389 lines
47 KiB
XML
389 lines
47 KiB
XML
<document id="446A8DD1ACED3D9E8C261E5E323DC8DA" ID-DOI="10.5281/zenodo.281871" ID-GBIF-Dataset="70a9e355-8064-4e10-ae64-8fb6a357db5b" ID-ISSN="1175-5326" ID-Zenodo-Dep="281871" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460502734697" checkinUser="plazi" docAuthor="Johnson, Chelséa B., Quah, Evan, Anuar, Shahrul, Wood, Perry L., Grismer, Jesse L., Greer, Lee F., Onn, Chan Kin & Ahmad, Norhayati" docDate="2012" docId="03F587A5FFEBFF8527D4FB76FABAF0AB" docLanguage="en" docName="zt03406p058.pdf" docOrigin="Zootaxa 3406" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cyrtodactylus payacola Johnson, Quah, Anuar, Wood, Grismer, Greer, Onn & Ahmad, 2012, sp. nov." docType="treatment" docVersion="8" lastPageNumber="57" masterDocId="FFCCFFDDFFE5FF972743FF87FFFDF22F" masterDocTitle="Phylogeography, geographic variation, and taxonomy of the Bent-toed Gecko Cyrtodactylus quadrivirgatus Taylor, 1962 from Peninsular Malaysia with the description of a new swamp dwelling species" masterLastPageNumber="58" masterPageNumber="39" pageNumber="53" updateTime="1698312202066" updateUser="plazi">
|
||
<mods:mods id="8308E65A4EA456D90968F916340C51C6" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="F0115D62BC2233852093E1F537770436">
|
||
<mods:title id="417CBDF33DF882FE19643FDD172A31A6">Phylogeography, geographic variation, and taxonomy of the Bent-toed Gecko Cyrtodactylus quadrivirgatus Taylor, 1962 from Peninsular Malaysia with the description of a new swamp dwelling species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="FC56F2A780733F380784DB36EDF1724E" type="personal">
|
||
<mods:role id="80A950C8C97563A35654CC870B01A276">
|
||
<mods:roleTerm id="0DA53886E1817119C650609B08C43EB7">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="DFF5501716649613017883042925064E">Johnson, Chelséa B.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="2D49C7CF612254696174AC3CB1BAB11B" type="personal">
|
||
<mods:role id="497D0589841188555F8004DA4140F501">
|
||
<mods:roleTerm id="B2F0CDEFE955C6966B496B80FF470CAB">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1CE1826D251721CC93344F7C16087000">Quah, Evan</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="CFFB52F2422E897A7DDDDE521DBA74B7" type="personal">
|
||
<mods:role id="E25C52B86B221F47B1ACD699FFE81BE3">
|
||
<mods:roleTerm id="D9AE37C114AD116D0CD56EDC609B6767">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="43402CC48A29C3689D24237FCA59309B">Anuar, Shahrul</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="AF2485A79F96504EBC68EAC5B1459FA3" type="personal">
|
||
<mods:role id="44B656DA581ABC7A735E427AE0F4113F">
|
||
<mods:roleTerm id="D827E81B83337E5CAA6DC6EF0FAF6F0E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="01DDBB7EADF560A29C9A8732213ABC88">Wood, Perry L.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="39D069C9490D2205567C16C11A69E136" type="personal">
|
||
<mods:role id="6234E5778F64A17AC821711AE60586E4">
|
||
<mods:roleTerm id="7E05CDE78C9CD78C16231FEDD2D238D8">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="89635CA79DC058116366B8D421BCCF3C">Grismer, Jesse L.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="8AC11EC6E9E4999B4234B3C1127A9858" type="personal">
|
||
<mods:role id="FD4E1CA29B8BCA4458BBCFA79C2C9F4D">
|
||
<mods:roleTerm id="A6344AAC170E8BB48A1EB6F666BEE650">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="BF7E1E0405638A8C229559F1A92168B7">Greer, Lee F.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="24BFE7F2CDF2878C1A5F810547993F4A" type="personal">
|
||
<mods:role id="D6E52B9505631322806C6D9AECB04FA2">
|
||
<mods:roleTerm id="BDB679606A8EEE3F8310FC03DEA1859A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="06ED7C2645F8BD8C5A289D96022DB40B">Onn, Chan Kin</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="E3FECBD863817E758B76ADFED341E061" type="personal">
|
||
<mods:role id="B0E0CA3C6934F7929F251B601AE0EABC">
|
||
<mods:roleTerm id="BF7F4639016630665FB614DDD1253AB1">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1CA3E2792982D68D2DCD7782E96956DB">Ahmad, Norhayati</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="A46C448EFA3658B9C595FDC6BDD213D0">text</mods:typeOfResource>
|
||
<mods:relatedItem id="A5C4E1CB68E56A2510C0CA156ED65528" type="host">
|
||
<mods:titleInfo id="2553D781D590935A454DC109D27E31AB">
|
||
<mods:title id="89DD9C5DB3966F6E9E9A4C577E71A9DC">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="1D1381083EFD371F73798B8D0F150663">
|
||
<mods:date id="554CE64EFB2645644940AC300D436226">2012</mods:date>
|
||
<mods:detail id="4D99C766AB235361C90F1EC1BB71D9FE" type="volume">
|
||
<mods:number id="2B3CB72E8F0843D729CAA2CDA2415A58">3406</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="7072B764F4F07874B677B010EBCC0944" unit="page">
|
||
<mods:start id="6606613B9798D5AE960397357C3BD9AC">39</mods:start>
|
||
<mods:end id="D63AAED0D7B8CBFCDEABD0DE3D60EABA">58</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="4CE355609890F7994C7255E44276F4B3">journal article</mods:classification>
|
||
<mods:identifier id="F1D308436B442A31A7768804403C2AC0" type="DOI">10.5281/zenodo.281871</mods:identifier>
|
||
<mods:identifier id="DBEF2D04BFF8FC1D0404266C54FDA76A" type="GBIF-Dataset">70a9e355-8064-4e10-ae64-8fb6a357db5b</mods:identifier>
|
||
<mods:identifier id="6C7A812C06D955038BBF05D24351AFD4" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="DB8BF28B067CB60F6C3EC81611A2D7E9" type="Zenodo-Dep">281871</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03F587A5FFEBFF8527D4FB76FABAF0AB" ID-DOI="http://doi.org/10.5281/zenodo.5690860" ID-GBIF-Taxon="119521743" ID-Zenodo-Dep="5690860" LSID="urn:lsid:plazi:treatment:03F587A5FFEBFF8527D4FB76FABAF0AB" httpUri="http://treatment.plazi.org/id/03F587A5FFEBFF8527D4FB76FABAF0AB" lastPageId="18" lastPageNumber="57" pageId="14" pageNumber="53">
|
||
<subSubSection id="C3466538FFEBFF9927D4FB76FF02F703" pageId="14" pageNumber="53" type="nomenclature">
|
||
<paragraph id="8BE336B3FFEBFF9927D4FB76FDE9F725" blockId="14.[151,532,1264,1324]" box="[151,532,1264,1291]" pageId="14" pageNumber="53">
|
||
<heading id="D0AB81DFFFEBFF9927D4FB76FDE9F725" bold="true" box="[151,532,1264,1291]" fontSize="11" level="1" pageId="14" pageNumber="53" reason="1">
|
||
<emphasis id="B928EAA1FFEBFF9927D4FB76FDE9F725" bold="true" box="[151,532,1264,1291]" pageId="14" pageNumber="53">
|
||
<taxonomicName id="4C5C4D30FFEBFF9927D4FB76FE4DF724" box="[151,432,1265,1291]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="53" phylum="Chordata" rank="species" species="payacola" status="sp. nov.">
|
||
<emphasis id="B928EAA1FFEBFF9927D4FB76FE4DF724" bold="true" box="[151,432,1265,1291]" italics="true" pageId="14" pageNumber="53">Cyrtodactylus payacola</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21B57DAFFEBFF9926F5FB77FDE9F725" box="[438,532,1264,1290]" pageId="14" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BE336B3FFEBFF9927D4FA93FF02F703" blockId="14.[151,532,1264,1324]" box="[151,255,1300,1324]" pageId="14" pageNumber="53">
|
||
<figureCitation id="13672A36FFEBFF9927D4FA93FF18F703" box="[151,229,1300,1324]" captionStart="FIGURE 6" captionStartId="15.[151,250,793,816]" captionTargetBox="[163,1436,193,779]" captionTargetId="figure@15.[163,1448,193,779]" captionTargetPageId="15" captionText="FIGURE 6. Type series of Cyrtodactylus payacola sp. nov. Holotype LSUHC 10074." httpUri="https://zenodo.org/record/281877/files/figure.png" pageId="14" pageNumber="53">Figs. 6</figureCitation>
|
||
,
|
||
<figureCitation id="13672A36FFEBFF9927B2FA92FF02F703" box="[241,255,1301,1324]" captionStart="FIGURE 7" captionStartId="16.[151,250,1363,1386]" captionTargetBox="[172,1424,176,1340]" captionTargetId="figure@16.[172,1424,163,1344]" captionTargetPageId="16" captionText="FIGURE 7. Cyrtodactylus payacola sp. nov. showing dark (upper; LSUHC 10074) and light (lower; LSUHC 10074) color phases." httpUri="https://zenodo.org/record/281878/files/figure.png" pageId="14" pageNumber="53">7</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3466538FFEBFF9927D4FADBFD5BF75E" box="[151,678,1371,1394]" pageId="14" pageNumber="53" type="reference_group">
|
||
<paragraph id="8BE336B3FFEBFF9927D4FADBFD5BF75E" blockId="14.[151,678,1371,1394]" box="[151,678,1371,1394]" pageId="14" pageNumber="53">
|
||
<treatmentCitationGroup id="AB4C119DFFEBFF9927D4FADBFD5BF75E" box="[151,678,1371,1394]" pageId="14" pageNumber="53">
|
||
<taxonomicName id="4C5C4D30FFEBFF9927D4FADBFD5FF75E" authority="Grismer 2011:429" authorityName="Grismer" authorityPageNumber="429" authorityYear="2011" box="[151,674,1371,1394]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="53" phylum="Chordata" rank="species" species="quadrivirgatus">
|
||
<emphasis id="B928EAA1FFEBFF9927D4FADBFED8F75D" box="[151,293,1372,1394]" italics="true" pageId="14" pageNumber="53">Cyrtodactylus</emphasis>
|
||
cf.
|
||
<emphasis id="B928EAA1FFEBFF99260FFADBFE1CF75D" box="[332,481,1372,1394]" italics="true" pageId="14" pageNumber="53">quadrivirgatus</emphasis>
|
||
<treatmentCitation id="0AFD10A2FFEBFF9926A4FADCFD5FF75E" author="Grismer" box="[487,674,1371,1393]" page="429" pageId="14" pageNumber="53" year="2011">
|
||
<bibRefCitation id="EFCD4B42FFEBFF9926A4FADCFD8BF75E" author="Grismer" box="[487,630,1371,1393]" pageId="14" pageNumber="53" refString="Grismer, L. L. (2011) Lizards of Peninsular Malaysia, Singapore and their Adjacent Archipelagos. Edition Chimaira, 728 pp." type="book" year="2011">Grismer 2011</bibRefCitation>
|
||
:429
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3466538FFEBFF9927D4FA18FC9FF49D" pageId="14" pageNumber="53" type="materials_examined">
|
||
<paragraph id="8BE336B3FFEBFF9927D4FA18FB98F7F5" blockId="14.[151,1437,1438,2002]" pageId="14" pageNumber="53">
|
||
<emphasis id="B928EAA1FFEBFF9927D4FA18FEF6F798" bold="true" box="[151,267,1439,1463]" pageId="14" pageNumber="53">
|
||
<typeStatus id="54E78811FFEBFF9927D4FA18FEFBF798" box="[151,262,1439,1463]" pageId="14" pageNumber="53" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult male (
|
||
<collectionCode id="ED4DAE76FFEBFF9926DEFA19FE07F799" box="[413,506,1438,1462]" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="14" pageNumber="53">LSUHC</collectionCode>
|
||
10074) collected by E. Quah S. H. on
|
||
<date id="FFE21073FFEBFF9924EEFA19FB85F799" box="[941,1144,1438,1462]" pageId="14" pageNumber="53" value="2011-09-04">4 September 2011</date>
|
||
from Bukit Panchor State Park, Penang, West
|
||
<collectingCountry id="F34B7623FFEBFF992634FA45FE23F7F5" box="[375,478,1474,1498]" name="Malaysia" pageId="14" pageNumber="53">Malaysia</collectingCountry>
|
||
<geoCoordinate id="EE685074FFEBFF9926A7FA45FD84F7F5" box="[484,633,1474,1498]" direction="north" orientation="latitude" pageId="14" pageNumber="53" precision="1" value="5.15775">05º09.465' N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE685074FFEBFF9925C6FA45FCDCF7F5" box="[645,801,1474,1498]" direction="east" orientation="longitude" pageId="14" pageNumber="53" precision="1" value="100.54808">100º32.885' E</geoCoordinate>
|
||
at an elevation of
|
||
<quantity id="4CA49B56FFEBFF9924B3FA45FBD7F7F5" box="[1008,1066,1474,1498]" metricMagnitude="1" metricUnit="m" metricValue="4.7" pageId="14" pageNumber="53" unit="m" value="47.0">47 m</quantity>
|
||
a.s.l.
|
||
</paragraph>
|
||
<paragraph id="8BE336B3FFEBFF992784FA60FCD6F469" blockId="14.[151,1437,1438,2002]" pageId="14" pageNumber="53">
|
||
<emphasis id="B928EAA1FFEBFF992784FA60FEBBF7D0" bold="true" box="[199,326,1511,1535]" pageId="14" pageNumber="53">
|
||
<typeStatus id="54E78811FFEBFF992784FA60FEBCF7D0" box="[199,321,1511,1535]" pageId="14" pageNumber="53" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
All paratopotypes were collected in various combinations by M. A. Muin, E. Quah S.H., S. Anuar, C. K. Onn, and L. L. Grismer from the same locality as the
|
||
<typeStatus id="54E78811FFEBFF992469F98DFC6DF40D" box="[810,912,1546,1570]" pageId="14" pageNumber="53" type="holotype">holotype</typeStatus>
|
||
.
|
||
<collectionCode id="ED4DAE76FFEBFF9924DAF98DFC08F40D" box="[921,1013,1546,1570]" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="14" pageNumber="53">LSUHC</collectionCode>
|
||
9982 was collected on
|
||
<date id="FFE21073FFEBFF9923BAF98DFA65F40D" box="[1273,1432,1546,1570]" pageId="14" pageNumber="53" value="2010-03-06">6 March 2010</date>
|
||
;
|
||
<collectionCode id="ED4DAE76FFEBFF9927D4F9A9FF0EF469" box="[151,243,1582,1606]" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="14" pageNumber="53">LSUHC</collectionCode>
|
||
10070 and 10071 were collected on
|
||
<date id="FFE21073FFEBFF9925CCF9A9FCD5F469" box="[655,808,1582,1606]" pageId="14" pageNumber="53" value="2011-06-29">29 June 2011</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE336B3FFEBFF992784F9D5FC9FF49D" blockId="14.[151,1437,1438,2002]" pageId="14" pageNumber="53">
|
||
<emphasis id="B928EAA1FFEBFF992784F9D5FDAFF444" bold="true" box="[199,594,1618,1643]" pageId="14" pageNumber="53">Additional specimens examined.</emphasis>
|
||
<collectionCode id="ED4DAE76FFEBFF99251DF9D5FD47F445" box="[606,698,1618,1642]" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="14" pageNumber="53">LSUHC</collectionCode>
|
||
10076 was collected from Shah Alam, Selangor (
|
||
<geoCoordinate id="EE685074FFEBFF992253F9D5FA6AF445" box="[1296,1431,1618,1642]" direction="north" orientation="latitude" pageId="14" pageNumber="53" precision="1" value="3.0877">3°05.262’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE685074FFEBFF9927D4F9F1FEC4F4A1" box="[151,313,1654,1678]" direction="east" orientation="longitude" pageId="14" pageNumber="53" precision="1" value="101.52444">101°31.466”E</geoCoordinate>
|
||
;
|
||
<quantity id="4CA49B56FFEBFF992605F9F1FE82F4A1" box="[326,383,1654,1678]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="53" unit="m" value="15.0">15 m</quantity>
|
||
a.s.l.) by Daicus Belabut.
|
||
<collectionCode id="ED4DAE76FFEBFF9925EAF9F1FD22F4A1" box="[681,735,1654,1678]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="14" pageNumber="53">ZRC</collectionCode>
|
||
2.1127 was collected from the Sungai Chikus Forest Reserve, Perak (4°
|
||
<geoCoordinate id="EE685074FFEBFF9927BFF91DFE99F49D" box="[252,356,1690,1714]" direction="north" orientation="latitude" pageId="14" pageNumber="53" precision="55" value="9.426">09.426N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE685074FFEBFF99262CF91DFDF2F49D" box="[367,527,1690,1714]" direction="east" orientation="longitude" pageId="14" pageNumber="53" precision="1" value="101.00937">101°00.562’E</geoCoordinate>
|
||
;
|
||
<quantity id="4CA49B56FFEBFF992558F91CFDBBF49D" box="[539,582,1691,1714]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="53" unit="m" value="7.0">7 m</quantity>
|
||
a.s.l.) during
|
||
<date id="FFE21073FFEBFF9925A2F91DFCA3F49D" box="[737,862,1690,1714]" pageId="14" pageNumber="53" value="1925-04">April 1925</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3466538FFEBFF992784F939FD6EF5FD" pageId="14" pageNumber="53" type="diagnosis">
|
||
<paragraph id="8BE336B3FFEBFF992784F939FD6EF5FD" blockId="14.[151,1437,1438,2002]" pageId="14" pageNumber="53">
|
||
<emphasis id="B928EAA1FFEBFF992784F939FEBCF4F8" bold="true" box="[199,321,1726,1751]" pageId="14" pageNumber="53">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C5C4D30FFEBFF99260FF939FDA7F4F9" box="[332,602,1726,1750]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="53" phylum="Chordata" rank="species" species="payacola" status="sp. nov.">
|
||
<emphasis id="B928EAA1FFEBFF99260FF939FDA7F4F9" box="[332,602,1726,1750]" italics="true" pageId="14" pageNumber="53">Cyrtodactylus payacola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EAA1FFEBFF992527F938FD3FF4F8" bold="true" box="[612,706,1727,1751]" pageId="14" pageNumber="53">
|
||
<taxonomicNameLabel id="A21B57DAFFEBFF992527F938FD3FF4F8" box="[612,706,1727,1751]" pageId="14" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is distinguished from all other Sundaland species by having a maximum SVL of
|
||
<quantity id="4CA49B56FFEBFF99262FF965FE29F4D5" box="[364,468,1762,1786]" metricMagnitude="-2" metricUnit="m" metricValue="6.77" pageId="14" pageNumber="53" unit="mm" value="67.7">67.7 mm</quantity>
|
||
; moderately sized, conical, keeled body tubercles; tubercles occurring on the occiput, forelimbs, hind limbs, and beyond base of tail; 44–51 ventral scales; no transversely enlarged, median subcaudal scales; 18–21 subdigital lamellae on fourth toe; abrupt transition between postfemoral and ventral femoral scales; no femoral pores; 11 or 12 contiguous, pore-bearing precloacal scales; shallow, longitudinal, precloacal groove; a pair of posteromedially elongate, triangular to semilunar-shaped, paravertebral blotches on nape prominently outlined in light yellow; no wide, dark, ventrolateral stripes on flanks; no white reticulum on top of head; paired, semitransversely arranged, dark blotches on body.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF23663BFFEAFF9827D4FC9EFBFDF100" box="[151,1024,793,816]" httpUri="https://zenodo.org/record/281877/files/figure.png" pageId="15" pageNumber="54" targetBox="[163,1436,193,779]" targetPageId="15">
|
||
<paragraph id="8BE336B3FFEAFF9827D4FC9EFBFDF100" blockId="15.[151,1024,793,816]" box="[151,1024,793,816]" pageId="15" pageNumber="54">
|
||
<emphasis id="B928EAA1FFEAFF9827D4FC9EFEEFF101" bold="true" box="[151,274,793,816]" pageId="15" pageNumber="54">FIGURE 6.</emphasis>
|
||
Type series of
|
||
<taxonomicName id="4C5C4D30FFEAFF9826EEFC9EFD63F100" box="[429,670,793,815]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="54" phylum="Chordata" rank="species" species="payacola" status="sp. nov.">
|
||
<emphasis id="B928EAA1FFEAFF9826EEFC9EFD63F100" box="[429,670,793,815]" italics="true" pageId="15" pageNumber="54">Cyrtodactylus payacola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EAA1FFEAFF9825E7FC9DFD08F100" bold="true" box="[676,757,794,815]" pageId="15" pageNumber="54">
|
||
<taxonomicNameLabel id="A21B57DAFFEAFF9825E7FC9DFD08F100" box="[676,757,794,815]" pageId="15" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Holotype LSUHC 10074.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3466538FFEAFF862784FCDCFA84F383" lastPageId="17" lastPageNumber="56" pageId="15" pageNumber="54" type="description">
|
||
<paragraph id="8BE336B3FFEAFF982784FCDCFF1CF7BF" blockId="15.[151,1437,859,2036]" pageId="15" pageNumber="54">
|
||
<emphasis id="B928EAA1FFEAFF982784FCDCFE1BF15B" bold="true" box="[199,486,859,884]" pageId="15" pageNumber="54">
|
||
Description of
|
||
<typeStatus id="54E78811FFEAFF98263BFCDBFE1CF15B" box="[376,481,860,884]" pageId="15" pageNumber="54" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult male SVL
|
||
<quantity id="4CA49B56FFEAFF9825F3FCDBFCE5F15B" box="[688,792,860,884]" metricMagnitude="-2" metricUnit="m" metricValue="6.77" pageId="15" pageNumber="54" unit="mm" value="67.7">67.7 mm</quantity>
|
||
; head moderate in length (HL/SVL 0.26), wide (HW/HL 0.59), flat (HD/HL 0.38), distinct from neck, triangular in dorsal profile, lores weakly inflated, prefrontal region deeply concave, canthus rostralis smoothly rounded; snout elongate (ES/HL 0.40), rounded in dorsal profile; eye large (ED/HL 0.23); ear opening elliptical, obliquely oriented, moderate in size (EL/HL 0.06), eye to eye distance greater than diameter of eye; rostral square, punctuated with pores, partially divided dorsally by three postrostral granular scales (=internasals), bordered posteriorly by large left and right supranasals, laterally by first supralabials; external nares bordered anteriorly by rostral, dorsally by a large anterior supranasal and small posterior supranasal, posteriorly by two large postnasals, and ventrally by first supralabial; 10 (R, L) rectangular supralabials extending to just beyond dorsal inflection of labial margins, tapering abruptly below midpoint of eye; nine (R, L) infralabials tapering smoothly posteriorly to below orbit; scales of rostrum and lores raised, slightly larger size than granular scales on top of head and occiput; scales of occiput intermixed with slightly enlarged tubercles; supraorbitals smooth; dorsal superciliaries elongate, smooth; mental triangular, bordered laterally by first infralabials, posteriorly by left and right rectangular postmentals contacting medially for 40% of their length; one row of slightly enlarged, elongate sublabials extending posteriorly to fifth infralabial; gular scales small, granular, grading posteriorly into slightly larger, flatter, throat scales which grade into larger, flat, smooth, imbricate, pectoral and ventral scales.
|
||
</paragraph>
|
||
<paragraph id="8BE336B3FFEAFF982784FA1BFB19F446" blockId="15.[151,1437,859,2036]" pageId="15" pageNumber="54">Body relatively short (AG/SVL 0.44) with well-defined ventrolateral folds; dorsal scales small, granular, interspersed with moderately sized, conical, semi-regularly arranged, keeled tubercles being most dense on flanks; tubercles extend from occiput to anterior one-sixth of tail; tubercles on occiput and nape relatively small, those on body largest; approximately 21 longitudinal rows of tubercles at midbody, 37 paravertebral tubercles on body; 51 flat, imbricate, ventral scales, ventral scales larger than dorsal scales; precloacal scales not large; 11 contiguous, pore-bearing precloacal scales forming a “V” bordering a longitudinal, shallow precloacal groove.</paragraph>
|
||
<paragraph id="8BE336B3FFEAFF982784F9F3FB08F5DB" blockId="15.[151,1437,859,2036]" pageId="15" pageNumber="54">Forelimbs moderate in stature, relatively short (FL/SVL 0.13); granular scales of forearm slightly larger than those on body, interspersed with large, keeled tubercles; palmar scales rounded, flat; digits well-developed, inflected at basal, interphalangeal joints; subdigital lamellae transversely expanded proximal to joint inflection, more granular distal to inflection; digits slightly more narrow distal to inflections; claws well-developed, sheathed by a dorsal and ventral scale; hind limbs more robust than forelimbs, moderate in length (TBL/SVL 0.14), covered dorsally by granular scales interspersed with large, conical tubercles, covered anteriorly by flat, slightly larger scales; ventral scales of thigh flat, imbricate, larger than dorsals; ventral tibial scales flat; femoral scales imbricate; small postfemoral scales form an abrupt union with large ventral scales on posteroventral margin of thigh; plantar scales low, flat; digits well-developed, inflected at basal, interphalangeal joints; subdigital lamellae transversely expanded proximal to inflections, more granular distal to inflections, digits more narrow distal to inflections; 19R, 18L subdigital lamellae on fourth toe; claws well-developed, sheathed by a dorsal and ventral scale.</paragraph>
|
||
<caption id="DF23663BFFF5FF8727D4FAD4FF1FF7A7" httpUri="https://zenodo.org/record/281878/files/figure.png" pageId="16" pageNumber="55" targetBox="[172,1424,176,1340]" targetPageId="16">
|
||
<paragraph id="8BE336B3FFF5FF8727D4FAD4FF1FF7A7" blockId="16.[151,1436,1363,1416]" pageId="16" pageNumber="55">
|
||
<emphasis id="B928EAA1FFF5FF8727D4FAD4FEE8F746" bold="true" box="[151,277,1363,1386]" pageId="16" pageNumber="55">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C5C4D30FFF5FF87265EFAD3FDECF745" box="[285,529,1364,1386]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="55" phylum="Chordata" rank="species" species="payacola" status="sp. nov.">
|
||
<emphasis id="B928EAA1FFF5FF87265EFAD3FDECF745" box="[285,529,1364,1386]" italics="true" pageId="16" pageNumber="55">Cyrtodactylus payacola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EAA1FFF5FF87255AFAD3FD91F746" bold="true" box="[537,620,1364,1385]" pageId="16" pageNumber="55">
|
||
<taxonomicNameLabel id="A21B57DAFFF5FF87255AFAD3FD91F746" box="[537,620,1364,1385]" pageId="16" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
showing dark (upper; LSUHC 10074) and light (lower; LSUHC 10074) color phases.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE336B3FFF5FF872784FA37FE6EF453" blockId="16.[151,1436,1456,2020]" pageId="16" pageNumber="55">
|
||
Tail moderate,
|
||
<quantity id="4CA49B56FFF5FF872630FA37FE24F7E7" box="[371,473,1456,1480]" metricMagnitude="-2" metricUnit="m" metricValue="9.07" pageId="16" pageNumber="55" unit="mm" value="90.7">90.7 mm</quantity>
|
||
in length, tapering to a point,
|
||
<quantity id="4CA49B56FFF5FF872479FA37FC6FF7E7" box="[826,914,1456,1480]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="16" pageNumber="55" unit="mm" value="5.3">5.3 mm</quantity>
|
||
in width at base; dorsal scales of base of tail granular becoming flatter posteriorly; no median row of transversely enlarged, subcaudal scales; subcaudal scales larger than dorsal caudal scales; one pair of paravertebral and dorsolateral tubercle rows on either side of midline; paravertebral rows not widely separated; caudal tubercles decrease in size posteriorly, extending approximately one-sixth length of tail; one enlarged, postcloacal tubercle on left and right base of tail on hemipenial swelling; all postcloacal scales flat.
|
||
</paragraph>
|
||
<paragraph id="8BE336B3FFF5FF87278AF900FF48F5CB" blockId="16.[151,1436,1456,2020]" pageId="16" pageNumber="55">
|
||
<emphasis id="B928EAA1FFF5FF87278AF900FE5DF48F" bold="true" box="[201,416,1671,1696]" pageId="16" pageNumber="55">Coloration in life.</emphasis>
|
||
Dorsal: ground color of head, neck, body, limbs, and tail light straw-yellow; light brown mottling on top of head and rostrum; diffused, light brown, postorbital patch; paired, symmetrical, triangular to semilunar-shaped, dark blotches on entire portion of nape prominently outline in ground color; seven dark brown rectangular-shaped bands extending from below nape to base of tail; bands counter shaded with lighter color; wide, dark, elongate markings above shoulders; irregularly shaped, dark blotches on flanks; body darkly speckled overall; brown blotching on limbs, obscure banding on hind limbs; dark body bands extend onto tail to form brown bands alternating with cream-colored bands neither of which encircle tail; anterior caudal bands transform into black and white, respectively, on posterior two-third of tail. Ventral: surfaces of head, body and limbs lightly stippled in gray; subcaudal region darkened by fine mottling; iris reddish. Lizards lighten considerably at night (
|
||
<figureCitation id="13672A36FFF5FF87222CF82FFF5AF5CB" captionStart="FIGURE 7" captionStartId="16.[151,250,1363,1386]" captionTargetBox="[172,1424,176,1340]" captionTargetId="figure@16.[172,1424,163,1344]" captionTargetPageId="16" captionText="FIGURE 7. Cyrtodactylus payacola sp. nov. showing dark (upper; LSUHC 10074) and light (lower; LSUHC 10074) color phases." httpUri="https://zenodo.org/record/281878/files/figure.png" pageId="16" pageNumber="55">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE336B3FFF4FF86278AFF10FA84F383" blockId="17.[151,1436,151,1039]" pageId="17" pageNumber="56">
|
||
<emphasis id="B928EAA1FFF4FF86278AFF10FEC3F29F" bold="true" box="[201,318,151,176]" pageId="17" pageNumber="56">Variation.</emphasis>
|
||
The paratypotypes (
|
||
<figureCitation id="13672A36FFF4FF862564FF10FD96F280" box="[551,619,151,175]" captionStart="FIGURE 6" captionStartId="15.[151,250,793,816]" captionTargetBox="[163,1436,193,779]" captionTargetId="figure@15.[163,1448,193,779]" captionTargetPageId="15" captionText="FIGURE 6. Type series of Cyrtodactylus payacola sp. nov. Holotype LSUHC 10074." httpUri="https://zenodo.org/record/281877/files/figure.png" pageId="17" pageNumber="56">Fig. 6</figureCitation>
|
||
) and the additional specimens examined approximate the
|
||
<typeStatus id="54E78811FFF4FF8623B9FF10FAA1F280" box="[1274,1372,151,175]" pageId="17" pageNumber="56" type="holotype">holotype</typeStatus>
|
||
in all aspects of coloration. The caudal bands in LSUHC 9982 encircle only the posterior one-half of the tail. The additional specimens examined (LSUHC 10076 from Shah Alam, Selangor and ZRC 2.1127 from the Chikus Forest Reserve, Perak) share the species-level diagnostic characters of the
|
||
<typeStatus id="54E78811FFF4FF8624DFFE83FC03F333" box="[924,1022,260,284]" pageId="17" pageNumber="56" type="holotype">holotype</typeStatus>
|
||
and paratopotypes and approximate the
|
||
<typeStatus id="54E78811FFF4FF862781FEA0FED9F310" box="[194,292,295,319]" pageId="17" pageNumber="56" type="holotype">holotype</typeStatus>
|
||
in general aspects of coloration although the dark, dorsal markings form semi-transversely arranged bands as opposed to forming more paired markings. Furthermore, LSUHC 10076 has paired, non-symmetrical, semilunar-shaped, dark blotches only on the upper portion of nape as opposed to extending along the entire nape of the neck and the caudal bands encircle the tail along its entire length. Meristic differences are shown in
|
||
<tableCitation id="C6DE0308FFF4FF86225CFE13FA88F383" box="[1311,1397,404,428]" captionStart="TABLE 6" captionStartId="11.[151,235,875,896]" captionText="TABLE 6. Meristic data and diagnostic characters of Cyrtodactylus payacola sp. nov. Measurements are in mm." pageId="17" pageNumber="56">Table 6</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3466538FFF4FF86278AFE30FF1BF18B" pageId="17" pageNumber="56" type="distribution">
|
||
<paragraph id="8BE336B3FFF4FF86278AFE30FEF5F0AB" blockId="17.[151,1436,151,1039]" pageId="17" pageNumber="56">
|
||
