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<document id="8E203A183377954CA38D6A18DFF2436B" ID-CLB-Dataset="31902" ID-DOI="10.11646/zootaxa.4329.2.1" ID-GBIF-Dataset="7e33ec7d-211b-4042-8d07-344364ee805d" ID-ISSN="1175-5326" ID-Zenodo-Dep="1001915" ID-ZooBank="8226E27D-E3A9-481A-B51B-558643Bbb03A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507106636447" checkinUser="plazi" docAuthor="Tarasov, Sergei" docDate="2017" docId="03C187E1AE4DB622DD9E42BCFD4C004C" docLanguage="en" docName="zootaxa.4329.2.1.pdf" docOrigin="Zootaxa 4329 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parachorius bolavensis S. Tarasov 2017, new species" docType="treatment" docUuid="A116D60D-8D17-434E-8AAD-84A0C331C23E" docUuidSource="ZooBank" docVersion="12" lastPageNumber="119" masterDocId="FFF8FF99AE5DB630DD09474FFFDB0270" masterDocTitle="A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data" masterLastPageNumber="149" masterPageNumber="101" pageNumber="117" updateTime="1698469310636" updateUser="ExternalLinkService">
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<mods:titleInfo id="BD6F89E55FF45D161F1E83397CBB4C23">
<mods:title id="8FDE226056F075FBC1200CF25F6639E4">A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data</mods:title>
</mods:titleInfo>
<mods:name id="F727E998100D5E026FD9B6A940A1A598" type="personal">
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<mods:roleTerm id="89432B6E7C51C955D502A07805ADA5FF">Author</mods:roleTerm>
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<mods:namePart id="E509664C6E66B9808878A14819683D5E">Tarasov, Sergei</mods:namePart>
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<mods:title id="198998131120B97EF58498F233C1CC38">Zootaxa</mods:title>
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<mods:part id="E37F59073905BA4BE2F738A3B8D7CBF9">
<mods:date id="76C3ACEB23F45A7AA88649E8DB05B226">2017</mods:date>
<mods:detail id="C22131414CB0DB6FE0AB3726C64115C3" type="pubDate">
<mods:number id="7FEB5D7BFCCBF41CFB4E9E23BE6366C5">2017-10-04</mods:number>
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<mods:number id="B5E7B34FC7763F9C31B75658C952ACD0">4329</mods:number>
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<mods:identifier id="C88B819E1CF4D1D31C2C73A18AD75C66" type="DOI">10.11646/zootaxa.4329.2.1</mods:identifier>
<mods:identifier id="289D497F787681A29CF52F0D04713435" type="GBIF-Dataset">7e33ec7d-211b-4042-8d07-344364ee805d</mods:identifier>
<mods:identifier id="52301D9334E4694421F0E1C418B8A349" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03C187E1AE4DB622DD9E42BCFD4C004C" ID-DOI="http://doi.org/10.5281/zenodo.6010810" ID-GBIF-Taxon="135106131" ID-Zenodo-Dep="6010810" LSID="urn:lsid:zoobank.org:act:A116D60D-8D17-434E-8AAD-84A0C331C23E" httpUri="http://treatment.plazi.org/id/03C187E1AE4DB622DD9E42BCFD4C004C" lastPageId="18" lastPageNumber="119" pageId="16" pageNumber="117">
<subSubSection id="C372657CAE4DB620DD9E42BCFEE9045F" pageId="16" pageNumber="117" type="nomenclature">
<paragraph id="8BD736F7AE4DB620DD9E42BCFD68047E" blockId="16.[151,981,1523,1619]" box="[151,691,1523,1550]" pageId="16" pageNumber="117">
<heading id="D09F819BAE4DB620DD9E42BCFD68047E" bold="true" box="[151,691,1523,1550]" fontSize="11" level="1" pageId="16" pageNumber="117" reason="1">
<emphasis id="B91CEAE5AE4DB620DD9E42BCFD68047E" bold="true" box="[151,691,1523,1550]" pageId="16" pageNumber="117">
<taxonomicName id="4C684D74AE4DB620DD9E42BCFDC6047E" ID-CoL="6TNM4" authority="Tarasov" authorityName="S. Tarasov" authorityYear="2017" box="[151,541,1523,1550]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="bolavensis" status="sp. nov.">
<emphasis id="B91CEAE5AE4DB620DD9E42BCFE6A047D" bold="true" box="[151,433,1523,1549]" italics="true" pageId="16" pageNumber="117">Parachorius bolavensis</emphasis>
Tarasov
</taxonomicName>
<taxonomicNameLabel id="A22F579EAE4DB620DF2D42BBFD68047E" box="[548,691,1524,1550]" pageId="16" pageNumber="117" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD736F7AE4DB620DD9E4159FEE9045F" blockId="16.[151,981,1523,1619]" box="[151,306,1558,1583]" pageId="16" pageNumber="117">
(
<figureCitation id="13532A72AE4DB620DD964159FF35045E" box="[159,238,1558,1583]" captionStart="FIGURE 7" captionStartId="17.[151,250,1493,1515]" captionTargetBox="[153,1407,193,1475]" captionTargetId="figure@17.[151,1436,193,1479]" captionTargetPageId="17" captionText="FIGURE 7. Morphology of Parachorius bolavensis. (A) aedeagus, lateral view (holotype). (B) aedeagus, lateral view (paratype). (C) aedeagus. (D) habitus (male, holotype). (E) protibia (male, paratype). (F) protibia (female, paratype). (G) left metafemur (male, paratype)." httpUri="https://zenodo.org/record/1001929/files/figure.png" pageId="16" pageNumber="117">Figs. 7</figureCitation>
,
<figureCitation id="13532A72AE4DB620DDF34159FEF1045F" box="[250,298,1558,1583]" captionStart="FIGURE 25" captionStartId="42.[151,250,1911,1933]" captionTargetBox="[151,1433,273,1824]" captionTargetId="figure@42.[151,1436,273,1824]" captionTargetPageId="42" captionText="FIGURE 25. Distribution maps of Parachorius species. (A) Parachorius asymmetricus. (B) Parachorius bolavensis. (C) Parachorius fukiensis. (D) Parachorius fungorum. (E) Parachorius globosus. (F) Parachorius gotoi. Type locality — type locality of species (holotype or lectotype); Type. Collection data — localities of type specimens associated with species (paratypes or paralectotypes); Collection data — localities of other specimens examined in this study." httpUri="https://zenodo.org/record/1001967/files/figure.png" pageId="16" pageNumber="117">25B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE4DB620DD9E4175FC0E0423" box="[151,981,1594,1619]" pageId="16" pageNumber="117" type="description">
<paragraph id="8BD736F7AE4DB620DD9E4175FC0E0423" blockId="16.[151,981,1523,1619]" box="[151,981,1594,1619]" pageId="16" pageNumber="117">urn:lsid:zoobank.org:act:A116D60D-8D17-434E-8AAD-84A0C331C23E</paragraph>
</subSubSection>
<subSubSection id="C372657CAE4DB620DD9E41CDFDED04EA" box="[151,566,1666,1691]" pageId="16" pageNumber="117" type="materials_examined">
<paragraph id="8BD736F7AE4DB620DD9E41CDFDED04EA" blockId="16.[151,1437,1666,2015]" box="[151,566,1666,1691]" pageId="16" pageNumber="117">
<emphasis id="B91CEAE5AE4DB620DD9E41CDFEF604EB" bold="true" box="[151,301,1666,1691]" pageId="16" pageNumber="117">Type locality</emphasis>
. Bolaven Plateau, Laos
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE4DB620DDCE41E9FA8704CE" box="[199,1372,1702,1727]" pageId="16" pageNumber="117" type="etymology">
<paragraph id="8BD736F7AE4DB620DDCE41E9FA8704CE" blockId="16.[151,1437,1666,2015]" box="[199,1372,1702,1727]" pageId="16" pageNumber="117">
<emphasis id="B91CEAE5AE4DB620DDCE41E9FE9504CF" bold="true" box="[199,334,1702,1727]" pageId="16" pageNumber="117">Etymology.</emphasis>
The name of this species is derived from its
<typeStatus id="54D38855AE4DB620DE4D41E8FCAF04CF" box="[836,884,1703,1727]" pageId="16" pageNumber="117">type</typeStatus>
locality in Bolaven plateau, southern
<collectingCountry id="F37F7667AE4DB620D81441E8FA8C04CE" box="[1309,1367,1703,1726]" name="Laos" pageId="16" pageNumber="117">Laos</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE4DB620DDCE4185FB42055B" pageId="16" pageNumber="117" type="distribution">
<paragraph id="8BD736F7AE4DB620DDCE4185FB42055B" blockId="16.[151,1437,1666,2015]" pageId="16" pageNumber="117">
<emphasis id="B91CEAE5AE4DB620DDCE4185FEBB0493" bold="true" box="[199,352,1738,1763]" pageId="16" pageNumber="117">Distribution.</emphasis>
The species is known only from the two localities on Bolaven plateau in southern
<collectingCountry id="F37F7667AE4DB620D8114184FA950492" box="[1304,1358,1739,1762]" name="Laos" pageId="16" pageNumber="117">Laos</collectingCountry>
where the
<typeStatus id="54D38855AE4DB620DDCD41A0FF2F0577" box="[196,244,1775,1799]" pageId="16" pageNumber="117">type</typeStatus>
series was collected. Presumably, this species is endemic to Bolaven plateau that has unique climatic conditions due to its elevation (average height is approximately
<quantity id="4C909B12AE4DB620DE6C405CFC60055B" box="[869,955,1811,1835]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="117" unit="m" value="1000.0">1000 m</quantity>
) over the lowlands.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE4DB620DDCE4079FC9F05E7" pageId="16" pageNumber="117" type="diagnosis">
<paragraph id="8BD736F7AE4DB620DDCE4079FC9F05E7" blockId="16.[151,1437,1666,2015]" pageId="16" pageNumber="117">
<emphasis id="B91CEAE5AE4DB620DDCE4079FE9A053F" bold="true" box="[199,321,1846,1871]" pageId="16" pageNumber="117">Diagnosis.</emphasis>
This species is most closely related to
<taxonomicName id="4C684D74AE4DB620DE014078FC5E053F" authorityName="Balthasar S. Tarasov" authorityYear="2017" baseAuthorityName="Balthasar" baseAuthorityYear="1960" box="[776,901,1846,1871]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="fukiensis">
<emphasis id="B91CEAE5AE4DB620DE014078FC5E053F" box="[776,901,1846,1871]" italics="true" pageId="16" pageNumber="117">P. fukiensis</emphasis>
</taxonomicName>
. It differs from
<taxonomicName id="4C684D74AE4DB620D94D4078FB1F053F" authorityName="Balthasar S. Tarasov" authorityYear="2017" baseAuthorityName="Balthasar" baseAuthorityYear="1960" box="[1092,1220,1846,1871]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="fukiensis">
<emphasis id="B91CEAE5AE4DB620D94D4078FB1F053F" box="[1092,1220,1846,1871]" italics="true" pageId="16" pageNumber="117">P. fukiensis</emphasis>
</taxonomicName>
and the remaining
<taxonomicName id="4C684D74AE4DB620DD9E4015FEF80503" box="[151,291,1882,1907]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAE5AE4DB620DD9E4015FEF80503" box="[151,291,1882,1907]" italics="true" pageId="16" pageNumber="117">Parachorius</emphasis>
</taxonomicName>
species by the shape of aedeagus (parameres widened basally, their apices acute) and by the widely rimmed middle dilation of the posterior metafemoral margin.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE4DB621DDCE40EDFBEE0583" lastPageId="17" lastPageNumber="118" pageId="16" pageNumber="117" type="description">
<paragraph id="8BD736F7AE4DB620DDCE40EDFB0A05CB" blockId="16.[151,1437,1666,2015]" box="[199,1233,1954,1979]" pageId="16" pageNumber="117">
<emphasis id="B91CEAE5AE4DB620DDCE40EDFE8205CB" bold="true" box="[199,345,1954,1979]" pageId="16" pageNumber="117">Description.</emphasis>
<emphasis id="B91CEAE5AE4DB620DC5640EDFE4405CB" box="[351,415,1954,1979]" italics="true" pageId="16" pageNumber="117">Body.</emphasis>
Body black, length
<quantity id="4C909B12AE4DB620DF8840EDFCD005CA" box="[641,779,1954,1979]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.5" metricValueMin="4.2" pageId="16" pageNumber="117" unit="mm" value="4.35" valueMax="4.5" valueMin="4.2">4.2–4.5 mm</quantity>
. Elytra uniformly colored, mostly black.
</paragraph>
<paragraph id="8BD736F7AE4DB620DDCE4089FBAD05AF" blockId="16.[151,1437,1666,2015]" box="[199,1142,1990,2015]" pageId="16" pageNumber="117">
<emphasis id="B91CEAE5AE4DB620DDCE4089FED005AF" box="[199,267,1990,2015]" italics="true" pageId="16" pageNumber="117">Head.</emphasis>
Clypeus with two teeth, outer margin of each tooth distinctly notched basally.
</paragraph>
<caption id="DF17667FAE4CB621DD9E429AFE630458" httpUri="https://zenodo.org/record/1001929/files/figure.png" pageId="17" pageNumber="118" targetBox="[153,1407,193,1475]" targetPageId="17">
<paragraph id="8BD736F7AE4CB621DD9E429AFE630458" blockId="17.[151,1436,1492,1576]" pageId="17" pageNumber="118">
<emphasis id="B91CEAE5AE4CB621DD9E429AFEC2079A" bold="true" box="[151,281,1493,1515]" pageId="17" pageNumber="118">FIGURE 7.</emphasis>
Morphology of
<taxonomicName id="4C684D74AE4CB621DCDB429BFD13079A" authorityName="S. Tarasov" authorityYear="2017" box="[466,712,1492,1514]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="bolavensis">
<emphasis id="B91CEAE5AE4CB621DCDB429BFD13079A" box="[466,712,1492,1514]" italics="true" pageId="17" pageNumber="118">Parachorius bolavensis</emphasis>
</taxonomicName>
. (A) aedeagus, lateral view (holotype). (B) aedeagus, lateral view (paratype). (C) aedeagus. (D) habitus (male, holotype). (E) protibia (male, paratype). (F) protibia (female, paratype). (G) left metafemur (male, paratype).
</paragraph>
</caption>
<paragraph id="8BD736F7AE4CB621DDCE411FFB5804FC" blockId="17.[151,1436,1615,2036]" pageId="17" pageNumber="118">
<emphasis id="B91CEAE5AE4CB621DDCE411FFEE60417" box="[199,317,1616,1639]" italics="true" pageId="17" pageNumber="118">Pronotum.</emphasis>
Pronotum rounded laterally. Frontal angles rounded. Disc covered with punctures separated by 1–2 puncture diameter(s). Pronotum in anterior-posterior direction notably shorter than elytra.
</paragraph>
<paragraph id="8BD736F7AE4CB621DDCE41D8FD1B04DF" blockId="17.[151,1436,1615,2036]" box="[199,704,1687,1712]" pageId="17" pageNumber="118">
<emphasis id="B91CEAE5AE4CB621DDCE41D8FECE04C0" box="[199,277,1687,1712]" italics="true" pageId="17" pageNumber="118">Elytra.</emphasis>
Elytra covered with sparse punctures.
</paragraph>
<paragraph id="8BD736F7AE4CB621DDCE41F2FD9404A4" blockId="17.[151,1436,1615,2036]" box="[199,591,1723,1748]" pageId="17" pageNumber="118">
<emphasis id="B91CEAE5AE4CB621DDCE41F2FEC904A4" box="[199,274,1725,1748]" italics="true" pageId="17" pageNumber="118">Wings.</emphasis>
Wings normally developed.
</paragraph>
<paragraph id="8BD736F7AE4CB621DDCE41AFFD6005DC" blockId="17.[151,1436,1615,2036]" pageId="17" pageNumber="118">
<emphasis id="B91CEAE5AE4CB621DDCE41AFFED80487" box="[199,259,1760,1783]" italics="true" pageId="17" pageNumber="118">Legs.</emphasis>
Profemur with anterior margin medially produced in angle or with almost straight or sinuate anterior margin. Protibia with apical spur approximately reaching middle of protarsomere 3. First tooth normal, notably modified in males, approximately twice as wide as second tooth; first and second teeth distinctly separated from each other, first tooth distinctly separated from denticles located between first and second teeth. Metafemoral posterior margin not serrate, dilated medially, dilation widely rimmed. Metatibia slightly sinuate, inner margin not denticulate, produced into a small angle apically.
</paragraph>
<paragraph id="8BD736F7AE4CB621DDCE40F7FBBD05A0" blockId="17.[151,1436,1615,2036]" box="[199,1126,1975,2000]" pageId="17" pageNumber="118">
<emphasis id="B91CEAE5AE4CB621DDCE40F7FEB905BF" box="[199,354,1976,1999]" italics="true" pageId="17" pageNumber="118">Metasternum.</emphasis>
Transverse basal ridge of metasternum reduced and extremely short.
</paragraph>
<paragraph id="8BD736F7AE4CB621DDCE4094FBEE0583" blockId="17.[151,1436,1615,2036]" box="[199,1077,2011,2036]" pageId="17" pageNumber="118">
<emphasis id="B91CEAE5AE4CB621DDCE4094FEE00584" box="[199,315,2011,2036]" italics="true" pageId="17" pageNumber="118">Aedeagus.</emphasis>
Parameres symmetrical, widened basally (lateral view), apex acute.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE4FB622DDCE47D8FD4C004C" pageId="18" pageNumber="119" type="materials_examined">
<paragraph id="8BD736F7AE4FB622DDCE47D8FB5402C0" blockId="18.[151,1436,151,573]" box="[199,1167,151,176]" pageId="18" pageNumber="119">
<emphasis id="B91CEAE5AE4FB622DDCE47D8FEE002C0" bold="true" box="[199,315,151,176]" pageId="18" pageNumber="119">
<typeStatus id="54D38855AE4FB622DDCE47D8FEED02C0" box="[199,310,151,176]" pageId="18" pageNumber="119" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male (
<figureCitation id="13532A72AE4FB622DC8247D8FE3E02C0" box="[395,485,151,176]" captionStart="FIGURE 7" captionStartId="17.[151,250,1493,1515]" captionTargetBox="[153,1407,193,1475]" captionTargetId="figure@17.[151,1436,193,1479]" captionTargetPageId="17" captionText="FIGURE 7. Morphology of Parachorius bolavensis. (A) aedeagus, lateral view (holotype). (B) aedeagus, lateral view (paratype). (C) aedeagus. (D) habitus (male, holotype). (E) protibia (male, paratype). (F) protibia (female, paratype). (G) left metafemur (male, paratype)." httpUri="https://zenodo.org/record/1001929/files/figure.png" pageId="18" pageNumber="119">Fig. 7D</figureCitation>
). Profemur with anterior margin medially produced in angle.
</paragraph>
<paragraph id="8BD736F7AE4FB622DDCE47F3FD4C004C" blockId="18.[151,1436,151,573]" pageId="18" pageNumber="119">
<materialsCitation id="3B003CAAAE4FB622DDCE47F3FB830288" ID-GBIF-Occurrence="1658411493" collectorName="S. Tarasov" country="Laos" county="Material" elevation="1200" location="Ban Houayteuay" longitude="15.077583" municipality="Bolaven" pageId="18" pageNumber="119" specimenCount="1" typeStatus="holotype">
<emphasis id="B91CEAE5AE4FB622DDCE47F3FDF002A5" bold="true" box="[199,555,188,213]" pageId="18" pageNumber="119">
<collectingCounty id="62B64E7BAE4FB622DDCE47F3FEF402A5" box="[199,303,188,213]" pageId="18" pageNumber="119">Material</collectingCounty>
examined.
<typeStatus id="54D38855AE4FB622DCB647F3FDF002A5" box="[447,555,188,213]" pageId="18" pageNumber="119" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="F37F7667AE4FB622DF3747F3FDAC02A5" box="[574,631,188,213]" name="Laos" pageId="18" pageNumber="119">
<emphasis id="B91CEAE5AE4FB622DF3747F3FDAC02A5" bold="true" box="[574,631,188,213]" pageId="18" pageNumber="119">Laos</emphasis>
</collectingCountry>
:
<emphasis id="B91CEAE5AE4FB622DF8347F3FD1F02A5" bold="true" box="[650,708,188,213]" pageId="18" pageNumber="119">male</emphasis>
, &quot;S[outhern]
<collectingCountry id="F37F7667AE4FB622DE6A47F2FC4702A4" box="[867,924,189,212]" name="Laos" pageId="18" pageNumber="119">Laos</collectingCountry>
,
<collectingMunicipality id="6BB3AC8DAE4FB622DEA547F3FBD202A5" box="[940,1033,188,213]" pageId="18" pageNumber="119">Bolaven</collectingMunicipality>
plateau,
<location id="8EB7602CAE4FB622D97F47F2FAEE02A5" LSID="urn:lsid:plazi:treatment:03C187E1AE4DB622DD9E42BCFD4C004C:8EB7602CAE4FB622D97F47F2FAEE02A5" box="[1142,1333,189,213]" country="Laos" county="Material" longitude="15.077583" municipality="Bolaven" name="Ban Houayteuay" pageId="18" pageNumber="119">Ban Houayteuay</location>
h
<elevation id="0045D1C4AE4FB622D84747F3FA4702A5" box="[1358,1436,188,213]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="18" pageNumber="119" unit="m" value="1200.0">
<quantity id="4C909B12AE4FB622D84747F3FA4702A5" box="[1358,1436,188,213]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="18" pageNumber="119" unit="m" value="1200.0">1200m</quantity>
</elevation>
N
<geoCoordinate id="EE5C5030AE4FB622DDA54790FEED0288" box="[172,310,223,248]" direction="east" orientation="longitude" pageId="18" pageNumber="119" precision="1" value="15.077583">15°4.655' E</geoCoordinate>
106°16.848'
<collectingMethod id="52294EE0AE4FB622DCC34790FCBC0287" box="[458,871,223,248]" pageId="18" pageNumber="119">disturb montane forest pitfall traps</collectingMethod>
leg.
<collectorName id="269D5321AE4FB622DEA547AFFBF30287" box="[940,1064,224,248]" pageId="18" pageNumber="119">S. Tarasov</collectorName>
6.5
</materialsCitation>
-
<date id="FFD61037AE4FB622D9694790FB080288" box="[1120,1235,223,248]" pageId="18" pageNumber="119" value="2008-06-14">14.6.2008</date>
/ HOLOTYPUS
<taxonomicName id="4C684D74AE4FB622DD9E464BFDFD036C" authority="S. Tarasov" authorityName="S. Tarasov" authorityYear="2017" box="[151,550,260,285]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" species="bolavensis">Parachorius bolavensis S. Tarasov</taxonomicName>
det. 2010&quot;, 15°4.655'N, 106°16.848'E (ZMUC).
<emphasis id="B91CEAE5AE4FB622D978464BFB32036D" bold="true" box="[1137,1257,260,285]" pageId="18" pageNumber="119">Paratypes</emphasis>
(
<emphasis id="B91CEAE5AE4FB622D9F6464BFAB7036D" bold="true" box="[1279,1388,260,285]" pageId="18" pageNumber="119">10 males</emphasis>
,
<emphasis id="B91CEAE5AE4FB622D889464BFF210330" bold="true" pageId="18" pageNumber="119">10 females)</emphasis>
:
<materialsCitation id="3B003CAAAE4FB622DC184668FBD90314" ID-GBIF-Occurrence="1658411516" collectionCode="FMHD" country="Laos" location="Xekatam" pageId="18" pageNumber="119" specimenCode="FMHD#2008-037" specimenCount="14" specimenCount-female="6" specimenCount-male="8" stateProvince="Champasak Province" typeStatus="paratype">
<emphasis id="B91CEAE5AE4FB622DC184668FD2D0330" bold="true" box="[273,758,295,320]" pageId="18" pageNumber="119">
<specimenCount id="9D6EFD7EAE4FB622DC184668FEB30330" box="[273,360,295,320]" pageId="18" pageNumber="119" type="male">8 male</specimenCount>
<typeStatus id="54D38855AE4FB622DC7E4668FE340330" box="[375,495,295,320]" pageId="18" pageNumber="119" type="paratype">paratypes</typeStatus>
,
<specimenCount id="9D6EFD7EAE4FB622DF0D4668FDAB0330" box="[516,624,295,320]" pageId="18" pageNumber="119" type="female">6 female</specimenCount>
<typeStatus id="54D38855AE4FB622DF894668FD2D0330" box="[640,758,295,320]" pageId="18" pageNumber="119" type="paratype">paratypes</typeStatus>
</emphasis>
,
<collectingRegion id="49ACF815AE4FB622DE054668FBDD0330" box="[780,1030,295,320]" country="Laos" name="Champasak" pageId="18" pageNumber="119">Champasak Province</collectingRegion>
: Bolaven Plateau, Ban Thôngvay (=
<location id="8EB7602CAE4FB622DDA44603FECC0315" LSID="urn:lsid:plazi:treatment:03C187E1AE4DB622DD9E42BCFD4C004C:8EB7602CAE4FB622DDA44603FECC0315" box="[173,279,332,357]" country="Laos" name="Xekatam" pageId="18" pageNumber="119" stateProvince="Champasak Province">Xékatam</location>
) vicinity, old logging road N of village,
<specimenCode id="DBCE9E8CAE4FB622DE3C4603FBD90314" box="[821,1026,332,356]" collectionCode="FMHD" pageId="18" pageNumber="119">FMHD#2008-037</specimenCode>
</materialsCitation>
, ANMT site 1231, 15°14.288'N, 106°31.891'E,
<quantity id="4C909B12AE4FB622DC4B4620FE4503F7" box="[322,414,367,392]" metricMagnitude="3" metricUnit="m" metricValue="1.095" pageId="18" pageNumber="119" unit="m" value="1095.0">1095 m</quantity>
, selectively logged forest, on
<date id="FFD61037AE4FB622DE0D4620FC4103F8" box="[772,922,367,392]" pageId="18" pageNumber="119" value="2008-06-08" valueMax="2008-06-16" valueMin="2008-06-08">8–16/vi/2008</date>
(A. Newton, M. Thayer) (FMNH);
<emphasis id="B91CEAE5AE4FB622D8414620FED903DD" bold="true" pageId="18" pageNumber="119">1 male paratype</emphasis>
,
<materialsCitation id="3B003CAAAE4FB622DC1B46DBFD4303A0" ID-GBIF-Occurrence="1658411507" collectionCode="FMHD" country="Laos" location="Xekatam" pageId="18" pageNumber="119" specimenCode="FMHD#2008-040" specimenCount="3" specimenCount-female="3" stateProvince="Champasak Province" typeStatus="paratype">
<emphasis id="B91CEAE5AE4FB622DC1B46DBFE2303DD" bold="true" box="[274,504,404,429]" pageId="18" pageNumber="119">
<specimenCount id="9D6EFD7EAE4FB622DC1B46DBFEA203DD" box="[274,377,404,429]" pageId="18" pageNumber="119" type="female">3 female</specimenCount>
<typeStatus id="54D38855AE4FB622DC8B46DBFE2303DD" box="[386,504,404,429]" pageId="18" pageNumber="119" type="paratype">paratypes</typeStatus>
</emphasis>
,
<collectingRegion id="49ACF815AE4FB622DF0046DBFD2603DD" box="[521,765,404,429]" country="Laos" name="Champasak" pageId="18" pageNumber="119">Champasak Province</collectingRegion>
: Bolaven Plateau, Ban Thôngvay (=
<location id="8EB7602CAE4FB622D99346DBFAD803DD" LSID="urn:lsid:plazi:treatment:03C187E1AE4DB622DD9E42BCFD4C004C:8EB7602CAE4FB622D99346DBFAD803DD" box="[1178,1283,404,429]" country="Laos" name="Xekatam" pageId="18" pageNumber="119" stateProvince="Champasak Province">Xékatam</location>
) vicinity, old logging road N of village,
<specimenCode id="DBCE9E8CAE4FB622DCC346F7FD4303A0" box="[458,664,440,464]" collectionCode="FMHD" pageId="18" pageNumber="119">FMHD#2008-040</specimenCode>
</materialsCitation>
, ANMT site 1232, 15°14.494'N, 106°31.807'E,
<quantity id="4C909B12AE4FB622D9B746F7FACC03BF" box="[1214,1303,440,464]" metricMagnitude="3" metricUnit="m" metricValue="1.17" pageId="18" pageNumber="119" unit="m" value="1170.0">1170 m</quantity>
, selectively logged forest, on
<date id="FFD61037AE4FB622DC784693FDD20385" box="[369,521,476,501]" pageId="18" pageNumber="119" value="2008-06-08" valueMax="2008-06-16" valueMin="2008-06-08">8–16/vi/2008</date>
, (A. Newton, M. Thayer) (FMNH);
<emphasis id="B91CEAE5AE4FB622DEC24693FB410385" bold="true" box="[971,1178,476,501]" pageId="18" pageNumber="119">1 male paratype</emphasis>
,
<materialsCitation id="3B003CAAAE4FB622D9B94693FD4C004C" ID-GBIF-Occurrence="1658411494" collectingDate="2008-05-06" collectingDateMax="2008-06-14" collectingDateMin="2008-05-06" collectorName="S. Tarasov" country="Laos" elevation="1200" latitude="15.077583" location="Ban Houayteuay" longLatPrecision="1" longitude="106.2808" municipality="Bolaven Plateau" pageId="18" pageNumber="119" specimenCount="1" specimenCount-female="1" stateProvince="Champasak Province" typeStatus="paratype">
<emphasis id="B91CEAE5AE4FB622D9B94693FA4E0385" bold="true" box="[1200,1429,476,501]" pageId="18" pageNumber="119">
<specimenCount id="9D6EFD7EAE4FB622D9B94693FAC00385" box="[1200,1307,476,501]" pageId="18" pageNumber="119" type="female">1 female</specimenCount>
<typeStatus id="54D38855AE4FB622D8234693FA4E0385" box="[1322,1429,476,501]" pageId="18" pageNumber="119" type="paratype">paratype</typeStatus>
</emphasis>
,
<collectingRegion id="49ACF815AE4FB622DD9E46B0FE500068" box="[151,395,511,536]" country="Laos" name="Champasak" pageId="18" pageNumber="119">Champasak Province</collectingRegion>
,
<collectingMunicipality id="6BB3AC8DAE4FB622DC9246B0FD810068" box="[411,602,511,536]" pageId="18" pageNumber="119">Bolaven Plateau</collectingMunicipality>
,
<location id="8EB7602CAE4FB622DF60454FFCF10068" LSID="urn:lsid:plazi:treatment:03C187E1AE4DB622DD9E42BCFD4C004C:8EB7602CAE4FB622DF60454FFCF10068" box="[617,810,512,536]" country="Laos" latitude="15.077583" longLatPrecision="1" longitude="106.2808" municipality="Bolaven Plateau" name="Ban Houayteuay" pageId="18" pageNumber="119" stateProvince="Champasak Province">Ban Houayteuay</location>
,
<geoCoordinate id="EE5C5030AE4FB622DE3346B0FC670068" box="[826,956,511,536]" direction="north" orientation="latitude" pageId="18" pageNumber="119" precision="1" value="15.077583">15°4.655'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE4FB622DEC546B0FBBD0068" box="[972,1126,511,536]" direction="east" orientation="longitude" pageId="18" pageNumber="119" precision="1" value="106.2808">106°16.848'E</geoCoordinate>
,
<elevation id="0045D1C4AE4FB622D97E454FFB090067" box="[1143,1234,512,536]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="18" pageNumber="119" unit="m" value="1200.0">
<quantity id="4C909B12AE4FB622D97E454FFB090067" box="[1143,1234,512,536]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="18" pageNumber="119" unit="m" value="1200.0">1200 m</quantity>
</elevation>
, disturb montane forest, on
<collectingDate id="EF92E9DFAE4FB622DC01456BFE68004D" box="[264,435,548,573]" pageId="18" pageNumber="119" value="2008-05-06" valueMax="2008-06-14" valueMin="2008-05-06">
<date id="FFD61037AE4FB622DC01456BFE68004D" box="[264,435,548,573]" pageId="18" pageNumber="119" value="2008-05-06" valueMax="2008-06-14" valueMin="2008-05-06">6/v–14/vi/2008</date>
</collectingDate>
(
<collectorName id="269D5321AE4FB622DCCD456BFDE0004C" box="[452,571,548,572]" pageId="18" pageNumber="119">S. Tarasov</collectorName>
) (CST).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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