageId="12" pageNumber="407">Fig. 9</figureCitation>
|
||
B–E).
|
||
</paragraph>
|
||
<paragraph id="8BFAF60FFFC0E5546680FD19FAC16597" blockId="12.[151,1437,150,1509]" pageId="12" pageNumber="407">Parapodia biramous. Notopodia thick with acicular tips emergent. Neuropodia distally truncate, acicular tips emergent. Branchial filaments present between successive parapodia. Dorsal cirri thin, long, reaching neurochaetal tips, cirri subdistally swollen, resembling antennae. Cirrophores robust. Nephridial papillae long, thick.</paragraph>
|
||
<paragraph id="8BFAF60FFFC0E5546680FD6DFB19645F" blockId="12.[151,1437,150,1509]" pageId="12" pageNumber="407">Notochaetae thin capillaries, long, with spinous surface, lower notochaetae shorter, slightly curved, serrated. Neurochaetae with upper region short, subdistally with long thin spines, tips entire, long, acute.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35FA584FFC0E5546680FCD5FC8B64C3" pageId="12" pageNumber="407" type="etymology">
|
||
<paragraph id="8BFAF60FFFC0E5546680FCD5FC8B64C3" blockId="12.[151,1437,150,1509]" pageId="12" pageNumber="407">
|
||
<emphasis id="B9312A1DFFC0E5546680FCD5FE93643B" bold="true" box="[199,334,835,861]" pageId="12" pageNumber="407">Etymology.</emphasis>
|
||
This species is named after Mrs. Leslie Harris, our long-term friend and teacher, in recognition of her efforts supporting taxonomy of polychaetes, and especially because she has been very generous by housing us during repetitive visits to Los Angeles during the last 20 years.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35FA584FFC0E5546680FC39FDD862FB" pageId="12" pageNumber="407" type="discussion">
|
||
<paragraph id="8BFAF60FFFC0E5546680FC39FDDB63C7" blockId="12.[151,1437,150,1509]" pageId="12" pageNumber="407">
|
||
<emphasis id="B9312A1DFFC0E5546680FC39FEE664AF" bold="true" box="[199,315,943,969]" pageId="12" pageNumber="407">Remarks.</emphasis>
|
||
<bibRefCitation id="EFD48BFEFFC0E5546705FC39FDDC64AF" author="Hartman, O." box="[322,513,943,969]" pageId="12" pageNumber="407" pagination="1 - 156" refId="ref13886" refString="Hartman, O. (1939 a) Polychaetous annelids, 1. Aphroditidae to Pisionidae. Allan Hancock Pacific Expeditions, 7, 1 - 156, pls. 1 - 28." type="journal article" year="1939">Hartman (1939a)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="4C458D8CFFC0E554643EFC39FBDD64AE" authorityName="Hartman" authorityYear="1939" box="[633,1024,943,969]" class="Polychaeta" family="Polynoidae" genus="Lepidonotus" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="407" phylum="Annelida" rank="subSpecies" species="pomareae" subSpecies="panamensis">
|
||
<emphasis id="B9312A1DFFC0E554643EFC39FBDD64AE" box="[633,1024,943,969]" italics="true" pageId="12" pageNumber="407">Lepidonotus pomareae panamensis</emphasis>
|
||
</taxonomicName>
|
||
and indicated its differences from the nominal form by having elytra “conspicuously covered with numerous large headed, high tubercles, the sixth pair of elytra is rectangular, not triangular, and the total length is over
|
||
<quantity id="4CBD5BEAFFC0E554657BFC60FC556377" box="[828,904,1014,1041]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="12" pageNumber="407" unit="mm" value="35.0">35 mm</quantity>
|
||
as against
|
||
<quantity id="4CBD5BEAFFC0E55465BAFC61FB976377" box="[1021,1098,1015,1041]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="407" unit="mm" value="13.0">13 mm</quantity>
|
||
specified for
|
||
<taxonomicName id="4C458D8CFFC0E554629AFC61FABB6376" baseAuthorityName="Kinberg" baseAuthorityYear="1856" box="[1245,1382,1015,1040]" class="Polychaeta" family="Polynoidae" genus="Lepidonotus" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="407" phylum="Annelida" rank="species" species="pomareae">
|
||