<emphasis id="B928EAA1FFF4FF86278AFE30FE9FF3FF" bold="true" box="[201,354,439,464]" pageId="17" pageNumber="56">Distribution.</emphasis>
|
||
<taxonomicName id="4C5C4D30FFF4FF86262AFE3FFD89F3FF" box="[361,628,440,464]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="56" phylum="Chordata" rank="species" species="payacola" status="sp. nov.">
|
||
<emphasis id="B928EAA1FFF4FF86262AFE3FFD89F3FF" box="[361,628,440,464]" italics="true" pageId="17" pageNumber="56">Cyrtodactylus payacola</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21B57DAFFF4FF86253FFE3FFD2BF3E0" box="[636,726,440,464]" pageId="17" pageNumber="56" rank="species">
|
||
<emphasis id="B928EAA1FFF4FF86253FFE3FFD33F3FF" bold="true" box="[636,718,440,464]" pageId="17" pageNumber="56">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is known from the Bukit Panchor State Park, Penang; the Sungai Chikus Forest Reserve, Perak; and Shah Alam, Selangor, Peninsular
|
||
<collectingCountry id="F34B7623FFF4FF86248CFE5BFBC8F3DB" box="[975,1077,476,500]" name="Malaysia" pageId="17" pageNumber="56">Malaysia</collectingCountry>
|
||
(
|
||
<figureCitation id="13672A36FFF4FF862305FE5BFB75F3DB" box="[1094,1160,476,500]" captionStart="FIGURE 1" captionStartId="1.[151,249,1642,1665]" captionTargetBox="[210,1331,193,1567]" captionTargetId="figure@1.[210,1358,193,1597]" captionTargetPageId="1" captionText="FIGURE 1. Closed red circles are known localities of Cyrtodactylus payacola sp. nov. Closed white circles are localities from which specimens were sampled for the molecular and morphological analysis of Cyrtodactylus quadrivirgatus. Closed black circle (Shah Alam, Selangor) was not sampled for the molecular analysis and closed black squares are the remaining known localities for C. quadrivirgatus (fide Grismer 2011) from which no specimens were examined. Numbers correspond to the following localities: 1. Pulau Langkawi, Kedah. 2. Pulau Tuba, Kedah. 3. Ulu Muda, Kedah. 4. Gunung Jerai, Kedah. 5. Sungai Sedim, Kedah. 6. Pulau Pinang, Penang. 7. Bukit Larut, Perak. 8. Pulau Pangkor, Perak. 9. Temengor Forest Reserve, Perak. 10. Sungai Pelus, Perak. 11. Pulau Perhentian Besar, Terengganu. 12. Gunung Lawit, Terengganu. 13. Sungai Tembak, Terengganu. 14. Kuala Aring, Terengganu. 15. Kuala Tahan, Pahang. 16. Sungai Lembing, Pahang. 17. Gunung Benom, Pahang. 18. Fraser’s Hill, Pahang. 19. Genting Highlands, Pahang. 20. Ulu Kali, Pahang. 21. Ulu Gombak, Selangor. 22. Lakum Forest Reserve, Pahang. 23. Bukit Rengit, Pahang. 24. Bukit Bau, Pahang. 25. Pasoh, Negri Sembilan. 26. Gunung Ledang, Johor. 27. Pulau Tulai, Johor. 28. Pulau Tioman, Pahang. 29. Endau – Rompin, Johor. 30. Gunung Panti, Johor. 31. Singapore. 32. Shah Alam, Selangor. 33. Ulu Temiang Forest Reserve, Kelantan. 34. Serasa Forest Reserve, Kelantan. 35. Empangan Tembat, Terengganu. 36. Templer’s Park, Selangor. 37. Bukit Panchor, Penang. 38. Sungai Chikus Forest Reserve, Perak." httpUri="https://zenodo.org/record/281872/files/figure.png" pageId="17" pageNumber="56">Fig. 1</figureCitation>
|
||
) and most likely ranges throughout all lowland coastal areas west of the Banjaran Bintang and Titiwangsa to as far south perhaps as
|
||
<collectingCountry id="F34B7623FFF4FF8627D4FDA3FF13F013" box="[151,238,548,572]" name="Malaysia" pageId="17" pageNumber="56">Melaka</collectingCountry>
|
||
,
|
||
<collectingCountry id="F34B7623FFF4FF8627BBFDA3FEB1F013" box="[248,332,548,572]" name="Malaysia" pageId="17" pageNumber="56">Melaka</collectingCountry>
|
||
where its sister species
|
||
<taxonomicName id="4C5C4D30FFF4FF86251BFDA2FD0BF013" box="[600,758,549,572]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="56" phylum="Chordata" rank="species" species="pantienesis">
|
||
<emphasis id="B928EAA1FFF4FF86251BFDA2FD0BF013" box="[600,758,549,572]" italics="true" pageId="17" pageNumber="56">C. pantienesis</emphasis>
|
||
</taxonomicName>
|
||
has also been reported to occur (Grismer
|
||
<emphasis id="B928EAA1FFF4FF86238AFDA2FAFDF013" box="[1225,1280,548,572]" italics="true" pageId="17" pageNumber="56">et al.</emphasis>
|
||
2008). To the north, it may not range beyond the
|
||
<collectingCountry id="F34B7623FFF4FF862566FDC0FDA5F070" box="[549,600,583,607]" name="Thailand" pageId="17" pageNumber="56">Thai</collectingCountry>
|
||
–Malaysian border owing to the potential dispersal barrier of the Banjaran Nakawan.
|
||
</paragraph>
|
||
<paragraph id="8BE336B3FFF4FF86278AFD17FF1BF18B" blockId="17.[151,1436,151,1039]" pageId="17" pageNumber="56">
|
||
<emphasis id="B928EAA1FFF4FF86278AFD17FE71F087" bold="true" box="[201,396,655,680]" pageId="17" pageNumber="56">Natural history.</emphasis>
|
||
The three localities from which
|
||
<taxonomicName id="4C5C4D30FFF4FF86245BFD17FBD5F087" box="[792,1064,656,680]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="56" phylum="Chordata" rank="species" species="payacola" status="sp. nov.">
|
||
<emphasis id="B928EAA1FFF4FF86245BFD17FBD5F087" box="[792,1064,656,680]" italics="true" pageId="17" pageNumber="56">Cyrtodactylus payacola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EAA1FFF4FF862377FD17FB6FF087" bold="true" box="[1076,1170,656,680]" pageId="17" pageNumber="56">
|
||
<taxonomicNameLabel id="A21B57DAFFF4FF862377FD17FB6FF087" box="[1076,1170,656,680]" pageId="17" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known are all wet, swampy, lowland forests no higher than
|
||
<quantity id="4CA49B56FFF4FF862518FD33FD69F0E3" box="[603,660,692,716]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="56" unit="m" value="15.0">15 m</quantity>
|
||
in elevation (
|
||
<figureCitation id="13672A36FFF4FF86246EFD33FC8CF0E3" box="[813,881,692,716]" captionStart="FIGURE 8" captionStartId="17.[151,250,1980,2003]" captionTargetBox="[153,1436,1093,1944]" captionTargetId="figure@17.[153,1438,1093,1945]" captionTargetPageId="17" captionText="FIGURE 8. Swampy microhabitat of Cyrtodactylus payacola sp. nov. at Bukit Panchor, Penang." httpUri="https://zenodo.org/record/281879/files/figure.png" pageId="17" pageNumber="56">Fig. 8</figureCitation>
|
||
). The lizards from Bukit Panchor and Shah Alam were collected at night from the surface of leaves up to
|
||
<quantity id="4CA49B56FFF4FF862448FD50FCCBF0C0" box="[779,822,727,751]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="56" unit="m" value="2.0">2 m</quantity>
|
||
above the forest floor. During this period this species’ coloration lightens considerably (
|
||
<figureCitation id="13672A36FFF4FF86254CFD7BFDAFF13B" box="[527,594,764,788]" captionStart="FIGURE 7" captionStartId="16.[151,250,1363,1386]" captionTargetBox="[172,1424,176,1340]" captionTargetId="figure@16.[172,1424,163,1344]" captionTargetPageId="16" captionText="FIGURE 7. Cyrtodactylus payacola sp. nov. showing dark (upper; LSUHC 10074) and light (lower; LSUHC 10074) color phases." httpUri="https://zenodo.org/record/281878/files/figure.png" pageId="17" pageNumber="56">Fig. 7</figureCitation>
|
||
) making them easily confused with
|
||
<taxonomicName id="4C5C4D30FFF4FF8624A8FD7AFB4CF13B" box="[1003,1201,764,788]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="56" phylum="Chordata" rank="species" species="quadrivirgatus">
|
||
<emphasis id="B928EAA1FFF4FF8624A8FD7AFB4CF13B" box="[1003,1201,764,788]" italics="true" pageId="17" pageNumber="56">C. quadrivirgatus</emphasis>
|
||
</taxonomicName>
|
||
with which it is narrowly sympatric at Bukit Panchor and Shah Alam. At Bukit Panchor,
|
||
<taxonomicName id="4C5C4D30FFF4FF8624E3FCA7FB9AF117" box="[928,1127,800,824]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="56" phylum="Chordata" rank="species" species="quadrivirgatus">
|
||
<emphasis id="B928EAA1FFF4FF8624E3FCA7FB9AF117" box="[928,1127,800,824]" italics="true" pageId="17" pageNumber="56">C. quadrivirgatus</emphasis>
|
||
</taxonomicName>
|
||
appears to occur only up to the edge of the swampy areas, remaining in the more elevated drier regions (LSUHC 10072–73). LSUHC 10070, a gravid female carrying two eggs, was collected on
|
||
<date id="FFE21073FFF4FF862595FCE0FCD0F1AF" box="[726,813,871,896]" pageId="17" pageNumber="56" value="1929-06">29 June</date>
|
||
indicating that breeding takes place at least during this month.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3466538FFF4FF85278AFC37FABAF0AB" lastPageId="18" lastPageNumber="57" pageId="17" pageNumber="56" type="etymology">
|
||
<paragraph id="8BE336B3FFF4FF86278AFC37FEACF620" blockId="17.[151,1436,151,1039]" pageId="17" pageNumber="56">
|
||
<emphasis id="B928EAA1FFF4FF86278AFC37FEB2F1E7" bold="true" box="[201,335,944,968]" pageId="17" pageNumber="56">Etymology.</emphasis>
|
||
The specific epithet
|
||
<taxonomicName id="4C5C4D30FFF4FF862578FC37FD5FF1E7" box="[571,674,944,968]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="56" phylum="Chordata" rank="species" species="payacola">
|
||
<emphasis id="B928EAA1FFF4FF862578FC37FD5FF1E7" box="[571,674,944,968]" italics="true" pageId="17" pageNumber="56">payacola</emphasis>
|
||
</taxonomicName>
|
||
is derived from the word
|
||
<emphasis id="B928EAA1FFF4FF862485FC36FC00F1E7" box="[966,1021,945,968]" italics="true" pageId="17" pageNumber="56">paya</emphasis>
|
||
meaning “swamp” in the Malay language and the Latin suffix
|
||
<emphasis id="B928EAA1FFF4FF862682FC53FE0CF1C3" box="[449,497,980,1004]" italics="true" pageId="17" pageNumber="56">colo</emphasis>
|
||
, which means to inhabit or dwell in and is in reference to microhabitat in which this species is found.
|
||
</paragraph>
|
||
<caption id="DF23663BFFF4FF8627D4F83BFB92F5FD" box="[151,1135,1980,2003]" httpUri="https://zenodo.org/record/281879/files/figure.png" pageId="17" pageNumber="56" targetBox="[153,1436,1093,1944]" targetPageId="17">
|
||
<paragraph id="8BE336B3FFF4FF8627D4F83BFB92F5FD" blockId="17.[151,1135,1980,2003]" box="[151,1135,1980,2003]" pageId="17" pageNumber="56">
|
||
<emphasis id="B928EAA1FFF4FF8627D4F83BFEEFF5FD" bold="true" box="[151,274,1980,2003]" pageId="17" pageNumber="56">FIGURE 8.</emphasis>
|
||
Swampy microhabitat of
|
||
<taxonomicName id="4C5C4D30FFF4FF86255AF83AFCF7F5FC" box="[537,778,1981,2003]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="56" phylum="Chordata" rank="species" species="payacola" status="sp. nov.">
|
||
<emphasis id="B928EAA1FFF4FF86255AF83AFCF7F5FC" box="[537,778,1981,2003]" italics="true" pageId="17" pageNumber="56">Cyrtodactylus payacola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EAA1FFF4FF862452F83AFC9CF5FD" bold="true" box="[785,865,1981,2002]" pageId="17" pageNumber="56">
|
||
<taxonomicNameLabel id="A21B57DAFFF4FF862452F83AFC9CF5FD" box="[785,865,1981,2002]" pageId="17" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
at Bukit Panchor, Penang.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE336B3FFF7FF85278AFF10FC2BF3E0" blockId="18.[151,1437,151,644]" pageId="18" pageNumber="57">
|
||
<emphasis id="B928EAA1FFF7FF85278AFF10FE92F29F" bold="true" box="[201,367,151,176]" pageId="18" pageNumber="57">Comparisons.</emphasis>
|
||
<taxonomicName id="4C5C4D30FFF7FF852634FF1FFD78F29F" box="[375,645,152,176]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="57" phylum="Chordata" rank="species" species="payacola" status="sp. nov.">
|
||
<emphasis id="B928EAA1FFF7FF852634FF1FFD78F29F" box="[375,645,152,176]" italics="true" pageId="18" pageNumber="57">Cyrtodactylus payacola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EAA1FFF7FF8525CEFF1FFD14F29F" bold="true" box="[653,745,152,176]" pageId="18" pageNumber="57">
|
||
<taxonomicNameLabel id="A21B57DAFFF7FF8525CEFF1FFD14F29F" box="[653,745,152,176]" pageId="18" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is readily differentiated from all other Sundaland species of
|
||
<taxonomicName id="4C5C4D30FFF7FF8527D4FF3BFEC9F2FB" box="[151,308,188,212]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="57" phylum="Chordata" rank="genus">
|
||
<emphasis id="B928EAA1FFF7FF8527D4FF3BFEC9F2FB" box="[151,308,188,212]" italics="true" pageId="18" pageNumber="57">Cyrtodactylus</emphasis>
|
||
</taxonomicName>
|
||
on the basis of color pattern and morphology (see
|
||
<bibRefCitation id="EFCD4B42FFF7FF852437FF3BFB75F2FB" author="Chan" box="[884,1160,188,212]" pageId="18" pageNumber="57" refString="Chan, K. O. and Norhayati, A. (2010) A new insular species of Cyrtodactylus (Squamata: Gekkonidae) from northeastern Peninsular Malaysia, Malaysia. Zootaxa, 2389, 47 - 56." type="journal article" year="2010">Chan & Norhayati 2010</bibRefCitation>
|
||
:
|
||
<tableCitation id="C6DE0308FFF7FF8523D7FF3BFB1BF2FB" box="[1172,1254,188,212]" captionStart="TABLE 1. A" captionStartId="3.[151,239,771,795]" captionText="TABLE 1. A collective list of taxa used in the molecular analyses, including voucher numbers, locality information, and" pageId="18" pageNumber="57">Table 1</tableCitation>
|
||
). It is separated from
|
||
<emphasis id="B928EAA1FFF7FF852797FF67FE5FF2D7" box="[212,418,224,248]" italics="true" pageId="18" pageNumber="57">
|
||
<taxonomicName id="4C5C4D30FFF7FF852797FF67FE60F2D7" box="[212,413,224,248]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="57" phylum="Chordata" rank="species" species="quadrivirgatus">C. quadrivirgatus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
a species to which it was provisionally ascribed (
|
||
<bibRefCitation id="EFCD4B42FFF7FF85248DFF58FB93F2D8" author="Grismer" box="[974,1134,223,247]" pageId="18" pageNumber="57" refString="Grismer, L. L. (2011) Lizards of Peninsular Malaysia, Singapore and their Adjacent Archipelagos. Edition Chimaira, 728 pp." type="book" year="2011">Grismer 2011</bibRefCitation>
|
||
), based on having 11 or 12 contiguous, pore-bearing precloacal scales along a shallow, longitudinal precloacal groove as opposed to having 0–5 pore-bearing scales and no groove and a pair of posteromedially elongate, triangular to semilunar-shaped, paravertebral blotches on the nape that are prominently outlined in light yellow as opposed to lacking such markings. It differs from its sister species,
|
||
<emphasis id="B928EAA1FFF7FF8526BEFEF7FD6BF3A7" box="[509,662,368,392]" italics="true" pageId="18" pageNumber="57">
|
||
<taxonomicName id="4C5C4D30FFF7FF8526BEFEF7FD6FF3A7" box="[509,658,368,392]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="57" phylum="Chordata" rank="species" species="pantiensis">C. pantiensis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
in having more precloacal pores (11 or 12 versus eight or nine); having fewer subdigital lamellae on the fourth toes (18–21 versus 22 or 23); and having a generally immaculate ground color as opposed to the dense, dark brown speckling seen in
|
||
<taxonomicName id="4C5C4D30FFF7FF85247EFE3FFC32F3FF" box="[829,975,440,464]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="57" phylum="Chordata" rank="species" species="pantiensis">
|
||
<emphasis id="B928EAA1FFF7FF85247EFE3FFC32F3FF" box="[829,975,440,464]" italics="true" pageId="18" pageNumber="57">C. pantiensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE336B3FFF7FF85278AFE5BFABAF0AB" blockId="18.[151,1437,151,644]" pageId="18" pageNumber="57">
|
||
The Shah Alam specimen (LSUHC 10076) is more similar to
|
||
<taxonomicName id="4C5C4D30FFF7FF8524C6FE5AFBF3F3DB" box="[901,1038,476,500]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="57" phylum="Chordata" rank="species" species="payacola" status="sp. nov.">
|
||
<emphasis id="B928EAA1FFF7FF8524C6FE5AFBF3F3DB" box="[901,1038,476,500]" italics="true" pageId="18" pageNumber="57">C. payacola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EAA1FFF7FF852356FE5BFB94F3DB" bold="true" box="[1045,1129,476,500]" pageId="18" pageNumber="57">
|
||
<taxonomicNameLabel id="A21B57DAFFF7FF852356FE5BFB94F3DB" box="[1045,1129,476,500]" pageId="18" pageNumber="57" rank="species">sp. nov</taxonomicNameLabel>
|
||
</emphasis>
|
||
in all aspects of coloration and morphology than it is to any other species of
|
||
<taxonomicName id="4C5C4D30FFF7FF8525FDFD87FCA6F037" box="[702,859,512,536]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="57" phylum="Chordata" rank="genus">
|
||
<emphasis id="B928EAA1FFF7FF8525FDFD87FCA6F037" box="[702,859,512,536]" italics="true" pageId="18" pageNumber="57">Cyrtodactylus</emphasis>
|
||
</taxonomicName>
|
||
except for having paired dark, dorsal markings and paired, non-symmetrical, semilunar-shaped, dark blotches on only the upper portion of nape. Although it is a female, there are 12 large, non-pore-bearing scales homologous to those pore-bearing scales in males. Therefore, we tentatively consider this specimen
|
||
<taxonomicName id="4C5C4D30FFF7FF852501FDEAFD34F0AB" box="[578,713,620,644]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="57" phylum="Chordata" rank="species" species="payacola" status="sp. nov.">
|
||
<emphasis id="B928EAA1FFF7FF852501FDEAFD34F0AB" box="[578,713,620,644]" italics="true" pageId="18" pageNumber="57">C. payacola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EAA1FFF7FF852593FDEBFCD7F0AB" bold="true" box="[720,810,620,644]" pageId="18" pageNumber="57">
|
||
<taxonomicNameLabel id="A21B57DAFFF7FF852593FDEBFCD7F0AB" box="[720,810,620,644]" pageId="18" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and await the acquisition of additional material.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |