treatments-xml/data/03/F2/87/03F287DFFF8AFFE43E27FECF95BB51ED.xml

<document id="5EDA9AE61494BB4BC4D2E854A6D3C9FC" ID-CLB-Dataset="6626" ID-DOI="10.5852/ejt.2021.750.1361" ID-GBIF-Dataset="f3d698f2-f653-4be9-8d49-9ac6502912c0" ID-ISSN="2118-9773" ID-Zenodo-Dep="4770536" ID-ZooBank="560648AD-81B8-464C-B408-6BA92BA086C8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621350363403" checkinUser="felipe" docAuthor="Wong, Man-Kwan, Lee, Mao-Ying &amp; Chen, Wei-Jen" docDate="2021" docId="03F287DFFF8AFFE43E27FECF95BB51ED" docLanguage="en" docName="ejt-750_52-69.pdf" docOrigin="European Journal of Taxonomy 750" docSource="http://dx.doi.org/10.5852/ejt.2021.750.1361" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Luciobrotula polylepis Wong &amp; Lee &amp; Chen 2021, sp. nov." docType="treatment" docUuid="E7C043DA-005E-494F-9E00-21454E6E61BA" docUuidSource="ZooBank" docVersion="8" lastPageNumber="63" masterDocId="FFCBFFA7FF8CFFEF3C46FF9D972E5206" masterDocTitle="Integrative taxonomy reveals a rare and new cusk-eel species of Luciobrotula (Teleostei, Ophidiidae) from the Solomon Sea, West Pacific" masterLastPageNumber="69" masterPageNumber="52" pageNumber="58" updateTime="1698939984431" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B448CD76DE7C803B684811DB1FB316D7">Integrative taxonomy reveals a rare and new cusk-eel species of Luciobrotula (Teleostei, Ophidiidae) from the Solomon Sea, West Pacific</mods:title>
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<mods:namePart id="64503C90EACECAA0E17E47ECCBFFE2F8">Wong, Man-Kwan</mods:namePart>
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<mods:affiliation id="C0A2E478D02BC6F46605FA67B9074CA4">Institute of Oceanography, National Taiwan University, No. 1, Sec. 4, Roosevelt Road, Taipei 10617, Taiwan. &amp; Jun.</mods:affiliation>
<mods:nameIdentifier id="1BA04AF4327999660EF9D78FAE92825A" type="email">Jun.mkwong@gmail.com</mods:nameIdentifier>
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<mods:namePart id="CC51523E9F56C2498EA53D44948D51F7">Lee, Mao-Ying</mods:namePart>
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<mods:affiliation id="84A32116F474B00CF0CE4AAE1ED9221C">Marine Fisheries Division, Fisheries Research Institute, Council of Agriculture, No. 199, Heyi Road, Keelung 202008, Taiwan.</mods:affiliation>
<mods:nameIdentifier id="17F2BF9D92E1EE13E7AAB57675AD58DF" type="email">coleopetera@gmail.com</mods:nameIdentifier>
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<mods:namePart id="E982A3A0C603592EED34C6EFE419F4A3">Chen, Wei-Jen</mods:namePart>
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<mods:affiliation id="1AE7F031D7B022A294346D676B3316A1">Institute of Oceanography, National Taiwan University, No. 1, Sec. 4, Roosevelt Road, Taipei 10617, Taiwan.</mods:affiliation>
<mods:nameIdentifier id="F1FD0843A121459E6989C047692831C7" type="email">wjchen.actinops@gmail.com</mods:nameIdentifier>
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<mods:title id="D8F6EB279EB6A30BFDDBB5D2A04D0504">European Journal of Taxonomy</mods:title>
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<treatment id="03F287DFFF8AFFE43E27FECF95BB51ED" ID-DOI="http://doi.org/10.5281/zenodo.4773705" ID-GBIF-Taxon="182237372" ID-Zenodo-Dep="4773705" LSID="urn:lsid:zoobank.org:act:E7C043DA-005E-494F-9E00-21454E6E61BA" httpUri="http://treatment.plazi.org/id/03F287DFFF8AFFE43E27FECF95BB51ED" lastPageId="11" lastPageNumber="63" pageId="6" pageNumber="58">
<subSubSection id="C3416542FF8AFFE93E27FECF94FD536B" box="[609,979,338,365]" pageId="6" pageNumber="58" type="nomenclature">
<paragraph id="8BE436C9FF8AFFE93E27FECF94FD536B" blockId="6.[370,1219,338,432]" box="[609,979,338,365]" pageId="6" pageNumber="58">
<heading id="D0AC81A5FF8AFFE93E27FECF94FD536B" box="[609,979,338,365]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="58" reason="2">
<taxonomicName id="4C5B4D4AFF8AFFE93E27FECF945F536B" authority="Wong &amp; Lee &amp; Chen, 2021" authorityName="Wong &amp; Lee &amp; Chen" authorityYear="2021" box="[609,881,338,365]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="polylepis" status="sp. nov.">
<emphasis id="B92FEADBFF8AFFE93E27FECF945F536B" bold="true" box="[609,881,338,365]" italics="true" pageId="6" pageNumber="58">Luciobrotula polylepis</emphasis>
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<taxonomicNameLabel id="A21C57A0FF8AFFE93F3EFECE94FD536B" box="[888,979,339,365]" pageId="6" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act: 
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<paragraph id="8BE436C9FF8AFFE93EF2FE0B94AE53B6" blockId="6.[370,1219,338,432]" box="[692,896,406,432]" pageId="6" pageNumber="58">
<figureCitation id="13602A4CFF8AFFE93EF2FE0B943353B6" box="[692,797,406,432]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="8.[189,232,1962,1988]" captionStartId-1="9.[190,233,1928,1954]" captionTargetBox-0="[189,1399,915,1918]" captionTargetBox-1="[216,1399,784,1857]" captionTargetId-0="figure-179@8.[189,1399,915,1918]" captionTargetId-1="figure-228@9.[189,1399,758,1888]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Fig. 3. Holotype of Luciobrotula polylepis sp. nov., 168.4 mm SL (NTUM 11915). A. Fresh specimen. B. Preserved specimen. C. Radiograph. Needle points to the lateral line end." captionText-1="Fig. 4. Luciobrotula polylepis sp. nov., holotype (NTUM 11915). A. Dorsal view of sensory pores on head. B. Ventral view of sensory pores on head. C. Median and ventral view of right sagitta. D. First gill raker (left side)." figureDoi-0="http://doi.org/10.5281/zenodo.4770544" figureDoi-1="http://doi.org/10.5281/zenodo.4770546" httpUri-0="https://zenodo.org/record/4770544/files/figure.png" httpUri-1="https://zenodo.org/record/4770546/files/figure.png" pageId="6" pageNumber="58">Figs 3–4</figureCitation>
; 
<tableCitation id="C6D90372FF8AFFE93F6FFE0B94AE53B6" box="[809,896,406,432]" captionStart="Table 2" captionStartId="7.[182,252,565,593]" captionTargetPageId="7" captionText="Table 2. Counts and measurements for species of Luciobrotula Smith &amp; Radcliffe, 1913." httpUri="http://table.plazi.org/id/DF246641FF8BFFE83CF0FDA893E55057" pageId="6" pageNumber="58" tableUuid="DF246641FF8BFFE83CF0FDA893E55057">Table 2</tableCitation>
</paragraph>
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<subSubSection id="C3416542FF8AFFE93CFBFE4B967755EE" pageId="6" pageNumber="58" type="diagnosis">
<paragraph id="8BE436C9FF8AFFE93CFBFE4B961853F6" blockId="6.[189,310,470,496]" box="[189,310,470,496]" pageId="6" pageNumber="58">
<heading id="D0AC81A5FF8AFFE93CFBFE4B961853F6" bold="true" box="[189,310,470,496]" fontSize="11" level="3" pageId="6" pageNumber="58" reason="3">
<emphasis id="B92FEADBFF8AFFE93CFBFE4B961853F6" bold="true" box="[189,310,470,496]" pageId="6" pageNumber="58">Diagnosis</emphasis>
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<paragraph id="8BE436C9FF8AFFE93CFBFD99933950A0" blockId="6.[189,1399,516,678]" pageId="6" pageNumber="58">
<taxonomicName id="4C5B4D4AFF8AFFE93CFBFD9996E05018" authority="Wong &amp; Lee &amp; Chen, 2021" authorityName="Wong &amp; Lee &amp; Chen" authorityYear="2021" box="[189,462,516,542]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="polylepis" status="sp. nov.">
<emphasis id="B92FEADBFF8AFFE93CFBFD9996E05018" box="[189,462,516,542]" italics="true" pageId="6" pageNumber="58">Luciobrotula polylepis</emphasis>
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<taxonomicNameLabel id="A21C57A0FF8AFFE93D98FD99956E5018" box="[478,576,516,542]" pageId="6" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
is morphologically distinct from all congeners by the following combination of characters: lateral line ending below 33 
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dorsal-fin ray; dorsal-fin rays 86, anal-fin rays 70, precaudal vertebrae 13, total vertebrae 50; gill rakers 17 (3 long rakers and 14 dentigerous plates); longest gill raker 2.1% SL; height of posterior margin of maxilla 3.2% SL; distance from the snout to end of lateral line 60% SL; one interorbital pore and four occipital pores.
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<emphasis id="B92FEADBFF8AFFE93CFBFD5196E750E0" bold="true" box="[189,457,716,742]" pageId="6" pageNumber="58">Differential diagnosis</emphasis>
</heading>
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The new species is most similar to 
<taxonomicName id="4C5B4D4AFF8AFFE93E23FD6794235115" authorityName="Nielsen" authorityYear="2009" box="[613,781,761,787]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="brasiliensis">
<emphasis id="B92FEADBFF8AFFE93E23FD6794235115" box="[613,781,761,787]" italics="true" pageId="6" pageNumber="58">L. brasiliensis</emphasis>
</taxonomicName>
because both share the low number of vertebrae. It differs from 
<taxonomicName id="4C5B4D4AFF8AFFE93D10FC8196D15133" authorityName="Nielsen" authorityYear="2009" box="[342,511,795,822]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="brasiliensis">
<emphasis id="B92FEADBFF8AFFE93D10FC8196485133" box="[342,358,796,821]" italics="true" pageId="6" pageNumber="58">L</emphasis>
. 
<emphasis id="B92FEADBFF8AFFE93D3EFC8696D15133" box="[376,511,795,821]" italics="true" pageId="6" pageNumber="58">brasiliensis</emphasis>
</taxonomicName>
by having a much longer lateral line (ending at the 33 
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dorsal-fin ray vs ending at the 2 
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dorsal-fin ray), a slightly more posterior position of the anal-fin origin (first anal ray below dorsal ray no. 22 vs first anal ray below dorsal ray no. 17), more pectoral-fin rays (32 vs 26), more gill rakers (17 vs 13–14), longer gill raker on first arch (2.1% SL vs 1.2% SL).
</paragraph>
<paragraph id="8BE436C9FF8AFFE93CFBFC5F952E5626" blockId="6.[189,1398,961,1056]" pageId="6" pageNumber="58">
<taxonomicName id="4C5B4D4AFF8AFFE93CFBFC5F96E851DA" authority="Wong &amp; Lee &amp; Chen, 2021" authorityName="Wong &amp; Lee &amp; Chen" authorityYear="2021" box="[189,454,962,988]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="polylepis" status="sp. nov.">
<emphasis id="B92FEADBFF8AFFE93CFBFC5F96E851DA" box="[189,454,962,988]" italics="true" pageId="6" pageNumber="58">Luciobrotula polylepis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C57A0FF8AFFE93D8DFC5F950B51DA" box="[459,549,962,988]" pageId="6" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
differs from 
<taxonomicName id="4C5B4D4AFF8AFFE93EF8FC5F941251DA" authorityName="Smith &amp; Radcliffe" authorityYear="1913" box="[702,828,962,988]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="bartschi">
<emphasis id="B92FEADBFF8AFFE93EF8FC5F941251DA" box="[702,828,962,988]" italics="true" pageId="6" pageNumber="58">L. bartschi</emphasis>
</taxonomicName>
(
<figureCitation id="13602A4CFF8AFFE93F0AFC5F949A51DA" box="[844,948,961,988]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="10.[189,232,1925,1951]" captionStartId-1="11.[189,232,1798,1824]" captionStartId-2="13.[189,232,1875,1901]" captionTargetBox-0="[191,1397,842,1875]" captionTargetBox-1="[207,1399,1141,1718]" captionTargetBox-2="[208,1399,1169,1800]" captionTargetId-0="figure-219@10.[191,1397,842,1878]" captionTargetId-1="figure-256@11.[189,1399,1141,1738]" captionTargetId-2="figure-364@13.[189,1399,1167,1808]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="13" captionText-0="Fig. 5. Luciobrotula bartschi Smith &amp; Radcliffe 1913. Voucher specimen, 393.7 mm SL (NTUM 16627). A. Fresh specimen. B. Preserved specimen. C. Radiograph. Second needle points to the end of the lateral line." captionText-1="Fig. 6. Luciobrotula bartschi Smith &amp; Radcliffe 1913. Voucher specimen, 301.7 mm SL (ASIZP 0068164). A. Preserved specimen. B. Radiograph. Needle points to the end of the lateral line." captionText-2="Fig. 7. Luciobrotula bartschi Smith &amp; Radcliffe 1913. Voucher specimen, 181.6 mm SL (ASIZP 0063749). A. Preserved specimen. B. Radiograph. Needle points to the end of the lateral line. Arrows point to fused vertebrae." figureDoi-0="http://doi.org/10.5281/zenodo.4770550" figureDoi-1="http://doi.org/10.5281/zenodo.4770552" figureDoi-2="http://doi.org/10.5281/zenodo.4770556" httpUri-0="https://zenodo.org/record/4770550/files/figure.png" httpUri-1="https://zenodo.org/record/4770552/files/figure.png" httpUri-2="https://zenodo.org/record/4770556/files/figure.png" pageId="6" pageNumber="58">Figs 5–7</figureCitation>
) in having a slightly longer lateral line (ending at the 33 
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dorsal-fin ray vs the 18 
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–26 
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dorsal-fin ray) and narrower posterior margin of maxilla (3.2% SL vs 3.6–4.7% SL).
</paragraph>
<paragraph id="8BE436C9FF8AFFE93CFBFBDB945A56C0" blockId="6.[189,1399,1094,1222]" pageId="6" pageNumber="58">
It differs from 
<taxonomicName id="4C5B4D4AFF8AFFE93D33FBDA96C85666" authorityName="Nielsen" authorityYear="2009" box="[373,486,1094,1120]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="coheni">
<emphasis id="B92FEADBFF8AFFE93D33FBDA96C85666" box="[373,486,1094,1120]" italics="true" pageId="6" pageNumber="58">L. coheni</emphasis>
</taxonomicName>
by having more anal-fin rays (70 vs 59–65), fewer total gill rakers (17 vs 21–26), a more anterior anal-fin origin (anterior anal-fin ray below 17 
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vertebra vs anterior anal-fin ray below 21 
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–22 
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vertebrae), a narrower interorbital space (3.5% SL vs 3.9–5.6% SL), and a narrower posterior margin of the maxilla (3.2% SL vs 3.9–4.9% SL).
</paragraph>
<paragraph id="8BE436C9FF8AFFE93CFBFB709556574D" blockId="6.[189,1399,1260,1355]" pageId="6" pageNumber="58">
It differs from 
<taxonomicName id="4C5B4D4AFF8AFFE93D2FFB7095155700" baseAuthorityName="Cohen" baseAuthorityYear="1964" box="[361,571,1260,1286]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="corethromycter">
<emphasis id="B92FEADBFF8AFFE93D2FFB7095155700" box="[361,571,1260,1286]" italics="true" pageId="6" pageNumber="58">L. corethromycter</emphasis>
</taxonomicName>
by having fewer dorsal-fin rays (86 vs 91–96), fewer gill rakers (17 vs 18–21), and anterior position of the anal-fin origin (first anal-fin ray below the 17 
<superScript id="7C2E9B81FF8AFFE9382AFA909357571D" attach="left" box="[1132,1145,1293,1307]" fontSize="6" pageId="6" pageNumber="58">th</superScript>
vertebra vs first analfin ray below the 20 
<superScript id="7C2E9B81FF8AFFE93DEEFAB2969B573B" attach="right" box="[424,437,1327,1341]" fontSize="6" pageId="6" pageNumber="58">th</superScript>
–22 
<superScript id="7C2E9B81FF8AFFE93DA7FAB296DC573B" attach="left" box="[481,498,1327,1341]" fontSize="6" pageId="6" pageNumber="58">nd</superScript>
vertebrae).
</paragraph>
<paragraph id="8BE436C9FF8AFFE93CFBFAEC931557F7" blockId="6.[189,1399,1392,1521]" pageId="6" pageNumber="58">
It differs from 
<taxonomicName id="4C5B4D4AFF8AFFE93D2FFAEC9692578C" authorityName="Cohen" authorityYear="1981" box="[361,444,1392,1418]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="nolfi">
<emphasis id="B92FEADBFF8AFFE93D2FFAEC9692578C" box="[361,444,1392,1418]" italics="true" pageId="6" pageNumber="58">L. nolfi</emphasis>
</taxonomicName>
by having a slightly longer lateral line (ending at the 33 
<superScript id="7C2E9B81FF8AFFE9380AFAED93745778" attach="left" box="[1100,1114,1392,1406]" fontSize="6" pageId="6" pageNumber="58">rd</superScript>
dorsal-fin ray vs ending at the 27 
<superScript id="7C2E9B81FF8AFFE93D61FA0F961A57A6" attach="none" box="[295,308,1426,1440]" fontSize="6" pageId="6" pageNumber="58">th</superScript>
–31 
<superScript id="7C2E9B81FF8AFFE93D27FA0F964257A6" attach="left" box="[353,364,1426,1440]" fontSize="6" pageId="6" pageNumber="58">st</superScript>
dorsal-fin ray), slightly more anterior position of the anal-fin origin (first anal-fin ray below the 17 
<superScript id="7C2E9B81FF8AFFE93D1EFA29964B57C4" attach="left" box="[344,357,1460,1474]" fontSize="6" pageId="6" pageNumber="58">th</superScript>
vertebra vs first anal-fin ray below the 19 
<superScript id="7C2E9B81FF8AFFE93F1BFA29944457C4" attach="none" box="[861,874,1460,1474]" fontSize="6" pageId="6" pageNumber="58">th</superScript>
–20 
<superScript id="7C2E9B81FF8AFFE93FD0FA29948D57C4" attach="left" box="[918,931,1460,1474]" fontSize="6" pageId="6" pageNumber="58">th</superScript>
vertebrae), smaller head (23.9% SL vs 24.5–28.0% SL), and relatively deeper body (16.3% SL vs 12.5–15.0% SL).
</paragraph>
<paragraph id="8BE436C9FF8AFFE93CFBF98A93C05491" blockId="6.[189,1399,1558,1687]" pageId="6" pageNumber="58">
It differs from 
<taxonomicName id="4C5B4D4AFF8AFFE93D2EF98596F95437" baseAuthorityName="Gosline" baseAuthorityYear="1954" box="[360,471,1559,1585]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="lineata">
<emphasis id="B92FEADBFF8AFFE93D2EF98596F95437" box="[360,471,1559,1585]" italics="true" pageId="6" pageNumber="58">L. lineata</emphasis>
</taxonomicName>
by having a much longer lateral line (ending at the 33 
<superScript id="7C2E9B81FF8AFFE9380BF98B93755422" attach="left" box="[1101,1115,1558,1572]" fontSize="6" pageId="6" pageNumber="58">rd</superScript>
dorsal-fin ray vs ending at the 2 
<superScript id="7C2E9B81FF8AFFE93D5FF9A596045440" attach="left" box="[281,298,1592,1606]" fontSize="6" pageId="6" pageNumber="58">nd</superScript>
dorsal-fin ray), fewer dorsal-fin rays (86 vs 92), more pectoral-fin rays (32 vs 26), shorter pelvic-fin rays (10.9% SL vs 15.0% SL) and longer gill raker on the first arch (2.1% SL vs 0.7% SL). A detailed comparison between the new species and other congeners is provided in 
<tableCitation id="C6D90372FF8AFFE938D6F9E093C45491" box="[1168,1258,1661,1687]" captionStart="Table 2" captionStartId="7.[182,252,565,593]" captionTargetPageId="7" captionText="Table 2. Counts and measurements for species of Luciobrotula Smith &amp; Radcliffe, 1913." httpUri="http://table.plazi.org/id/DF246641FF8BFFE83CF0FDA893E55057" pageId="6" pageNumber="58" tableUuid="DF246641FF8BFFE83CF0FDA893E55057">Table 2</tableCitation>
.
</paragraph>
<paragraph id="8BE436C9FF8AFFE93CFBF920967755EE" blockId="6.[188,1399,1725,2025]" pageId="6" pageNumber="58">
Along the 
<emphasis id="B92FEADBFF8AFFE93D7CF920964354D1" box="[314,365,1725,1751]" italics="true" pageId="6" pageNumber="58">COI</emphasis>
gene, the following apomorphic sites are unique nucleotides from the only specimen of 
<taxonomicName id="4C5B4D4AFF8AFFE93CFBF97D966B54FF" authority="Wong &amp; Lee &amp; Chen, 2021" authorityName="Wong &amp; Lee &amp; Chen" authorityYear="2021" box="[189,325,1759,1785]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="polylepis" status="sp. nov.">
<emphasis id="B92FEADBFF8AFFE93CFBF97D966B54FF" box="[189,325,1759,1785]" italics="true" pageId="6" pageNumber="58">L. polylepis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C57A0FF8AFFE93D0DF97D968B54FC" box="[331,421,1760,1786]" pageId="6" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
examined here; these nucleotide sites can be used for the molecular diagnosis of the species to differentiate it from 
<taxonomicName id="4C5B4D4AFF8AFFE93E6CF89F95B6551D" authorityName="Nielsen" authorityYear="2009" box="[554,664,1793,1819]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="coheni">
<emphasis id="B92FEADBFF8AFFE93E6CF89F95B6551D" box="[554,664,1793,1819]" italics="true" pageId="6" pageNumber="58">L. coheni</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5B4D4AFF8AFFE93E94F89F947C551D" authorityName="Smith &amp; Radcliffe" authorityYear="1913" box="[722,850,1793,1819]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="bartschi">
<emphasis id="B92FEADBFF8AFFE93E94F89F947C551D" box="[722,850,1793,1819]" italics="true" pageId="6" pageNumber="58">L. bartschi</emphasis>
</taxonomicName>
examined in this study. Nos. 97 (C vs T), 120 (A vs G), 147 (G vs A), 177 (G vs A), 180 (C vs T), 198 (C vs T), 219 (T vs C), 225 (C vs T), 294 (C vs T), 321 (A vs C), 324 (G vs A), 330 (A vs G), 336 (A vs C), 348 (G vs A or C), 363 (G vs T or C), 369 (T vs C), 372 (A vs C or T), 375 (C vs T), 381 (G vs A), 387 (C vs T), 390 (T vs C), 405 (C vs T), 420 (A vs G or C), 426 (T vs C), 465 (A vs C or G), 477 (A vs G), 540 (A vs G), 555 (G vs A), 565 (T vs C), 597 (T vs C), 603 (T vs A), 615 (C vs A), 648 (T vs C), 682 (C vs A or G), 675 (C vs T), 684 (G vs A), 687 (T vs C).
</paragraph>
</subSubSection>
<caption id="DF246641FF8BFFE83CF0FDA893E55057" ID-Table-UUID="DF246641FF8BFFE83CF0FDA893E55057" box="[182,1227,565,593]" httpUri="http://table.plazi.org/id/DF246641FF8BFFE83CF0FDA893E55057" pageId="7" pageNumber="59" startId="7.[182,252,565,593]" targetBox="[183,1924,628,1611]" targetIsTable="true" targetPageId="7">
<paragraph id="8BE436C9FF8BFFE83CF0FDA893E55057" blockId="7.[182,1227,565,593]" box="[182,1227,565,593]" pageId="7" pageNumber="59">
<emphasis id="B92FEADBFF8BFFE83CF0FDA896375057" bold="true" box="[182,281,565,593]" pageId="7" pageNumber="59">Table 2.</emphasis>
Counts and measurements for species of 
<taxonomicName id="4C5B4D4AFF8BFFE83F47FDA893EA5057" authority="Smith &amp; Radcliffe, 1913" authorityName="Smith &amp; Radcliffe" authorityYear="1913" box="[769,1220,565,593]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="59" phylum="Chordata" rank="genus">
<emphasis id="B92FEADBFF8BFFE83F47FDA894B55057" box="[769,923,565,593]" italics="true" pageId="7" pageNumber="59">Luciobrotula</emphasis>
Smith &amp; 
<bibRefCitation id="EFCA4B38FF8BFFE83848FDA893EA5057" author="Radcliffe L." box="[1038,1220,565,593]" pageId="7" pageNumber="59" pagination="135 - 176" refId="ref9137" refString="Radcliffe L. 1913. Descriptions of seven new genera and thirty-one new species of fishes of the families Brotulidae and Carapidae from the Philippine Islands and the Dutch East Indies. Proceedings of the United States National Museum 44 (1948): 135 - 176. https: // doi. org / 10.5479 / si. 00963801.44 - 1948.135" type="journal article" year="1913">Radcliffe, 1913</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BE436C9FF8BFFE83E2CFDE99059544D" pageId="7" pageNumber="59">
<table id="F95BC469FF8B00103CF1FDE990AA544D" box="[183,1924,628,1611]" gridcols="9" gridrows="5" pageId="7" pageNumber="59">
<tr id="356B348BFF8B00103CF1FDE990AA50A8" box="[183,1924,628,686]" gridrow="0" pageId="7" pageNumber="59" rowspan-0="1">
<th id="76BA5DF7FF8B00103E2CFDE995C750A8" box="[618,745,628,686]" gridcol="1" gridrow="0" pageId="7" pageNumber="59">
<emphasis id="B92FEADBFF8BFFE83E2CFDE995E350A8" bold="true" pageId="7" pageNumber="59">
<taxonomicName id="4C5B4D4AFF8BFFE83E2CFDE995C75088" authority="Wong &amp; Lee &amp; Chen, 2021" authorityName="Wong &amp; Lee &amp; Chen" authorityYear="2021" box="[618,745,628,654]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="polylepis" status="sp. nov.">
<emphasis id="B92FEADBFF8BFFE83E2CFDE995C75088" bold="true" box="[618,745,628,654]" italics="true" pageId="7" pageNumber="59">L. polylepis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C57A0FF8BFFE83E31FD0995E350A8" box="[631,717,660,686]" pageId="7" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="B92FEADBFF8BFFE83E8BFD0E95F350A4" box="[717,733,659,674]" italics="true" pageId="7" pageNumber="59">**</emphasis>
</th>
<th id="76BA5DF7FF8B00103F43FDE994A350A8" box="[773,909,628,686]" gridcol="2" gridrow="0" pageId="7" pageNumber="59">
<taxonomicName id="4C5B4D4AFF8BFFE83F43FD1994535098" authorityName="Smith &amp; Radcliffe" authorityYear="1913" box="[773,893,644,670]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="bartschi">
<emphasis id="B92FEADBFF8BFFE83F43FD1994535098" bold="true" box="[773,893,644,670]" italics="true" pageId="7" pageNumber="59">L. bartschi</emphasis>
</taxonomicName>
<emphasis id="B92FEADBFF8BFFE83F3BFD1E94A35094" box="[893,909,643,658]" italics="true" pageId="7" pageNumber="59">**</emphasis>
</th>
<th id="76BA5DF7FF8B00103FE6FDE9930E50A8" box="[928,1056,628,686]" gridcol="3" gridrow="0" pageId="7" pageNumber="59">
<taxonomicName id="4C5B4D4AFF8BFFE83FE6FD1993365098" authorityName="Smith &amp; Radcliffe" authorityYear="1913" box="[928,1048,644,670]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="bartschi">
<emphasis id="B92FEADBFF8BFFE83FE6FD1993365098" bold="true" box="[928,1048,644,670]" italics="true" pageId="7" pageNumber="59">L. bartschi</emphasis>
</taxonomicName>
*
</th>
<th id="76BA5DF7FF8B00103879FDE993CD50A8" box="[1087,1251,628,686]" gridcol="4" gridrow="0" pageId="7" pageNumber="59">
<taxonomicName id="4C5B4D4AFF8BFFE83879FD1993F55098" authorityName="Nielsen" authorityYear="2009" box="[1087,1243,644,670]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="brasiliensis">
<emphasis id="B92FEADBFF8BFFE83879FD1993F55098" bold="true" box="[1087,1243,644,670]" italics="true" pageId="7" pageNumber="59">L. brasiliensis</emphasis>
</taxonomicName>
*
</th>
<th id="76BA5DF7FF8B001038BAFDE992AC50A8" box="[1276,1410,628,686]" gridcol="5" gridrow="0" pageId="7" pageNumber="59">
<taxonomicName id="4C5B4D4AFF8BFFE83941FD19925E5098" authorityName="Nielsen" authorityYear="2009" box="[1287,1392,644,670]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="coheni">
<emphasis id="B92FEADBFF8BFFE83941FD19925E5098" bold="true" box="[1287,1392,644,670]" italics="true" pageId="7" pageNumber="59">L. coheni</emphasis>
</taxonomicName>
*
</th>
<th id="76BA5DF7FF8B001039DEFDE9914650A8" box="[1432,1640,628,686]" gridcol="6" gridrow="0" pageId="7" pageNumber="59">
<taxonomicName id="4C5B4D4AFF8BFFE839DEFD1991715098" baseAuthorityName="Cohen" baseAuthorityYear="1964" box="[1432,1631,644,670]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="corethromycter">
<emphasis id="B92FEADBFF8BFFE839DEFD1991715098" bold="true" box="[1432,1631,644,670]" italics="true" pageId="7" pageNumber="59">L. corethromycter</emphasis>
</taxonomicName>
*
</th>
<th id="76BA5DF7FF8B00103ACDFDE991D050A8" box="[1675,1790,628,686]" gridcol="7" gridrow="0" pageId="7" pageNumber="59">
<taxonomicName id="4C5B4D4AFF8BFFE83ACDFD1991DB5098" baseAuthorityName="Gosline" baseAuthorityYear="1954" box="[1675,1781,644,670]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="lineata">
<emphasis id="B92FEADBFF8BFFE83ACDFD1991DB5098" bold="true" box="[1675,1781,644,670]" italics="true" pageId="7" pageNumber="59">L. lineata</emphasis>
</taxonomicName>
*
</th>
<th id="76BA5DF7FF8B00103B5FFDE990AA50A8" box="[1817,1924,628,686]" gridcol="8" gridrow="0" pageId="7" pageNumber="59">
<taxonomicName id="4C5B4D4AFF8BFFE83B64FD19905D5098" authorityName="Cohen" authorityYear="1981" box="[1826,1907,644,670]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="nolfi">
<emphasis id="B92FEADBFF8BFFE83B64FD19905D5098" bold="true" box="[1826,1907,644,670]" italics="true" pageId="7" pageNumber="59">L. nolfi</emphasis>
</taxonomicName>
*
</th>
</tr>
<tr id="356B348BFF8B00103CF1FD2490AA50D5" box="[183,1924,697,723]" gridrow="1" pageId="7" pageNumber="59" rowspan-0="1">
<td id="76BA5DF7FF8B00103E2CFD2495C750D5" box="[618,745,697,723]" gridcol="1" gridrow="1" pageId="7" pageNumber="59">
<emphasis id="B92FEADBFF8BFFE83E33FD2495F050D5" bold="true" box="[629,734,697,723]" pageId="7" pageNumber="59">Holotype</emphasis>
</td>
<td id="76BA5DF7FF8B00103F43FD2494A350D5" box="[773,909,697,723]" gridcol="2" gridrow="1" pageId="7" pageNumber="59">
<emphasis id="B92FEADBFF8BFFE83F75FD24944B50D5" bold="true" box="[819,869,697,723]" pageId="7" pageNumber="59">n= 6</emphasis>
</td>
<td id="76BA5DF7FF8B00103FE6FD24930E50D5" box="[928,1056,697,723]" gridcol="3" gridrow="1" pageId="7" pageNumber="59">
<emphasis id="B92FEADBFF8BFFE83F80FD2494D750D5" bold="true" box="[966,1017,697,723]" pageId="7" pageNumber="59">n = 9</emphasis>
</td>
<td id="76BA5DF7FF8B00103879FD2493CD50D5" box="[1087,1251,697,723]" gridcol="4" gridrow="1" pageId="7" pageNumber="59">
<emphasis id="B92FEADBFF8BFFE8383EFD24938450D5" bold="true" box="[1144,1194,697,723]" pageId="7" pageNumber="59">n= 1</emphasis>
</td>
<td id="76BA5DF7FF8B001038BAFD2492AC50D5" box="[1276,1410,697,723]" gridcol="5" gridrow="1" pageId="7" pageNumber="59">
<emphasis id="B92FEADBFF8BFFE83960FD24927650D5" bold="true" box="[1318,1368,697,723]" pageId="7" pageNumber="59">n= 9</emphasis>
</td>
<td id="76BA5DF7FF8B001039DEFD24914650D5" box="[1432,1640,697,723]" gridcol="6" gridrow="1" pageId="7" pageNumber="59">
<emphasis id="B92FEADBFF8BFFE839A1FD24913750D5" bold="true" box="[1511,1561,697,723]" pageId="7" pageNumber="59">n= 9</emphasis>
</td>
<td id="76BA5DF7FF8B00103ACDFD2491D050D5" box="[1675,1790,697,723]" gridcol="7" gridrow="1" pageId="7" pageNumber="59">
<emphasis id="B92FEADBFF8BFFE83AEDFD2491F350D5" bold="true" box="[1707,1757,697,723]" pageId="7" pageNumber="59">n =1</emphasis>
</td>
<td id="76BA5DF7FF8B00103B5FFD2490AA50D5" box="[1817,1924,697,723]" gridcol="8" gridrow="1" pageId="7" pageNumber="59">
<emphasis id="B92FEADBFF8BFFE83B73FD24904650D5" bold="true" box="[1845,1896,697,723]" pageId="7" pageNumber="59">n = 8</emphasis>
</td>
</tr>
<tr id="356B348BFF8B00103CF1FD7990AA50F8" box="[183,1924,740,766]" gridrow="2" pageId="7" pageNumber="59">
<th id="76BA5DF7FF8B00103CF1FD79956150F8" box="[183,591,740,766]" gridcol="0" gridrow="2" pageId="7" pageNumber="59">Standard length (SL, mm)</th>
<td id="76BA5DF7FF8B00103E2CFD7995C750F8" box="[618,745,740,766]" gridcol="1" gridrow="2" pageId="7" pageNumber="59">168.4</td>
<td id="76BA5DF7FF8B00103F43FD7994A350F8" box="[773,909,740,766]" gridcol="2" gridrow="2" pageId="7" pageNumber="59">97.9–393.7</td>
<td id="76BA5DF7FF8B00103FE6FD79930E50F8" box="[928,1056,740,766]" gridcol="3" gridrow="2" pageId="7" pageNumber="59">85.0–405.0</td>
<td id="76BA5DF7FF8B00103879FD7993CD50F8" box="[1087,1251,740,766]" gridcol="4" gridrow="2" pageId="7" pageNumber="59">275.0</td>
<td id="76BA5DF7FF8B001038BAFD7992AC50F8" box="[1276,1410,740,766]" gridcol="5" gridrow="2" pageId="7" pageNumber="59">164.0–455.0</td>
<td id="76BA5DF7FF8B001039DEFD79914650F8" box="[1432,1640,740,766]" gridcol="6" gridrow="2" pageId="7" pageNumber="59">123.0–534.0</td>
<td id="76BA5DF7FF8B00103ACDFD7991D050F8" box="[1675,1790,740,766]" gridcol="7" gridrow="2" pageId="7" pageNumber="59">267.0</td>
<td id="76BA5DF7FF8B00103B5FFD7990AA50F8" box="[1817,1924,740,766]" gridcol="8" gridrow="2" pageId="7" pageNumber="59">137.0–</td>
</tr>
<tr id="356B348BFF8B00103CF1FC9990AA56A5" box="[183,1924,772,1187]" gridrow="3" pageId="7" pageNumber="59">
<th id="76BA5DF7FF8B00103CF1FC99956156A5" box="[183,591,772,1187]" gridcol="0" gridrow="3" pageId="7" pageNumber="59">
<emphasis id="B92FEADBFF8BFFE83CF1FC8696275133" bold="true" box="[183,265,795,821]" pageId="7" pageNumber="59">Counts</emphasis>
Dorsal-fin rays Caudal-fin rays Anal-fin rays Pectoral-fin rays Pelvic-fin rays Precaudal vertebrae Total vertebrae Pseudobranchial filaments Ant. dorsal ray above vertebra no. Ant. anal ray below dorsal ray no. Ant. anal-fin ray below vertebra no. End of lateral line below dorsal ray no. Total gill rakers Long gill rakers
</th>
<td id="76BA5DF7FF8B00103E2CFC9995C756A5" box="[618,745,772,1187]" gridcol="1" gridrow="3" pageId="7" pageNumber="59">86 11 70 32 2 13 50 NA 8 22 17 33 17 3</td>
<td id="76BA5DF7FF8B00103F43FC9994A356A5" box="[773,909,772,1187]" gridcol="2" gridrow="3" pageId="7" pageNumber="59">85–92 10–11 68–75 25–27 2 (14#)15–16 (49#)52–53 NA 7–9 19–24 17–20 18–26 14–16 3</td>
<td id="76BA5DF7FF8B00103FE6FC99930E56A5" box="[928,1056,772,1187]" gridcol="3" gridrow="3" pageId="7" pageNumber="59">87–94 11–12 69–75 25–27 2 15–16 53–55 2 7–9 20–24 18–20 19–26 16–17 3</td>
<td id="76BA5DF7FF8B00103879FC9993CD56A5" box="[1087,1251,772,1187]" gridcol="4" gridrow="3" pageId="7" pageNumber="59">84 12 69 26 2 13 51 2 8 17 17 2 13–14 3</td>
<td id="76BA5DF7FF8B001038BAFC9992AC56A5" box="[1276,1410,772,1187]" gridcol="5" gridrow="3" pageId="7" pageNumber="59">81–89 11 59–65 27–31 2 15 53–54 2 6–8 25–29 21–22 28–37 21–26 3</td>
<td id="76BA5DF7FF8B001039DEFC99914656A5" box="[1432,1640,772,1187]" gridcol="6" gridrow="3" pageId="7" pageNumber="59">91–96 10 68–74 26–27 2 16 56–57 2 7–9 24–28 20–22 24–32 18–21 3</td>
<td id="76BA5DF7FF8B00103ACDFC9991D056A5" box="[1675,1790,772,1187]" gridcol="7" gridrow="3" pageId="7" pageNumber="59">92 12 76 26 2 15 56 2–3 11 20 20 2 16–17 3</td>
<td id="76BA5DF7FF8B00103B5FFC9990AA56A5" box="[1817,1924,772,1187]" gridcol="8" gridrow="3" pageId="7" pageNumber="59">590.0 86–95 10–11 66–70 27–28 2 15–16 53–55 2 6–8 24–26 19–20 27–31 16–19 3</td>
</tr>
<tr id="356B348BFF8B00103CF1FB3590AA544D" box="[183,1924,1192,1611]" gridrow="4" pageId="7" pageNumber="59">
<th id="76BA5DF7FF8B00103CF1FB359561544D" box="[183,591,1192,1611]" gridcol="0" gridrow="4" pageId="7" pageNumber="59">
<emphasis id="B92FEADBFF8BFFE83CF1FB3596CD56C4" bold="true" box="[183,483,1192,1218]" pageId="7" pageNumber="59">Measurements in % of SL</emphasis>
Head Depth at origin of anal fin Upper jaw Dia. of eye window Interorbital Postorbital Height of posterior margin of maxilla Snout to end of lateral line Preanal Predorsal Base of pelvic to origin of anal fin Pectoral fin Pelvic-fin longest ray Longest gill filament on anterior arch Longest gill rakers on anterior arch
</th>
<td id="76BA5DF7FF8B00103E2CFB3595C7544D" box="[618,745,1192,1611]" gridcol="1" gridrow="4" pageId="7" pageNumber="59">23.9 16.3 10.6 2.9 3.5 14.3 3.2 59.6 48.2 31.1 33.2 9.1 10.9 1.1 2.1</td>
<td id="76BA5DF7FF8B00103F43FB3594A3544D" box="[773,909,1192,1611]" gridcol="2" gridrow="4" pageId="7" pageNumber="59">22.6–24.9 13.4–17.6 10.2–12.6 2.6–3.2 3.4–3.9 14.1–15.9 3.7–4.5 47.5–50.7 48.9–52.1 28.0–32.9 31.7–36.8 9.0–11.1 10.5–14.2 1.3–1.8 2.0–3.2</td>
<td id="76BA5DF7FF8B00103FE6FB35930E544D" box="[928,1056,1192,1611]" gridcol="3" gridrow="4" pageId="7" pageNumber="59">22.5–26.5 12.5–16.5 11.0–13.5 2.4–3.1 3.4–4.5 15.0–16.5 3.6–4.7 48.0–56.0 48.0–54.0 30.0–34.5 30.0–35.0 9.1–12.0 11.5–16.0 1.3–2.5 2.2–2.7</td>
<td id="76BA5DF7FF8B00103879FB3593CD544D" box="[1087,1251,1192,1611]" gridcol="4" gridrow="4" pageId="7" pageNumber="59">22.5 14.5 12.0 2.5 3.8 15.0 4.5 34.0 48.5 32.5 32.0 11.5 14.5 1.1 1.2</td>
<td id="76BA5DF7FF8B001038BAFB3592AC544D" box="[1276,1410,1192,1611]" gridcol="5" gridrow="4" pageId="7" pageNumber="59">24.5–30.5 14.5–17.5 12.0–15.0 2.7–3.3 3.9–5.6 16.5–19.5 3.9–4.9 60.0–71.0 52.0–62.0 26.0–32.5 36.5–40.5 9.1–12.0 11.5–15.0 2.8–3.6 1.8–3.1</td>
<td id="76BA5DF7FF8B001039DEFB359146544D" box="[1432,1640,1192,1611]" gridcol="6" gridrow="4" pageId="7" pageNumber="59">21.0–25.0 12.0–15.5 11.0–13.0 2.4–2.9 3.8–4.5 13.0–16.5 3.3–4.9 45.5–57.0 47.0–59.0 26.5–33.0 32.5–41.0 9.6–12.0 7.9–9.6 1.3–1.8 1.9–2.5</td>
<td id="76BA5DF7FF8B00103ACDFB3591D0544D" box="[1675,1790,1192,1611]" gridcol="7" gridrow="4" pageId="7" pageNumber="59">22.5 15.0 11.0 3.3 3.3 15.0 NA 33.0 49.5 32.0 29.0 10.5 15.0 1.0 0.7</td>
<td id="76BA5DF7FF8B00103B5FFB3590AA544D" box="[1817,1924,1192,1611]" gridcol="8" gridrow="4" pageId="7" pageNumber="59">24.5–28.0 12.5–15.0 11.5–12.5 3.0–3.5 4.0–5.0 15.0–16.5 3.9–5.0 51.0–55.0 47.0–58.0 28.5–35.5 32.0–33.5 11.0–12.5 11.0–12.5 1.4–2.0 2.1–2.4</td>
</tr>
</table>
</paragraph>
<footnote id="E8402AC7FF8BFFE83CF1F9EE949154C5" pageId="7" pageNumber="59">
<paragraph id="8BE436C9FF8BFFE83CF1F9EE96FD548C" blockId="7.[183,959,1651,1731]" box="[183,467,1651,1674]" pageId="7" pageNumber="59">
* Data from 
<bibRefCitation id="EFCA4B38FF8BFFE83D7BF9EE96E1548C" author="Nielsen J. G." box="[317,463,1651,1674]" pageId="7" pageNumber="59" pagination="141 - 156" refId="ref8790" refString="Nielsen J. G. 2009. A revision of the bathyal genus Luciobrotula (Teleostei, Ophidiidae) with two new species. Galathea Report 22: 141 - 156." type="book chapter" year="2009">Nielsen (2009)</bibRefCitation>
.
</paragraph>
<paragraph id="8BE436C9FF8BFFE83CF1F90D95AF54A1" blockId="7.[183,959,1651,1731]" box="[183,641,1680,1703]" pageId="7" pageNumber="59">** Data from examined specimens of this study</paragraph>
<paragraph id="8BE436C9FF8BFFE83CF1F931949154C5" blockId="7.[183,959,1651,1731]" box="[183,959,1708,1731]" pageId="7" pageNumber="59"># Data (in the parentheses) from an abnormal vertebrae development specimen.</paragraph>
</footnote>
<subSubSection id="C3416542FF84FFE73CFBFE94936D5391" pageId="8" pageNumber="60" type="etymology">
<paragraph id="8BE436C9FF84FFE73CFBFE94966B5325" blockId="8.[189,325,265,291]" box="[189,325,265,291]" pageId="8" pageNumber="60">
<heading id="D0AC81A5FF84FFE73CFBFE94966B5325" bold="true" box="[189,325,265,291]" fontSize="11" level="3" pageId="8" pageNumber="60" reason="3">
<emphasis id="B92FEADBFF84FFE73CFBFE94966B5325" bold="true" box="[189,325,265,291]" pageId="8" pageNumber="60">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BE436C9FF84FFE73CFBFEAA936D5391" blockId="8.[189,1399,311,408]" pageId="8" pageNumber="60">
The name 
<taxonomicName id="4C5B4D4AFF84FFE73D04FEAA96835357" authorityName="Wong &amp; Lee &amp; Chen" authorityYear="2021" box="[322,429,311,337]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="polylepis">
<emphasis id="B92FEADBFF84FFE73D04FEAA96835357" box="[322,429,311,337]" italics="true" pageId="8" pageNumber="60">polylepis</emphasis>
</taxonomicName>
is derived from the Greek ‘ 
<emphasis id="B92FEADBFF84FFE73F49FEAA946C5357" box="[783,834,311,337]" italics="true" pageId="8" pageNumber="60">poly</emphasis>
’, meaning ‘many’ or ‘numerous’, and ‘ 
<emphasis id="B92FEADBFF84FFE7396BFEAA924B5357" box="[1325,1381,311,337]" italics="true" pageId="8" pageNumber="60">lepis</emphasis>
’, meaning ‘scales’, in reference to the much longer lateral line and therefore more lateral line scales compared with 
<taxonomicName id="4C5B4D4AFF84FFE73D35FEE396DA5391" authorityName="Smith &amp; Radcliffe" authorityYear="1913" box="[371,500,381,407]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="bartschi">
<emphasis id="B92FEADBFF84FFE73D35FEE396DA5391" box="[371,500,381,407]" italics="true" pageId="8" pageNumber="60">L. bartschi</emphasis>
</taxonomicName>
, the only congener distributed in the West Pacific.
</paragraph>
</subSubSection>
<subSubSection id="C3416542FF84FFE73CFBFE2296455093" pageId="8" pageNumber="60" type="materials_examined">
<paragraph id="8BE436C9FF84FFE73CFBFE22964353DF" blockId="8.[189,365,447,473]" box="[189,365,447,473]" pageId="8" pageNumber="60">
<heading id="D0AC81A5FF84FFE73CFBFE22964353DF" bold="true" box="[189,365,447,473]" fontSize="11" level="3" pageId="8" pageNumber="60" reason="3">
<emphasis id="B92FEADBFF84FFE73CFBFE22964353DF" bold="true" box="[189,365,447,473]" pageId="8" pageNumber="60">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BE436C9FF84FFE73CACFE739673500E" blockId="8.[189,1400,494,661]" box="[234,349,494,520]" pageId="8" pageNumber="60">
<emphasis id="B92FEADBFF84FFE73CACFE739673500E" bold="true" box="[234,349,494,520]" pageId="8" pageNumber="60">
<typeStatus id="54E0886BFF84FFE73CACFE739673500E" box="[234,349,494,520]" pageId="8" pageNumber="60" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BE436C9FF84FFE73CFBFD8C96455093" blockId="8.[189,1400,494,661]" pageId="8" pageNumber="60">
<materialsCitation id="3B333C94FF84FFE73CFBFD8C96455093" ID-GBIF-Occurrence="3114129304" accessionNumber="MW218670" collectingDate="2014-05-06" collectionCode="R, V, ALIS, MW" country="Papua New Guinea" county="West Pacific" elevation="525" latitude="-6.133333" location="Ainto Bay" longLatPrecision="1305" longitude="149.16667" municipality="Solomon Sea" pageId="8" pageNumber="60" specimenCode="PNG2363, NTUM 11915" specimenCount="1" stateProvince="West New Britain" typeStatus="holotype">
SOLOMON SEA • 
<quantity id="4CA39B2CFF84FFE73DE1FD8D950E502D" box="[423,544,528,555]" metricMagnitude="-1" metricUnit="m" metricValue="1.684" pageId="8" pageNumber="60" unit="mm" value="168.4">168.4 mm</quantity>
SL, sample ID: 
<specimenCode id="DBFD9EB2FF84FFE73EA3FD8D9470502C" box="[741,862,528,554]" pageId="8" pageNumber="60">PNG2363</specimenCode>
; 
<location id="8E846012FF84FFE73F2FFD8C94CB502D" LSID="urn:lsid:plazi:treatment:03F287DFFF8AFFE43E27FECF95BB51ED:8E846012FF84FFE73F2FFD8C94CB502D" box="[873,997,529,555]" country="Papua New Guinea" county="West Pacific" latitude="-6.133333" longLatPrecision="1305" longitude="149.16667" municipality="Solomon Sea" name="Ainto Bay" pageId="8" pageNumber="60" stateProvince="West New Britain">Ainto Bay</location>
, 
<locationDeviation id="9A562FEBFF84FFE73FB4FD8C920C502D" box="[1010,1314,529,555]" pageId="8" pageNumber="60">SE of New Britain Island</locationDeviation>
, 
<collectingCountry id="F34C7659FF84FFE73969FD8C96795048" name="Papua New Guinea" pageId="8" pageNumber="60">Papua New Guinea</collectingCountry>
, 
<location id="8E846012FF84FFE73D23FDA9952B5048" LSID="urn:lsid:plazi:treatment:03F287DFFF8AFFE43E27FECF95BB51ED:8E846012FF84FFE73D23FDA9952B5048" box="[357,517,564,590]" country="Papua New Guinea" county="West Pacific" latitude="-6.133333" longLatPrecision="1305" longitude="149.16667" municipality="Solomon Sea" name="Solomon Sea" pageId="8" pageNumber="60" stateProvince="West New Britain">Solomon Sea</location>
, 
<location id="8E846012FF84FFE73E55FDA995845048" LSID="urn:lsid:plazi:treatment:03F287DFFF8AFFE43E27FECF95BB51ED:8E846012FF84FFE73E55FDA995845048" box="[531,682,564,590]" country="Papua New Guinea" county="West Pacific" latitude="-6.133333" longLatPrecision="1305" longitude="149.16667" municipality="Solomon Sea" name="West Pacific" pageId="8" pageNumber="60" stateProvince="West New Britain">West Pacific</location>
, stn CP4334; 
<geoCoordinate id="EE6F500EFF84FFE73F12FDAF94805048" box="[852,942,562,590]" degrees="6" direction="south" minutes="08" orientation="latitude" pageId="8" pageNumber="60" precision="925" value="-6.133333">6°08′ S</geoCoordinate>
, 
<geoCoordinate id="EE6F500EFF84FFE73FFBFDAF93195048" box="[957,1079,562,590]" degrees="149" direction="east" minutes="10" orientation="longitude" pageId="8" pageNumber="60" precision="925" value="149.16667">149°10′ E</geoCoordinate>
; 
<elevation id="0076D1FAFF84FFE73800FDA992355048" box="[1094,1307,564,590]" metricMagnitude="2" metricUnit="m" metricValue="5.25" metricValueMax="6.2" metricValueMin="4.3" pageId="8" pageNumber="60" unit="m" value="525.0" valueMax="620.0" valueMin="430.0">
<quantity id="4CA39B2CFF84FFE73800FDA993E35048" box="[1094,1229,564,590]" metricMagnitude="2" metricUnit="m" metricValue="5.25" metricValueMax="6.2" metricValueMin="4.3" pageId="8" pageNumber="60" unit="m" value="525.0" valueMax="620.0" valueMin="430.0">430–620 m</quantity>
depth
</elevation>
; 
<date id="FFE51009FF84FFE7396FFDA997D55077" pageId="8" pageNumber="60" value="2014-05-06">
<collectingDate id="EFA1E9E1FF84FFE7396FFDA997D55077" pageId="8" pageNumber="60" value="2014-05-06">6 May 2014</collectingDate>
</date>
; R/V 
<emphasis id="B92FEADBFF84FFE73D16FDCA96A55077" box="[336,395,599,625]" italics="true" pageId="8" pageNumber="60">ALIS</emphasis>
; French beam trawl; MADEEP expedition; GenBank registration: 
<accessionNumber id="9408AB2AFF84FFE738A7FDCA925C5077" box="[1249,1394,599,625]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW218670" pageId="8" pageNumber="60">MW218670</accessionNumber>
; 
<specimenCode id="DBFD9EB2FF84FFE73CFBFDE696495093" box="[189,359,634,661]" collectionCode="NTUM" pageId="8" pageNumber="60">NTUM 11915</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3416542FF84FFE43CFBFD2694A25013" lastPageId="11" lastPageNumber="63" pageId="8" pageNumber="60" type="description">
<paragraph id="8BE436C9FF84FFE73CFBFD26966150D3" blockId="8.[189,335,699,725]" box="[189,335,699,725]" pageId="8" pageNumber="60">
<heading id="D0AC81A5FF84FFE73CFBFD26966150D3" bold="true" box="[189,335,699,725]" fontSize="11" level="3" pageId="8" pageNumber="60" reason="3">
<emphasis id="B92FEADBFF84FFE73CFBFD26966150D3" bold="true" box="[189,335,699,725]" pageId="8" pageNumber="60">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BE436C9FF84FFE63CFBFD769412503B" blockId="8.[188,1398,746,843]" lastBlockId="9.[189,1401,265,573]" lastPageId="9" lastPageNumber="61" pageId="8" pageNumber="60">
Measurements and counts of the 
<typeStatus id="54E0886BFF84FFE73E7BFD77958A5102" box="[573,676,746,772]" pageId="8" pageNumber="60" type="holotype">holotype</typeStatus>
given in 
<tableCitation id="C6D90372FF84FFE73F56FD7794445102" box="[784,874,746,772]" captionStart="Table 2" captionStartId="7.[182,252,565,593]" captionTargetPageId="7" captionText="Table 2. Counts and measurements for species of Luciobrotula Smith &amp; Radcliffe, 1913." httpUri="http://table.plazi.org/id/DF246641FF8BFFE83CF0FDA893E55057" pageId="8" pageNumber="60" tableUuid="DF246641FF8BFFE83CF0FDA893E55057">Table 2</tableCitation>
. Body elongate with tapering caudal portion, snout and head slightly depressed; eye small and round, horizontal eye diameter about half of snout length. Mouth large, oblique; upper jaw reaching a vertical through the posterior margin of orbit, posterior part vertically much extended, slightly protruding beyond lower jaw when mouth closed. Boomerang-formed vomer; palatine, and upper and lower jaw with many small, close-set, rather blunt teeth in several irregular rows; fang-like teeth absent in both jaws. One median and a pair of two large basibranchial tooth patches. Anterior nostril with low rim and placed midway between upper lip and posterior nostril, with small rounded flap rising from anterior rim. Posterior margins of preopercle, interopercle, and subopercle rounded, without spine. First gill arch with four finely dentigerous plates on upper branch, one long raker on the angle, and lower branch with two long rakers interspaced with 10 dentigerous plates (
<figureCitation id="13602A4CFF85FFE63D83FE62950C501F" box="[453,546,511,537]" captionStart="Fig" captionStartId="9.[190,233,1928,1954]" captionTargetBox="[216,1399,784,1857]" captionTargetId="figure-228@9.[189,1399,758,1888]" captionTargetPageId="9" captionText="Fig. 4. Luciobrotula polylepis sp. nov., holotype (NTUM 11915). A. Dorsal view of sensory pores on head. B. Ventral view of sensory pores on head. C. Median and ventral view of right sagitta. D. First gill raker (left side)." figureDoi="http://doi.org/10.5281/zenodo.4770546" httpUri="https://zenodo.org/record/4770546/files/figure.png" pageId="9" pageNumber="61">Fig. 4D</figureCitation>
); gill filaments ca 100, the longest about half as long as longest gill raker; pseudobranchial filament damaged, unavailable count.
</paragraph>
<caption id="DF246641FF84FFE73CFBF837931255EE" ID-DOI="http://doi.org/10.5281/zenodo.4770544" ID-Zenodo-Dep="4770544" httpUri="https://zenodo.org/record/4770544/files/figure.png" pageId="8" pageNumber="60" startId="8.[189,232,1962,1988]" targetBox="[189,1399,915,1918]" targetPageId="8">
<paragraph id="8BE436C9FF84FFE73CFBF837931255EE" blockId="8.[189,1399,1962,2024]" pageId="8" pageNumber="60">
<emphasis id="B92FEADBFF84FFE73CFBF837962555C2" bold="true" box="[189,267,1962,1988]" pageId="8" pageNumber="60">Fig. 3.</emphasis>
Holotype of 
<taxonomicName id="4C5B4D4AFF84FFE73DE1F837959A55C2" authority="Wong &amp; Lee &amp; Chen, 2021" authorityName="Wong &amp; Lee &amp; Chen" authorityYear="2021" box="[423,692,1962,1988]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="polylepis" status="sp. nov.">
<emphasis id="B92FEADBFF84FFE73DE1F837959A55C2" box="[423,692,1962,1988]" italics="true" pageId="8" pageNumber="60">Luciobrotula polylepis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C57A0FF84FFE73EFAF836943755C3" box="[700,793,1963,1989]" pageId="8" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
, 168.4 mm SL (NTUM 11915). 
<emphasis id="B92FEADBFF84FFE738D2F837938755C2" bold="true" box="[1172,1193,1962,1988]" pageId="8" pageNumber="60">A</emphasis>
. Fresh specimen. 
<emphasis id="B92FEADBFF84FFE73CFBF85397FF55EE" bold="true" box="[189,209,1998,2024]" pageId="8" pageNumber="60">B</emphasis>
. Preserved specimen. 
<emphasis id="B92FEADBFF84FFE73D9EF85096C355E1" bold="true" box="[472,493,1997,2023]" pageId="8" pageNumber="60">C</emphasis>
. Radiograph. Needle points to the lateral line end.
</paragraph>
</caption>
<paragraph id="8BE436C9FF85FFE53CFBFDF995F8536C" blockId="9.[189,1399,612,709]" lastBlockId="10.[189,1399,265,362]" lastPageId="10" lastPageNumber="62" pageId="9" pageNumber="61">
Sensory pores are found all over head (
<figureCitation id="13602A4CFF85FFE63ECAFDF994205078" box="[652,782,612,638]" captionStart="Fig" captionStartId="9.[190,233,1928,1954]" captionTargetBox="[216,1399,784,1857]" captionTargetId="figure-228@9.[189,1399,758,1888]" captionTargetPageId="9" captionText="Fig. 4. Luciobrotula polylepis sp. nov., holotype (NTUM 11915). A. Dorsal view of sensory pores on head. B. Ventral view of sensory pores on head. C. Median and ventral view of right sagitta. D. First gill raker (left side)." figureDoi="http://doi.org/10.5281/zenodo.4770546" httpUri="https://zenodo.org/record/4770546/files/figure.png" pageId="9" pageNumber="61">Fig. 4A–B</figureCitation>
). Supraorbital with group of eight pores behind eye, five pores immediately above eye, and five small pores in a row on tip of snout, larger pore between flaps on tip of snout, and above each nostril, one interorbital pore, four occipital pores, six suborbital pores and four mandibular pores, 10 small pores close to lower jaw, between this row and mandibular having four small pores, and finally a row of six pores above posterior mandibular, two pores behind posterior end of maxilla, and preopercle with six pores.
</paragraph>
<caption id="DF246641FF85FFE63CF8F815965755EF" ID-DOI="http://doi.org/10.5281/zenodo.4770546" ID-Zenodo-Dep="4770546" httpUri="https://zenodo.org/record/4770546/files/figure.png" pageId="9" pageNumber="61" startId="9.[190,233,1928,1954]" targetBox="[216,1399,784,1857]" targetPageId="9">
<paragraph id="8BE436C9FF85FFE63CF8F815965755EF" blockId="9.[188,1398,1928,2025]" pageId="9" pageNumber="61">
<emphasis id="B92FEADBFF85FFE63CF8F815962255A4" bold="true" box="[190,268,1928,1954]" pageId="9" pageNumber="61">Fig. 4.</emphasis>
<taxonomicName id="4C5B4D4AFF85FFE63D53F815950D55A4" authority="Wong &amp; Lee &amp; Chen, 2021" authorityName="Wong &amp; Lee &amp; Chen" authorityYear="2021" box="[277,547,1928,1954]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="polylepis" status="sp. nov.">
<emphasis id="B92FEADBFF85FFE63D53F815950D55A4" box="[277,547,1928,1954]" italics="true" pageId="9" pageNumber="61">Luciobrotula polylepis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C57A0FF85FFE63E6BF81495A455A5" box="[557,650,1929,1955]" pageId="9" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (NTUM 11915). 
<emphasis id="B92FEADBFF85FFE63F96F81594CB55A4" bold="true" box="[976,997,1928,1954]" pageId="9" pageNumber="61">A</emphasis>
. Dorsal view of sensory pores on head. 
<emphasis id="B92FEADBFF85FFE63D44F836963855C3" bold="true" box="[258,278,1963,1989]" pageId="9" pageNumber="61">B</emphasis>
. Ventral view of sensory pores on head. 
<emphasis id="B92FEADBFF85FFE63EAAF836942F55C3" bold="true" box="[748,769,1963,1989]" pageId="9" pageNumber="61">C</emphasis>
. Median and ventral view of right sagitta. 
<emphasis id="B92FEADBFF85FFE638A9F836922A55C3" bold="true" box="[1263,1284,1963,1989]" pageId="9" pageNumber="61">D</emphasis>
. First gill raker (left side).
</paragraph>
</caption>
<paragraph id="8BE436C9FF86FFE53CFBFE0C956A53F7" blockId="10.[189,1399,401,498]" pageId="10" pageNumber="62">
Sagittal otolith is elongate and thin, about 2.5 times as long as high. Sulcus divided into ostium and cauda. Cauda is about ⅔ of ostium (
<figureCitation id="13602A4CFF86FFE53E27FE29959053C8" box="[609,702,436,462]" captionStart="Fig" captionStartId="9.[190,233,1928,1954]" captionTargetBox="[216,1399,784,1857]" captionTargetId="figure-228@9.[189,1399,758,1888]" captionTargetPageId="9" captionText="Fig. 4. Luciobrotula polylepis sp. nov., holotype (NTUM 11915). A. Dorsal view of sensory pores on head. B. Ventral view of sensory pores on head. C. Median and ventral view of right sagitta. D. First gill raker (left side)." figureDoi="http://doi.org/10.5281/zenodo.4770546" httpUri="https://zenodo.org/record/4770546/files/figure.png" pageId="10" pageNumber="62">Fig. 4C</figureCitation>
). Due to the damaged anterior rim, the presence of an ostial channel could not be ascertained.
</paragraph>
<paragraph id="8BE436C9FF86FFE53CFBFD8495BD50E5" blockId="10.[189,1399,537,739]" pageId="10" pageNumber="62">
Body, top of head, and opercle covered with small cycloid scales, with ca 72 scales in oblique line from origin of anal fin forwards and ca 111 scales from upper part of gill slit to base of caudal fin; single lateral line originating at upper angle of opercle and extending posteriorly in straight line placed about midway between midline and profile of body, ending below 33 
<superScript id="7C2E9B81FF86FFE53FE4FD1C949E5089" attach="left" box="[930,944,641,655]" fontSize="6" pageId="10" pageNumber="62">rd</superScript>
dorsal-fin ray. Dorsal-fin origin above end of pectoral fin; anal-fin origin at about mid-body of fish, pectoral fin placed medially and pelvic fin reaching one third from base to anal fin.
</paragraph>
<caption id="DF246641FF86FFE53CFBF81896C555E0" ID-DOI="http://doi.org/10.5281/zenodo.4770550" ID-Zenodo-Dep="4770550" httpUri="https://zenodo.org/record/4770550/files/figure.png" pageId="10" pageNumber="62" startId="10.[189,232,1925,1951]" targetBox="[191,1397,842,1875]" targetPageId="10">
<paragraph id="8BE436C9FF86FFE53CFBF81896C555E0" blockId="10.[189,1399,1925,2022]" pageId="10" pageNumber="62">
<emphasis id="B92FEADBFF86FFE53CFBF81896345599" bold="true" box="[189,282,1925,1951]" pageId="10" pageNumber="62">Fig. 5.</emphasis>
<taxonomicName id="4C5B4D4AFF86FFE53D76F81894B15599" authority="Smith &amp; Radcliffe 1913" authorityName="Smith &amp; Radcliffe" authorityYear="1913" box="[304,927,1925,1951]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="10" pageNumber="62" phylum="Chordata" rank="species" species="bartschi">
<emphasis id="B92FEADBFF86FFE53D76F818956E5599" box="[304,576,1925,1951]" italics="true" pageId="10" pageNumber="62">Luciobrotula bartschi</emphasis>
Smith &amp; 
<bibRefCitation id="EFCA4B38FF86FFE53E98F81894B15599" author="Radcliffe L." box="[734,927,1925,1951]" pageId="10" pageNumber="62" pagination="135 - 176" refId="ref9137" refString="Radcliffe L. 1913. Descriptions of seven new genera and thirty-one new species of fishes of the families Brotulidae and Carapidae from the Philippine Islands and the Dutch East Indies. Proceedings of the United States National Museum 44 (1948): 135 - 176. https: // doi. org / 10.5479 / si. 00963801.44 - 1948.135" type="journal article" year="1913">Radcliffe 1913</bibRefCitation>
</taxonomicName>
. Voucher specimen, 393.7 mm SL (NTUM 16627). 
<emphasis id="B92FEADBFF86FFE53DCCF83596B155C4" bold="true" box="[394,415,1960,1986]" pageId="10" pageNumber="62">A</emphasis>
. Fresh specimen. 
<emphasis id="B92FEADBFF86FFE53E3DF83495A355C5" bold="true" box="[635,653,1961,1987]" pageId="10" pageNumber="62">B</emphasis>
. Preserved specimen. 
<emphasis id="B92FEADBFF86FFE53FDAF835948155C4" bold="true" box="[924,943,1960,1986]" pageId="10" pageNumber="62">C</emphasis>
. Radiograph. Second needle points to the end of the lateral line.
</paragraph>
</caption>
<paragraph id="8BE436C9FF87FFE43CFBFE9494E5538A" blockId="11.[189,1398,265,397]" pageId="11" pageNumber="63">
Third neural spine pointed, length of first spine half as long as second spine (
<figureCitation id="13602A4CFF87FFE43822FE9493EC5325" box="[1124,1218,265,291]" captionStart="Fig" captionStartId="8.[189,232,1962,1988]" captionTargetBox="[189,1399,915,1918]" captionTargetId="figure-179@8.[189,1399,915,1918]" captionTargetPageId="8" captionText="Fig. 3. Holotype of Luciobrotula polylepis sp. nov., 168.4 mm SL (NTUM 11915). A. Fresh specimen. B. Preserved specimen. C. Radiograph. Needle points to the lateral line end." figureDoi="http://doi.org/10.5281/zenodo.4770544" httpUri="https://zenodo.org/record/4770544/files/figure.png" pageId="11" pageNumber="63">Fig. 3C</figureCitation>
), neural spines of posterior 10 pre-caudal vertebrae with blunt tips and broad bases, 4 
<superScript id="7C2E9B81FF87FFE43845FEB6933E533F" attach="none" box="[1027,1040,299,313]" fontSize="6" pageId="11" pageNumber="63">th</superScript>
–11 
<superScript id="7C2E9B81FF87FFE4387DFEB69366533F" attach="left" box="[1083,1096,299,313]" fontSize="6" pageId="11" pageNumber="63">th</superScript>
precaudal vertebrae with broad bases and depressed neural spines, 7 
<superScript id="7C2E9B81FF87FFE43E86FED295E3535B" attach="none" box="[704,717,335,349]" fontSize="6" pageId="11" pageNumber="63">th</superScript>
–13 
<superScript id="7C2E9B81FF87FFE43EBFFED29428535B" attach="left" box="[761,774,335,349]" fontSize="6" pageId="11" pageNumber="63">th</superScript>
precaudal vertebrae with parapophyses, and pleural ribs on 3 
<superScript id="7C2E9B81FF87FFE43D62FEEF961C5386" attach="right" box="[292,306,370,384]" fontSize="6" pageId="11" pageNumber="63">rd</superScript>
–6 
<superScript id="7C2E9B81FF87FFE43D09FEEF96725386" attach="left" box="[335,348,370,384]" fontSize="6" pageId="11" pageNumber="63">th</superScript>
precaudal vertebrae. Epipleural ribs hard to observe.
</paragraph>
<paragraph id="8BE436C9FF87FFE43CFBFE2994A25013" blockId="11.[189,1399,436,533]" pageId="11" pageNumber="63">
Head brown; body brownish-yellow with bluish-brown abdomen (
<figureCitation id="13602A4CFF87FFE43F8AFE29930553C8" box="[972,1067,436,462]" captionStart="Fig" captionStartId="8.[189,232,1962,1988]" captionTargetBox="[189,1399,915,1918]" captionTargetId="figure-179@8.[189,1399,915,1918]" captionTargetPageId="8" captionText="Fig. 3. Holotype of Luciobrotula polylepis sp. nov., 168.4 mm SL (NTUM 11915). A. Fresh specimen. B. Preserved specimen. C. Radiograph. Needle points to the lateral line end." figureDoi="http://doi.org/10.5281/zenodo.4770544" httpUri="https://zenodo.org/record/4770544/files/figure.png" pageId="11" pageNumber="63">Fig. 3A</figureCitation>
). Dorsal, pectoral, anal, and caudal fins black. Color of preserved specimen similar to that of fresh specimens, the head and body uniformly brown with dark bluish-brown abdomen (
<figureCitation id="13602A4CFF87FFE43F64FE6794505013" box="[802,894,506,533]" captionStart="Fig" captionStartId="8.[189,232,1962,1988]" captionTargetBox="[189,1399,915,1918]" captionTargetId="figure-179@8.[189,1399,915,1918]" captionTargetPageId="8" captionText="Fig. 3. Holotype of Luciobrotula polylepis sp. nov., 168.4 mm SL (NTUM 11915). A. Fresh specimen. B. Preserved specimen. C. Radiograph. Needle points to the lateral line end." figureDoi="http://doi.org/10.5281/zenodo.4770544" httpUri="https://zenodo.org/record/4770544/files/figure.png" pageId="11" pageNumber="63">Fig. 3B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3416542FF87FFE43CFBFDA1922950AE" pageId="11" pageNumber="63" type="distribution">
<paragraph id="8BE436C9FF87FFE43CFBFDA196765050" blockId="11.[189,344,572,598]" box="[189,344,572,598]" pageId="11" pageNumber="63">
<heading id="D0AC81A5FF87FFE43CFBFDA196765050" bold="true" box="[189,344,572,598]" fontSize="11" level="3" pageId="11" pageNumber="63" reason="3">
<emphasis id="B92FEADBFF87FFE43CFBFDA196765050" bold="true" box="[189,344,572,598]" pageId="11" pageNumber="63">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BE436C9FF87FFE43CFBFDF7922950AE" blockId="11.[189,1398,618,680]" pageId="11" pageNumber="63">
Possibly endemic to waters off 
<collectingCountry id="F34C7659FF87FFE43E77FDF694365082" box="[561,792,618,645]" name="Papua New Guinea" pageId="11" pageNumber="63">Papua New Guinea</collectingCountry>
; the only known specimen was collected on the SE continental slope of 
<collectingRegion id="499FF82BFF87FFE43DEAFD13951250A1" box="[428,572,653,680]" country="Papua New Guinea" name="West New Britain" pageId="11" pageNumber="63">New Britain</collectingRegion>
Island, 
<collectingCountry id="F34C7659FF87FFE43EDCFD1394AE50A1" box="[666,896,653,680]" name="Papua New Guinea" pageId="11" pageNumber="63">Papua New Guinea</collectingCountry>
, at depths of 
<quantity id="4CA39B2CFF87FFE4385AFD10938F50AE" box="[1052,1185,653,680]" metricMagnitude="2" metricUnit="m" metricValue="5.25" metricValueMax="6.2" metricValueMin="4.3" pageId="11" pageNumber="63" unit="m" value="525.0" valueMax="620.0" valueMin="430.0">430–620 m</quantity>
(
<figureCitation id="13602A4CFF87FFE438F6FD1093DB50AE" box="[1200,1269,653,680]" captionStart="Fig" captionStartId="2.[189,232,1961,1987]" captionTargetBox="[194,1392,1062,1918]" captionTargetId="figure-216@2.[189,1399,1041,1922]" captionTargetPageId="2" captionText="Fig. 1. Distribution of Luciobrotula polylepis sp. nov. (red star) and L. bartschi Smith &amp; Radcliffe 1913 (solid circles) based on specimens examined in this study." figureDoi="http://doi.org/10.5281/zenodo.4770538" httpUri="https://zenodo.org/record/4770538/files/figure.png" pageId="11" pageNumber="63">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3416542FF87FFE43CFBFD5295BB51ED" pageId="11" pageNumber="63" type="description">
<paragraph id="8BE436C9FF87FFE43CFBFD5296E450EF" blockId="11.[189,458,719,745]" box="[189,458,719,745]" pageId="11" pageNumber="63">
<heading id="D0AC81A5FF87FFE43CFBFD5296E450EF" bold="true" box="[189,458,719,745]" fontSize="11" level="3" pageId="11" pageNumber="63" reason="3">
<emphasis id="B92FEADBFF87FFE43CFBFD5296E450EF" bold="true" box="[189,458,719,745]" pageId="11" pageNumber="63">Accompanying fauna</emphasis>
</heading>
</paragraph>
<paragraph id="8BE436C9FF87FFE43CFBFD6395BB51ED" blockId="11.[189,1399,765,1003]" pageId="11" pageNumber="63">
<taxonomicName id="4C5B4D4AFF87FFE43CFBFD6396A1511E" box="[189,399,766,792]" class="Actinopterygii" family="Ophidiidae" genus="Monomitopus" kingdom="Animalia" order="Ophidiiformes" pageId="11" pageNumber="63" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B92FEADBFF87FFE43CFBFD63964E5111" box="[189,352,766,791]" italics="true" pageId="11" pageNumber="63">Monomitopus</emphasis>
sp.
</taxonomicName>
and 
<taxonomicName id="4C5B4D4AFF87FFE43D90FD609316511E" authority="Smith &amp; Radcliffe, 1913" authorityName="Smith &amp; Radcliffe" authorityYear="1913" box="[470,1080,765,792]" class="Actinopterygii" family="Ophidiidae" genus="Glyptophidium" kingdom="Animalia" order="Ophidiiformes" pageId="11" pageNumber="63" phylum="Chordata" rank="species" species="lucidum">
<emphasis id="B92FEADBFF87FFE43D90FD6095DA5111" box="[470,756,765,791]" italics="true" pageId="11" pageNumber="63">Glyptophidium lucidum</emphasis>
Smith &amp; 
<bibRefCitation id="EFCA4B38FF87FFE43F3DFD609316511E" author="Radcliffe L." box="[891,1080,765,792]" pageId="11" pageNumber="63" pagination="135 - 176" refId="ref9137" refString="Radcliffe L. 1913. Descriptions of seven new genera and thirty-one new species of fishes of the families Brotulidae and Carapidae from the Philippine Islands and the Dutch East Indies. Proceedings of the United States National Museum 44 (1948): 135 - 176. https: // doi. org / 10.5479 / si. 00963801.44 - 1948.135" type="journal article" year="1913">Radcliffe, 1913</bibRefCitation>
</taxonomicName>
were the only two other ophidiids collected along with 
<taxonomicName id="4C5B4D4AFF87FFE43E7EFCBF95E9513D" authority="Wong &amp; Lee &amp; Chen, 2021" authorityName="Wong &amp; Lee &amp; Chen" authorityYear="2021" box="[568,711,801,827]" class="Actinopterygii" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="11" pageNumber="63" phylum="Chordata" rank="species" species="polylepis" status="sp. nov.">
<emphasis id="B92FEADBFF87FFE43E7EFCBF95E9513D" box="[568,711,801,827]" italics="true" pageId="11" pageNumber="63">L. polylepis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C57A0FF87FFE43E95FCBC941B513D" box="[723,821,801,827]" pageId="11" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
, in addition to 
<taxonomicName id="4C5B4D4AFF87FFE43FB0FCBF962F5159" authority="(Gilbert, 1905)" baseAuthorityName="Gilbert" baseAuthorityYear="1905" class="Actinopterygii" family="Epigonidae" genus="Epigonus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="63" phylum="Chordata" rank="species" species="atherinoides">
<emphasis id="B92FEADBFF87FFE43FB0FCBF9228513D" box="[1014,1286,801,827]" italics="true" pageId="11" pageNumber="63">Epigonus atherinoides</emphasis>
(Gilbert, 1905)
</taxonomicName>
(
<taxonomicName id="4C5B4D4AFF87FFE43D5DFCD9956D5159" authority="Poey, 1861" authorityName="Poey" authorityYear="1861" box="[283,579,836,863]" class="Actinopterygii" family="Epigonidae" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="63" phylum="Chordata" rank="family">Epigonidae Poey, 1861</taxonomicName>
) (
<bibRefCitation id="EFCA4B38FF87FFE43E20FCD994AE5158" author="Okamoto M. &amp; Chen W. - J. &amp; Shinohara G." box="[614,896,836,862]" pageId="11" pageNumber="63" pagination="141 - 150" refId="ref9008" refString="Okamoto M., Chen W. - J. &amp; Shinohara G. 2018. Epigonus okamotoi (Perciformes: Epigonidae), a junior synonym of E. draco, with new distributional records for E. atherinoides and E. lifouensis in the West Pacific. Zootaxa 4476 (1): 141 - 150. https: // doi. org / 10.11646 / zootaxa. 4476.1.13" type="journal article" year="2018">
Okamoto 
<emphasis id="B92FEADBFF87FFE43EA1FCD894035158" box="[743,813,836,862]" italics="true" pageId="11" pageNumber="63">et al.</emphasis>
2018
</bibRefCitation>
). The mud bottom living invertebrates collected from the same site included sea cradles, sea snails, sea stars, deep-sea barnacles, decapods (https://expeditions.mnhn.fr/campaign/madeep/event/cp4334#les_photos), and a recently described deep-sea spider crab, 
<taxonomicName id="4C5B4D4AFF87FFE43D9BFC3393CA51CE" authority="Lee, Richer de Forges &amp; Ng 2019" authorityName="Lee, Richer de Forges &amp; Ng" authorityYear="2019" box="[477,1252,941,968]" class="Malacostraca" family="Epialtidae" genus="Tunepugettia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="63" phylum="Arthropoda" rank="species" species="corbariae">
<emphasis id="B92FEADBFF87FFE43D9BFC3395D151C1" box="[477,767,941,967]" italics="true" pageId="11" pageNumber="63">Tunepugettia corbariae</emphasis>
Lee, Richer de Forges &amp; Ng 2019
</taxonomicName>
(
<taxonomicName id="4C5B4D4AFF87FFE438B8FC30965D51ED" authority="MacLeay, 1838" authorityName="MacLeay" authorityYear="1838" class="Malacostraca" family="Epialtidae" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="63" phylum="Arthropoda" rank="family">Epialtidae MacLeay, 1838</taxonomicName>
) (Lee B.-Y. 
<emphasis id="B92FEADBFF87FFE43E40FC4F951451ED" box="[518,570,977,1003]" italics="true" pageId="11" pageNumber="63">et al</emphasis>
. 2019).
</paragraph>
</subSubSection>
</treatment>
</document>