<emphasis id="B9312A1DFFC0E554629AFC61FABB6376" box="[1245,1382,1015,1040]" italics="true" pageId="12" pageNumber="407">L. pomareae</emphasis>
|
||
</taxonomicName>
|
||
.” As indicated below, the
|
||
<typeStatus id="54FE48ADFFC0E554673DFB8DFE076353" box="[378,474,1051,1077]" pageId="12" pageNumber="407" type="holotype">holotype</typeStatus>
|
||
of this subspecies, later raised to species level as
|
||
<taxonomicName id="4C458D8CFFC0E55465BAFB8DFB416352" authorityName=": Hartman" authorityYear="1948" box="[1021,1180,1051,1076]" class="Polychaeta" family="Polynoidae" genus="Lepidonotus" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="407" phylum="Annelida" rank="species" species="panamensis">
|
||
<emphasis id="B9312A1DFFC0E55465BAFB8DFB416352" box="[1021,1180,1051,1076]" italics="true" pageId="12" pageNumber="407">L. panamensis</emphasis>
|
||
</taxonomicName>
|
||
, is regarded as a junior synonym of
|
||
<taxonomicName id="4C458D8CFFC0E5546765FBA9FDBD633F" authority="(Treadwell, 1937)" baseAuthorityName="Treadwell" baseAuthorityYear="1937" box="[290,608,1087,1113]" class="Polychaeta" family="Polynoidae" genus="Lepidonotus" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="407" phylum="Annelida" rank="species" species="pilosus">
|
||
<emphasis id="B9312A1DFFC0E5546765FBA9FE4D633F" box="[290,400,1087,1113]" italics="true" pageId="12" pageNumber="407">C. pilosus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD48BFEFFC0E55467D8FBA9FD85633F" author="Treadwell, A. L." box="[415,600,1087,1113]" pageId="12" pageNumber="407" pagination="139 - 159" refId="ref16030" refString="Treadwell, A. L. (1937) The Templeton Crocker Expedition, 8. Polychaetous annelids from the West coast of Lower California, the Gulf of California and Clarion Island. Zoologica, Scientific Contributions of the New York Zoological Society, 22, 139 - 159, Pls 1 - 2." type="journal article" year="1937">Treadwell, 1937</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. However, the specimen investigated here belongs neither to
|
||
<taxonomicName id="4C458D8CFFC0E5546342FBA9FAAD633F" baseAuthorityName="Treadwell" baseAuthorityYear="1937" box="[1285,1392,1087,1113]" class="Polychaeta" family="Polynoidae" genus="Lepidonotus" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="407" phylum="Annelida" rank="species" species="pilosus">
|
||
<emphasis id="B9312A1DFFC0E5546342FBA9FAAD633F" box="[1285,1392,1087,1113]" italics="true" pageId="12" pageNumber="407">L. pilosus</emphasis>
|
||
</taxonomicName>
|
||
nor to
|
||
<taxonomicName id="4C458D8CFFC0E55466F4FBF5FEE1631A" baseAuthorityName="Kinberg" baseAuthorityYear="1856" box="[179,316,1123,1148]" class="Polychaeta" family="Polynoidae" genus="Lepidonotus" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="407" phylum="Annelida" rank="species" species="pomareae">
|
||
<emphasis id="B9312A1DFFC0E55466F4FBF5FEE1631A" box="[179,316,1123,1148]" italics="true" pageId="12" pageNumber="407">L. pomareae</emphasis>
|
||
</taxonomicName>
|
||
, and is described here as
|
||
<taxonomicName id="4C458D8CFFC0E5546415FBF5FD13631B" authority="Salazar-Silva & López-Sánchez & Salazar-Vallejo, 2020" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" box="[594,718,1123,1149]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="407" phylum="Annelida" rank="species" species="harrisae" status="sp. nov.">
|
||
<emphasis id="B9312A1DFFC0E5546415FBF5FD13631B" box="[594,718,1123,1149]" italics="true" pageId="12" pageNumber="407">C. harrisae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2029766FFC0E5546492FBF5FCD2631B" box="[725,783,1123,1149]" pageId="12" pageNumber="407" rank="species">n. sp.</taxonomicNameLabel>
|
||
The material was erroneously referred from station 254-34, but it comes from station 245-34.
|
||
</paragraph>
|
||
<paragraph id="8BFAF60FFFC0E5546680FB3DFD686257" blockId="12.[151,1437,150,1509]" pageId="12" pageNumber="407">
|
||
Further,
|
||
<taxonomicName id="4C458D8CFFC0E5546762FB3DFE7E63A3" authority="Salazar-Silva & López-Sánchez & Salazar-Vallejo, 2020" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" box="[293,419,1195,1221]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="407" phylum="Annelida" rank="species" species="harrisae" status="sp. nov.">
|
||
<emphasis id="B9312A1DFFC0E5546762FB3DFE7E63A3" box="[293,419,1195,1221]" italics="true" pageId="12" pageNumber="407">C. harrisae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2029766FFC0E55467ECFB3DFE3B63A3" box="[427,486,1195,1221]" pageId="12" pageNumber="407" rank="species">n. sp.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C458D8CFFC0E554643BFB3DFD3063A3" baseAuthorityName="Treadwell" baseAuthorityYear="1937" box="[636,749,1195,1221]" class="Polychaeta" family="Polynoidae" genus="Lepidonotus" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="407" phylum="Annelida" rank="species" species="pilosus">
|
||
<emphasis id="B9312A1DFFC0E554643BFB3DFD3063A3" box="[636,749,1195,1221]" italics="true" pageId="12" pageNumber="407">C. pilosus</emphasis>
|
||
</taxonomicName>
|
||
because its elytra do not have pedunculated tubercles, in the first pair of elytra of
|
||
<taxonomicName id="4C458D8CFFC0E55467CCFB59FDD0638F" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" box="[395,525,1231,1257]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="407" phylum="Annelida" rank="species" species="harrisae">
|
||
<emphasis id="B9312A1DFFC0E55467CCFB59FDD0638F" box="[395,525,1231,1257]" italics="true" pageId="12" pageNumber="407">C. harrisae</emphasis>
|
||
</taxonomicName>
|
||
the macrotubercles are mainly globular, scarce and concentrated along the area corresponding to the elytrophoral plug. In elytra of median and posterior segments, the macrotubercles are smaller, and the microtubercles bear long, tapered peaks.
|
||
</paragraph>
|
||
<paragraph id="8BFAF60FFFC0E5546680FAADFDD862FB" blockId="12.[151,1437,150,1509]" pageId="12" pageNumber="407">
|
||
On the other hand, members of
|
||
<taxonomicName id="4C458D8CFFC0E554646DFAADFD776233" authority="Salazar-Silva & López-Sánchez & Salazar-Vallejo, 2020" authorityName="Salazar-Silva & López-Sánchez & Salazar-Vallejo" authorityYear="2020" box="[554,682,1339,1365]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="407" phylum="Annelida" rank="species" species="harrisae" status="sp. nov.">
|
||
<emphasis id="B9312A1DFFC0E554646DFAADFD776233" box="[554,682,1339,1365]" italics="true" pageId="12" pageNumber="407">C. harrisae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2029766FFC0E55464F6FAADFD306233" box="[689,749,1339,1365]" pageId="12" pageNumber="407" rank="species">n. sp.</taxonomicNameLabel>
|
||
also differ from members of
|
||
<taxonomicName id="4C458D8CFFC0E5546272FAADFA5D6233" authority="(Kinberg, 1856)" baseAuthorityName="Kinberg" baseAuthorityYear="1856" box="[1077,1408,1338,1365]" class="Polychaeta" family="Polynoidae" genus="Lepidonotus" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="407" phylum="Annelida" rank="species" species="pomareae">
|
||
<emphasis id="B9312A1DFFC0E5546272FAADFB956233" box="[1077,1096,1339,1365]" italics="true" pageId="12" pageNumber="407">C</emphasis>
|
||
.
|
||
<emphasis id="B9312A1DFFC0E5546211FAADFB186232" box="[1110,1221,1339,1364]" italics="true" pageId="12" pageNumber="407">pomareae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD48BFEFFC0E5546294FAADFAAA6233" author="Kinberg, J. G. H." box="[1235,1399,1338,1365]" pageId="12" pageNumber="407" pagination="381 - 388" refId="ref14336" refString="Kinberg, J. G. H. (1856) Nya slagten och arter af Annelider. Ofversigt af Kongliga Vetenskaps-Akademiens Forhhandlingar Stockholm, 12 (9 - 10), 381 - 388." type="journal article" year="1856">Kinberg, 1856</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
in that the latter lacks the long conical tubercles scattered along the posterior elytral half area, and only the anterior elytra have long macrotubercles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35FA584FFC0E5546680FA31FC9862A7" box="[199,837,1446,1473]" pageId="12" pageNumber="407" type="materials_examined">
|
||
<paragraph id="8BFAF60FFFC0E5546680FA31FC9862A7" blockId="12.[151,1437,150,1509]" box="[199,837,1446,1473]" pageId="12" pageNumber="407">
|
||
<materialsCitation id="3B2DFC52FFC0E5546680FA31FC9C62A7" ID-GBIF-Occurrence="2989844308" box="[199,833,1446,1473]" country="Panama" elevation="59" location="Pacora Island" pageId="12" pageNumber="407" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B9312A1DFFC0E5546680FA31FEBB62A6" bold="true" box="[199,358,1446,1473]" pageId="12" pageNumber="407">
|
||
<typeStatus id="54FE48ADFFC0E5546680FA31FEDF62A7" box="[199,258,1447,1473]" pageId="12" pageNumber="407">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<location id="8E9AA0D4FFC0E554672BFA31FDD962A7" LSID="urn:lsid:plazi:treatment:03EC4719FFC7E55466D0F9F6FCD36283:8E9AA0D4FFC0E554672BFA31FDD962A7" box="[364,516,1447,1473]" country="Panama" name="Pacora Island" pageId="12" pageNumber="407">Pacora Island</location>
|
||
,
|
||
<collectingCountry id="F352B69FFFC0E5546455FA31FDB362A7" box="[530,622,1447,1473]" name="Panama" pageId="12" pageNumber="407">Panamá</collectingCountry>
|
||
, in
|
||
<elevation id="0068113CFFC0E55464D2FA31FC9C62A7" box="[661,833,1447,1473]" metricMagnitude="1" metricUnit="m" metricValue="5.85" metricValueMax="9.0" metricValueMin="2.7" pageId="12" pageNumber="407" unit="m" value="58.5" valueMax="90.0" valueMin="27.0">
|
||
<quantity id="4CBD5BEAFFC0E55464D2FA31FD2562A7" box="[661,760,1447,1473]" metricMagnitude="1" metricUnit="m" metricValue="5.85" metricValueMax="9.0" metricValueMin="2.7" pageId="12" pageNumber="407" unit="m" value="58.5" valueMax="90.0" valueMin="27.0">27–90 m</quantity>
|
||
depth
|
||
</elevation>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35FA584FFC0E5546680FA5CFCD36283" box="[199,782,1482,1509]" pageId="12" pageNumber="407" type="distribution">
|
||
<paragraph id="8BFAF60FFFC0E5546680FA5CFCD36283" blockId="12.[151,1437,150,1509]" box="[199,782,1482,1509]" pageId="12" pageNumber="407">
|
||
<emphasis id="B9312A1DFFC0E5546680FA5CFEBC6282" bold="true" box="[199,353,1482,1508]" pageId="12" pageNumber="407">Distribution.</emphasis>
|
||
Only known from the original record.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |