303 lines
35 KiB
XML
303 lines
35 KiB
XML
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<mods:title id="822317ECF67A764F9B4078974B0E1FFC">An annotated checklist of the South African Acanthodrilidae (Oligochaeta: Acanthodrilidae: Acanthodrilinae, Benhamiinae)</mods:title>
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<mods:namePart id="4FA9062A0A01FCCBCF4FB897151ECF6A">Plisko, Jadwiga Danuta</mods:namePart>
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<mods:title id="B42259A6768EFC976B19BBD482D63192">Zootaxa</mods:title>
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<mods:date id="FABD4E90E54E3A7F832B9BD6426DA3D9">2012</mods:date>
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<mods:number id="7BFE1A191694C4F48C8A015032650F71">3458</mods:number>
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<treatment id="03F54361FFB0FF86E68BD2B0BDAF6C66" ID-DOI="http://doi.org/10.5281/zenodo.6171101" ID-GBIF-Taxon="119524940" ID-Zenodo-Dep="6171101" LSID="urn:lsid:plazi:treatment:03F54361FFB0FF86E68BD2B0BDAF6C66" httpUri="http://treatment.plazi.org/id/03F54361FFB0FF86E68BD2B0BDAF6C66" lastPageId="9" lastPageNumber="12" pageId="8" pageNumber="12">
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<subSubSection id="C346A1FCFFB0FF87E68BD2B0BF5C6AC8" box="[151,549,1183,1209]" pageId="8" pageNumber="12" type="nomenclature">
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<paragraph id="8BE3F277FFB0FF87E68BD2B0BF5C6AC8" blockId="8.[151,549,1183,1209]" box="[151,549,1183,1209]" pageId="8" pageNumber="12">
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<heading id="D0AB451BFFB0FF87E68BD2B0BF5C6AC8" bold="true" box="[151,549,1183,1209]" fontSize="11" level="1" pageId="8" pageNumber="12" reason="1">
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<taxonomicName id="4C5C89F4FFB0FF87E68BD2B0BF5C6AC8" authority="Beddard, 1897" authorityName="Beddard" authorityYear="1897" box="[151,549,1183,1209]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="12" phylum="Annelida" rank="species" species="purcelli">
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<emphasis id="B9282E65FFB0FF87E68BD2B0BF5C6AC8" bold="true" box="[151,549,1183,1209]" pageId="8" pageNumber="12">
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<emphasis id="B9282E65FFB0FF87E68BD2B0BC2D6AC8" bold="true" box="[151,340,1183,1209]" italics="true" pageId="8" pageNumber="12">Chilota purcelli</emphasis>
|
||
(
|
||
<bibRefCitation id="EFCD8F86FFB0FF87E77FD2B0BF646AC8" author="Beddard" box="[355,541,1183,1209]" pageId="8" pageNumber="12" refString="Beddard, F. E. (1897) On a collection of earthworms from South Africa belonging to the genus Acanthodrilus. With an Appendix: on a new genus of earthworms belonging to the family Eudrilidae. Proceedings of the Zoological Society of London, 1897, 336 - 349." type="proceedings paper" year="1897">Beddard, 1897</bibRefCitation>
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||
)
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||
</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C346A1FCFFB0FF87E68BD2C8BF176BA2" pageId="8" pageNumber="12" type="reference_group">
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<paragraph id="8BE3F277FFB0FF87E68BD2C8BF366A8C" blockId="8.[151,1437,1254,1491]" box="[151,591,1254,1277]" pageId="8" pageNumber="12">
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<treatmentCitationGroup id="AB4CD559FFB0FF87E68BD2C8BF366A8C" box="[151,591,1254,1277]" pageId="8" pageNumber="12">
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<taxonomicName id="4C5C89F4FFB0FF87E68BD2C8BF306A8D" ID-CoL="2a207b47-077f-4a78-b370-db46bd13cdb9" authority="Beddard, 1897: 337" authorityName="Beddard" authorityPageNumber="337" authorityYear="1897" box="[151,585,1254,1277]" class="Clitellata" family="Acanthodrilidae" genus="Acanthodrilus" kingdom="Animalia" order="Crassiclitellata" pageId="8" pageNumber="12" phylum="Annelida" rank="species" species="purcelli">
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<emphasis id="B9282E65FFB0FF87E68BD2C8BC036A8C" box="[151,378,1255,1277]" italics="true" pageId="8" pageNumber="12">Acanthodrilus purcelli</emphasis>
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<treatmentCitation id="0AFDD466FFB0FF87E79DD2C9BF306A8D" author="Beddard" box="[385,585,1254,1276]" page="337" pageId="8" pageNumber="12" year="1897">
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<bibRefCitation id="EFCD8F86FFB0FF87E79DD2C9BF606A8D" author="Beddard" box="[385,537,1254,1276]" pageId="8" pageNumber="12" refString="Beddard, F. E. (1897) On a collection of earthworms from South Africa belonging to the genus Acanthodrilus. With an Appendix: on a new genus of earthworms belonging to the family Eudrilidae. Proceedings of the Zoological Society of London, 1897, 336 - 349." type="proceedings paper" year="1897">Beddard, 1897</bibRefCitation>
|
||
: 337
|
||
</treatmentCitation>
|
||
</taxonomicName>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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<paragraph id="8BE3F277FFB0FF87E68BD32ABE286B6A" blockId="8.[151,1437,1254,1491]" box="[151,849,1285,1307]" pageId="8" pageNumber="12">
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||
<treatmentCitationGroup id="AB4CD559FFB0FF87E68BD32ABE286B6A" box="[151,849,1285,1307]" pageId="8" pageNumber="12">
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<taxonomicName id="4C5C89F4FFB0FF87E68BD32ABFD76B6A" ID-CoL="2a207b47-077f-4a78-b370-db46bd13cdb9" authority="Reynolds & Cook 1976: 160" authorityName="Reynolds & Cook" authorityPageNumber="160" authorityYear="1976" box="[151,686,1285,1307]" class="Clitellata" family="Acanthodrilidae" genus="Acanthodrilus" kingdom="Animalia" order="Crassiclitellata" pageId="8" pageNumber="12" phylum="Annelida" rank="species" species="purcelli">
|
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<emphasis id="B9282E65FFB0FF87E68BD32ABC036B6A" box="[151,378,1285,1307]" italics="true" pageId="8" pageNumber="12">Acanthodrilus purcelli</emphasis>
|
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:
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<treatmentCitation id="0AFDD466FFB0FF87E794D32ABFD76B6A" author="Reynolds" box="[392,686,1285,1307]" page="160" pageId="8" pageNumber="12" year="1976">
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<bibRefCitation id="EFCD8F86FFB0FF87E794D32ABF026B6A" author="Reynolds" box="[392,635,1285,1307]" pageId="8" pageNumber="12" refString="Reynolds J. W. & Cook, D. G. (1976) Nomenclatura Oligochaetologica. A catalogue of names, descriptions and type specimens of the Oligochaeta. Ottawa: Runge Press. 217 pp." type="book" year="1976">Reynolds & Cook 1976</bibRefCitation>
|
||
: 160
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||
</treatmentCitation>
|
||
</taxonomicName>
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||
; BMNH; ZMH.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BE3F277FFB0FF87E68BD30BBF5A6B48" blockId="8.[151,1437,1254,1491]" box="[151,547,1315,1338]" pageId="8" pageNumber="12">
|
||
<treatmentCitationGroup id="AB4CD559FFB0FF87E68BD30BBF5A6B48" box="[151,547,1315,1338]" pageId="8" pageNumber="12">
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<taxonomicName id="4C5C89F4FFB0FF87E68BD30BBF666B48" authority="Michaelsen 1913a: 517" authorityName="Michaelsen" authorityPageNumber="517" authorityYear="1913" box="[151,543,1315,1338]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="12" phylum="Annelida" rank="species" species="priesti">
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<emphasis id="B9282E65FFB0FF87E68BD30BBC536B4B" box="[151,298,1316,1338]" italics="true" pageId="8" pageNumber="12">Chilota priesti</emphasis>
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<treatmentCitation id="0AFDD466FFB0FF87E72CD30CBF666B48" author="Michaelsen" box="[304,543,1315,1337]" page="517" pageId="8" pageNumber="12" year="1913">
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<bibRefCitation id="EFCD8F86FFB0FF87E72CD30CBC946B48" author="Michaelsen" box="[304,493,1315,1337]" pageId="8" pageNumber="12" refString="Michaelsen, W. (1913 a) Die Oligochaeten des Kaplandes. Zoologische Jahrbucher, Abteilung fur Systematik, 34, 473 - 556." type="journal article" year="1913" yearSuffix="a">Michaelsen 1913a</bibRefCitation>
|
||
: 517
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||
</treatmentCitation>
|
||
</taxonomicName>
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||
.
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||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BE3F277FFB0FF87E68BD36DBFFB6B29" blockId="8.[151,1437,1254,1491]" box="[151,642,1346,1368]" pageId="8" pageNumber="12">
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<treatmentCitationGroup id="AB4CD559FFB0FF87E68BD36DBFFB6B29" box="[151,642,1346,1368]" pageId="8" pageNumber="12">
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<taxonomicName id="4C5C89F4FFB0FF87E68BD36DBC536B29" box="[151,298,1346,1368]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="12" phylum="Annelida" rank="species" species="priesti">
|
||
<emphasis id="B9282E65FFB0FF87E68BD36DBC536B29" box="[151,298,1346,1368]" italics="true" pageId="8" pageNumber="12">Chilota priesti</emphasis>
|
||
</taxonomicName>
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||
f.
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<emphasis id="B9282E65FFB0FF87E759D36CBCF46B29" box="[325,397,1347,1368]" italics="true" pageId="8" pageNumber="12">typicus</emphasis>
|
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<treatmentCitation id="0AFDD466FFB0FF87E78FD36DBF046B29" author="Michaelsen" box="[403,637,1346,1368]" page="54" pageId="8" pageNumber="12" year="1913">
|
||
<bibRefCitation id="EFCD8F86FFB0FF87E78FD36DBF216B29" author="Michaelsen" box="[403,600,1346,1368]" pageId="8" pageNumber="12" refString="Michaelsen, W. (1913 b) Report upon the Oligochaeta in the South African Museum at Cape Town. Annals of the South African Museum, 13, 43 - 62." type="journal article" year="1913" yearSuffix="b">Michaelsen, 1913b</bibRefCitation>
|
||
: 54
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BE3F277FFB0FF87E68BD34EBFFB6B06" blockId="8.[151,1437,1254,1491]" box="[151,642,1377,1399]" pageId="8" pageNumber="12">
|
||
<treatmentCitationGroup id="AB4CD559FFB0FF87E68BD34EBFFB6B06" box="[151,642,1377,1399]" pageId="8" pageNumber="12">
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||
<taxonomicName id="4C5C89F4FFB0FF87E68BD34EBC536B06" box="[151,298,1377,1399]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="12" phylum="Annelida" rank="species" species="priesti">
|
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<emphasis id="B9282E65FFB0FF87E68BD34EBC536B06" box="[151,298,1377,1399]" italics="true" pageId="8" pageNumber="12">Chilota priesti</emphasis>
|
||
</taxonomicName>
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||
f.
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||
<emphasis id="B9282E65FFB0FF87E759D34DBCF86B06" box="[325,385,1378,1399]" italics="true" pageId="8" pageNumber="12">minor</emphasis>
|
||
<treatmentCitation id="0AFDD466FFB0FF87E794D34EBF046B06" author="Michaelsen" box="[392,637,1377,1399]" page="518" pageId="8" pageNumber="12" year="1913">
|
||
<bibRefCitation id="EFCD8F86FFB0FF87E794D34EBF326B06" author="Michaelsen" box="[392,587,1377,1399]" pageId="8" pageNumber="12" refString="Michaelsen, W. (1913 a) Die Oligochaeten des Kaplandes. Zoologische Jahrbucher, Abteilung fur Systematik, 34, 473 - 556." type="journal article" year="1913" yearSuffix="a">Michaelsen, 1913a</bibRefCitation>
|
||
: 518
|
||
</treatmentCitation>
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||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BE3F277FFB0FF87E68BD3AFBF0F6BE7" blockId="8.[151,1437,1254,1491]" box="[151,630,1408,1430]" pageId="8" pageNumber="12">
|
||
<treatmentCitationGroup id="AB4CD559FFB0FF87E68BD3AFBF0F6BE7" box="[151,630,1408,1430]" pageId="8" pageNumber="12">
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<taxonomicName id="4C5C89F4FFB0FF87E68BD3AFBC536BE7" box="[151,298,1408,1430]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="12" phylum="Annelida" rank="species" species="priesti">
|
||
<emphasis id="B9282E65FFB0FF87E68BD3AFBC536BE7" box="[151,298,1408,1430]" italics="true" pageId="8" pageNumber="12">Chilota priesti</emphasis>
|
||
</taxonomicName>
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||
f.
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||
<emphasis id="B9282E65FFB0FF87E759D3AEBCF86BE7" box="[325,385,1409,1430]" italics="true" pageId="8" pageNumber="12">minor</emphasis>
|
||
<treatmentCitation id="0AFDD466FFB0FF87E794D3AFBF0B6BE7" author="Michaelsen" box="[392,626,1408,1430]" page="55" pageId="8" pageNumber="12" year="1913">
|
||
<bibRefCitation id="EFCD8F86FFB0FF87E794D3AFBF346BE7" author="Michaelsen" box="[392,589,1408,1430]" pageId="8" pageNumber="12" refString="Michaelsen, W. (1913 b) Report upon the Oligochaeta in the South African Museum at Cape Town. Annals of the South African Museum, 13, 43 - 62." type="journal article" year="1913" yearSuffix="b">Michaelsen, 1913b</bibRefCitation>
|
||
: 55
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BE3F277FFB0FF87E68BD3B0BF176BA2" blockId="8.[151,1437,1254,1491]" pageId="8" pageNumber="12">
|
||
<treatmentCitationGroup id="AB4CD559FFB0FF87E68BD3B0BF176BA2" pageId="8" pageNumber="12">
|
||
<taxonomicName id="4C5C89F4FFB0FF87E68BD3B0BF4C6BC5" authority="Michaelsen 1900: 147" authorityName="Michaelsen" authorityPageNumber="147" authorityYear="1900" box="[151,565,1438,1461]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="12" phylum="Annelida" rank="species" species="purcelli">
|
||
<emphasis id="B9282E65FFB0FF87E68BD3B0BC436BC4" box="[151,314,1439,1461]" italics="true" pageId="8" pageNumber="12">Chilota purcelli</emphasis>
|
||
:
|
||
<treatmentCitation id="0AFDD466FFB0FF87E757D3B1BF4C6BC5" author="Michaelsen" box="[331,565,1438,1460]" page="147" pageId="8" pageNumber="12" year="1900">
|
||
<bibRefCitation id="EFCD8F86FFB0FF87E757D3B1BF796BC5" author="Michaelsen" box="[331,512,1438,1460]" pageId="8" pageNumber="12" refString="Michaelsen, W. (1900) Oligochaeta. In: Das Tierreich. Lief. 10. Berlin: R. Friedlander & Sohn, 575 pp." type="book" year="1900">Michaelsen 1900</bibRefCitation>
|
||
: 147
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
,
|
||
<treatmentCitation id="0AFDD466FFB0FF87E45FD3B1BE416BC5" author="Michaelsen" box="[579,824,1438,1460]" page="521" pageId="8" pageNumber="12" year="1913">
|
||
<bibRefCitation id="EFCD8F86FFB0FF87E45FD3B1BE7A6BC5" author="Michaelsen" box="[579,771,1438,1460]" pageId="8" pageNumber="12" refString="Michaelsen, W. (1913 a) Die Oligochaeten des Kaplandes. Zoologische Jahrbucher, Abteilung fur Systematik, 34, 473 - 556." type="journal article" year="1913" yearSuffix="a">Michaelsen 1913a</bibRefCitation>
|
||
: 521
|
||
</treatmentCitation>
|
||
, 1913b: 55;
|
||
<treatmentCitation id="0AFDD466FFB0FF87E5DCD3B1B98D6BC5" author="Reynolds" box="[960,1268,1438,1460]" page="160" pageId="8" pageNumber="12" year="1976">
|
||
<bibRefCitation id="EFCD8F86FFB0FF87E5DCD3B1B9C46BC5" author="Reynolds" box="[960,1213,1438,1460]" pageId="8" pageNumber="12" refString="Reynolds J. W. & Cook, D. G. (1976) Nomenclatura Oligochaetologica. A catalogue of names, descriptions and type specimens of the Oligochaeta. Ottawa: Runge Press. 217 pp." type="book" year="1976">Reynolds & Cook 1976</bibRefCitation>
|
||
: 160
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AFDD466FFB0FF87E31DD3B1BD976BA2" author="Pickford" page="453" pageId="8" pageNumber="12" year="1937">
|
||
<bibRefCitation id="EFCD8F86FFB0FF87E31DD3B1B8E16BC5" author="Pickford" box="[1281,1432,1438,1460]" pageId="8" pageNumber="12" refString="Pickford, G. E. (1937) A monograph of the Acanthodriline earthworms of South Africa. Cambridge: W. Heffer & Sons Ltd. 612 pp." type="book" year="1937">Pickford 1937</bibRefCitation>
|
||
: 453
|
||
</treatmentCitation>
|
||
–475;
|
||
<treatmentCitation id="0AFDD466FFB0FF87E731D392BF686BA2" author="Sciacchitano" box="[301,529,1469,1491]" page="10" pageId="8" pageNumber="12" year="1960">
|
||
<bibRefCitation id="EFCD8F86FFB0FF87E731D392BC926BA2" author="Sciacchitano" box="[301,491,1469,1491]" pageId="8" pageNumber="12" refString="Sciacchitano, I. (1960) Oligochaeta part. In: Hanstrom, B., Brinck, P. & Rudebeck, G. (Eds), South African animal life: results of the Lund University Expedition in 1950 - 1951. Uppsala, Almqvist & Wiksells, 7, 9 - 12." type="journal article" year="1960">Sciacchitano 1960</bibRefCitation>
|
||
: 10
|
||
</treatmentCitation>
|
||
; BMNH.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346A1FCFFB0FF87E68BD02FBDA6682E" pageId="8" pageNumber="12" type="materials_examined">
|
||
<paragraph id="8BE3F277FFB0FF87E68BD02FBDA6682E" blockId="8.[151,1437,1535,2028]" pageId="8" pageNumber="12">
|
||
<emphasis id="B9282E65FFB0FF87E68BD02FBC576869" bold="true" box="[151,302,1535,1560]" pageId="8" pageNumber="12">
|
||
<typeStatus id="54E74CD5FFB0FF87E68BD02FBDA96869" box="[151,208,1536,1560]" pageId="8" pageNumber="12">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B9282E65FFB0FF87E721D3D0BC146869" bold="true" box="[317,365,1535,1560]" pageId="8" pageNumber="12">WC</emphasis>
|
||
, Cape Peninsula: ‘under rotten log in the woods on the Newlands slope…’ In the sample were ‘ten specimens but only one of these ten was sexually mature; this individual was the largest of the series’ (
|
||
<bibRefCitation id="EFCD8F86FFB0FF87E320D00BBDAA682E" author="Beddard" pageId="8" pageNumber="12" refString="Beddard, F. E. (1897) On a collection of earthworms from South Africa belonging to the genus Acanthodrilus. With an Appendix: on a new genus of earthworms belonging to the family Eudrilidae. Proceedings of the Zoological Society of London, 1897, 336 - 349." type="proceedings paper" year="1897">Beddard 1897</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346A1FCFFB0FF87E6DBD044BF9B68BD" pageId="8" pageNumber="12" type="distribution">
|
||
<paragraph id="8BE3F277FFB0FF87E6DBD044BF9B68BD" blockId="8.[151,1437,1535,2028]" pageId="8" pageNumber="12">
|
||
<emphasis id="B9282E65FFB0FF87E6DBD044BC2068F5" bold="true" box="[199,345,1643,1668]" pageId="8" pageNumber="12">Distribution</emphasis>
|
||
.
|
||
<taxonomicName id="4C5C89F4FFB0FF87E771D043BFA468F5" authority="Beddard, 1897" authorityName="Beddard" authorityYear="1897" box="[365,733,1644,1668]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="12" phylum="Annelida" rank="species" species="purcelli">
|
||
<emphasis id="B9282E65FFB0FF87E771D043BF5C68F5" box="[365,549,1644,1668]" italics="true" pageId="8" pageNumber="12">Chilota purcelli</emphasis>
|
||
<bibRefCitation id="EFCD8F86FFB0FF87E42ED043BFA468F5" author="Beddard" box="[562,733,1644,1668]" pageId="8" pageNumber="12" refString="Beddard, F. E. (1897) On a collection of earthworms from South Africa belonging to the genus Acanthodrilus. With an Appendix: on a new genus of earthworms belonging to the family Eudrilidae. Proceedings of the Zoological Society of London, 1897, 336 - 349." type="proceedings paper" year="1897">Beddard, 1897</bibRefCitation>
|
||
</taxonomicName>
|
||
with its five subspecies (see below) is the most common megadrile in the Cape Peninsula and its neighbourhood. It is recorded also from distant localities in the coastal-belt area bordering Western and Eastern Cape provinces.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346A1FCFFB0FF86E6DBD0F7BDAF6C66" lastPageId="9" lastPageNumber="13" pageId="8" pageNumber="12" type="discussion">
|
||
<paragraph id="8BE3F277FFB0FF86E6DBD0F7BDAF6C66" blockId="8.[151,1437,1535,2028]" lastBlockId="9.[151,1436,151,535]" lastPageId="9" lastPageNumber="13" pageId="8" pageNumber="12">
|
||
<emphasis id="B9282E65FFB0FF87E6DBD0F7BC4D6881" bold="true" box="[199,308,1752,1776]" pageId="8" pageNumber="12">Remarks</emphasis>
|
||
. One adult specimen BMNH 1904.10.5.888 labelled as ‘type’ is probably the
|
||
<typeStatus id="54E74CD5FFB0FF87E2BDD0F8B87F689E" box="[1185,1286,1751,1775]" pageId="8" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
, although the inserted label is disintegrating, and it is difficult to establish the collection locality. BMNH 1904.10.5.905–907 with three immature specimens collected at ‘Cape of Good Hope’, labelled
|
||
<emphasis id="B9282E65FFB0FF87E5C1D10FB9996949" box="[989,1248,1824,1848]" italics="true" pageId="8" pageNumber="12">
|
||
<taxonomicName id="4C5C89F4FFB0FF87E5C1D10FB9A26949" box="[989,1243,1824,1848]" class="Clitellata" family="Acanthodrilidae" genus="Acanthodrilus" kingdom="Animalia" order="Crassiclitellata" pageId="8" pageNumber="12" phylum="Annelida" rank="species" species="purcelli">Acanthodrilus purcelli</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
redetermined by Pickford as
|
||
<taxonomicName id="4C5C89F4FFB0FF87E739D16BBCA3692D" box="[293,474,1860,1884]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="12" phylum="Annelida" rank="species" species="purcelli">
|
||
<emphasis id="B9282E65FFB0FF87E739D16BBCA3692D" box="[293,474,1860,1884]" italics="true" pageId="8" pageNumber="12">Chilota purcelli</emphasis>
|
||
</taxonomicName>
|
||
, is probably also from Beddard’s
|
||
<typeStatus id="54E74CD5FFB0FF87E573D16ABEE6692C" box="[879,927,1861,1885]" pageId="8" pageNumber="12">type</typeStatus>
|
||
series material. Reynolds & Cook's (1976) indication of ‘ZMUH 7832’ being the
|
||
<typeStatus id="54E74CD5FFB0FF87E445D146BFF069F0" box="[601,649,1897,1921]" pageId="8" pageNumber="12">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="4C5C89F4FFB0FF87E506D147BEB769F1" box="[794,974,1896,1920]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="12" phylum="Annelida" rank="species" species="purcelli">
|
||
<emphasis id="B9282E65FFB0FF87E506D147BEB769F1" box="[794,974,1896,1920]" italics="true" pageId="8" pageNumber="12">Chilota purcelli</emphasis>
|
||
</taxonomicName>
|
||
is confirmed here. The presently found ZMH V-7832 specimen of
|
||
<taxonomicName id="4C5C89F4FFB0FF87E7DFD1A3BE1569D5" authority="Beddard, 1897" authorityName="Beddard" authorityYear="1897" box="[451,876,1932,1956]" class="Clitellata" family="Acanthodrilidae" genus="Acanthodrilus" kingdom="Animalia" order="Crassiclitellata" pageId="8" pageNumber="12" phylum="Annelida" rank="species" species="purcelli">
|
||
<emphasis id="B9282E65FFB0FF87E7DFD1A3BFC769D5" box="[451,702,1932,1956]" italics="true" pageId="8" pageNumber="12">Acanthodrilus purcelli</emphasis>
|
||
<bibRefCitation id="EFCD8F86FFB0FF87E4D8D1A3BE1569D5" author="Beddard" box="[708,876,1932,1956]" pageId="8" pageNumber="12" refString="Beddard, F. E. (1897) On a collection of earthworms from South Africa belonging to the genus Acanthodrilus. With an Appendix: on a new genus of earthworms belonging to the family Eudrilidae. Proceedings of the Zoological Society of London, 1897, 336 - 349." type="proceedings paper" year="1897">Beddard, 1897</bibRefCitation>
|
||
</taxonomicName>
|
||
, collected at ‘Südwestliches Kapland, Table Mtn’, is the specimen of the
|
||
<typeStatus id="54E74CD5FFB0FF87E78CD19EBCB969B8" box="[400,448,1969,1993]" pageId="8" pageNumber="12">type</typeStatus>
|
||
series identified originally by Beddard.
|
||
<bibRefCitation id="EFCD8F86FFB0FF87E59ED19FB94A69B9" author="Pickford" box="[898,1075,1968,1992]" pageId="8" pageNumber="12" refString="Pickford, G. E. (1937) A monograph of the Acanthodriline earthworms of South Africa. Cambridge: W. Heffer & Sons Ltd. 612 pp." type="book" year="1937">Pickford (1937)</bibRefCitation>
|
||
studied Beddard’s material kept at the BMNH and further samples, altogether more than one hundred, collected in the Western Cape and the southeastern part of the Eastern Cape, at various time, from various biotopes, by numerous collectors. The comprehensive results were monographed (
|
||
<bibRefCitation id="EFCD8F86FFB1FF86E491D693BE4D6EA5" author="Pickford" box="[653,820,188,212]" pageId="9" pageNumber="13" refString="Pickford, G. E. (1937) A monograph of the Acanthodriline earthworms of South Africa. Cambridge: W. Heffer & Sons Ltd. 612 pp." type="book" year="1937">Pickford 1937</bibRefCitation>
|
||
) in detailed descriptions of the species' external and internal characters (
|
||
<emphasis id="B9282E65FFB1FF86E767D6CFBCD76E89" box="[379,430,224,248]" italics="true" pageId="9" pageNumber="13">ibid.</emphasis>
|
||
p. 453–475), the ‘geographical distribution and sub-speciation in
|
||
<taxonomicName id="4C5C89F4FFB1FF86E2B7D6CFB8266E89" box="[1195,1375,224,248]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="13" phylum="Annelida" rank="species" species="purcelli">
|
||
<emphasis id="B9282E65FFB1FF86E2B7D6CFB8266E89" box="[1195,1375,224,248]" italics="true" pageId="9" pageNumber="13">Chilota purcelli</emphasis>
|
||
</taxonomicName>
|
||
’, the ‘key to the sub-species of
|
||
<taxonomicName id="4C5C89F4FFB1FF86E7D9D72BBF016F6D" box="[453,632,260,284]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="13" phylum="Annelida" rank="species" species="purcelli">
|
||
<emphasis id="B9282E65FFB1FF86E7D9D72BBF016F6D" box="[453,632,260,284]" italics="true" pageId="9" pageNumber="13">Chilota purcelli</emphasis>
|
||
</taxonomicName>
|
||
’, including also differentiations of traits (p. 476–477). Pages 477–500 (
|
||
<bibRefCitation id="EFCD8F86FFB1FF86E682D708BC326F4E" author="Pickford" box="[158,331,295,319]" pageId="9" pageNumber="13" refString="Pickford, G. E. (1937) A monograph of the Acanthodriline earthworms of South Africa. Cambridge: W. Heffer & Sons Ltd. 612 pp." type="book" year="1937">Pickford 1937</bibRefCitation>
|
||
) contain extensive descriptions of the five subspecies, recognizing also ‘series’, ‘forms’, ‘variations’, and ‘transitions’. This exhaustive study based on remarkably large material is followed by a detailed revision of
|
||
<taxonomicName id="4C5C89F4FFB1FF86E701D75FBFC66FF6" authority="Michaelsen, 1905" authorityName="Michaelsen" authorityYear="1905" box="[285,703,367,392]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="13" phylum="Annelida" rank="species" species="montanus">
|
||
<emphasis id="B9282E65FFB1FF86E701D75FBC906FF9" box="[285,489,368,392]" italics="true" pageId="9" pageNumber="13">Chilota montanus</emphasis>
|
||
<bibRefCitation id="EFCD8F86FFB1FF86E7E8D740BFC66FF6" author="Michaelsen" box="[500,703,367,391]" pageId="9" pageNumber="13" refString="Michaelsen, W. (1905) Die Oligochaeten der deutschen Sudpolar-Expedition 1901 - 1903. Deutsche Sudpolar-Expedition 1901 - 1903, 9, Zoologie I, 1 - 58." type="journal article" year="1905">Michaelsen, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
(put already in synonymy with
|
||
<taxonomicName id="4C5C89F4FFB1FF86E226D75FB80C6FF6" authority="Beddard, 1897" authorityName="Beddard" authorityYear="1897" box="[1082,1397,367,392]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="13" phylum="Annelida" rank="species" species="purcelli">
|
||
<emphasis id="B9282E65FFB1FF86E226D75FB9BB6FF9" box="[1082,1218,368,392]" italics="true" pageId="9" pageNumber="13">
|
||
<collectingCountry id="F34BB2E7FFB1FF86E226D75FB9276FF9" box="[1082,1118,368,392]" name="Switzerland" pageId="9" pageNumber="13">Ch</collectingCountry>
|
||
. purcelli
|
||
</emphasis>
|
||
<bibRefCitation id="EFCD8F86FFB1FF86E2D1D740B80C6FF6" author="Beddard" box="[1229,1397,367,391]" pageId="9" pageNumber="13" refString="Beddard, F. E. (1897) On a collection of earthworms from South Africa belonging to the genus Acanthodrilus. With an Appendix: on a new genus of earthworms belonging to the family Eudrilidae. Proceedings of the Zoological Society of London, 1897, 336 - 349." type="proceedings paper" year="1897">Beddard, 1897</bibRefCitation>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFCD8F86FFB1FF86E68BD7BBBC046FDD" author="Michaelsen" box="[151,381,404,428]" pageId="9" pageNumber="13" refString="Michaelsen, W. (1913 a) Die Oligochaeten des Kaplandes. Zoologische Jahrbucher, Abteilung fur Systematik, 34, 473 - 556." type="journal article" year="1913" yearSuffix="a">Michaelsen (1913a)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C5C89F4FFB1FF86E793D7BBBE6C6FDD" authority="Michaelsen 1913" authorityName="Michaelsen" authorityYear="1913" box="[399,789,404,428]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="13" phylum="Annelida" rank="species" species="priesti">
|
||
<emphasis id="B9282E65FFB1FF86E793D7BBBF4F6FDD" box="[399,566,404,428]" italics="true" pageId="9" pageNumber="13">Chilota priesti</emphasis>
|
||
<bibRefCitation id="EFCD8F86FFB1FF86E45FD7BBBE6C6FDD" author="Michaelsen" box="[579,789,404,428]" pageId="9" pageNumber="13" refString="Michaelsen, W. (1913 a) Die Oligochaeten des Kaplandes. Zoologische Jahrbucher, Abteilung fur Systematik, 34, 473 - 556." type="journal article" year="1913" yearSuffix="a">Michaelsen 1913a</bibRefCitation>
|
||
</taxonomicName>
|
||
and its forms
|
||
<emphasis id="B9282E65FFB1FF86E5D7D7BAB9696FDD" box="[971,1040,405,428]" italics="true" pageId="9" pageNumber="13">typica</emphasis>
|
||
and
|
||
<emphasis id="B9282E65FFB1FF86E24DD7BAB9EB6FDD" box="[1105,1170,405,428]" italics="true" pageId="9" pageNumber="13">minor</emphasis>
|
||
.
|
||
<collectingCountry id="F34BB2E7FFB1FF86E2B9D7BBB9BD6FDD" box="[1189,1220,404,428]" name="American Samoa" pageId="9" pageNumber="13">As</collectingCountry>
|
||
a result, Pickford recognizes five subspecies:
|
||
<taxonomicName id="4C5C89F4FFB1FF86E7C3D797BEFD6FBE" authority="Beddard, 1897" authorityName="Beddard" authorityYear="1897" box="[479,900,439,464]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="13" phylum="Annelida" rank="subSpecies" species="purcelli" subSpecies="purcelli">
|
||
<emphasis id="B9282E65FFB1FF86E7C3D797BFB26FA1" box="[479,715,440,464]" italics="true" pageId="9" pageNumber="13">
|
||
<collectingCountry id="F34BB2E7FFB1FF86E7C3D797BF7A6FA1" box="[479,515,440,464]" name="Switzerland" pageId="9" pageNumber="13">Ch</collectingCountry>
|
||
. purcelli purcelli
|
||
</emphasis>
|
||
<bibRefCitation id="EFCD8F86FFB1FF86E4C4D798BEFD6FBE" author="Beddard" box="[728,900,439,463]" pageId="9" pageNumber="13" refString="Beddard, F. E. (1897) On a collection of earthworms from South Africa belonging to the genus Acanthodrilus. With an Appendix: on a new genus of earthworms belonging to the family Eudrilidae. Proceedings of the Zoological Society of London, 1897, 336 - 349." type="proceedings paper" year="1897">Beddard, 1897</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C89F4FFB1FF86E589D797B81D6FBE" authority="Pickford, 1937" authorityName="Pickford" authorityYear="1937" box="[917,1380,439,464]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="13" phylum="Annelida" rank="subSpecies" species="purcelli" subSpecies="longinquus">
|
||
<emphasis id="B9282E65FFB1FF86E589D797B9D06FA1" box="[917,1193,440,464]" italics="true" pageId="9" pageNumber="13">
|
||
<collectingCountry id="F34BB2E7FFB1FF86E589D797BEC06FA1" box="[917,953,440,464]" name="Switzerland" pageId="9" pageNumber="13">Ch</collectingCountry>
|
||
. purcelli longinquus
|
||
</emphasis>
|
||
<bibRefCitation id="EFCD8F86FFB1FF86E2A9D798B81D6FBE" author="Pickford" box="[1205,1380,439,463]" pageId="9" pageNumber="13" refString="Pickford, G. E. (1937) A monograph of the Acanthodriline earthworms of South Africa. Cambridge: W. Heffer & Sons Ltd. 612 pp." type="book" year="1937">Pickford, 1937</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C89F4FFB1FF86E368D797BF746F85" authority="Pickford, 1937" authorityName="Pickford" authorityYear="1937" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="13" phylum="Annelida" rank="subSpecies" species="purcelli" subSpecies="spinosus">
|
||
<emphasis id="B9282E65FFB1FF86E368D797BC2E6F85" italics="true" pageId="9" pageNumber="13">
|
||
<collectingCountry id="F34BB2E7FFB1FF86E368D797B8E16FA1" box="[1396,1432,440,464]" name="Switzerland" pageId="9" pageNumber="13">Ch</collectingCountry>
|
||
. purcelli spinosus
|
||
</emphasis>
|
||
<bibRefCitation id="EFCD8F86FFB1FF86E77DD7F3BF746F85" author="Pickford" box="[353,525,476,500]" pageId="9" pageNumber="13" refString="Pickford, G. E. (1937) A monograph of the Acanthodriline earthworms of South Africa. Cambridge: W. Heffer & Sons Ltd. 612 pp." type="book" year="1937">Pickford, 1937</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5C89F4FFB1FF86E407D7F3BE886F85" authority="Michaelsen, 1905" authorityName="Michaelsen" authorityYear="1905" box="[539,1009,476,500]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="13" phylum="Annelida" rank="subSpecies" species="purcelli" subSpecies="montanus">
|
||
<emphasis id="B9282E65FFB1FF86E407D7F3BF456F85" box="[539,572,476,500]" italics="true" pageId="9" pageNumber="13">
|
||
<collectingCountry id="F34BB2E7FFB1FF86E407D7F3BF456F85" box="[539,572,476,500]" name="Switzerland" pageId="9" pageNumber="13">Ch</collectingCountry>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B9282E65FFB1FF86E451D7F3BE636F85" box="[589,794,476,500]" italics="true" pageId="9" pageNumber="13">purcelli montanus</emphasis>
|
||
<bibRefCitation id="EFCD8F86FFB1FF86E538D7F3BE886F85" author="Michaelsen" box="[804,1009,476,500]" pageId="9" pageNumber="13" refString="Michaelsen, W. (1905) Die Oligochaeten der deutschen Sudpolar-Expedition 1901 - 1903. Deutsche Sudpolar-Expedition 1901 - 1903, 9, Zoologie I, 1 - 58." type="journal article" year="1905">Michaelsen, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C5C89F4FFB1FF86E22DD7F3BDAB6C66" authority="Michaelsen, 1913" authorityName="Michaelsen" authorityYear="1913" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="13" phylum="Annelida" rank="subSpecies" species="purcelli" subSpecies="priesti">
|
||
<emphasis id="B9282E65FFB1FF86E22DD7F3B8736F85" box="[1073,1290,476,500]" italics="true" pageId="9" pageNumber="13">
|
||
<collectingCountry id="F34BB2E7FFB1FF86E22DD7F3B92C6F85" box="[1073,1109,476,500]" name="Switzerland" pageId="9" pageNumber="13">Ch</collectingCountry>
|
||
. purcelli priesti
|
||
</emphasis>
|
||
Michaelsen, 1913
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |