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<document id="189958B7F2F3471F6AF7D0D137150843" ID-CLB-Dataset="25313" ID-DOI="10.11646/zootaxa.4679.3.7" ID-GBIF-Dataset="0a75e36a-4254-44be-9a1a-2c2bc6f115fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="3471264" ID-ZooBank="1FF46982-622E-4C27-AB8E-8F8A993A419A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1570099354004" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2019" docId="03F287B9FFDC1B19FF10FB710DFC2646" docLanguage="en" docName="zootaxa.4679.3.7.pdf" docOrigin="Zootaxa 4679 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Microscydmus (Microscydmus) nishimeanus Jałoszyński 2019, sp. n." docType="treatment" docVersion="7" lastPageNumber="544" masterDocId="FFCBFFC1FFDF1B1CFF87FFF80864257D" masterDocTitle="Unexpected diversity of Microscydmus Saulcy &amp; Croissandeau in Japan (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="552" masterPageNumber="539" pageNumber="542" updateTime="1698757590553" updateUser="ExternalLinkService">
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<mods:title id="670E50FBA2976E674C5DF91199E44C63">Unexpected diversity of Microscydmus Saulcy &amp; Croissandeau in Japan (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:namePart id="5AC58DA26638E8182790C4AA1C1385E3">Jałoszyński, Paweł</mods:namePart>
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<mods:date id="6D3EB26E3EA2D8F2A9F7FB213CF2EC29">2019</mods:date>
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<mods:number id="7F83585459A9D17B7F6C0D5F748B86C8">2019-10-03</mods:number>
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<mods:number id="EA3AEF77E24AECF09F3E6CC77D0AC969">4679</mods:number>
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<mods:classification id="D8BA4B64BF3B44DD359B2E7ACCFC96D7">journal article</mods:classification>
<mods:identifier id="CA2C4C0BD70645BD3F6C4FB395D4AC45" type="CLB-Dataset">25313</mods:identifier>
<mods:identifier id="29379C5841EDAEB83E6F29F6A86920EC" type="DOI">10.11646/zootaxa.4679.3.7</mods:identifier>
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<mods:identifier id="9D397FE6A6ADCA35AEA882DE32F91B53" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03F287B9FFDC1B19FF10FB710DFC2646" ID-DOI="http://doi.org/10.5281/zenodo.5925428" ID-GBIF-Taxon="159628632" ID-Zenodo-Dep="5925428" LSID="urn:lsid:plazi:treatment:03F287B9FFDC1B19FF10FB710DFC2646" httpUri="http://treatment.plazi.org/id/03F287B9FFDC1B19FF10FB710DFC2646" lastPageId="5" lastPageNumber="544" pageId="3" pageNumber="542">
<subSubSection id="C3416524FFDC1B1FFF10FB710A9A21DE" box="[151,766,1161,1188]" pageId="3" pageNumber="542" type="nomenclature">
<paragraph id="8BE436AFFFDC1B1FFF10FB710A9A21DE" blockId="3.[151,766,1161,1223]" box="[151,766,1161,1188]" pageId="3" pageNumber="542">
<heading id="D0AC81C3FFDC1B1FFF10FB710A9A21DE" bold="true" box="[151,766,1161,1188]" fontSize="11" level="1" pageId="3" pageNumber="542" reason="1">
<emphasis id="B92FEABDFFDC1B1FFF10FB710A9A21DE" bold="true" box="[151,766,1161,1188]" pageId="3" pageNumber="542">
<taxonomicName id="4C5B4D2CFFDC1B1FFF10FB710AD021D9" ID-CoL="8Y6FF" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[151,692,1161,1188]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="species" species="nishimeanus" status="sp. nov." subGenus="Microscydmus">
<emphasis id="B92FEABDFFDC1B1FFF10FB71092321D9" bold="true" box="[151,327,1161,1188]" italics="true" pageId="3" pageNumber="542">Microscydmus</emphasis>
(
<emphasis id="B92FEABDFFDC1B1FFEDFFB710A6C21D9" bold="true" box="[344,520,1161,1188]" italics="true" pageId="3" pageNumber="542">Microscydmus</emphasis>
)
<emphasis id="B92FEABDFFDC1B1FFD9EFB710AD021D9" bold="true" box="[537,692,1161,1188]" italics="true" pageId="3" pageNumber="542">nishimeanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C57C6FFDC1B1FFD3BFB710A9A21DE" box="[700,766,1161,1187]" pageId="3" pageNumber="542" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFDC1B1FFF10FB55090921BA" box="[151,365,1197,1223]" pageId="3" pageNumber="542" type="description">
<paragraph id="8BE436AFFFDC1B1FFF10FB55090921BA" blockId="3.[151,766,1161,1223]" box="[151,365,1197,1223]" pageId="3" pageNumber="542">
(
<figureCitation id="13602A2AFFDC1B1FFF18FB55088321BA" box="[159,231,1197,1223]" captionStart="FIGURES 1–8" captionStartId="4.[151,264,1753,1778]" captionTargetBox="[170,1420,187,1717]" captionTargetId="figure@4.[151,1436,174,1734]" captionTargetPageId="4" captionText="FIGURES 1–8. Dorsal habitus of males. Microscydmus okinawanus sp. n. (1); M. oharaensis sp. n. (2); M. nishimeanus sp. n. (3); M. kochiensis sp. n. (4); M. elytratus sp. n. (5); M. tibialis sp. n. (6); M. yusuharanus sp. n. (7); M. omogonus sp. n. (8)." figureDoi="http://doi.org/10.5281/zenodo.3471266" httpUri="https://zenodo.org/record/3471266/files/figure.png" pageId="3" pageNumber="542">Figs 3</figureCitation>
,
<figureCitation id="13602A2AFFDC1B1FFF73FB55097621BA" box="[244,274,1197,1223]" captionStart="FIGURES 9–16" captionStartId="7.[151,264,1648,1673]" captionTargetBox="[151,1436,769,1629]" captionTargetId="figure@7.[151,1436,769,1629]" captionTargetPageId="7" captionText="FIGURES 9–16. Heads of males in anterodorsal view. Microscydmus okinawanus sp. n. (9); M. oharaensis sp. n. (10); M. nishimeanus sp. n. (11); M. kochiensis sp. n. (12); M. elytratus sp. n. (13); M. tibialis sp. n. (14); M. yusuharanus sp. n. (15); M. omogonus sp. n. (16). Arrowheads show posterolateral vertexal setae." figureDoi="http://doi.org/10.5281/zenodo.3471268" httpUri="https://zenodo.org/record/3471268/files/figure.png" pageId="3" pageNumber="542">11</figureCitation>
,
<figureCitation id="13602A2AFFDC1B1FFE99FB55095821BA" box="[286,316,1197,1223]" captionStart="FIGURES 17–24" captionStartId="9.[151,264,1941,1966]" captionTargetBox="[155,1427,699,1921]" captionTargetId="figure@9.[151,1436,697,1921]" captionTargetPageId="9" captionText="FIGURES 17–24. Apices of male protibia in dorsal (17–21, 22–24) and ventral (22) views. Microscydmus okinawanus sp. n. (17); M. oharaensis sp. n. (18); M. nishimeanus sp. n. (19); M. kochiensis sp. n. (20); M. elytratus sp. n. (21); M. tibialis sp. n. (22); M. yusuharanus sp. n. (23); M. omogonus sp. n. (24)." figureDoi="http://doi.org/10.5281/zenodo.3471270" httpUri="https://zenodo.org/record/3471270/files/figure.png" pageId="3" pageNumber="542">19</figureCitation>
,
<figureCitation id="13602A2AFFDC1B1FFECFFB55090121BA" box="[328,357,1197,1223]" captionStart="FIGURES 25–32" captionStartId="11.[151,264,1484,1509]" captionTargetBox="[156,1424,237,1452]" captionTargetId="figure@11.[151,1436,229,1465]" captionTargetPageId="11" captionText="FIGURES 25–32. Aedeagus in ventral view. Microscydmus okinawanus sp. n. (25); M. oharaensis sp. n. (26); M. nishimeanus sp. n. (27); M. kochiensis sp. n. (28); M. elytratus sp. n. (29); M. tibialis sp. n. (30); M. yusuharanus sp. n. (31); M. omogonus sp. n. (32)." figureDoi="http://doi.org/10.5281/zenodo.3471272" httpUri="https://zenodo.org/record/3471272/files/figure.png" pageId="3" pageNumber="542">27</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFDC1B1FFF10FB0D089220BE" pageId="3" pageNumber="542" type="materials_examined">
<paragraph id="8BE436AFFFDC1B1FFF10FB0D089220BE" blockId="3.[151,1437,1268,2015]" pageId="3" pageNumber="542">
<emphasis id="B92FEABDFFDC1B1FFF10FB0D09232072" bold="true" box="[151,327,1268,1295]" pageId="3" pageNumber="542">Type material.</emphasis>
<materialsCitation id="3B333CF2FFDC1B1FFEC9FB0D0C002006" ID-GBIF-Occurrence="2423812453" collectingDate="2001-04-20" collectionCode="NSMT" collectorName="S. Karasawa" country="Japan" county="Mt. Nishimedake" determinerName="P. Jaloszynski" location="Ryukyus" municipality="Kunigami-son" pageId="3" pageNumber="542" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="holotype">
<emphasis id="B92FEABDFFDC1B1FFEC9FB0D0A9D2072" bold="true" box="[334,761,1268,1295]" pageId="3" pageNumber="542">
<typeStatus id="54E0880DFFDC1B1FFEC9FB0D09DB2072" box="[334,447,1269,1295]" pageId="3" pageNumber="542" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F34C763FFFDC1B1FFE4BFB0C0A4C2072" box="[460,552,1268,1295]" name="Japan" pageId="3" pageNumber="542">JAPAN</collectingCountry>
(
<collectingRegion id="499FF84DFFDC1B1FFDB1FB0C0A8F2072" box="[566,747,1268,1295]" country="Japan" name="Okinawa" pageId="3" pageNumber="542">Okinawa Pref.</collectingRegion>
):
</emphasis>
<specimenCount id="9D5DFD26FFDC1B1FFC87FB0D0B732072" box="[768,791,1269,1295]" pageId="3" pageNumber="542" type="male">♂</specimenCount>
, three labels: &quot;
<collectingCounty id="62854E23FFDC1B1FFC3CFB0D0CE52072" box="[955,1153,1269,1295]" pageId="3" pageNumber="542">Mt. Nishimedake</collectingCounty>
(No. 1) / (epiphytic fern: litter) /
<collectingMunicipality id="6B80ACD5FFDC1B1FFF6BFAE109EB204E" box="[236,399,1305,1331]" pageId="3" pageNumber="542">Kunigami-son</collectingMunicipality>
, Okinawa Is. /
<emphasis id="B92FEABDFFDC1B1FFDBEFAE10B46204E" bold="true" box="[569,802,1304,1331]" pageId="3" pageNumber="542">
[
<location id="8E846074FFDC1B1FFDC7FAE10AD4204E" LSID="urn:lsid:plazi:treatment:03F287B9FFDC1B19FF10FB710DFC2646:8E846074FFDC1B1FFDC7FAE10AD4204E" box="[576,688,1305,1331]" country="Japan" county="Mt. Nishimedake" municipality="Kunigami-son" name="Ryukyus" pageId="3" pageNumber="542" stateProvince="Okinawa">Ryukyus</location>
:
<collectingCountry id="F34C763FFFDC1B1FFD3AFAE00B7F204E" box="[701,795,1304,1331]" name="Japan" pageId="3" pageNumber="542">JAPAN</collectingCountry>
]
</emphasis>
/
<date id="FFE5106FFFDC1B1FFCBEFAE10BD7204E" box="[825,947,1305,1331]" pageId="3" pageNumber="542" value="2001-04-20">
<collectingDate id="EFA1E987FFDC1B1FFCBEFAE10BD7204E" box="[825,947,1305,1331]" pageId="3" pageNumber="542" value="2001-04-20">20.iv.2001</collectingDate>
</date>
,
<collectorName id="26AE5379FFDC1B1FFC38FAE10C2B204E" box="[959,1103,1305,1331]" pageId="3" pageNumber="542">S. Karasawa</collectorName>
&quot; [white, printed]; &quot;
<emphasis id="B92FEABDFFDC1B1FFAAFFAE0094D202A" bold="true" pageId="3" pageNumber="542">Epiphytic Fern Survey</emphasis>
/
<taxonomicName id="4C5B4D2CFFDC1B1FFEBFFAC509BF202A" authorityName="Leach" authorityYear="1815" box="[312,475,1341,1367]" class="Insecta" family="Scydmaenidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="family">Scydmaenidae</taxonomicName>
/
<taxonomicName id="4C5B4D2CFFDC1B1FFE6BFAC50AEB202A" authorityName="Saulcy &amp; Croissandeau" authorityYear="1893" box="[492,655,1341,1367]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="genus">
<emphasis id="B92FEABDFFDC1B1FFE6BFAC50AEB202A" box="[492,655,1341,1367]" italics="true" pageId="3" pageNumber="542">Microscydmus</emphasis>
</taxonomicName>
sp. 1 /
<determinerName id="7E38DA6BFFDC1B1FFD52FAC50B14202A" box="[725,880,1341,1367]" pageId="3" pageNumber="542">P. Jałoszyński</determinerName>
det.,2005&quot; [printed, creamy], &quot;
<taxonomicName id="4C5B4D2CFFDC1B1FFB41FAC50DF9202A" authorityName="Saulcy &amp; Croissandeau" authorityYear="1893" box="[1222,1437,1341,1367]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="genus">
<emphasis id="B92FEABDFFDC1B1FFB41FAC50DF9202A" box="[1222,1437,1341,1367]" italics="true" pageId="3" pageNumber="542">MICROSCYDMUS</emphasis>
</taxonomicName>
(s. str.) /
<emphasis id="B92FEABDFFDC1B1FFF7FFA9909E02006" box="[248,388,1377,1403]" italics="true" pageId="3" pageNumber="542">nishimeanus</emphasis>
m. / P.
<bibRefCitation id="EFCA4B5EFFDC1B1FFE50FA990AF92006" author="Jaloszynski, P." box="[471,669,1377,1403]" pageId="3" pageNumber="542" pagination="205 - 220" refId="ref9055" refString="Jaloszynski, P. (2019) Taxonomy of the &quot; Afroeudesis group &quot; of glandulariine ant-like stone beetles (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4612 (2), 205 - 220. https: // doi. org / 10.11646 / zootaxa. 4612.2.4" type="journal article" year="2019">Jałoszyński, 2019</bibRefCitation>
/
<typeStatus id="54E0880DFFDC1B1FFD37FA990B3E2006" box="[688,858,1377,1403]" pageId="3" pageNumber="542" type="holotype">HOLOTYPUS</typeStatus>
&quot; [red, printed] (
<collectionCode id="ED4AAE6AFFDC1B1FFB8FFA990C382006" box="[1032,1116,1377,1403]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="3" pageNumber="542">NSMT</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B333CF2FFDC1B1FFBE9FA990C0D20E2" ID-GBIF-Occurrence="2423812460" collectingDate="2001-04-20" collectionCode="NSMT" collectorName="S. Karasawa" country="Japan" county="Mt. Nishimedake" determinerName="P. Jaloszynski" location="Ryukyus" municipality="Kunigami-son" pageId="3" pageNumber="542" specimenCount="8" specimenCount-female="8" stateProvince="Okinawa" typeStatus="paratype">
<emphasis id="B92FEABDFFDC1B1FFBE9FA990D3D2006" bold="true" box="[1134,1369,1376,1403]" pageId="3" pageNumber="542">
<typeStatus id="54E0880DFFDC1B1FFBE9FA990C822006" box="[1134,1254,1377,1403]" pageId="3" pageNumber="542" type="paratype">Paratypes</typeStatus>
(11 exx.):
</emphasis>
<specimenCount id="9D5DFD26FFDC1B1FFAD9FA990DF82006" box="[1374,1436,1377,1403]" pageId="3" pageNumber="542" type="female">8 ♀♀</specimenCount>
and 3 exx. of unknown sex (undissected), same data as for holotype, except for yellow
</materialsCitation>
<materialsCitation id="3B333CF2FFDC1B1FFBF6FA7D089620BE" ID-GBIF-Occurrence="2423812451" collectionCode="NSMT" country="Japan" pageId="3" pageNumber="542" specimenCount="1" stateProvince="Okinawa" typeStatus="paratype">
<typeStatus id="54E0880DFFDC1B1FFBF6FA7D0D7020E2" box="[1137,1300,1413,1439]" pageId="3" pageNumber="542" type="paratype">PARATYPUS</typeStatus>
labels (cPJ,
<collectionCode id="ED4AAE6AFFDC1B1FFF10FA51088E20BE" box="[151,234,1449,1475]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="3" pageNumber="542">NSMT</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFDC1B1FFF40FA34089723E6" pageId="3" pageNumber="542" type="diagnosis">
<paragraph id="8BE436AFFFDC1B1FFF40FA34089723E6" blockId="3.[151,1437,1268,2015]" pageId="3" pageNumber="542">
<emphasis id="B92FEABDFFDC1B1FFF40FA340926209A" bold="true" box="[199,322,1484,1511]" pageId="3" pageNumber="542">Diagnosis.</emphasis>
Body length
<quantity id="4CA39B4AFFDC1B1FFE5EFA350A1A209A" box="[473,638,1485,1511]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" metricValueMax="9.8" metricValueMin="9.0" pageId="3" pageNumber="542" unit="mm" value="0.94" valueMax="0.98" valueMin="0.9">0.90–0.98 mm</quantity>
; posteromedian portion of vertex with punctures larger and denser than those on surrounding areas; bristles on tempora and vertex short and sparse, barely discernible; protibia strongly thickened in apical region, with apical comb of six curved teeth that occupy only apical margin, proximal teeth of the same length and shape; aedeagus with a pair of short, transverse subapical structures connected at middle, and with strongly elongated, asymmetrical lateral components flanking a long tubular structure; parameres nearly straight.
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFDC1B19FF40F95C096B27B2" lastPageId="5" lastPageNumber="544" pageId="3" pageNumber="542" type="description">
<paragraph id="8BE436AFFFDC1B1FFF40F95C0949239E" blockId="3.[151,1437,1268,2015]" pageId="3" pageNumber="542">
<emphasis id="B92FEABDFFDC1B1FFF40F95C093D23C2" bold="true" box="[199,345,1700,1727]" pageId="3" pageNumber="542">Description.</emphasis>
Body of male (
<figureCitation id="13602A2AFFDC1B1FFD80F95D0A2D23C2" box="[519,585,1701,1727]" captionStart="FIGURES 1–8" captionStartId="4.[151,264,1753,1778]" captionTargetBox="[170,1420,187,1717]" captionTargetId="figure@4.[151,1436,174,1734]" captionTargetPageId="4" captionText="FIGURES 1–8. Dorsal habitus of males. Microscydmus okinawanus sp. n. (1); M. oharaensis sp. n. (2); M. nishimeanus sp. n. (3); M. kochiensis sp. n. (4); M. elytratus sp. n. (5); M. tibialis sp. n. (6); M. yusuharanus sp. n. (7); M. omogonus sp. n. (8)." figureDoi="http://doi.org/10.5281/zenodo.3471266" httpUri="https://zenodo.org/record/3471266/files/figure.png" pageId="3" pageNumber="542">Fig. 3</figureCitation>
) slender, slightly flattened, pigmentation dark brown, vestiture light brown; BL
<quantity id="4CA39B4AFFDC1B1FFF45F930094D239E" box="[194,297,1736,1763]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="3" pageNumber="542" unit="mm" value="0.95">0.95 mm</quantity>
.
</paragraph>
<paragraph id="8BE436AFFFDC1B1FFF40F9150A1C22A2" blockId="3.[151,1437,1268,2015]" pageId="3" pageNumber="542">
Head (
<figureCitation id="13602A2AFFDC1B1FFE94F9150900227A" box="[275,356,1773,1799]" captionStart="FIGURES 9–16" captionStartId="7.[151,264,1648,1673]" captionTargetBox="[151,1436,769,1629]" captionTargetId="figure@7.[151,1436,769,1629]" captionTargetPageId="7" captionText="FIGURES 9–16. Heads of males in anterodorsal view. Microscydmus okinawanus sp. n. (9); M. oharaensis sp. n. (10); M. nishimeanus sp. n. (11); M. kochiensis sp. n. (12); M. elytratus sp. n. (13); M. tibialis sp. n. (14); M. yusuharanus sp. n. (15); M. omogonus sp. n. (16). Arrowheads show posterolateral vertexal setae." figureDoi="http://doi.org/10.5281/zenodo.3471268" httpUri="https://zenodo.org/record/3471268/files/figure.png" pageId="3" pageNumber="542">Fig. 11</figureCitation>
) broadest at eyes, HL
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, HW
<quantity id="4CA39B4AFFDC1B1FFC95F9150B1E227A" box="[786,890,1773,1799]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="542" unit="mm" value="0.23">0.23 mm</quantity>
; tempora weakly arcuate, about as long as eyes; posterior margin of vertex weakly arcuate, vertex and frons confluent and convex; supraantennal tubercles weakly elevated but well-defined; eyes large, strongly convex and coarsely faceted. Frons and vertex covered with small and shallow but dense and distinct punctures, which on posteromesal portion of vertex are slightly larger than those on surrounding areas; setae sparse, short and suberect, bristles on tempora and vertex barely discernible; vertex with a pair of long posterolateral setae. Antennae slender, AnL
<quantity id="4CA39B4AFFDC1B1FFCA7F8590BE222C6" box="[800,902,1953,1979]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="3" pageNumber="542" unit="mm" value="0.38">0.38 mm</quantity>
, antennomeres I and II strongly elongate, III–X each transverse, XI about as long as broad.
</paragraph>
<caption id="DF246627FFDB1B18FF10F9210DED226B" ID-DOI="http://doi.org/10.5281/zenodo.3471266" ID-Zenodo-Dep="3471266" httpUri="https://zenodo.org/record/3471266/files/figure.png" pageId="4" pageNumber="543" startId="4.[151,264,1753,1778]" targetBox="[170,1420,187,1717]" targetPageId="4">
<paragraph id="8BE436AFFFDB1B18FF10F9210DED226B" blockId="4.[151,1436,1753,1814]" pageId="4" pageNumber="543">
<emphasis id="B92FEABDFFDB1B18FF10F921095E238F" bold="true" box="[151,314,1753,1778]" pageId="4" pageNumber="543">FIGURES 1–8.</emphasis>
Dorsal habitus of males.
<taxonomicName id="4C5B4D2CFFDB1B18FDBCF9220B2A238F" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[571,846,1754,1778]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="543" phylum="Arthropoda" rank="species" species="okinawanus" status="sp. nov.">
<emphasis id="B92FEABDFFDB1B18FDBCF9220B2A238F" box="[571,846,1754,1778]" italics="true" pageId="4" pageNumber="543">Microscydmus okinawanus</emphasis>
</taxonomicName>
<emphasis id="B92FEABDFFDB1B18FCD3F9220BE9238F" bold="true" box="[852,909,1754,1778]" pageId="4" pageNumber="543">
<taxonomicNameLabel id="A21C57C6FFDB1B18FCD3F9220BE9238F" box="[852,909,1754,1778]" pageId="4" pageNumber="543" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(1);
<taxonomicName id="4C5B4D2CFFDB1B18FC3AF9230C2A238F" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[957,1102,1754,1778]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="543" phylum="Arthropoda" rank="species" species="oharaensis" status="sp. nov.">
<emphasis id="B92FEABDFFDB1B18FC3AF9230C2A238F" box="[957,1102,1754,1778]" italics="true" pageId="4" pageNumber="543">M. oharaensis</emphasis>
</taxonomicName>
<emphasis id="B92FEABDFFDB1B18FBD3F9220CE9238F" bold="true" box="[1108,1165,1754,1778]" pageId="4" pageNumber="543">
<taxonomicNameLabel id="A21C57C6FFDB1B18FBD3F9220CE9238F" box="[1108,1165,1754,1778]" pageId="4" pageNumber="543" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(2);
<taxonomicName id="4C5B4D2CFFDB1B18FB3AF9230D3A238F" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[1213,1374,1754,1778]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="543" phylum="Arthropoda" rank="species" species="nishimeanus" status="sp. nov.">
<emphasis id="B92FEABDFFDB1B18FB3AF9230D3A238F" box="[1213,1374,1754,1778]" italics="true" pageId="4" pageNumber="543">M. nishimeanus</emphasis>
</taxonomicName>
<emphasis id="B92FEABDFFDB1B18FAE3F9220DF8238F" bold="true" box="[1380,1436,1754,1778]" pageId="4" pageNumber="543">
<taxonomicNameLabel id="A21C57C6FFDB1B18FAE3F9220DF8238F" box="[1380,1436,1754,1778]" pageId="4" pageNumber="543" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(3);
<taxonomicName id="4C5B4D2CFFDB1B18FF45F907092A226B" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[194,334,1790,1814]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="543" phylum="Arthropoda" rank="species" species="kochiensis" status="sp. nov.">
<emphasis id="B92FEABDFFDB1B18FF45F907092A226B" box="[194,334,1790,1814]" italics="true" pageId="4" pageNumber="543">M. kochiensis</emphasis>
</taxonomicName>
<emphasis id="B92FEABDFFDB1B18FED3F90609E9226B" bold="true" box="[340,397,1790,1814]" pageId="4" pageNumber="543">
<taxonomicNameLabel id="A21C57C6FFDB1B18FED3F90609E9226B" box="[340,397,1790,1814]" pageId="4" pageNumber="543" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(4);
<taxonomicName id="4C5B4D2CFFDB1B18FE39F9070A5D226B" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[446,569,1790,1814]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="543" phylum="Arthropoda" rank="species" species="elytratus" status="sp. nov.">
<emphasis id="B92FEABDFFDB1B18FE39F9070A5D226B" box="[446,569,1790,1814]" italics="true" pageId="4" pageNumber="543">M. elytratus</emphasis>
</taxonomicName>
<emphasis id="B92FEABDFFDB1B18FDB8F9060A1C226B" bold="true" box="[575,632,1790,1814]" pageId="4" pageNumber="543">
<taxonomicNameLabel id="A21C57C6FFDB1B18FDB8F9060A1C226B" box="[575,632,1790,1814]" pageId="4" pageNumber="543" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(5);
<taxonomicName id="4C5B4D2CFFDB1B18FD2EF9070B75226B" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[681,785,1790,1814]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="543" phylum="Arthropoda" rank="species" species="tibialis" status="sp. nov.">
<emphasis id="B92FEABDFFDB1B18FD2EF9070B75226B" box="[681,785,1790,1814]" italics="true" pageId="4" pageNumber="543">M. tibialis</emphasis>
</taxonomicName>
<emphasis id="B92FEABDFFDB1B18FC9FF9060B35226B" bold="true" box="[792,849,1790,1814]" pageId="4" pageNumber="543">
<taxonomicNameLabel id="A21C57C6FFDB1B18FC9FF9060B35226B" box="[792,849,1790,1814]" pageId="4" pageNumber="543" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(6);
<taxonomicName id="4C5B4D2CFFDB1B18FC05F9070C41226B" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[898,1061,1790,1814]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="543" phylum="Arthropoda" rank="species" species="yusuharanus" status="sp. nov.">
<emphasis id="B92FEABDFFDB1B18FC05F9070C41226B" box="[898,1061,1790,1814]" italics="true" pageId="4" pageNumber="543">M. yusuharanus</emphasis>
</taxonomicName>
<emphasis id="B92FEABDFFDB1B18FBACF9060C00226B" bold="true" box="[1067,1124,1790,1814]" pageId="4" pageNumber="543">
<taxonomicNameLabel id="A21C57C6FFDB1B18FBACF9060C00226B" box="[1067,1124,1790,1814]" pageId="4" pageNumber="543" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(7);
<taxonomicName id="4C5B4D2CFFDB1B18FB11F9070D7B226B" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[1174,1311,1791,1814]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="543" phylum="Arthropoda" rank="species" species="omogonus" status="sp. nov.">
<emphasis id="B92FEABDFFDB1B18FB11F9070D7B226B" box="[1174,1311,1791,1814]" italics="true" pageId="4" pageNumber="543">M. omogonus</emphasis>
</taxonomicName>
<emphasis id="B92FEABDFFDB1B18FAA1F9060D3B226B" bold="true" box="[1318,1375,1790,1814]" pageId="4" pageNumber="543">
<taxonomicNameLabel id="A21C57C6FFDB1B18FAA1F9060D3B226B" box="[1318,1375,1790,1814]" pageId="4" pageNumber="543" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(8).
</paragraph>
</caption>
<paragraph id="8BE436AFFFDB1B18FF40F8BD0CF52292" blockId="4.[151,1437,1860,2031]" pageId="4" pageNumber="543">
Pronotum in dorsal view nearly circular, broadest at middle, PL
<quantity id="4CA39B4AFFDB1B18FC0CF8BC0B942222" box="[907,1008,1860,1887]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="543" unit="mm" value="0.25">0.25 mm</quantity>
, PW
<quantity id="4CA39B4AFFDB1B18FBADF8BD0CEB2222" box="[1066,1167,1861,1887]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="4" pageNumber="543" unit="mm" value="0.28">0.28 mm</quantity>
; anterior margin weakly arcuate; anterior corners not marked; sides strongly rounded, except for posterior 1/5, where margins are distinctly sinuate; posterior pronotal corners distinct, obtuse-angled; pronotal base with a well-defined, short transverse groove, additionally with one tiny pit near each hind corner. Punctures on disc fine and inconspicuous, smaller and shallower than those on head; setae sparse, short, suberect; lateral bristles short and sparse.
</paragraph>
<paragraph id="8BE436AFFFDA1B19FF40FF610CAB2462" blockId="5.[151,1437,152,827]" pageId="5" pageNumber="544">
Elytra together oval, broadest slightly in front of middle, EL
<quantity id="4CA39B4AFFDA1B19FC03FF600B8825CE" box="[900,1004,152,179]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="544" unit="mm" value="0.5">0.50 mm</quantity>
, EW
<quantity id="4CA39B4AFFDA1B19FBAAFF610CF125CE" box="[1069,1173,153,179]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="5" pageNumber="544" unit="mm" value="0.38">0.38 mm</quantity>
, EI 1.33; humeral calli distinct; elytral base with large and deep basal impression that extends posterad as a broad, shallow and diffuse groove; narrow adsutural region in anterior half of each elytron slightly impressed; apices of elytra separately rounded. Punctures as indistinct as those on pronotum; setae short, sparse and nearly recumbent.
</paragraph>
<paragraph id="8BE436AFFFDA1B19FF40FED10B50241A" blockId="5.[151,1437,152,827]" pageId="5" pageNumber="544">
Protibia (
<figureCitation id="13602A2AFFDA1B19FEB6FED109E0243E" box="[305,388,297,323]" captionStart="FIGURES 17–24" captionStartId="9.[151,264,1941,1966]" captionTargetBox="[155,1427,699,1921]" captionTargetId="figure@9.[151,1436,697,1921]" captionTargetPageId="9" captionText="FIGURES 17–24. Apices of male protibia in dorsal (17–21, 22–24) and ventral (22) views. Microscydmus okinawanus sp. n. (17); M. oharaensis sp. n. (18); M. nishimeanus sp. n. (19); M. kochiensis sp. n. (20); M. elytratus sp. n. (21); M. tibialis sp. n. (22); M. yusuharanus sp. n. (23); M. omogonus sp. n. (24)." figureDoi="http://doi.org/10.5281/zenodo.3471270" httpUri="https://zenodo.org/record/3471270/files/figure.png" pageId="5" pageNumber="544">Fig. 19</figureCitation>
) strongly thickened in apical region, with apical comb of six curved teeth that occupy only apical margin, proximal teeth of the same length and shape.
</paragraph>
<paragraph id="8BE436AFFFDA1B19FF40FE890BDF2485" blockId="5.[151,1437,152,827]" pageId="5" pageNumber="544">
Aedeagus (
<figureCitation id="13602A2AFFDA1B19FEC4FE8909F624F6" box="[323,402,369,395]" captionStart="FIGURES 25–32" captionStartId="11.[151,264,1484,1509]" captionTargetBox="[156,1424,237,1452]" captionTargetId="figure@11.[151,1436,229,1465]" captionTargetPageId="11" captionText="FIGURES 25–32. Aedeagus in ventral view. Microscydmus okinawanus sp. n. (25); M. oharaensis sp. n. (26); M. nishimeanus sp. n. (27); M. kochiensis sp. n. (28); M. elytratus sp. n. (29); M. tibialis sp. n. (30); M. yusuharanus sp. n. (31); M. omogonus sp. n. (32)." figureDoi="http://doi.org/10.5281/zenodo.3471272" httpUri="https://zenodo.org/record/3471272/files/figure.png" pageId="5" pageNumber="544">Fig. 27</figureCitation>
) moderately slender, AeL
<quantity id="4CA39B4AFFDA1B19FD32FE890B7D24F6" box="[693,793,369,395]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="544" unit="mm" value="0.13">0.13 mm</quantity>
, endophallus with long median tubular component flanked by a pair of shorter elongate and clearly asymmetrical sclerites in submedian region of median lobe, in subapical region the tubular component connected to a pair of short, oval transverse sclerites; parameres nearly straight, each presumably with one apical seta (seta broken off, but its socket visible).
</paragraph>
<paragraph id="8BE436AFFFDA1B19FF40FDF9096B27B2" blockId="5.[151,1437,152,827]" pageId="5" pageNumber="544">
Female. Externally similar to male, also with thickened protibiae, but lacking apical combs, most specimens lighter brown than male, punctures on vertex and frons variable, from similar to those in male to more evenly distributed and less conspicuous; BL
<quantity id="4CA39B4AFFDA1B19FDC2FDB10A8F271E" box="[581,747,585,611]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" metricValueMax="9.8" metricValueMin="9.0" pageId="5" pageNumber="544" unit="mm" value="0.94" valueMax="0.98" valueMin="0.9">0.90–0.98 mm</quantity>
(mean
<quantity id="4CA39B4AFFDA1B19FCC4FDB10BC9271E" box="[835,941,585,611]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" pageId="5" pageNumber="544" unit="mm" value="0.94">0.94 mm</quantity>
); HL
<quantity id="4CA39B4AFFDA1B19FC76FDB10CF2271E" box="[1009,1174,585,611]" metricMagnitude="-4" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="5" pageNumber="544" unit="mm" value="0.195" valueMax="0.2" valueMin="0.19">0.19–0.20 mm</quantity>
(mean
<quantity id="4CA39B4AFFDA1B19FB69FDB10D3C271E" box="[1262,1368,585,611]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="544" unit="mm" value="0.2">0.20 mm</quantity>
), HW
<quantity id="4CA39B4AFFDA1B19FF10FD95095827FA" box="[151,316,621,647]" metricMagnitude="-4" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="5" pageNumber="544" unit="mm" value="0.215" valueMax="0.23" valueMin="0.2">0.20–0.23 mm</quantity>
(mean
<quantity id="4CA39B4AFFDA1B19FE13FD95099A27FA" box="[404,510,621,647]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="5" pageNumber="544" unit="mm" value="0.22">0.22 mm</quantity>
), AnL
<quantity id="4CA39B4AFFDA1B19FDD7FD940A9127FA" box="[592,757,620,647]" metricMagnitude="-4" metricUnit="m" metricValue="3.3999999999999995" metricValueMax="3.5" metricValueMin="3.3" pageId="5" pageNumber="544" unit="mm" value="0.33999999999999997" valueMax="0.35" valueMin="0.33">0.33–0.35 mm</quantity>
(mean
<quantity id="4CA39B4AFFDA1B19FCCBFD950BD327FA" box="[844,951,621,647]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="5" pageNumber="544" unit="mm" value="0.34">0.34 mm</quantity>
); PL
<quantity id="4CA39B4AFFDA1B19FC71FD940CFF27FA" box="[1014,1179,620,647]" metricMagnitude="-4" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="5" pageNumber="544" unit="mm" value="0.245" valueMax="0.25" valueMin="0.24">0.24–0.25 mm</quantity>
(mean
<quantity id="4CA39B4AFFDA1B19FB74FD940D3927FA" box="[1267,1373,620,647]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="544" unit="mm" value="0.25">0.25 mm</quantity>
), PW
<quantity id="4CA39B4AFFDA1B19FF10FD68095A27D6" box="[151,318,656,683]" metricMagnitude="-4" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="5" pageNumber="544" unit="mm" value="0.265" valueMax="0.28" valueMin="0.25">0.25–0.28 mm</quantity>
(mean
<quantity id="4CA39B4AFFDA1B19FE1FFD690A6727D6" box="[408,515,657,683]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="5" pageNumber="544" unit="mm" value="0.27">0.27 mm</quantity>
); EL
<quantity id="4CA39B4AFFDA1B19FDC0FD680A8927D6" box="[583,749,656,683]" metricMagnitude="-4" metricUnit="m" metricValue="5.05" metricValueMax="5.3" metricValueMin="4.8" pageId="5" pageNumber="544" unit="mm" value="0.505" valueMax="0.53" valueMin="0.48">0.48–0.53 mm</quantity>
(mean
<quantity id="4CA39B4AFFDA1B19FCCFFD680BD727D6" box="[840,947,656,683]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="544" unit="mm" value="0.5">0.50 mm</quantity>
), EW
<quantity id="4CA39B4AFFDA1B19FC78FD680CC227D6" box="[1023,1190,656,683]" metricMagnitude="-4" metricUnit="m" metricValue="3.3999999999999995" metricValueMax="3.5" metricValueMin="3.3" pageId="5" pageNumber="544" unit="mm" value="0.33999999999999997" valueMax="0.35" valueMin="0.33">0.33–0.35 mm</quantity>
(mean
<quantity id="4CA39B4AFFDA1B19FA87FD690D0827D6" box="[1280,1388,657,683]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="5" pageNumber="544" unit="mm" value="0.34">0.34 mm</quantity>
), EI 1.41–1.50.
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFDA1B19FF40FD200A39278E" box="[199,605,728,755]" pageId="5" pageNumber="544" type="distribution">
<paragraph id="8BE436AFFFDA1B19FF40FD200A39278E" blockId="5.[151,1437,152,827]" box="[199,605,728,755]" pageId="5" pageNumber="544">
<emphasis id="B92FEABDFFDA1B19FF40FD200905278E" bold="true" box="[199,353,728,755]" pageId="5" pageNumber="544">Distribution.</emphasis>
<collectingCountry id="F34C763FFFDA1B19FEE0FD2109CE278E" box="[359,426,729,755]" name="Japan" pageId="5" pageNumber="544">Japan</collectingCountry>
, Okinawa-jima.
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFDA1B19FF40FD050BE32665" box="[199,903,765,792]" pageId="5" pageNumber="544" type="etymology">
<paragraph id="8BE436AFFFDA1B19FF40FD050BE32665" blockId="5.[151,1437,152,827]" box="[199,903,765,792]" pageId="5" pageNumber="544">
<emphasis id="B92FEABDFFDA1B19FF40FD05092B266A" bold="true" box="[199,335,765,791]" pageId="5" pageNumber="544">Etymology.</emphasis>
Locotypical, after Mt. Nishime (= Nishime-dake).
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFDA1B19FF40FCD90DFC2646" box="[199,1432,801,827]" pageId="5" pageNumber="544" type="discussion">
<paragraph id="8BE436AFFFDA1B19FF40FCD90DFC2646" blockId="5.[151,1437,152,827]" box="[199,1432,801,827]" pageId="5" pageNumber="544">
<emphasis id="B92FEABDFFDA1B19FF40FCD9095F2646" bold="true" box="[199,315,801,827]" pageId="5" pageNumber="544">Remarks.</emphasis>
This species is most similar to
<taxonomicName id="4C5B4D2CFFDA1B19FD1CFCDA0B232646" authorityName="Jałoszyński" authorityYear="2019" box="[667,839,801,827]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="544" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="B92FEABDFFDA1B19FD1CFCDA0B232646" box="[667,839,801,827]" italics="true" pageId="5" pageNumber="544">M. okinawanus</emphasis>
</taxonomicName>
; differences were discussed in remarks for the latter.
</paragraph>
</subSubSection>
</treatment>
</document>
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