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<document id="5BD56787496869FD46C0518E00F73613" ID-CLB-Dataset="30368" ID-DOI="10.5852/ejt.2018.418" ID-GBIF-Dataset="ff41552a-7cc5-42ba-9336-7a61f2845b41" ID-Zenodo-Dep="1210533" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1522513246312" checkinUser="plazi" docAuthor="Wesołowska, Wanda" docDate="2018" docId="03C987FA3370871FFE6EFEDE0066F8AC" docLanguage="en" docName="ejt-418_wesolowska.pdf.imf" docOrigin="European Journal of Taxonomy 418" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pochytoides Berland &amp; Millot 1941" docType="treatment" docVersion="12" lastPageNumber="3" masterDocId="FFF0FF823372871DFF8CFFD5036EFFBE" masterDocTitle="A revision of the genus Pochytoides Berland &amp; Millot, 1941 (Araneae: Salticidae), with descriptions of six new species" masterLastPageNumber="26" masterPageNumber="1" pageNumber="3" updateTime="1698535849002" updateUser="ExternalLinkService">
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<mods:title id="C42DCC3B65BF96FB6BBC4AB7889BB7FD">A revision of the genus Pochytoides Berland &amp; Millot, 1941 (Araneae: Salticidae), with descriptions of six new species</mods:title>
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<mods:date id="D854902454528482A393E7844E72524E">2018</mods:date>
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<subSubSection id="C37A65673370871FFE6EFEDE0272FED9" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BDF36EC3370871FFE6EFEDE073CFE98" blockId="2.[482,1106,267,294]" box="[482,1106,267,294]" pageId="2" pageNumber="3">
<heading id="D09781803370871FFE6EFEDE073CFE98" box="[482,1106,267,294]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
Genus
<taxonomicName id="4C604D6F3370871FFDB8FEDE008CFE9B" ID-CoL="6T9G" authority="Berland &amp; Millot, 1941" authorityName="Berland &amp; Millot" authorityYear="1941" box="[564,994,267,294]" class="Arachnida" family="Salticidae" genus="Pochytoides" higherTaxonomySource="CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B914EAFE3370871FFDB8FEDE01ABFE98" bold="true" box="[564,709,267,294]" italics="true" pageId="2" pageNumber="3">Pochytoides</emphasis>
<bibRefCitation id="EFF14B1D3370871FFD40FEDE008CFE9B" author="Berland L. &amp; Millot J." box="[716,994,267,293]" journalOrPublisher="Memoires du Musee d'Histoire naturelle" pageId="2" pageNumber="3" pagination="297 - 424" part="12" refId="ref8621" refString="Berland L. &amp; Millot J. 1941. Les araignees de l'Afrique occidentale francaise. I. Les Salticides. Memoires du Musee d'Histoire naturelle 12: 297 - 424." title="Les araignees de l'Afrique occidentale francaise. I. Les Salticides" type="journal article" year="1941">Berland &amp; Millot, 1941</bibRefCitation>
</taxonomicName>
stat. nov.
</heading>
</paragraph>
<paragraph id="8BDF36EC3370871FFF31FE980272FED9" blockId="2.[189,284,333,359]" box="[189,284,333,359]" pageId="2" pageNumber="3">
<heading id="D09781803370871FFF31FE980272FED9" bold="true" box="[189,284,333,359]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B914EAFE3370871FFF31FE980272FED9" bold="true" box="[189,284,333,359]" pageId="2" pageNumber="3">Gender</emphasis>
</heading>
</paragraph>
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<subSubSection id="C37A65673370871FFF31FEAE0242FE2B" box="[189,300,379,405]" pageId="2" pageNumber="3" type="description">
<paragraph id="8BDF36EC3370871FFF31FEAE0242FE2B" blockId="2.[189,300,379,405]" box="[189,300,379,405]" pageId="2" pageNumber="3">Feminine</paragraph>
</subSubSection>
<subSubSection id="C37A65673370871FFF31FE6801CDFDBB" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="8BDF36EC3370871FFF31FE680236FE69" blockId="2.[189,675,445,517]" box="[189,344,445,471]" pageId="2" pageNumber="3">
<heading id="D09781803370871FFF31FE680236FE69" bold="true" box="[189,344,445,471]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B914EAFE3370871FFF31FE680236FE69" bold="true" box="[189,344,445,471]" pageId="2" pageNumber="3">Type species</emphasis>
</heading>
</paragraph>
<paragraph id="8BDF36EC3370871FFF31FE3E01CDFDBB" blockId="2.[189,675,445,517]" box="[189,675,491,517]" pageId="2" pageNumber="3">
<taxonomicName id="4C604D6F3370871FFF31FE3E01CDFDBB" authority="Berland &amp; Millot, 1941" authorityName="Berland &amp; Millot" authorityYear="1941" box="[189,675,491,517]" class="Arachnida" family="Salticidae" genus="Pochyta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="poissoni">
<emphasis id="B914EAFE3370871FFF31FE3E0273FDBB" box="[189,285,491,517]" italics="true" pageId="2" pageNumber="3">Pochyta</emphasis>
<emphasis id="B914EAFE3370871FFEA8FE3902E8FDBB" box="[292,390,492,517]" italics="true" pageId="2" pageNumber="3">poissoni</emphasis>
<bibRefCitation id="EFF14B1D3370871FFE02FE3E01CDFDBB" author="Berland L. &amp; Millot J." box="[398,675,491,517]" journalOrPublisher="Memoires du Musee d'Histoire naturelle" pageId="2" pageNumber="3" pagination="297 - 424" part="12" refId="ref8621" refString="Berland L. &amp; Millot J. 1941. Les araignees de l'Afrique occidentale francaise. I. Les Salticides. Memoires du Musee d'Histoire naturelle 12: 297 - 424." title="Les araignees de l'Afrique occidentale francaise. I. Les Salticides" type="journal article" year="1941">Berland &amp; Millot, 1941</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C37A65673370871FFF31FDF80659FCF7" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="8BDF36EC3370871FFF31FDF80258FDF9" blockId="2.[189,1399,557,841]" box="[189,310,557,583]" pageId="2" pageNumber="3">
<heading id="D09781803370871FFF31FDF80258FDF9" bold="true" box="[189,310,557,583]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B914EAFE3370871FFF31FDF80258FDF9" bold="true" box="[189,310,557,583]" pageId="2" pageNumber="3">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BDF36EC3370871FFF31FD8E0659FCF7" blockId="2.[189,1399,557,841]" pageId="2" pageNumber="3">The genus is characterized by the structure of the genitalia. The male palps have a retrolateral tibial apophysis which is broad and straight, with a blunt tip and a dorsal apophysis which is long, thin and perpendicular to the palpal axis. The cymbium is narrow and the bulb is short, more or less triangular, with a well separated basal part and a large embolic division. The presence of a small, spike-like tegular apophysis on the prolateral side of the bulb is a distinctive feature. The epigynes have copulatory openings hidden in a deep and extremely sclerotized cup-shaped atria. The internal structure is simple with seminal ducts more or less short and straight and the spermathecae ovoid or spherical. The genitalic pattern is consistent throughout the whole genus.</paragraph>
</subSubSection>
<subSubSection id="C37A65673370871FFF31FCA50100FAE1" pageId="2" pageNumber="3" type="description">
<paragraph id="8BDF36EC3370871FFF31FCA50220FC34" blockId="2.[189,1399,880,1375]" box="[189,334,880,906]" pageId="2" pageNumber="3">
<heading id="D09781803370871FFF31FCA50220FC34" bold="true" box="[189,334,880,906]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B914EAFE3370871FFF31FCA50220FC34" bold="true" box="[189,334,880,906]" pageId="2" pageNumber="3">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BDF36EC3370871FFF31FC4B0100FAE1" blockId="2.[189,1399,880,1375]" pageId="2" pageNumber="3">
Small to medium spiders, ranging from about
<quantity id="4C989B093370871FFD42FC4A002FFC07" box="[718,833,926,953]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="2" pageNumber="3" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3 to 5 mm</quantity>
in length. Carapace brown, oval, high and broad, with steep posterior slope beginning just behind the eye field. Eyes on small tubercles, with black rings around each (except as noted), eye field trapezoid, anterior row of eyes slightly wider than the posterior row.Anterior median eyes large. Fovea visible, sulciform, lying on small concave area. Clypeus vertical, low. Chelicerae large and long, fang relatively short, two teeth on promargin and a single large tooth on retromargin (
<figureCitation id="135B2A693370871FFEF0FB9B02B2FBD7" box="[380,476,1102,1129]" captionStart="Fig. 9" captionStartId="15.[189,232,1870,1896]" captionTargetBox="[201,1397,277,1805]" captionTargetId="figure@15.[186,1397,266,1812]" captionTargetPageId="15" captionText="Fig. 9. Pochytoides poissoni (Berland &amp; Millot, 1941) comb. nov., Ƌ. A. General appearance, lateral view. B. Carapace, dorsolateral view. C. Cheliceral dentition. D. First leg. E–K. Palpal organ. E. Ventroprolateral view. F, I. Ventral view. G, J. Retrolateral view. H. Dorsal view. Scale bars:A–B, D = 1.0 mm, C, E–K = 0.3 mm." httpUri="https://zenodo.org/record/1210551/files/figure.png" pageId="2" pageNumber="3">Figs 9C</figureCitation>
,
<figureCitation id="135B2A693370871FFE66FB9A0173FBD7" box="[490,541,1103,1129]" captionStart="Fig.11" captionStartId="18.[189,232,1911,1937]" captionTargetBox="[219,1356,661,1874]" captionTargetId="figure@18.[219,1368,650,1887]" captionTargetPageId="18" captionText="Fig.11.Pochytoides securissp. nov.,paratypes,Ƌ.A.Generalappearance, dorsalview.B.Frontalview. C.Cheliceraldentition.D–H.Palpalorgan.D–E.Ventralview.F.Ventroretrolateralview.G.Retrolateral view. H. Dorsal view. Scalebars:A–B = 1.0 mm, C–H = 0.3 mm." httpUri="https://zenodo.org/record/1210555/files/figure.png" pageId="2" pageNumber="3">11C</figureCitation>
). Endites slightly convergent. Labium trapezoid. Sternum shield-shaped. Pedicel short, not visible in dorsal view. Abdomen ovoid, slightly narrower than carapace. Legs more or less subequal, anterior pair slightly longer than others, especially in males. Spination of leg I in male: femur dorsally 0–1–1 and 0–0–1 prolaterally, patella 1 prolaterally, tibia ventrally 2–2–2–2 and prolaterally 1–1–0 or 1–0, metatarsus ventrally 2–2 and 2 apically on dorsal surface. Ventral spines on tibia and metatarsus extremely long (
<figureCitation id="135B2A693370871FFDE2FB2B01A2FAA7" box="[622,716,1278,1305]" captionStart="Fig. 9" captionStartId="15.[189,232,1870,1896]" captionTargetBox="[201,1397,277,1805]" captionTargetId="figure@15.[186,1397,266,1812]" captionTargetPageId="15" captionText="Fig. 9. Pochytoides poissoni (Berland &amp; Millot, 1941) comb. nov., Ƌ. A. General appearance, lateral view. B. Carapace, dorsolateral view. C. Cheliceral dentition. D. First leg. E–K. Palpal organ. E. Ventroprolateral view. F, I. Ventral view. G, J. Retrolateral view. H. Dorsal view. Scale bars:A–B, D = 1.0 mm, C, E–K = 0.3 mm." httpUri="https://zenodo.org/record/1210551/files/figure.png" pageId="2" pageNumber="3">Fig. 9D</figureCitation>
). Spination of leg I in female similar, but spines on patella and prolateral surface of tibia absent. Female palp with single retrolateral spine on tarsus (
<figureCitation id="135B2A693370871FFB76FAF70607FA82" box="[1274,1385,1314,1340]" captionStart="Fig. 10" captionStartId="16.[189,232,1909,1935]" captionTargetBox="[218,1357,850,1845]" captionTargetId="figure@16.[197,1391,839,1865]" captionTargetPageId="16" captionText="Fig. 10. Pochytoides poissoni (Berland &amp; Millot, 1941) comb. nov., holotype, ♀ (B, D, G), specimen from Ziéla (A, C, E, H) and specimen from Fouenyi (F). A. Tip of female’s palp. B–F. Epigyne. G–H. Internal structureof epigyne. Scale bars:A= 0.3 mm, B–H= 0.2 mm." httpUri="https://zenodo.org/record/1210553/files/figure.png" pageId="2" pageNumber="3">Fig. 10A</figureCitation>
). Structure of genitalia: see Diagnosis.
</paragraph>
</subSubSection>
<subSubSection id="C37A65673370871FFF31FA520066F8AC" pageId="2" pageNumber="3" type="discussion">
<paragraph id="8BDF36EC3370871FFF31FA520241FA1F" blockId="2.[189,303,1415,1441]" box="[189,303,1415,1441]" pageId="2" pageNumber="3">
<heading id="D09781803370871FFF31FA520241FA1F" bold="true" box="[189,303,1415,1441]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B914EAFE3370871FFF31FA520241FA1F" bold="true" box="[189,303,1415,1441]" pageId="2" pageNumber="3">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BDF36EC3370871FFF31FA6001D3F987" blockId="2.[189,1398,1461,1593]" pageId="2" pageNumber="3">
A large part of the bulb is enveloped by the cymbium, so that the study of this structure, and understanding its functional morphology, requires separation of the bulb from the cymbium. Unfortunately, insufficient numbers of male specimens prevented such study and the detailed bulb structure of only one species,
<taxonomicName id="4C604D6F3370871FFF31F9CA0254F986" authority="Wesołowska, 2018" authorityName="Wesołowska" authorityYear="2018" box="[189,314,1567,1592]" class="Arachnida" family="Salticidae" genus="Pochytoides" higherTaxonomySource="CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="spiniger" status="sp. nov.">
<emphasis id="B914EAFE3370871FFF31F9CA0254F986" box="[189,314,1567,1592]" italics="true" pageId="2" pageNumber="3">P. spiniger</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757853370871FFECDF9CA02F0F987" box="[321,414,1567,1593]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
, is shown (
<figureCitation id="135B2A693370871FFDAEF9CB01DEF987" box="[546,688,1566,1593]" captionStart="Fig.13" captionStartId="20.[189,232,1921,1947]" captionTargetBox="[221,1365,712,1848]" captionTargetId="figure@20.[189,1399,691,1863]" captionTargetPageId="20" captionText="Fig.13.Pochytoidesspinigersp.nov.,paratypes,Ƌ.A–C.Palpalorgan.A.Ventralview.B.Anteroventral view. C. Retrolateral view. D. Palpal apophyses, dorsal view. E–G. Separated bulb. E. Ventral view. F. Retrolateral view. G. Prolateral view. Scale bar = 0.3 mm." httpUri="https://zenodo.org/record/1210559/files/figure.png" pageId="2" pageNumber="3">Fig. 13E–G</figureCitation>
).
</paragraph>
<paragraph id="8BDF36EC3370871FFF31F9B50241F9C4" blockId="2.[189,303,1632,1658]" box="[189,303,1632,1658]" pageId="2" pageNumber="3">
<heading id="D09781803370871FFF31F9B50241F9C4" bold="true" box="[189,303,1632,1658]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B914EAFE3370871FFF31F9B50241F9C4" bold="true" box="[189,303,1632,1658]" pageId="2" pageNumber="3">Affinities</emphasis>
</heading>
</paragraph>
<paragraph id="8BDF36EC3370871FFF31F95B0066F8AC" blockId="2.[189,1399,1678,1810]" pageId="2" pageNumber="3">
The morphological characters mentioned in the introduction clearly show that
<taxonomicName id="4C604D6F3370871FFBDBF95B0788F916" authorityName="Berland &amp; Millot" authorityYear="1941" box="[1111,1254,1678,1704]" class="Arachnida" family="Salticidae" genus="Pochytoides" higherTaxonomySource="CoL" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B914EAFE3370871FFBDBF95B0788F916" box="[1111,1254,1678,1704]" italics="true" pageId="2" pageNumber="3">Pochytoides</emphasis>
</taxonomicName>
is related to other genera included in the subtribe
<taxonomicName id="4C604D6F3370871FFDE3F9670072F972" box="[623,796,1714,1740]" class="Arachnida" family="Hahniidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subTribe" subTribe="Thiratoscirtina">Thiratoscirtina</taxonomicName>
<bibRefCitation id="EFF14B1D3370871FFCAFF9670733F972" author="Bodner M. R. &amp; Maddison W. P." box="[803,1117,1714,1740]" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="2" pageNumber="3" pagination="213 - 240" part="65" publicationUrl="https://doi.org/10.1016/j.ympev.2012.06.005" refId="ref8653" refString="Bodner M. R. &amp; Maddison W. P. 2012. The biogeography and age of salticid spider radiations (Araneae: Salticidae). Molecular Phylogenetics and Evolution 65: 213 - 240. https: // doi. org / 10.1016 / j. ympev. 2012.06.005" title="The biogeography and age of salticid spider radiations (Araneae: Salticidae)" type="journal article" year="2012">
Bodner &amp; Maddison,
<date id="FFDE102C3370871FFBAEF9670733F972" box="[1058,1117,1714,1740]" pageId="2" pageNumber="3" value="2012">2012</date>
</bibRefCitation>
(sensu
<bibRefCitation id="EFF14B1D3370871FFB3AF9670600F972" author="Maddison W. P." box="[1206,1390,1713,1740]" journalOrPublisher="Journal of Arachnology" pageId="2" pageNumber="3" pagination="231 - 292" part="43" publicationUrl="https://doi.org/10.1636/arac-43-03-231-292" refId="ref8704" refString="Maddison W. P. 2015. A phylogenetic classification of jumping spiders (Araneae: Salticidae). Journal of Arachnology 43: 231 - 292. https: // doi. org / 10.1636 / arac- 43 - 03 - 231 - 292" title="A phylogenetic classification of jumping spiders (Araneae: Salticidae)" type="journal article" year="2015">
Maddison
<date id="FFDE102C3370871FFABEF9640600F972" box="[1330,1390,1713,1740]" pageId="2" pageNumber="3" value="2015">2015</date>
</bibRefCitation>
) in the subfamily
<taxonomicName id="4C604D6F3370871FFE04F90001A5F951" authority="Blackwall, 1841" authorityName="Blackwall" authorityYear="1841" box="[392,715,1749,1775]" class="Arachnida" family="Salticidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Salticinae">Salticinae Blackwall, 1841</taxonomicName>
. At the moment, closer relationships of the genus remain unresolved and demand further molecular studies.
</paragraph>
</subSubSection>
</treatment>
</document>
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