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<document id="79A30BAB017632BE3CD032B7801F3F8C" ID-CLB-Dataset="21808" ID-DOI="10.5852/ejt.2014.100" ID-GBIF-Dataset="5329bfc4-61b2-48f5-a96a-caf4ba31039b" ID-ISSN="2118-9773" ID-Zenodo-Dep="3860450" ID-ZooBank="B3E6C489-6D96-4AF5-A33D-EE8329A9321B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1590595554426" checkinUser="carolina" docAuthor="Enghoff, Henrik" docDate="2014" docId="334F2769C17BFF901C03F37EFC03FC6E" docLanguage="en" docName="ejt-2014-100.pdf.imf" docOrigin="European Journal of Taxonomy 100" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Chaleponcus quasimodo Enghoff 2014, sp. nov." docType="treatment" docUuid="37A2F533-120D-45C5-8C49-B77A4C2DA088" docUuidSource="ZooBank" docVersion="7" lastPageNumber="25" masterDocId="CF765F11C16DFF881E57F137FFA0FFA3" masterDocTitle="A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)" masterLastPageNumber="75" masterPageNumber="1" pageNumber="23" updateTime="1698841637397" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="79B0F52C0801B6C7B1127F8997A6420F">A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)</mods:title>
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<mods:namePart id="4A1F490A80C6AC2686C9C6C0348D5747">Enghoff, Henrik</mods:namePart>
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<mods:title id="B47A09521F46DFF89819A1D6BD2AB15D">European Journal of Taxonomy</mods:title>
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<mods:date id="DD019EC0B7E2FEDB1AB3FC29CCD72EE8">2014</mods:date>
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<treatment id="334F2769C17BFF901C03F37EFC03FC6E" ID-DOI="http://doi.org/10.5281/zenodo.3861189" ID-GBIF-Taxon="164362767" ID-Zenodo-Dep="3861189" LSID="urn:lsid:zoobank.org:act:37A2F533-120D-45C5-8C49-B77A4C2DA088" httpUri="http://treatment.plazi.org/id/334F2769C17BFF901C03F37EFC03FC6E" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23">
<subSubSection id="F3FCC5F4C17BFF9E1C03F37EFC40FDC7" box="[596,992,585,612]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="BB59967FC17BFF9E1C03F37EFC40FDC7" blockId="22.[366,1222,585,682]" box="[596,992,585,612]" pageId="22" pageNumber="23">
<heading id="E0112113C17BFF9E1C03F37EFC40FDC7" box="[596,992,585,612]" centered="true" fontSize="11" level="2" pageId="22" pageNumber="23" reason="2">
<taxonomicName id="7CE6EDFCC17BFF9E1C03F37EFCDEFDC7" ID-CoL="TH7B" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[596,894,585,612]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="quasimodo" status="sp. nov.">
<emphasis id="89924A6DC17BFF9E1C03F37EFCDEFDC7" bold="true" box="[596,894,585,612]" italics="true" pageId="22" pageNumber="23">Chaleponcus quasimodo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C17BFF9E1DD2F37DFC40FDC7" box="[901,992,586,612]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="F3FCC5F4C17BFF9E1F39F35AFCF4FD09" pageId="22" pageNumber="23" type="description">
<paragraph id="BB59967FC17BFF9E1F39F35AFB66FD2B" blockId="22.[366,1222,585,682]" box="[366,1222,621,648]" pageId="22" pageNumber="23">
<uri id="CF779A7DC17BFF9E1F39F35AFB66FD2B" box="[366,1222,621,648]" pageId="22" pageNumber="23">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="BB59967FC17BFF9E1C88F3A7FCF4FD09" blockId="22.[366,1222,585,682]" box="[735,852,656,682]" pageId="22" pageNumber="23">
<figureCitation id="23DD8AFAC17BFF9E1C88F3A7FC8AFD09" box="[735,810,656,682]" captionStart="Fig" captionStartId="15.[187,230,1816,1842]" captionTargetBox="[247,1356,507,1717]" captionTargetId="figure@15.[200,1385,488,1723]" captionTargetPageId="15" captionText="Fig. 9. Chaleponcus spp., outline of coxite of left gonopod, posterior view. (The transverse plane of the gonopod is usually slightly at angles with the exact transverse plane of the millipede, so it may be necessary to tilt the gonopod slightly in order to obtain the view shown on the drawings.) The stippled lines indicate the place where the telopodital basomere emerges from the coxal cavity. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3860472" httpUri="https://zenodo.org/record/3860472/files/figure.png" pageId="22" pageNumber="23">Figs 9</figureCitation>
,
<figureCitation id="23DD8AFAC17BFF9E1D60F3A7FCF4FD09" box="[823,852,656,682]" captionStart="Fig" captionStartId="23.[189,232,1896,1922]" captionTargetBox="[189,1398,289,1853]" captionTargetId="figure@23.[189,1398,289,1853]" captionTargetPageId="23" captionText="Fig. 13. Chaleponcus quasimodo sp. nov., left gonopod. A. Anterior view. B. Anterior-mesal view. C. Ventral view. D. Posterior view. al = anterior lamella of telomere, mf = metaplical flange, mp = mesal metaplical process, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior lamella of telomere, ps = proximal spine of solenomere. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860480" httpUri="https://zenodo.org/record/3860480/files/figure.png" pageId="22" pageNumber="23">13</figureCitation>
</paragraph>
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<subSubSection id="F3FCC5F4C17BFF9E1EEAF3E6FC48FCC3" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="BB59967FC17BFF9E1EEAF3E6FE96FD48" blockId="22.[189,310,721,747]" box="[189,310,721,747]" pageId="22" pageNumber="23">
<heading id="E0112113C17BFF9E1EEAF3E6FE96FD48" bold="true" box="[189,310,721,747]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="89924A6DC17BFF9E1EEAF3E6FE96FD48" bold="true" box="[189,310,721,747]" pageId="22" pageNumber="23">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="BB59967FC17BFF9E1EEAF237FC48FCC3" blockId="22.[189,1399,768,865]" pageId="22" pageNumber="23">
Medium-sized. Differs from all other group members except
<taxonomicName id="7CE6EDFCC17BFF9E1DF1F237FBFFFCB9" authorityName="Kraus" authorityYear="1958" box="[934,1119,768,794]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="dabagaensis">
<emphasis id="89924A6DC17BFF9E1DF1F237FBFFFCB9" box="[934,1119,768,794]" italics="true" pageId="22" pageNumber="23">C. dabagaensis</emphasis>
</taxonomicName>
by the hunch-backed profile of the gonopod coxa (
<figureCitation id="23DD8AFAC17BFF9E1C43F214FDFDFC9E" box="[532,605,803,829]" captionStart="Fig" captionStartId="15.[187,230,1816,1842]" captionTargetBox="[247,1356,507,1717]" captionTargetId="figure@15.[200,1385,488,1723]" captionTargetPageId="15" captionText="Fig. 9. Chaleponcus spp., outline of coxite of left gonopod, posterior view. (The transverse plane of the gonopod is usually slightly at angles with the exact transverse plane of the millipede, so it may be necessary to tilt the gonopod slightly in order to obtain the view shown on the drawings.) The stippled lines indicate the place where the telopodital basomere emerges from the coxal cavity. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3860472" httpUri="https://zenodo.org/record/3860472/files/figure.png" pageId="22" pageNumber="23">Fig. 9</figureCitation>
); differs from
<taxonomicName id="7CE6EDFCC17BFF9E1D5EF213FC1FFC9E" authorityName="Kraus" authorityYear="1958" box="[777,959,803,830]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="dabagaensis">
<emphasis id="89924A6DC17BFF9E1D5EF213FC1FFC9E" box="[777,959,803,830]" italics="true" pageId="22" pageNumber="23">C. dabagaensis</emphasis>
</taxonomicName>
by the profile of the coxal tip and by the absence of a thumblike process on the main stem of the telomere.
</paragraph>
</subSubSection>
<subSubSection id="F3FCC5F4C17BFF9E1EEAF2BFFD0FFC57" pageId="22" pageNumber="23" type="etymology">
<paragraph id="BB59967FC17BFF9E1EEAF2BFFEE4FC01" blockId="22.[189,324,904,930]" box="[189,324,904,930]" pageId="22" pageNumber="23">
<heading id="E0112113C17BFF9E1EEAF2BFFEE4FC01" bold="true" box="[189,324,904,930]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="89924A6DC17BFF9E1EEAF2BFFEE4FC01" bold="true" box="[189,324,904,930]" pageId="22" pageNumber="23">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="BB59967FC17BFF9E1EEAF281FD0FFC57" blockId="22.[189,1399,950,1012]" pageId="22" pageNumber="23">The name is a noun, here in apposition, and refers to Quasimodo, the hunch-back of Notre-Dame, title figure of the novel by Victor Hugo (1839).</paragraph>
</subSubSection>
<subSubSection id="F3FCC5F4C17BFF9E1EEAF52CFCF1FAF6" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="BB59967FC17BFF9E1EEAF52CFD8EFB95" blockId="22.[189,558,1051,1078]" box="[189,558,1051,1078]" pageId="22" pageNumber="23">
<emphasis id="89924A6DC17BFF9E1EEAF52CFE2DFB96" bold="true" box="[189,397,1051,1077]" pageId="22" pageNumber="23">Material studied</emphasis>
(total:
<specimenCount id="ADE05DF6C17BFF9E1FB5F52CFD85FB95" box="[482,549,1051,1078]" pageId="22" pageNumber="23" type="male">9 ♂♂</specimenCount>
)
</paragraph>
<materialsCitation id="0B8E9C22C17BFF9E1EBDF57DFCDFFB09" ID-GBIF-Occurrence="2626747313" collectingDate="1984-10-08" collectingDateMax="1984-10-10" collectingDateMin="1984-10-08" collectingMethod="pitfall traps" collectionCode="ZMUC" collectorName="N. Scharff" country="Tanzania" county="Mufindi District" elevation="1700" location="Kigogo" municipality="Udzungwa Mts" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1" stateProvince="Iringa Region" typeStatus="holotype">
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<emphasis id="89924A6DC17BFF9E1EBDF57DFEFCFBC7" bold="true" box="[234,348,1098,1124]" pageId="22" pageNumber="23">
<typeStatus id="645D28DDC17BFF9E1EBDF57DFEFCFBC7" box="[234,348,1098,1124]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="BB59967FC17BFF9E1EEAF55AFCDFFB09" blockId="22.[189,1399,1098,1195]" pageId="22" pageNumber="23">
<collectingCountry id="C3F1D6EFC17BFF9E1EEAF55AFEF7FB24" box="[189,343,1133,1159]" name="Tanzania" pageId="22" pageNumber="23">TANZANIA</collectingCountry>
:
<specimenCount id="ADE05DF6C17BFF9E1F36F55AFED9FB24" box="[353,377,1133,1159]" pageId="22" pageNumber="23" type="male"></specimenCount>
,
<collectingRegion id="7922589DC17BFF9E1FD2F55AFD8AFB24" box="[389,554,1133,1159]" country="Tanzania" name="Iringa" pageId="22" pageNumber="23">Iringa Region</collectingRegion>
,
<collectingCounty id="5238EEF3C17BFF9E1C62F55AFD56FB24" box="[565,758,1133,1159]" pageId="22" pageNumber="23">Mufindi District</collectingCounty>
,
<collectingMunicipality id="5B3D0C05C17BFF9E1D56F55AFC17FB24" box="[769,951,1133,1159]" pageId="22" pageNumber="23">Udzungwa Mts</collectingMunicipality>
,
<location id="BE39C0A4C17BFF9E1D94F55AFBBBFB24" LSID="urn:lsid:plazi:treatment:334F2769C17BFF901C03F37EFC03FC6E:BE39C0A4C17BFF9E1D94F55AFBBBFB24" box="[963,1051,1133,1159]" country="Tanzania" county="Mufindi District" municipality="Udzungwa Mts" name="Kigogo" pageId="22" pageNumber="23" stateProvince="Iringa Region">Kigogo</location>
FR,
<quantity id="7C1E3B9AC17BFF9E1A03F55AFB0EFB24" box="[1108,1198,1133,1159]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="22" pageNumber="23" unit="m" value="1700.0">
<elevation id="30CB714CC17BFF9E1A03F55AFB0EFB24" box="[1108,1198,1133,1159]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="22" pageNumber="23" unit="m" value="1700.0">1700 m</elevation>
</quantity>
,
<date id="CF58B0BFC17BFF9E1AEDF55AFAD2FB24" box="[1210,1394,1133,1159]" pageId="22" pageNumber="23" value="1984-10-08" valueMax="1984-10-10" valueMin="1984-10-08">
<collectingDate id="DF1C4957C17BFF9E1AEDF55AFAD2FB24" box="[1210,1394,1133,1159]" pageId="22" pageNumber="23" value="1984-10-08" valueMax="1984-10-10" valueMin="1984-10-08">810 Oct. 1984</collectingDate>
</date>
,
<collectingMethod id="62A7EE68C17BFF9E1EEAF5A7FEE3FB08" box="[189,323,1168,1195]" pageId="22" pageNumber="23">pitfall traps</collectingMethod>
in montane rain forest.
<collectorName id="1613F3A9C17BFF9E1C0BF5A6FD76FB09" box="[604,726,1168,1195]" pageId="22" pageNumber="23">N. Scharff</collectorName>
leg. (
<collectionCode id="DDF70EBAC17BFF9E1D40F5A7FCD3FB09" box="[791,883,1168,1194]" collectionName="Denmark, Kobenhavn [= Copenhagen], University of Copenhagen, Zoological Museum" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="22" pageNumber="23">ZMUC</collectionCode>
).
</paragraph>
</materialsCitation>
<paragraph id="BB59967FC17BFF9E1EBDF5E5FEC7FB4F" blockId="22.[189,1399,1234,1366]" box="[234,359,1234,1260]" pageId="22" pageNumber="23">
<emphasis id="89924A6DC17BFF9E1EBDF5E5FEC7FB4F" bold="true" box="[234,359,1234,1260]" pageId="22" pageNumber="23">
<typeStatus id="645D28DDC17BFF9E1EBDF5E5FEC7FB4F" box="[234,359,1234,1260]" pageId="22" pageNumber="23" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="BB59967FC17BFF9E1EEAF5C2FCF1FAF6" blockId="22.[189,1399,1234,1366]" pageId="22" pageNumber="23">
All from
<collectingCountry id="C3F1D6EFC17BFF9E1F7EF5C2FE61FAAC" box="[297,449,1269,1295]" name="Tanzania" pageId="22" pageNumber="23">TANZANIA</collectingCountry>
,
<collectingRegion id="7922589DC17BFF9E1F9AF5C2FDD2FAAC" box="[461,626,1269,1295]" country="Tanzania" name="Iringa" pageId="22" pageNumber="23">Iringa Region</collectingRegion>
, Mufindi District, Udzungwa Mts, Kigogo FR, all in ZMUC:
<specimenCount id="ADE05DF6C17BFF9E1B14F5C2FAD1FAAC" box="[1347,1393,1269,1295]" pageId="22" pageNumber="23" type="male">1 ♂</specimenCount>
,
<quantity id="7C1E3B9AC17BFF9E1EEAF42FFEB8FA91" box="[189,280,1304,1330]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="22" pageNumber="23" unit="m" value="1700.0">1700 m</quantity>
,
<date id="CF58B0BFC17BFF9E1F72F42FFE7EFA91" box="[293,478,1304,1330]" pageId="22" pageNumber="23" value="1984-10-08" valueMax="1984-10-10" valueMin="1984-10-08">810 Oct. 1984</date>
, pitfall traps in montane rain forest. N. Scharff leg.;
<specimenCount id="ADE05DF6C17BFF9E1A1EF42FFB2DFA91" box="[1097,1165,1304,1330]" pageId="22" pageNumber="23" type="male">7 ♂♂</specimenCount>
,
<quantity id="7C1E3B9AC17BFF9E1ACDF42FFB55FA91" box="[1178,1269,1304,1330]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="22" pageNumber="23" unit="m" value="1900.0">1900 m</quantity>
,
<date id="CF58B0BFC17BFF9E1B55F42FFF5AFAF6" pageId="22" pageNumber="23" value="1984-10-07" valueMax="1984-10-15" valueMin="1984-10-07">715 Oct. 1984</date>
, pitfall traps in montane rain forest. N. Scharff leg.
</paragraph>
</subSubSection>
<subSubSection id="F3FCC5F4C17BFF9E1EEAF44AFAD7FA66" pageId="22" pageNumber="23" type="distribution">
<paragraph id="BB59967FC17BFF9E1EEAF44AFEFDFA35" blockId="22.[189,349,1404,1431]" box="[189,349,1404,1431]" pageId="22" pageNumber="23">
<heading id="E0112113C17BFF9E1EEAF44AFEFDFA35" bold="true" box="[189,349,1404,1431]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="89924A6DC17BFF9E1EEAF44AFEFDFA35" bold="true" box="[189,349,1404,1431]" pageId="22" pageNumber="23">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="BB59967FC17BFF9E1EEAF49CFAD7FA66" blockId="22.[189,1399,1451,1478]" box="[189,1399,1451,1478]" pageId="22" pageNumber="23">
<collectingCountry id="C3F1D6EFC17BFF9E1EEAF49CFEF5FA66" box="[189,341,1451,1477]" name="Tanzania" pageId="22" pageNumber="23">TANZANIA</collectingCountry>
,
<collectingRegion id="7922589DC17BFF9E1F37F49CFDA4FA66" box="[352,516,1451,1477]" country="Tanzania" name="Iringa" pageId="22" pageNumber="23">Iringa Region</collectingRegion>
, Mufindi District, Udzungwa Mts, Kigogo FR,
<quantity id="7C1E3B9AC17BFF9E1A7FF49CFB22FA65" box="[1064,1154,1451,1478]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="22" pageNumber="23" unit="m" value="1700.0">1700 m</quantity>
, montane rain forest.
</paragraph>
</subSubSection>
<subSubSection id="F3FCC5F4C17BFF901EEAF4DBFDA0FD04" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23" type="description">
<paragraph id="BB59967FC17BFF9E1EEAF4DBFE02F9A4" blockId="22.[189,418,1516,1543]" box="[189,418,1516,1543]" pageId="22" pageNumber="23">
<emphasis id="89924A6DC17BFF9E1EEAF4DBFEFFF9A5" bold="true" box="[189,351,1516,1542]" pageId="22" pageNumber="23">Description (</emphasis>
male)
</paragraph>
<paragraph id="BB59967FC17BFF9E1EEAF72CFD67F996" blockId="22.[189,711,1563,1589]" box="[189,711,1563,1589]" pageId="22" pageNumber="23">
DIAMETER.
<quantity id="7C1E3B9AC17BFF9E1F6BF72CFE6CF996" box="[316,460,1563,1589]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="22" pageNumber="23" unit="mm" value="2.2" valueMax="2.3" valueMin="2.1">2.12.3 mm</quantity>
, 4749 podous rings.
</paragraph>
<paragraph id="BB59967FC17BFF9E1EEAF76BFB09F93A" blockId="22.[189,1398,1628,1690]" pageId="22" pageNumber="23">COLOUR. After 30 years in alcohol almost completely bleached, with traces of paramedian darker bands flanking a broad middorsal light stripe, sometimes a thin dark line in middle of stripe.</paragraph>
<paragraph id="BB59967FC17BFF9E1EEAF7F6FCB0F95D" blockId="22.[189,1399,1729,1791]" pageId="22" pageNumber="23">ANAL VALVES. Each with a long, slightly curving dorsal spine and a small, triangular ventral one; marginal rim raised, setiferous tubercles on small ravelins.</paragraph>
<paragraph id="BB59967FC17BFF9E1EEAF611FA84F89C" blockId="22.[189,1316,1829,1856]" box="[189,1316,1829,1856]" pageId="22" pageNumber="23">LIMBUS. With broadly rounded lobes, as long as broad or slightly longer, striate on outer surface.</paragraph>
<paragraph id="BB59967FC17BFF9E1EEAF65FFE4AF822" blockId="22.[189,490,1895,1922]" box="[189,490,1895,1922]" pageId="22" pageNumber="23">TARSAL SETATION. Normal.</paragraph>
<paragraph id="BB59967FC17BFF901EEAF69EFC68FE5B" blockId="22.[189,1398,1960,2022]" lastBlockId="24.[189,1399,267,505]" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23">
GONOPOD COXA (
<figureCitation id="23DD8AFAC17BFF9E1FD4F69FFE72F860" box="[387,466,1960,1987]" captionStart="Fig" captionStartId="15.[187,230,1816,1842]" captionTargetBox="[247,1356,507,1717]" captionTargetId="figure@15.[200,1385,488,1723]" captionTargetPageId="15" captionText="Fig. 9. Chaleponcus spp., outline of coxite of left gonopod, posterior view. (The transverse plane of the gonopod is usually slightly at angles with the exact transverse plane of the millipede, so it may be necessary to tilt the gonopod slightly in order to obtain the view shown on the drawings.) The stippled lines indicate the place where the telopodital basomere emerges from the coxal cavity. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3860472" httpUri="https://zenodo.org/record/3860472/files/figure.png" pageId="22" pageNumber="23">Figs 9</figureCitation>
,
<figureCitation id="23DD8AFAC17BFF9E1FB4F69EFDA3F860" box="[483,515,1961,1987]" captionStart="Fig" captionStartId="23.[189,232,1896,1922]" captionTargetBox="[189,1398,289,1853]" captionTargetId="figure@23.[189,1398,289,1853]" captionTargetPageId="23" captionText="Fig. 13. Chaleponcus quasimodo sp. nov., left gonopod. A. Anterior view. B. Anterior-mesal view. C. Ventral view. D. Posterior view. al = anterior lamella of telomere, mf = metaplical flange, mp = mesal metaplical process, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior lamella of telomere, ps = proximal spine of solenomere. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860480" httpUri="https://zenodo.org/record/3860480/files/figure.png" pageId="22" pageNumber="23">13</figureCitation>
). About 3 × as long as wide. Lateral margin straight up to level of
<emphasis id="89924A6DC17BFF9E1B18F69EFAD0F861" box="[1359,1392,1961,1986]" italics="true" pageId="22" pageNumber="23">ms</emphasis>
, then curving mesad and then lateral, leaving a deep, rounded incision and giving the entire coxa a hunchback profile; apical margin slightly convex, irregularly undulate, forming
<emphasis id="89924A6DC175FF901AD9F03BFB0AFE86" box="[1166,1194,268,293]" italics="true" pageId="24" pageNumber="25">ca</emphasis>
. 70° angles with lateral margin. Mesal margin of proplica with two mesad processes apically, one above, one below metaplical shelf-spine (
<emphasis id="89924A6DC175FF901F8BF064FDA8FECF" box="[476,520,339,364]" italics="true" pageId="24" pageNumber="25">mss</emphasis>
). Metaplical flange (
<emphasis id="89924A6DC175FF901D46F065FC8EFECF" box="[785,814,338,364]" italics="true" pageId="24" pageNumber="25">mf</emphasis>
) ending in approximately right-angled-rounded corner, hardly projecting; metaplical mesal margin with rounded-triangular process (
<emphasis id="89924A6DC175FF901A97F041FB44FE2C" box="[1216,1252,374,399]" italics="true" pageId="24" pageNumber="25">mp</emphasis>
) just below level of arculus, distally with triangular incision with coarsely nodose margin. Metaplical shelf (
<emphasis id="89924A6DC175FF901B7BF0AEFAEDFE11" box="[1324,1357,409,434]" italics="true" pageId="24" pageNumber="25">ms</emphasis>
) of moderate size, projecting slightly posteriad as well as mesad, continuing without transition in metaplical shelf-spine (
<emphasis id="89924A6DC175FF901F18F0E8FEDBFE5B" box="[335,379,479,504]" italics="true" pageId="24" pageNumber="25">mss</emphasis>
);
<emphasis id="89924A6DC175FF901FC3F0E8FE60FE5B" box="[404,448,479,504]" italics="true" pageId="24" pageNumber="25">mss</emphasis>
of medium length, curving obliquely basad.
</paragraph>
<caption id="EF99C6F7C17AFF9F1EEAF65FFC2BF84F" ID-DOI="http://doi.org/10.5281/zenodo.3860480" ID-Zenodo-Dep="3860480" httpUri="https://zenodo.org/record/3860480/files/figure.png" pageId="23" pageNumber="24" startId="23.[189,232,1896,1922]" targetBox="[189,1398,289,1853]" targetPageId="23">
<paragraph id="BB59967FC17AFF9F1EEAF65FFC2BF84F" blockId="23.[189,1399,1896,2028]" pageId="23" pageNumber="24">
<emphasis id="89924A6DC17AFF9F1EEAF65FFEBDF821" bold="true" box="[189,285,1896,1922]" pageId="23" pageNumber="24">Fig. 13.</emphasis>
<taxonomicName id="7CE6EDFCC17AFF9F1F7EF65FFDEFF821" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[297,591,1896,1922]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="quasimodo" status="sp. nov.">
<emphasis id="89924A6DC17AFF9F1F7EF65FFDEFF821" box="[297,591,1896,1922]" italics="true" pageId="23" pageNumber="24">Chaleponcus quasimodo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C17AFF9F1C0DF65EFD1BF820" box="[602,699,1897,1923]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
, left gonopod.
<emphasis id="89924A6DC17AFF9F1D20F65FFC2CF821" bold="true" box="[887,908,1896,1922]" pageId="23" pageNumber="24">A</emphasis>
. Anterior view.
<emphasis id="89924A6DC17AFF9F1A00F65EFBCBF820" bold="true" box="[1111,1131,1897,1923]" pageId="23" pageNumber="24">B</emphasis>
. Anterior-mesal view.
<emphasis id="89924A6DC17AFF9F1EEAF6BCFF72F806" bold="true" box="[189,210,1931,1957]" pageId="23" pageNumber="24">C</emphasis>
. Ventral view.
<emphasis id="89924A6DC17AFF9F1FDAF6BBFE02F805" bold="true" box="[397,418,1932,1958]" pageId="23" pageNumber="24">D</emphasis>
. Posterior view.
<emphasis id="89924A6DC17AFF9F1C24F6BBFD2AF805" box="[627,650,1932,1958]" italics="true" pageId="23" pageNumber="24">al</emphasis>
= anterior lamella of telomere,
<emphasis id="89924A6DC17AFF9F1A43F6BBFB91F805" box="[1044,1073,1932,1958]" italics="true" pageId="23" pageNumber="24">mf</emphasis>
= metaplical flange,
<emphasis id="89924A6DC17AFF9F1B60F6BBFAFBF806" box="[1335,1371,1932,1957]" italics="true" pageId="23" pageNumber="24">mp</emphasis>
= mesal metaplical process,
<emphasis id="89924A6DC17AFF9F1FA5F687FDB3F86A" box="[498,531,1968,1993]" italics="true" pageId="23" pageNumber="24">ms</emphasis>
= metaplical shelf,
<emphasis id="89924A6DC17AFF9F1CABF687FC88F86A" box="[764,808,1968,1993]" italics="true" pageId="23" pageNumber="24">mss</emphasis>
= metaplical shelf-spine,
<emphasis id="89924A6DC17AFF9F1A0EF698FBD0F86A" box="[1113,1136,1967,1993]" italics="true" pageId="23" pageNumber="24">pl</emphasis>
= posterior lamella of telomere,
<emphasis id="89924A6DC17AFF9F1F65F6E4FEECF84F" box="[306,332,2003,2028]" italics="true" pageId="23" pageNumber="24">ps</emphasis>
= proximal spine of solenomere. Scales 0.1 mm.
</paragraph>
</caption>
<paragraph id="BB59967FC175FF901EEAF317FC14FDFE" blockId="24.[189,1398,544,679]" pageId="24" pageNumber="25">
GONOPOD TELOPODITE (
<figureCitation id="23DD8AFAC175FF901F91F317FD80FD99" box="[454,544,544,570]" captionStart="Fig" captionStartId="23.[189,232,1896,1922]" captionTargetBox="[189,1398,289,1853]" captionTargetId="figure@23.[189,1398,289,1853]" captionTargetPageId="23" captionText="Fig. 13. Chaleponcus quasimodo sp. nov., left gonopod. A. Anterior view. B. Anterior-mesal view. C. Ventral view. D. Posterior view. al = anterior lamella of telomere, mf = metaplical flange, mp = mesal metaplical process, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior lamella of telomere, ps = proximal spine of solenomere. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860480" httpUri="https://zenodo.org/record/3860480/files/figure.png" pageId="24" pageNumber="25">Fig. 13</figureCitation>
). Solenomere with medium long, almost straight proximal spine (
<emphasis id="89924A6DC175FF901B1CF316FAC5FD99" box="[1355,1381,545,570]" italics="true" pageId="24" pageNumber="25">ps</emphasis>
). Telomere distally with two relatively small and simple branches:
</paragraph>
<paragraph id="BB59967FC175FF901EEAF350FDA2FD22" blockId="24.[189,1398,544,679]" box="[189,514,614,641]" pageId="24" pageNumber="25">
• an anterior lamella (
<emphasis id="89924A6DC175FF901F8DF351FE51FD23" box="[474,497,614,640]" italics="true" pageId="24" pageNumber="25">al</emphasis>
),
</paragraph>
<paragraph id="BB59967FC175FF901EEAF3BAFDA0FD04" blockId="24.[189,1398,544,679]" box="[189,512,653,679]" pageId="24" pageNumber="25">
• a posterior lamella (
<emphasis id="89924A6DC175FF901F8FF3BAFE4FFD04" box="[472,495,653,679]" italics="true" pageId="24" pageNumber="25">pl</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="F3FCC5F4C175FF901EEAF3F9FEA8FC99" pageId="24" pageNumber="25" type="multiple">
<paragraph id="BB59967FC175FF901EEAF3F9FE4EFD4B" blockId="24.[189,494,718,744]" box="[189,494,718,744]" pageId="24" pageNumber="25">
<heading id="E0112113C175FF901EEAF3F9FE4EFD4B" bold="true" box="[189,494,718,744]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="89924A6DC175FF901EEAF3F9FE4EFD4B" bold="true" box="[189,494,718,744]" pageId="24" pageNumber="25">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="BB59967FC175FF901EEAF3CAFEA8FC99" blockId="24.[189,1399,765,826]" pageId="24" pageNumber="25">
Known only from Kigogo FR, Udzungwa Mts. Altitudinal range:
<quantity id="7C1E3B9AC175FF901D93F3CAFBCAFCB4" box="[964,1130,765,791]" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="1.9" metricValueMin="1.7" pageId="24" pageNumber="25" unit="m" value="1800.0" valueMax="1900.0" valueMin="1700.0">17001900 m</quantity>
. Habitat: montane rain forest.
</paragraph>
</subSubSection>
<subSubSection id="F3FCC5F4C175FF901EEAF256FC03FC6E" pageId="24" pageNumber="25" type="discussion">
<paragraph id="BB59967FC175FF901EEAF256FE3EFCD8" blockId="24.[189,414,865,891]" box="[189,414,865,891]" pageId="24" pageNumber="25">
<heading id="E0112113C175FF901EEAF256FE3EFCD8" bold="true" box="[189,414,865,891]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="89924A6DC175FF901EEAF256FE3EFCD8" bold="true" box="[189,414,865,891]" pageId="24" pageNumber="25">Coexisting species</emphasis>
</heading>
</paragraph>
<paragraph id="BB59967FC175FF901EEAF2A7FC03FC6E" blockId="24.[189,1398,912,974]" pageId="24" pageNumber="25">
All four species known from Kigogo FR, i.e.,
<taxonomicName id="7CE6EDFCC175FF901C8BF2A7FC20FC09" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[732,896,912,938]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="quasimodo" status="sp. nov.">
<emphasis id="89924A6DC175FF901C8BF2A7FC20FC09" box="[732,896,912,938]" italics="true" pageId="24" pageNumber="25">C. quasimodo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C175FF901DD0F2A7FC42FC09" box="[903,994,912,938]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="7CE6EDFCC175FF901DA6F2A7FBC3FC09" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1009,1123,912,938]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="scopus" status="sp. nov.">
<emphasis id="89924A6DC175FF901DA6F2A7FBC3FC09" box="[1009,1123,912,938]" italics="true" pageId="24" pageNumber="25">C. scopus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C175FF901A3CF2A7FB66FC09" box="[1131,1222,912,938]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="7CE6EDFCC175FF901A83F2A7FAEDFC09" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1236,1357,912,938]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="howelli" status="sp. nov.">
<emphasis id="89924A6DC175FF901A83F2A7FAEDFC09" box="[1236,1357,912,938]" italics="true" pageId="24" pageNumber="25">C. howelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C175FF901B02F2A7FF4FFC6D" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="7CE6EDFCC175FF901F70F283FE14FC6E" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[295,436,947,974]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="mwabvui" status="sp. nov.">
<emphasis id="89924A6DC175FF901F70F283FE14FC6E" box="[295,436,947,974]" italics="true" pageId="24" pageNumber="25">C. mwabvui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C175FF901FECF283FDB8FC6D" box="[443,536,948,974]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
, were found in the same samples.
</paragraph>
</subSubSection>
</treatment>
</document>