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<mods:title id="6D942663E9C62540E0DAD4F004C77891">Two new species and one new record of Willowsia Shoebotham, 1917 (Collembola: Entomobryidae) from the Eastern Himalayan region of India</mods:title>
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<taxonomicName id="4C68F6A43461FFCE102DFBA7BA505E29" ID-CoL="5BZNT" authority="Pan, Zhang, &amp; Chen, 2006" authorityName="Pan, Zhang, &amp; Chen" authorityYear="2006" box="[538,1050,1038,1065]" class="Collembola" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Entomobryomorpha" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="shiae">
<emphasis id="B91C51353461FFCE102DFBA7BC925E28" bold="true" box="[538,728,1038,1065]" italics="true" pageId="14" pageNumber="115">Willowsia shiae</emphasis>
Pan, Zhang, &amp; Chen, 2006
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<figureCitation id="135391A23461FFCE1097FB9BBD435E4D" box="[672,777,1074,1100]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="15.[189,232,1982,2008]" captionStartId-1="16.[189,232,1928,1954]" captionStartId-2="17.[189,232,1963,1989]" captionTargetBox-0="[308,1335,356,1926]" captionTargetBox-1="[282,1302,541,1866]" captionTargetBox-2="[512,1063,707,1888]" captionTargetId-0="figure-32@15.[281,1306,1090,1943]" captionTargetId-1="figure-106@16.[231,1356,517,1887]" captionTargetId-2="figure-141@17.[429,1158,673,1923]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionText-0="Fig. 7. Willowsia shiae Pan, Zhang &amp; Chen, 2006, ♀ (reg. no. 3382/H14). Scale bars = 500 µm." captionText-1="Fig. 8. Willowsia shiae Pan, Zhang &amp; Chen, 2006, ♀ (reg. no. 3385/H14). A. Scales. B. Dorsal head chaetotaxy (left side). C. Labrum. D. Lateral processes of labial palp (right side). E. Dorsal chaetotaxy of Th II–III. F. Trochanteral organ. Scale bars: A = 20 μm; B, E = 100 μm; C, F = 50 μm; D = 25 μm." captionText-2="Fig. 9. Willowsia shiae Pan, Zhang &amp; Chen, 2006, ♀ (reg. no. 3384/H14). Abd. I– Abd. IV chaetotaxy. Scale bar = 200 μm." figureDoi-0="http://doi.org/10.5281/zenodo.11277736" figureDoi-1="http://doi.org/10.5281/zenodo.11277740" figureDoi-2="http://doi.org/10.5281/zenodo.11277742" httpUri-0="https://zenodo.org/record/11277736/files/figure.png" httpUri-1="https://zenodo.org/record/11277740/files/figure.png" httpUri-2="https://zenodo.org/record/11277742/files/figure.png" pageId="14" pageNumber="115">Figs 7–9</figureCitation>
, 
<tableCitation id="C6EAB89C3461FFCE1123FB9BBDDE5E4D" box="[788,916,1074,1100]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="3.[189,259,558,586]" captionStartId-1="8.[189,259,265,291]" captionTargetPageId-0="3" captionTargetPageId-1="8" captionText-0="Table 1(continued on nextpage). Comparison of morphological charactersbetweenthe newspecies and relatedworld speciesofWillowsia Shoebotham, 1917." captionText-1="Table 2. Chaetotaxy comparison between the closely related world species of Willowsia Shoebotham, 1917." httpUri-0="http://table.plazi.org/id/DF17DDAF346CFFC3128AFD87BEB7586C" httpUri-1="http://table.plazi.org/id/DF17DDAF3467FFC8128AFEA0BF4A5B47" pageId="14" pageNumber="115" tableUuid-0="DF17DDAF346CFFC3128AFD87BEB7586C" tableUuid-1="DF17DDAF3467FFC8128AFEA0BF4A5B47">Tables 1–2</tableCitation>
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<emphasis id="B91C51353461FFCE128AFBDABFE75E8C" bold="true" box="[189,429,1139,1165]" pageId="14" pageNumber="115">Material examined</emphasis>
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leg.; reg no. 3382/H14; 
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(on slide), 14 specs in alcohol; 
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<collectingDate id="EF92520F3461FFCE11F1FA87BA2F5F48" box="[966,1125,1326,1353]" pageId="14" pageNumber="115" value="2023-03-26">26 Mar. 2023</collectingDate>
</date>
; 
<collectorName id="269DE8F13461FFCE1646FA86BAAC5F49" box="[1137,1254,1326,1353]" pageId="14" pageNumber="115">P. Mandal</collectorName>
leg.; reg no. 3384/H14; 
<collectionCode id="ED7915E23461FFCE1375FAFBBF375F6D" box="[322,381,1362,1388]" pageId="14" pageNumber="115">NZC</collectionCode>
</materialsCitation>
• 
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<specimenCount id="9D6E46AE3461FFCE13ADFAFBBF8D5F6D" box="[410,455,1362,1388]" count="1" pageId="14" pageNumber="115" type="female">1 ♀</specimenCount>
(on slide), 20 specs in alcohol; 
<collectingMunicipality id="6BB3175D3461FFCE1177FAFBBDAD5F6D" box="[832,999,1362,1388]" pageId="14" pageNumber="115">West Kameng</collectingMunicipality>
, near 
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; 
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, 
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, 
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a.s.l.
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; 
<date id="FFD6ABE73461FFCE101DFADCBC815F8E" box="[554,715,1397,1423]" pageId="14" pageNumber="115" value="2023-03-26">
<collectingDate id="EF92520F3461FFCE101DFADCBC815F8E" box="[554,715,1397,1423]" pageId="14" pageNumber="115" value="2023-03-26">26 Mar. 2023</collectingDate>
</date>
; 
<collectorName id="269DE8F13461FFCE10E0FADCBD225F8E" box="[727,872,1397,1423]" pageId="14" pageNumber="115">K.K. Suman</collectorName>
leg.; reg no. 3385/H14; 
<collectionCode id="ED7915E23461FFCE16BFFADCBA8C5F8E" box="[1160,1222,1397,1423]" pageId="14" pageNumber="115">NZC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C372DEAC3461FFD1128AFA1FBF0C5B47" lastPageId="17" lastPageNumber="118" pageId="14" pageNumber="115" type="diagnosis">
<paragraph id="8BD78D273461FFCE128AFA1FBF7D5FD1" blockId="14.[189,311,1462,1488]" box="[189,311,1462,1488]" pageId="14" pageNumber="115">
<heading id="D09F3A4B3461FFCE128AFA1FBF7D5FD1" bold="true" box="[189,311,1462,1488]" fontSize="11" level="3" pageId="14" pageNumber="115" reason="3">
<emphasis id="B91C51353461FFCE128AFA1FBF7D5FD1" bold="true" box="[189,311,1462,1488]" pageId="14" pageNumber="115">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD78D273461FFD1128AFA4CBF0C5B47" blockId="14.[189,1401,1509,1993]" lastBlockId="17.[189,1398,265,326]" lastPageId="17" lastPageNumber="118" pageId="14" pageNumber="115">
Average adult body length 
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. General body colour pale straw yellow, blue pigment patches scattered all over body, Abd. III–V almost completely pigmented, rest of segments become darker at lateral edges, head with diffused pigment; antennae and legs evenly pigmented (
<figureCitation id="135391A23461FFCE16A5F982BB5D5C44" box="[1170,1303,1579,1605]" captionStart="Fig" captionStartId="15.[189,232,1982,2008]" captionTargetBox="[308,1335,356,1926]" captionTargetId="figure-32@15.[281,1306,1090,1943]" captionTargetPageId="15" captionText="Fig. 7. Willowsia shiae Pan, Zhang &amp; Chen, 2006, ♀ (reg. no. 3382/H14). Scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.11277736" httpUri="https://zenodo.org/record/11277736/files/figure.png" pageId="14" pageNumber="115">Fig. 7A–B</figureCitation>
). Scales spinulate 
<typeStatus id="54D333853461FFCE1307F9E6BF295C68" box="[304,355,1615,1641]" pageId="14" pageNumber="115">type</typeStatus>
(
<figureCitation id="135391A23461FFCE1342F9E7BF9F5C68" box="[373,469,1614,1641]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[282,1302,541,1866]" captionTargetId="figure-106@16.[231,1356,517,1887]" captionTargetPageId="16" captionText="Fig. 8. Willowsia shiae Pan, Zhang &amp; Chen, 2006, ♀ (reg. no. 3385/H14). A. Scales. B. Dorsal head chaetotaxy (left side). C. Labrum. D. Lateral processes of labial palp (right side). E. Dorsal chaetotaxy of Th II–III. F. Trochanteral organ. Scale bars: A = 20 μm; B, E = 100 μm; C, F = 50 μm; D = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.11277740" httpUri="https://zenodo.org/record/11277740/files/figure.png" pageId="14" pageNumber="115">Fig. 8A</figureCitation>
), present on body, antennae, ventral side of manubrium. Legs without scales. Antennal segment ratio as I: II: III: IV = 1: 2: 1.3–1.9: 2–3.4. Ant. IV with bilobed apical bulb. Ant. III organ with two rod-like sense organ. Head with five antennal (An), four median (M 1 –M 4), eight sutural (S 
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–S 
<subScript id="17EC8F623461FFCE12C8F962BF465CD8" attach="left" box="[255,268,1739,1753]" fontSize="6" pageId="14" pageNumber="115">6,</subScript>
S 
<subScript id="17EC8F623461FFCE1316F962BF655CD8" attach="left" box="[289,303,1739,1753]" fontSize="6" pageId="14" pageNumber="115">4i</subScript>
, S 
<subScript id="17EC8F623461FFCE1367F962BF175CD8" attach="left" box="[336,349,1739,1753]" fontSize="6" pageId="14" pageNumber="115">5i</subScript>
) setae (
<figureCitation id="135391A23461FFCE1389F911BC565CD3" box="[446,540,1720,1746]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[282,1302,541,1866]" captionTargetId="figure-106@16.[231,1356,517,1887]" captionTargetPageId="16" captionText="Fig. 8. Willowsia shiae Pan, Zhang &amp; Chen, 2006, ♀ (reg. no. 3385/H14). A. Scales. B. Dorsal head chaetotaxy (left side). C. Labrum. D. Lateral processes of labial palp (right side). E. Dorsal chaetotaxy of Th II–III. F. Trochanteral organ. Scale bars: A = 20 μm; B, E = 100 μm; C, F = 50 μm; D = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.11277740" httpUri="https://zenodo.org/record/11277740/files/figure.png" pageId="14" pageNumber="115">Fig. 8B</figureCitation>
). Blunt and conical labral papilla without setae; labral setae formula as 4/4, 5, 4, prelabral setae ciliate, others smooth (
<figureCitation id="135391A23461FFCE10C8F972BD165CF4" box="[767,860,1755,1781]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[282,1302,541,1866]" captionTargetId="figure-106@16.[231,1356,517,1887]" captionTargetPageId="16" captionText="Fig. 8. Willowsia shiae Pan, Zhang &amp; Chen, 2006, ♀ (reg. no. 3385/H14). A. Scales. B. Dorsal head chaetotaxy (left side). C. Labrum. D. Lateral processes of labial palp (right side). E. Dorsal chaetotaxy of Th II–III. F. Trochanteral organ. Scale bars: A = 20 μm; B, E = 100 μm; C, F = 50 μm; D = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.11277740" httpUri="https://zenodo.org/record/11277740/files/figure.png" pageId="14" pageNumber="115">Fig. 8C</figureCitation>
). Labial triangle with five finely ciliated and five smooth setae. Lateral process of labial palp not reaching the base (
<figureCitation id="135391A23461FFCE163DF957BA205D18" box="[1034,1130,1790,1817]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[282,1302,541,1866]" captionTargetId="figure-106@16.[231,1356,517,1887]" captionTargetPageId="16" captionText="Fig. 8. Willowsia shiae Pan, Zhang &amp; Chen, 2006, ♀ (reg. no. 3385/H14). A. Scales. B. Dorsal head chaetotaxy (left side). C. Labrum. D. Lateral processes of labial palp (right side). E. Dorsal chaetotaxy of Th II–III. F. Trochanteral organ. Scale bars: A = 20 μm; B, E = 100 μm; C, F = 50 μm; D = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.11277740" httpUri="https://zenodo.org/record/11277740/files/figure.png" pageId="14" pageNumber="115">Fig. 8D</figureCitation>
). Dorsal chaetotaxy of Th. II as shown in the 
<figureCitation id="135391A23461FFCE13F4F88BBC535D3D" box="[451,537,1826,1852]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[282,1302,541,1866]" captionTargetId="figure-106@16.[231,1356,517,1887]" captionTargetPageId="16" captionText="Fig. 8. Willowsia shiae Pan, Zhang &amp; Chen, 2006, ♀ (reg. no. 3385/H14). A. Scales. B. Dorsal head chaetotaxy (left side). C. Labrum. D. Lateral processes of labial palp (right side). E. Dorsal chaetotaxy of Th II–III. F. Trochanteral organ. Scale bars: A = 20 μm; B, E = 100 μm; C, F = 50 μm; D = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.11277740" httpUri="https://zenodo.org/record/11277740/files/figure.png" pageId="14" pageNumber="115">Fig. 8E</figureCitation>
with two medio-median (m2), four medio-lateral (m4, m4i, m4p, m5), 19 posterior mac and two sens. Th. III with 16 median, eight lateral mac and two sens (
<figureCitation id="135391A23461FFCE1699F8ECBB415D5E" box="[1198,1291,1861,1887]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[282,1302,541,1866]" captionTargetId="figure-106@16.[231,1356,517,1887]" captionTargetPageId="16" captionText="Fig. 8. Willowsia shiae Pan, Zhang &amp; Chen, 2006, ♀ (reg. no. 3385/H14). A. Scales. B. Dorsal head chaetotaxy (left side). C. Labrum. D. Lateral processes of labial palp (right side). E. Dorsal chaetotaxy of Th II–III. F. Trochanteral organ. Scale bars: A = 20 μm; B, E = 100 μm; C, F = 50 μm; D = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.11277740" httpUri="https://zenodo.org/record/11277740/files/figure.png" pageId="14" pageNumber="115">Fig. 8E</figureCitation>
). Unguis with one outer, paired inner and one unpaired teeth. Unguiculus lanceolate with a serreted edge. Tenent hair with a clavate tip and slightly larger than unguis in length. Trochanteral organ with 37–40 smooth spiny setae (
<figureCitation id="135391A23461FFCE1363F807BFE55DC8" box="[340,431,1966,1993]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[282,1302,541,1866]" captionTargetId="figure-106@16.[231,1356,517,1887]" captionTargetPageId="16" captionText="Fig. 8. Willowsia shiae Pan, Zhang &amp; Chen, 2006, ♀ (reg. no. 3385/H14). A. Scales. B. Dorsal head chaetotaxy (left side). C. Labrum. D. Lateral processes of labial palp (right side). E. Dorsal chaetotaxy of Th II–III. F. Trochanteral organ. Scale bars: A = 20 μm; B, E = 100 μm; C, F = 50 μm; D = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.11277740" httpUri="https://zenodo.org/record/11277740/files/figure.png" pageId="14" pageNumber="115">Fig. 8F</figureCitation>
). Dorsal mac position as shown in 
<figureCitation id="135391A23461FFCE116DF807BDE95DC8" box="[858,931,1966,1993]" captionStart="Fig" captionStartId="17.[189,232,1963,1989]" captionTargetBox="[512,1063,707,1888]" captionTargetId="figure-141@17.[429,1158,673,1923]" captionTargetPageId="17" captionText="Fig. 9. Willowsia shiae Pan, Zhang &amp; Chen, 2006, ♀ (reg. no. 3384/H14). Abd. I– Abd. IV chaetotaxy. Scale bar = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.11277742" httpUri="https://zenodo.org/record/11277742/files/figure.png" pageId="14" pageNumber="115">Fig. 9</figureCitation>
. Abd. I with 4 mac (m2–m4, m4i) and one sens, M3 arch of Abd. II with four setae (m3, m3ep, m3e, m3ea), inner area with one seta (a3), m5 present and two sens. Abd. III three central (a2–a3, m3) and five lateral setae (am6, pm6, p6, m7a, p7), three sens present. Abd. IV with nine dorso-central (A3–A4, A6, B3–B6, Sm, Si),12–16 lateral setae and 6 S-setae. Ventral tube with 5+5 anterior large and other small ciliated setae, 5+5 smooth and 12–14 ciliated setae on lateral flap, posterior side with 1+1 smooth setae. Tenaculum quadridented. Length ratio of manubrium: dens as 1:1.26; dens uncrenulated part: mucro as 1.4: 1. Mucro bidented with a basal spine.
</paragraph>
</subSubSection>
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<paragraph id="8BD78D273460FFCF128AF817BB6F5DD8" blockId="15.[189,1317,1982,2009]" box="[189,1317,1982,2009]" pageId="15" pageNumber="116">
<emphasis id="B91C51353460FFCF128AF817BF415DD9" bold="true" box="[189,267,1982,2008]" pageId="15" pageNumber="116">Fig. 7.</emphasis>
<taxonomicName id="4C68F6A43460FFCF1324F817BD415DD9" authority="Pan, Zhang &amp; Chen, 2006" authorityName="Pan, Zhang, &amp; Chen" authorityYear="2006" box="[275,779,1982,2009]" class="Collembola" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Entomobryomorpha" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="shiae">
<emphasis id="B91C51353460FFCF1324F817BF855DD9" box="[275,463,1982,2008]" italics="true" pageId="15" pageNumber="116">Willowsia shiae</emphasis>
<bibRefCitation id="EFF9F0D63460FFCF13E2F816BD415DD9" author="Pan J. L. &amp; Zhang F. &amp; Chen J. X." box="[469,779,1982,2009]" pageId="15" pageNumber="116" pagination="261 - 266" refId="ref7986" refString="Pan J. L., Zhang F. &amp; Chen J. X. 2006. A new Chinese species of the genus Willowsia from Tibet (Collembola: Entomobryidae). Journal of the Kansas Entomological Society 79 (3): 261 - 266." type="journal article" year="2006">Pan, Zhang &amp; Chen, 2006</bibRefCitation>
</taxonomicName>
, ♀ (reg. no. 3382/H14). Scale bars = 500 µm.
</paragraph>
</caption>
<caption id="DF17DDAF347FFFD0128AF821BB265DE8" ID-DOI="http://doi.org/10.5281/zenodo.11277740" ID-Zenodo-Dep="11277740" httpUri="https://zenodo.org/record/11277740/files/figure.png" pageId="16" pageNumber="117" startId="16.[189,232,1928,1954]" targetBox="[282,1302,541,1866]" targetPageId="16">
<paragraph id="8BD78D27347FFFD0128AF821BB265DE8" blockId="16.[189,1398,1928,2025]" pageId="16" pageNumber="117">
<emphasis id="B91C5135347FFFD0128AF821BF475DA3" bold="true" box="[189,269,1928,1954]" pageId="16" pageNumber="117">Fig. 8.</emphasis>
<taxonomicName id="4C68F6A4347FFFD01321F821BD535DA3" authority="Pan, Zhang &amp; Chen, 2006" authorityName="Pan, Zhang, &amp; Chen" authorityYear="2006" box="[278,793,1928,1954]" class="Collembola" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Entomobryomorpha" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="shiae">
<emphasis id="B91C5135347FFFD01321F821BF9E5DA3" box="[278,468,1928,1954]" italics="true" pageId="16" pageNumber="117">Willowsia shiae</emphasis>
<bibRefCitation id="EFF9F0D6347FFFD013EBF821BD535DA3" author="Pan J. L. &amp; Zhang F. &amp; Chen J. X." box="[476,793,1928,1954]" pageId="16" pageNumber="117" pagination="261 - 266" refId="ref7986" refString="Pan J. L., Zhang F. &amp; Chen J. X. 2006. A new Chinese species of the genus Willowsia from Tibet (Collembola: Entomobryidae). Journal of the Kansas Entomological Society 79 (3): 261 - 266." type="journal article" year="2006">Pan, Zhang &amp; Chen, 2006</bibRefCitation>
</taxonomicName>
, ♀ (reg. no. 3385/H14). 
<emphasis id="B91C5135347FFFD01675F821BA1D5DA3" bold="true" box="[1090,1111,1928,1954]" pageId="16" pageNumber="117">A</emphasis>
. Scales. 
<emphasis id="B91C5135347FFFD016F2F821BA9D5DA3" bold="true" box="[1221,1239,1928,1954]" pageId="16" pageNumber="117">B</emphasis>
. Dorsal head chaetotaxy (left side). 
<emphasis id="B91C5135347FFFD013F3F802BF935DC4" bold="true" box="[452,473,1963,1989]" pageId="16" pageNumber="117">C</emphasis>
. Labrum. 
<emphasis id="B91C5135347FFFD01063F802BC235DC4" bold="true" box="[596,617,1963,1989]" pageId="16" pageNumber="117">D</emphasis>
. Lateral processes of labial palp (right side). 
<emphasis id="B91C5135347FFFD01648F802BAD85DC4" bold="true" box="[1151,1170,1963,1989]" pageId="16" pageNumber="117">E</emphasis>
. Dorsal chaetotaxy of Th II–III. 
<emphasis id="B91C5135347FFFD01365F867BF2E5DE9" bold="true" box="[338,356,1998,2024]" pageId="16" pageNumber="117">F</emphasis>
. Trochanteral organ. Scale bars: A = 20 μm; B, E = 100 μm; C, F = 50 μm; D = 25 μm.
</paragraph>
</caption>
<subSubSection id="C372DEAC347EFFD1128AFEC7BC5D5867" pageId="17" pageNumber="118" type="discussion">
<paragraph id="8BD78D27347EFFD1128AFEC7BF7A5B89" blockId="17.[189,304,366,392]" box="[189,304,366,392]" pageId="17" pageNumber="118">
<heading id="D09F3A4B347EFFD1128AFEC7BF7A5B89" bold="true" box="[189,304,366,392]" fontSize="11" level="3" pageId="17" pageNumber="118" reason="3">
<emphasis id="B91C5135347EFFD1128AFEC7BF7A5B89" bold="true" box="[189,304,366,392]" pageId="17" pageNumber="118">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD78D27347EFFD1128AFE34BC5D5867" blockId="17.[188,1399,412,615]" pageId="17" pageNumber="118">
Most of the characters of the Indian specimens of 
<taxonomicName id="4C68F6A4347EFFD11137FE34BD295BB7" authorityName="Pan, Zhang, &amp; Chen" authorityYear="2006" box="[768,867,412,438]" class="Collembola" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Entomobryomorpha" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="shiae">
<emphasis id="B91C5135347EFFD11137FE34BD295BB7" box="[768,867,412,438]" italics="true" pageId="17" pageNumber="118">W. shiae</emphasis>
</taxonomicName>
are similar as per the description of 
<bibRefCitation id="EFF9F0D6347EFFD1173EFE34BF455BD8" author="Pan J. L. &amp; Zhang F. &amp; Chen J. X." pageId="17" pageNumber="118" pagination="261 - 266" refId="ref7986" refString="Pan J. L., Zhang F. &amp; Chen J. X. 2006. A new Chinese species of the genus Willowsia from Tibet (Collembola: Entomobryidae). Journal of the Kansas Entomological Society 79 (3): 261 - 266." type="journal article" year="2006">
Pan 
<emphasis id="B91C5135347EFFD1170CFE34BB3C5BB7" box="[1339,1398,412,438]" italics="true" pageId="17" pageNumber="118">et al.</emphasis>
(2006)
</bibRefCitation>
; however, we have found that there are differences between them in dorsal head chaetotaxy; Th. II with two mac in Gr. I (T 
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), three mac in Gr. II (T 
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<figureCitation id="135391A2347EFFD11635FE4BBA105BFC" box="[1026,1114,482,509]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[282,1302,541,1866]" captionTargetId="figure-106@16.[231,1356,517,1887]" captionTargetPageId="16" captionText="Fig. 8. Willowsia shiae Pan, Zhang &amp; Chen, 2006, ♀ (reg. no. 3385/H14). A. Scales. B. Dorsal head chaetotaxy (left side). C. Labrum. D. Lateral processes of labial palp (right side). E. Dorsal chaetotaxy of Th II–III. F. Trochanteral organ. Scale bars: A = 20 μm; B, E = 100 μm; C, F = 50 μm; D = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.11277740" httpUri="https://zenodo.org/record/11277740/files/figure.png" pageId="17" pageNumber="118">Fig. 8E</figureCitation>
) and four mac in the p1 row of Th. II, p5 also present. Two sens present in each Th. segment. We have also named the mac of Th. II–III and Abd. I–IV following 
<bibRefCitation id="EFF9F0D6347EFFD11056FD80BD035842" author="Zhang F. &amp; Bellini B. C. &amp; Soto-Adames F. N." box="[609,841,553,579]" pageId="17" pageNumber="118" pagination="249 - 278" refId="ref8545" refString="Zhang F., Bellini B. C. &amp; Soto-Adames F. N. 2019. New insights into the systematics of Entomobryoidea (Collembola: Entomobryomorpha): first instar chaetotaxy, homology and classification. Zoological Systematics 44 (4): 249 - 278. https: // doi. org / 10.11865 / zs. 201926" type="journal article" year="2019">
Zhang 
<emphasis id="B91C5135347EFFD11084FD83BCBA5842" box="[691,752,553,579]" italics="true" pageId="17" pageNumber="118">et al.</emphasis>
(2019)
</bibRefCitation>
. Collected from the leaf litter of pine and other trees of tropical alpine forest.
</paragraph>
</subSubSection>
</treatment>
</document>