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<document id="51F70E5072AD193593685623106FB9DD" ID-CLB-Dataset="293777" ID-DOI="10.35463/j.apr.2023.02.01" ID-GBIF-Dataset="fe1ffee7-1fea-43d4-aa15-3302495459da" ID-ISSN="1842-371x" ID-Zenodo-Dep="10520925" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1705452171074" checkinUser="felipe" docAuthor="SCHLAGINTWEIT, FELIX &amp; YAZDI-MOGHADAM, MOHSEN" docDate="2023" docId="AA52B01C0807FFB3D5B83DAA08F09A7B" docLanguage="en" docName="ActaPalaeontolRom.19.2.3-13.pdf" docOrigin="Acta Palaeontologica Romaniae 19 (2)" docSource="http://dx.doi.org/10.35463/j.apr.2023.02.01" docStyle="DocumentStyle:CA474AAD92E97C533D63E0C5C0F159F1.1:ActaPalaeontolRom.2019-.journal_article" docStyleId="CA474AAD92E97C533D63E0C5C0F159F1" docStyleName="ActaPalaeontolRom.2019-.journal_article" docStyleVersion="1" docTitle="Pseudocyclammina sarvakensis SCHLAGINTWEIT &amp; YAZDI-MOGHADAM 2023, sp. nov." docType="treatment" docVersion="5" lastPageNumber="7" masterDocId="566BC8640803FFB7D52F3E6E0B7B9C03" masterDocTitle="Pseudocyclammina Sarvakensis Sp. Nov. And Pseudotextulariella Brevicamerata Sp. Nov.: Further Evidence For The Cenomanian Megadiversity Of Larger Benthic Foraminifera From The Sarvak Formation Of Sw Iran" masterLastPageNumber="13" masterPageNumber="3" pageNumber="7" updateTime="1713187070002" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="CFF7153E775E2BF760125AB7C8C8980F">Pseudocyclammina Sarvakensis Sp. Nov. And Pseudotextulariella Brevicamerata Sp. Nov.: Further Evidence For The Cenomanian Megadiversity Of Larger Benthic Foraminifera From The Sarvak Formation Of Sw Iran</mods:title>
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<mods:name id="102B6B90733784C810393F352E20526C" type="personal">
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<mods:namePart id="A9DC7C8E9D1779C30BF936380DC119AF">SCHLAGINTWEIT, FELIX</mods:namePart>
<mods:affiliation id="44B58453C3FDDBACF4E7494C3C5209EB">Lerchenauerstr. 167, 80935 Munich, Germany</mods:affiliation>
<mods:nameIdentifier id="39E0C6A4EAA00596FCC19D23F98A3091" type="email">felix.schlagintweit@gmx.de</mods:nameIdentifier>
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<mods:roleTerm id="E909AB36F7F2CB9F9CB26F467B0F030D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9C7E3ABE96F4E1E5CFBB2D367E0A6EFD">YAZDI-MOGHADAM, MOHSEN</mods:namePart>
<mods:affiliation id="876B9F9B284EFEA17A236D2C45C033D7">National Iranian Oil Company Exploration Directorate, Sheikh Bahayi square, Seoul street, 1994814695, Tehran, Iran &amp; School of Earth Sciences, Damghan University, Damghan 3671641167, Iran.</mods:affiliation>
</mods:name>
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<mods:title id="9E1973B1A257E5BF02B084285B4A4C7C">Acta Palaeontologica Romaniae</mods:title>
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<mods:date id="BA0E5B41C7A12021F9581F82733497D5">2023</mods:date>
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<mods:number id="F224CFCD6C95B7F01A63CD28DA24B19B">2023-02-03</mods:number>
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<treatment id="AA52B01C0807FFB3D5B83DAA08F09A7B" ID-DOI="http://doi.org/10.5281/zenodo.10974252" ID-Zenodo-Dep="10974252" LSID="urn:lsid:plazi:treatment:AA52B01C0807FFB3D5B83DAA08F09A7B" httpUri="http://treatment.plazi.org/id/AA52B01C0807FFB3D5B83DAA08F09A7B" lastPageNumber="7" pageId="4" pageNumber="7">
<subSubSection id="6AE152810807FFB3D5B83DAA09359FDF" box="[151,590,964,988]" pageId="4" pageNumber="7" type="nomenclature">
<paragraph id="2244010A0807FFB3D5B83DAA09359FDF" blockId="4.[151,771,732,1021]" box="[151,590,964,988]" pageId="4" pageNumber="7">
<heading id="790CB6660807FFB3D5B83DAA09359FDF" box="[151,590,964,988]" fontSize="10" level="2" pageId="4" pageNumber="7" reason="5">
<taxonomicName id="E5FB7A890807FFB3D5B83DAA0A899FDF" authority="SCHLAGINTWEIT &amp; YAZDI-MOGHADAM, 2023" authorityName="SCHLAGINTWEIT &amp; YAZDI-MOGHADAM" authorityYear="2023" box="[151,498,964,988]" class="Globothalamea" family="Cyclamminidae" genus="Pseudocyclammina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="sarvakensis" status="sp. nov.">
<emphasis id="108FDD180807FFB3D5B83DAA0A899FDF" bold="true" box="[151,498,964,988]" italics="true" pageId="4" pageNumber="7">Pseudocyclammina sarvakensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BBC60630807FFB3D4D63DAB09359FDF" box="[505,590,965,988]" pageId="4" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6AE152810807FFB3D5B83D8B0B8C9FFE" box="[151,247,997,1021]" pageId="4" pageNumber="7" type="description">
<paragraph id="2244010A0807FFB3D5B83D8B0B8C9FFE" blockId="4.[151,771,732,1021]" box="[151,247,997,1021]" pageId="4" pageNumber="7">
<figureCitation id="BAC01D8F0807FFB3D5B83D8B0BAE9FFE" box="[151,213,997,1021]" captionStart="Fig" captionStartId="5.[206,241,1726,1747]" captionTargetBox="[205,1379,165,1684]" captionTargetId="figure-33@5.[205,1379,165,1684]" captionTargetPageId="5" captionText="Fig. 4 Upper Cretaceous species of Pseudocyclammina Yabe &amp; Hanzawa, 1926; note the different scales. a-g Pseudocyclammina sarvakensis sp. nov., middle Cenomanian Sarvak Formation of SW Iran, Tang-e Darbast section, thin-section RAP 181 (b, d-f), and Shahneshin section, thin-section AKHS 97 (c, g). a Equatorial section, microspheric specimen? (from Sampò,1969 illustrated as Cyclammina sp. (?) in pl. XLV, fig. 2). b Equatorial section, holotype. c, e Oblique sections. d Subaxial section. f Oblique tangential section. g Oblique equatorial section, megalospheric specimen? h Pseudocyclammina massiliensis Maync, early-middle Campanian of Brač Island, Croatia. i Pseudocylammina rugosa (Orbigny), equatorial section, Cenomanian of W France (from Maync, 1952, pl. 12, fig. 10). j-m Pseudocyclammina sphaeroidea Gendrot, Upper Cretaceous (?Coniacian) of Austria (from Schlagintweit, 1992, pl. 1, figs. 13-16). Subaxial section (j), equatorial section of a specimen with uncoiling final stage (k), equatorial section (l), equatorial section slightly oblique (m)." figureDoi="http://doi.org/10.5281/zenodo.10520933" httpUri="https://zenodo.org/record/10520933/files/figure.png" pageId="4" pageNumber="7">Fig. 4</figureCitation>
a-g
</paragraph>
</subSubSection>
<subSubSection id="6AE152810807FFB3D5B83A4F0BBB98BE" pageId="4" pageNumber="7" type="reference_group">
<paragraph id="2244010A0807FFB3D5B83A4F09C1983A" blockId="4.[151,771,1057,1213]" box="[151,698,1057,1081]" pageId="4" pageNumber="7">
1969
<taxonomicName id="E5FB7A890807FFB3D5FB3A4F0A04983B" box="[212,383,1057,1081]" class="Globothalamea" family="Cyclamminidae" genus="Cyclammina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="undetermined">
<emphasis id="108FDD180807FFB3D5FB3A4F0A21983A" box="[212,346,1057,1081]" italics="true" pageId="4" pageNumber="7">Cyclammina</emphasis>
sp.
</taxonomicName>
(?) – Sampò, pl. XLV, fig. 2.
</paragraph>
<paragraph id="2244010A0807FFB3D5B83A2C0BD09878" blockId="4.[151,771,1057,1213]" pageId="4" pageNumber="7">
2018
<taxonomicName id="E5FB7A890807FFB3D5FB3A2C0A8F9859" baseAuthorityName="d'Orbigny" baseAuthorityYear="1850" box="[212,500,1090,1114]" class="Globothalamea" family="Cyclamminidae" genus="Pseudocyclammina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="rugosa">
<emphasis id="108FDD180807FFB3D5FB3A2C0A8F9859" box="[212,500,1090,1114]" italics="true" pageId="4" pageNumber="7">Pseudocyclammina rugosa</emphasis>
</taxonomicName>
– Omidi et al., pl. 2, fig. 5.
</paragraph>
<paragraph id="2244010A0807FFB3D5B83AEA0BBB98BE" blockId="4.[151,771,1057,1213]" pageId="4" pageNumber="7">
?2022
<taxonomicName id="E5FB7A890807FFB3D5CE3AEA097B989F" baseAuthorityName="d'Orbigny" baseAuthorityYear="1850" box="[225,512,1156,1180]" class="Globothalamea" family="Cyclamminidae" genus="Pseudocyclammina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="rugose">
<emphasis id="108FDD180807FFB3D5CE3AEA097B989F" box="[225,512,1156,1180]" italics="true" pageId="4" pageNumber="7">Pseudocyclammina rugose</emphasis>
</taxonomicName>
– Esfandyari et al., fig.
<specimenCount id="34FDCA830807FFB3D5B83ACB0BBB98BE" box="[151,192,1189,1213]" count="23" pageId="4" pageNumber="7" type="generic">23l.</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="6AE152810807FFB3D5B83A8F0BA59938" pageId="4" pageNumber="7" type="etymology">
<paragraph id="2244010A0807FFB3D5B83A8F0BA59938" blockId="4.[151,772,1249,2067]" pageId="4" pageNumber="7">
<emphasis id="108FDD180807FFB3D5B83A8F0A1498FA" bold="true" box="[151,367,1249,1273]" pageId="4" pageNumber="7">Derivatio nominis:</emphasis>
The species name refers to the Sarvak Formation cropping out in southwestern
<collectingCountry id="5AEC419A0807FFB3D7523B6C09D3991A" box="[637,680,1282,1305]" name="Iran" pageId="4" pageNumber="7">Iran</collectingCountry>
(Zagros Zone).
</paragraph>
</subSubSection>
<subSubSection id="6AE152810807FFB3D5B83B2A0AA399BC" pageId="4" pageNumber="7" type="materials_examined">
<paragraph id="2244010A0807FFB3D5B83B2A0AD7997E" blockId="4.[151,772,1249,2067]" pageId="4" pageNumber="7">
<materialsCitation id="92930B570807FFB3D5B83B2A0AD7997E" collectionCode="RAP" pageId="4" pageNumber="7" specimenCode="RAP 181" specimenCount="1" typeStatus="holotype">
<emphasis id="108FDD180807FFB3D5B83B2A0A73995F" bold="true" box="[151,264,1348,1372]" pageId="4" pageNumber="7">
<typeStatus id="FD40BFA80807FFB3D5B83B2A0A78995F" box="[151,259,1348,1372]" pageId="4" pageNumber="7" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="34FDCA830807FFB3D43C3B2A0A07995F" box="[275,380,1348,1372]" pageId="4" pageNumber="7" type="generic">Specimen</specimenCount>
illustrated in
<figureCitation id="BAC01D8F0807FFB3D7333B2A09F2995F" box="[540,649,1348,1372]" captionStart="Fig" captionStartId="5.[206,241,1726,1747]" captionTargetBox="[205,1379,165,1684]" captionTargetId="figure-33@5.[205,1379,165,1684]" captionTargetPageId="5" captionText="Fig. 4 Upper Cretaceous species of Pseudocyclammina Yabe &amp; Hanzawa, 1926; note the different scales. a-g Pseudocyclammina sarvakensis sp. nov., middle Cenomanian Sarvak Formation of SW Iran, Tang-e Darbast section, thin-section RAP 181 (b, d-f), and Shahneshin section, thin-section AKHS 97 (c, g). a Equatorial section, microspheric specimen? (from Sampò,1969 illustrated as Cyclammina sp. (?) in pl. XLV, fig. 2). b Equatorial section, holotype. c, e Oblique sections. d Subaxial section. f Oblique tangential section. g Oblique equatorial section, megalospheric specimen? h Pseudocyclammina massiliensis Maync, early-middle Campanian of Brač Island, Croatia. i Pseudocylammina rugosa (Orbigny), equatorial section, Cenomanian of W France (from Maync, 1952, pl. 12, fig. 10). j-m Pseudocyclammina sphaeroidea Gendrot, Upper Cretaceous (?Coniacian) of Austria (from Schlagintweit, 1992, pl. 1, figs. 13-16). Subaxial section (j), equatorial section of a specimen with uncoiling final stage (k), equatorial section (l), equatorial section slightly oblique (m)." figureDoi="http://doi.org/10.5281/zenodo.10520933" httpUri="https://zenodo.org/record/10520933/files/figure.png" pageId="4" pageNumber="7">Figure 4b</figureCitation>
, equatorial section, sample
<specimenCode id="725DA9710807FFB3D46D3B0B0ADF997E" box="[322,420,1381,1405]" pageId="4" pageNumber="7">RAP 181</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="2244010A0807FFB3D5B83BE90AA399BC" blockId="4.[151,772,1249,2067]" pageId="4" pageNumber="7">
<emphasis id="108FDD180807FFB3D5B83BE90A4D999C" bold="true" box="[151,310,1415,1439]" pageId="4" pageNumber="7">Type-locality:</emphasis>
<materialsCitation id="92930B570807FFB3D4683BE90AA899BC" location="Tang-e Darbast" pageId="4" pageNumber="7" specimenCount="1" stateProvince="Fars">
<location id="272457D10807FFB3D4683BE90A8C999D" LSID="urn:lsid:plazi:treatment:AA52B01C0807FFB3D5B83DAA08F09A7B:272457D10807FFB3D4683BE90A8C999D" box="[327,503,1414,1439]" name="Tang-e Darbast" pageId="4" pageNumber="7" stateProvince="Fars">Tang-e Darbast</location>
section, Interior
<collectingRegion id="E03FCFE80807FFB3D7E13BE90985999D" box="[718,766,1415,1438]" country="Iran" name="Fars" pageId="4" pageNumber="7">Fars</collectingRegion>
,
<location id="272457D10807FFB3D5B83BC60A2199BC" LSID="urn:lsid:plazi:treatment:AA52B01C0807FFB3D5B83DAA08F09A7B:272457D10807FFB3D5B83BC60A2199BC" box="[151,346,1447,1472]" name="Zagros Mountains" pageId="4" pageNumber="7" stateProvince="Fars">Zagros Mountains</location>
(
<figureCitation id="BAC01D8F0807FFB3D4473BC90AB399BC" box="[360,456,1447,1471]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="2.[192,227,1890,1911]" captionStartId-1="3.[212,247,1816,1837]" captionTargetBox-0="[192,1393,166,1848]" captionTargetBox-1="[213,1374,168,1772]" captionTargetId-0="figure-30@2.[192,1396,166,1848]" captionTargetId-1="figure-33@3.[212,1375,159,1778]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Fig. 2 a Simplified geological map of Iran (after Schlagintweit &amp; Yazdi-Moghadam, 2021) showing the main tectonic subdivisions. b Road map of the Tang-e Darbast section (type-locality of Iraqia ultima Yazdi-Moghadam &amp; Schlagintweit, 2022 and Simplorabanitina simplex Consorti, Schlagintweit, and Yazdi-Moghadam, 2022). c Road map of the Shahneshin section. d Position of Arman well-1. e Tectono-stratigraphic units of the Zagros belt (modified after Yazdi-Moghadam &amp; Schlagintweit, 2021). The position of the studied sections is marked by an asterisk. Abbreviations: BF Balarud Fault, CEIM Central East Iran Microplate, HZF High Zagros Fault, KZF Kazerun Fault, MFF Mountain Front Fault, MZRF Main Zagros Revers Fault, MZT Main Zagros Thrust, SSZ Sanandaj-Sirjan Zone, UDMA Uromia Dokhtar Magmatic Arc, ZFTB Zagros Fold" captionText-1="Fig. 3 Lithostratigraphic log of the Tang-e Darbast section, the type locality of Pseudocyclammina sarvakensis sp. nov and Pseudotextulariella brevicamerata sp. nov., showing the distribution of some (larger) benthic foraminifera. a Neoiraqia insolita (Decrouez &amp; Moullade), b Orbitolinopsis cenomaniensis Schlagintweit &amp; Yazdi-Moghadam, c Conicorbitolina cf. conica (d’Archiac), d Mesorbitolina aperta (Erman), e Orbitolina ex gr. concava (Lamarck), f Rotorbinella mesogeensis (Tronchetti), g Nezzazata gr. gyra-conica (Smout), h Decastroia cf. serrakieli Vicedo &amp; Piuz, i Myriastyla omanensis Piuz, Meister &amp; Vicedo, j Chrysalidina gradata Orbigny, k Iraqia ultima Yazdi- Moghadam &amp; Schlagintweit." figureDoi-0="http://doi.org/10.5281/zenodo.10520929" figureDoi-1="http://doi.org/10.5281/zenodo.10520931" httpUri-0="https://zenodo.org/record/10520929/files/figure.png" httpUri-1="https://zenodo.org/record/10520931/files/figure.png" pageId="4" pageNumber="7">Figs. 2-3</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6AE152810807FFB3D5B83BA60FBB9D28" pageId="4" pageNumber="7" type="description">
<paragraph id="2244010A0807FFB3D5B83BA60AFD9A02" blockId="4.[151,772,1249,2067]" pageId="4" pageNumber="7">
<emphasis id="108FDD180807FFB3D5B83BA60A4199E3" bold="true" box="[151,314,1480,1504]" pageId="4" pageNumber="7">Type-horizon:</emphasis>
Lower part of the middle Cenomanian of the Sarvak Formation.
</paragraph>
<paragraph id="2244010A0807FFB3D5B838650FBB9D28" blockId="4.[151,772,1249,2067]" lastBlockId="4.[816,1437,176,1656]" pageId="4" pageNumber="7">
<emphasis id="108FDD180807FFB3D5B838650A5F9A20" bold="true" box="[151,292,1547,1571]" pageId="4" pageNumber="7">Description:</emphasis>
Medium-sized (max. diameter
<quantity id="E503ACEF0807FFB3D7B9386409879A21" box="[662,764,1546,1570]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="4" pageNumber="7" unit="mm" value="1.35">1.35 mm</quantity>
) species of
<taxonomicName id="E5FB7A890807FFB3D42138420AA99A47" box="[270,466,1580,1604]" genus="Pseudocylammina" higherTaxonomySource="GBIF" kingdom="Chromista" pageId="4" pageNumber="7" phylum="Foraminifera" rank="genus">
<emphasis id="108FDD180807FFB3D42138420AA99A47" box="[270,466,1580,1604]" italics="true" pageId="4" pageNumber="7">Pseudocylammina</emphasis>
</taxonomicName>
with a planispiral, involute test displaying a rounded periphery. The spiral stage is well developed, and a tendency to uncoil in the final stage has not been observed. In subaxial sections, the test is slightly compressed (nautiloid morphology). Test surface as seen in equatorial section is smooth with much reduced to almost no sutural depressions. The chambers following the subspherical proloculus are arranged in 2.5 to 3.5 whorls displaying 14 to 16 chambers in the final whorl. The dimensions for the four measured proloculi diameters (
<quantity id="E503ACEF0807FFB3D5B139190A749B8D" box="[158,271,1911,1935]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="7" unit="mm" value="0.11">0.110 mm</quantity>
,
<quantity id="E503ACEF0807FFB3D43539190AF29B8D" box="[282,393,1911,1935]" metricMagnitude="-4" metricUnit="m" metricValue="1.25" pageId="4" pageNumber="7" unit="mm" value="0.125">0.125 mm</quantity>
,
<quantity id="E503ACEF0807FFB3D4BA3919097A9B8D" box="[405,513,1911,1935]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="7" unit="mm" value="0.14">0.140 mm</quantity>
and 0.300 mm) suggest a test dimorphism that needs further clarification. The early postembryonic chambers are subrhombohedric later with increasing width become recurved. The chamber lumen is markedly higher as the thickness of the septa and also of the wall. Multiple and tiny foramina (diameter 20-35 µm), with parallel arrangement in axial sections and septal spaces about twice the width of the former (
<figureCitation id="BAC01D8F0807FFB3D6173F7D08F89D28" box="[824,899,275,299]" captionStart="Fig" captionStartId="5.[206,241,1726,1747]" captionTargetBox="[205,1379,165,1684]" captionTargetId="figure-33@5.[205,1379,165,1684]" captionTargetPageId="5" captionText="Fig. 4 Upper Cretaceous species of Pseudocyclammina Yabe &amp; Hanzawa, 1926; note the different scales. a-g Pseudocyclammina sarvakensis sp. nov., middle Cenomanian Sarvak Formation of SW Iran, Tang-e Darbast section, thin-section RAP 181 (b, d-f), and Shahneshin section, thin-section AKHS 97 (c, g). a Equatorial section, microspheric specimen? (from Sampò,1969 illustrated as Cyclammina sp. (?) in pl. XLV, fig. 2). b Equatorial section, holotype. c, e Oblique sections. d Subaxial section. f Oblique tangential section. g Oblique equatorial section, megalospheric specimen? h Pseudocyclammina massiliensis Maync, early-middle Campanian of Brač Island, Croatia. i Pseudocylammina rugosa (Orbigny), equatorial section, Cenomanian of W France (from Maync, 1952, pl. 12, fig. 10). j-m Pseudocyclammina sphaeroidea Gendrot, Upper Cretaceous (?Coniacian) of Austria (from Schlagintweit, 1992, pl. 1, figs. 13-16). Subaxial section (j), equatorial section of a specimen with uncoiling final stage (k), equatorial section (l), equatorial section slightly oblique (m)." figureDoi="http://doi.org/10.5281/zenodo.10520933" httpUri="https://zenodo.org/record/10520933/files/figure.png" pageId="4" pageNumber="7">Fig. 4f</figureCitation>
). Wall agglutinated, alveolar.
</paragraph>
</subSubSection>
<subSubSection id="6AE152810807FFB3D61F3F5B08F09A7B" pageId="4" pageNumber="7" type="diagnosis">
<paragraph id="2244010A0807FFB3D61F3F5B08F09A7B" blockId="4.[816,1437,176,1656]" pageId="4" pageNumber="7">
<emphasis id="108FDD180807FFB3D61F3F5B08AA9D4E" bold="true" box="[816,977,309,333]" pageId="4" pageNumber="7">Comparisons:</emphasis>
As can be deduced from the synonymy, the new species has been confounded with
<taxonomicName id="E5FB7A890807FFB3D0013F380FB89D8C" authority="(d'Orbigny, 1850)" baseAuthorityName="d'Orbigny" baseAuthorityYear="1850" class="Globothalamea" family="Cyclamminidae" genus="Pseudocyclammina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="rugosa">
<emphasis id="108FDD180807FFB3D0013F3808959D8C" italics="true" pageId="4" pageNumber="7">Pseudocyclammina rugosa</emphasis>
(d’Orbigny, 1850)
</taxonomicName>
. These assignments however must have been made on the basis simplicity as
<taxonomicName id="E5FB7A890807FFB3D61F3FD408E79DD2" baseAuthorityName="d'Orbigny" baseAuthorityYear="1850" box="[816,924,442,465]" class="Globothalamea" family="Cyclamminidae" genus="Pseudocyclammina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="rugosa">
<emphasis id="108FDD180807FFB3D61F3FD408E79DD2" box="[816,924,442,465]" italics="true" pageId="4" pageNumber="7">P. rugosa</emphasis>
</taxonomicName>
represented the only species of the genus described (until now) from Cenomanian strata. However, with respect to the traditional specific criteria,
<taxonomicName id="E5FB7A890807FFB3D0623F9208F19E37" authority="SCHLAGINTWEIT &amp; YAZDI-MOGHADAM, 2023" authorityName="SCHLAGINTWEIT &amp; YAZDI-MOGHADAM" authorityYear="2023" class="Globothalamea" family="Cyclamminidae" genus="Pseudocyclammina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="sarvakensis" status="nov. sp.">
<emphasis id="108FDD180807FFB3D0623F9208F19E37" italics="true" pageId="4" pageNumber="7">P. sarvakensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BBC60630807FFB3D6BC3C7308929E37" box="[915,1001,541,564]" pageId="4" pageNumber="7" rank="species">nov. sp.</taxonomicNameLabel>
and
<taxonomicName id="E5FB7A890807FFB3D10F3C730FF39E37" baseAuthorityName="d'Orbigny" baseAuthorityYear="1850" box="[1056,1160,541,564]" class="Globothalamea" family="Cyclamminidae" genus="Pseudocyclammina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="rugosa">
<emphasis id="108FDD180807FFB3D10F3C730FF39E37" box="[1056,1160,541,564]" italics="true" pageId="4" pageNumber="7">P. rugosa</emphasis>
</taxonomicName>
are by no means comparable. According to
<bibRefCitation id="466A7CFB0807FFB3D1213C500FD09E56" author="Maync, W." box="[1038,1195,573,598]" pageId="4" pageNumber="7" pagination="35 - 53" refId="ref5289" refString="Maync, W., 1952. Critical taxonomic study and nomenclatural revision of the family Lituolidae based upon the prototype of the family, Lituola nautiloidea Lamarck, 1804. Contributions from the Cushman Foundation for Foraminiferal Research, 3: 35 - 53." type="journal article" year="1952">Maync (1952)</bibRefCitation>
,
<taxonomicName id="E5FB7A890807FFB3D1963C500E589E56" baseAuthorityName="d'Orbigny" baseAuthorityYear="1850" box="[1209,1315,574,597]" class="Globothalamea" family="Cyclamminidae" genus="Pseudocyclammina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="rugosa">
<emphasis id="108FDD180807FFB3D1963C500E589E56" box="[1209,1315,574,597]" italics="true" pageId="4" pageNumber="7">P. rugosa</emphasis>
</taxonomicName>
attains
<quantity id="E503ACEF0807FFB3D0543C5308D39E75" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="3.8" metricValueMin="3.1" pageId="4" pageNumber="7" unit="mm" value="3.45" valueMax="3.8" valueMin="3.1">3.1 to 3.8 mm</quantity>
millimeters in length, displaying five to six large inflated chambers in the last whorl (
<figureCitation id="BAC01D8F0807FFB3D0253CEE0E2E9E9B" box="[1290,1365,640,664]" captionStart="Fig" captionStartId="5.[206,241,1726,1747]" captionTargetBox="[205,1379,165,1684]" captionTargetId="figure-33@5.[205,1379,165,1684]" captionTargetPageId="5" captionText="Fig. 4 Upper Cretaceous species of Pseudocyclammina Yabe &amp; Hanzawa, 1926; note the different scales. a-g Pseudocyclammina sarvakensis sp. nov., middle Cenomanian Sarvak Formation of SW Iran, Tang-e Darbast section, thin-section RAP 181 (b, d-f), and Shahneshin section, thin-section AKHS 97 (c, g). a Equatorial section, microspheric specimen? (from Sampò,1969 illustrated as Cyclammina sp. (?) in pl. XLV, fig. 2). b Equatorial section, holotype. c, e Oblique sections. d Subaxial section. f Oblique tangential section. g Oblique equatorial section, megalospheric specimen? h Pseudocyclammina massiliensis Maync, early-middle Campanian of Brač Island, Croatia. i Pseudocylammina rugosa (Orbigny), equatorial section, Cenomanian of W France (from Maync, 1952, pl. 12, fig. 10). j-m Pseudocyclammina sphaeroidea Gendrot, Upper Cretaceous (?Coniacian) of Austria (from Schlagintweit, 1992, pl. 1, figs. 13-16). Subaxial section (j), equatorial section of a specimen with uncoiling final stage (k), equatorial section (l), equatorial section slightly oblique (m)." figureDoi="http://doi.org/10.5281/zenodo.10520933" httpUri="https://zenodo.org/record/10520933/files/figure.png" pageId="4" pageNumber="7">Fig. 4i</figureCitation>
). It is worth mentioning that already
<bibRefCitation id="466A7CFB0807FFB3D1A73CCF0E5F9EBA" author="Henson, F. R. S." box="[1160,1316,673,697]" pageId="4" pageNumber="7" refId="ref5055" refString="Henson, F. R. S., 1948. Larger Imperforate Foraminifera of South-Western Asia. British Museum (Natural History), London, xi + 127 pp." type="book" year="1948">Henson (1948</bibRefCitation>
, p. 10 and 15) mentioned that the wall structure of ‘
<taxonomicName id="E5FB7A890807FFB3D1DB3CAC08E49EF8" authority="d'Orbigny" authorityName="d'Orbigny" authorityYear="1850" class="Globothalamea" family="Lituolidae" genus="Lituola" kingdom="Chromista" order="Lituolida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="rugosa">
<emphasis id="108FDD180807FFB3D1DB3CAC0E3A9ED9" box="[1268,1345,706,730]" italics="true" pageId="4" pageNumber="7">Lituola</emphasis>
’
<emphasis id="108FDD180807FFB3D07D3CAD0EE79ED9" box="[1362,1436,707,730]" italics="true" pageId="4" pageNumber="7">rugosa</emphasis>
d’Orbigny
</taxonomicName>
and
<taxonomicName id="E5FB7A890807FFB3D6F53C8D0FD29EF8" authorityName="Yabe &amp; Hanzawa" authorityYear="1926" box="[986,1193,739,763]" class="Globothalamea" family="Cyclamminidae" genus="Pseudocyclammina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="genus">
<emphasis id="108FDD180807FFB3D6F53C8D0FD29EF8" box="[986,1193,739,763]" italics="true" pageId="4" pageNumber="7">Pseudocyclammina</emphasis>
</taxonomicName>
“
<emphasis id="108FDD180807FFB3D1903C8A08829F1F" italics="true" pageId="4" pageNumber="7">are essentially similar in construction</emphasis>
” and that “
<emphasis id="108FDD180807FFB3D15A3D6B0F2E9F3E" italics="true" pageId="4" pageNumber="7">separation of the two genera is somewhat uncertain</emphasis>
”. Another upper Cretaceous representative of
<taxonomicName id="E5FB7A890807FFB3D6D63D280FB39F5D" authorityName="Yabe &amp; Hanzawa" authorityYear="1926" box="[1017,1224,838,862]" class="Globothalamea" family="Cyclamminidae" genus="Pseudocyclammina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="genus">
<emphasis id="108FDD180807FFB3D6D63D280FB39F5D" box="[1017,1224,838,862]" italics="true" pageId="4" pageNumber="7">Pseudocyclammina</emphasis>
</taxonomicName>
is
<taxonomicName id="E5FB7A890807FFB3D1D83D290EE79F5D" authorityName="Maync" authorityYear="1959" box="[1271,1436,838,862]" class="Globothalamea" family="Cyclamminidae" genus="Pseudocyclammina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="massiliensis">
<emphasis id="108FDD180807FFB3D1D83D290EE79F5D" box="[1271,1436,838,862]" italics="true" pageId="4" pageNumber="7">P. massiliensis</emphasis>
</taxonomicName>
described by
<bibRefCitation id="466A7CFB0807FFB3D6ED3D060F209F7C" author="Maync, W." box="[962,1115,871,896]" pageId="4" pageNumber="7" pagination="179 - 189" refId="ref5374" refString="Maync, W., 1959. Deux nouvelles especes Cretacees du genre Pseudocyclammina (Foraminiferes). Revue de Micropaleontologie, 1: 179 - 189." type="journal article" year="1959">Maync (1959)</bibRefCitation>
from the Santonian of southeastern
<collectingCountry id="5AEC419A0807FFB3D6AA3DE708B69FA3" box="[901,973,905,928]" name="France" pageId="4" pageNumber="7">France</collectingCountry>
(
<figureCitation id="BAC01D8F0807FFB3D6F13DE60F559FA3" box="[990,1070,904,928]" captionStart="Fig" captionStartId="5.[206,241,1726,1747]" captionTargetBox="[205,1379,165,1684]" captionTargetId="figure-33@5.[205,1379,165,1684]" captionTargetPageId="5" captionText="Fig. 4 Upper Cretaceous species of Pseudocyclammina Yabe &amp; Hanzawa, 1926; note the different scales. a-g Pseudocyclammina sarvakensis sp. nov., middle Cenomanian Sarvak Formation of SW Iran, Tang-e Darbast section, thin-section RAP 181 (b, d-f), and Shahneshin section, thin-section AKHS 97 (c, g). a Equatorial section, microspheric specimen? (from Sampò,1969 illustrated as Cyclammina sp. (?) in pl. XLV, fig. 2). b Equatorial section, holotype. c, e Oblique sections. d Subaxial section. f Oblique tangential section. g Oblique equatorial section, megalospheric specimen? h Pseudocyclammina massiliensis Maync, early-middle Campanian of Brač Island, Croatia. i Pseudocylammina rugosa (Orbigny), equatorial section, Cenomanian of W France (from Maync, 1952, pl. 12, fig. 10). j-m Pseudocyclammina sphaeroidea Gendrot, Upper Cretaceous (?Coniacian) of Austria (from Schlagintweit, 1992, pl. 1, figs. 13-16). Subaxial section (j), equatorial section of a specimen with uncoiling final stage (k), equatorial section (l), equatorial section slightly oblique (m)." figureDoi="http://doi.org/10.5281/zenodo.10520933" httpUri="https://zenodo.org/record/10520933/files/figure.png" pageId="4" pageNumber="7">Fig. 4h</figureCitation>
). This is also a comparably large species commonly displaying uncoiled elongated tests (up to
<quantity id="E503ACEF0807FFB3D6533DA508A99FE0" box="[892,978,971,995]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="7" unit="mm" value="2.3">2.3 mm</quantity>
) with five to seven chambers in the final whorl. So far no specimen of
<taxonomicName id="E5FB7A890807FFB3D1AA3D820E599800" authorityName="SCHLAGINTWEIT &amp; YAZDI-MOGHADAM" authorityYear="2023" box="[1157,1314,1003,1027]" class="Globothalamea" family="Cyclamminidae" genus="Pseudocyclammina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="sarvakensis">
<emphasis id="108FDD180807FFB3D1AA3D820E599800" box="[1157,1314,1003,1027]" italics="true" pageId="4" pageNumber="7">P. sarvakensis</emphasis>
</taxonomicName>
displaying uncoiled test has been observed. The number of chambers in the final whorl might differentiate
<taxonomicName id="E5FB7A890807FFB3D1EA3A410E1B9845" authorityName="SCHLAGINTWEIT &amp; YAZDI-MOGHADAM" authorityYear="2023" box="[1221,1376,1070,1094]" class="Globothalamea" family="Cyclamminidae" genus="Pseudocyclammina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="sarvakensis">
<emphasis id="108FDD180807FFB3D1EA3A410E1B9845" box="[1221,1376,1070,1094]" italics="true" pageId="4" pageNumber="7">P. sarvakensis</emphasis>
</taxonomicName>
from all other Upper Jurassic–Lower Cretaceous
<taxonomicName id="E5FB7A890807FFB3D0023A210894988B" pageId="4" pageNumber="7">
<emphasis id="108FDD180807FFB3D0023A2108E2988B" italics="true" pageId="4" pageNumber="7">Pseudocyclammina</emphasis>
species
</taxonomicName>
(
<bibRefCitation id="466A7CFB0807FFB3D6D03A1E0FE9988B" author="Maync, W." box="[1023,1170,1136,1160]" pageId="4" pageNumber="7" pagination="179 - 189" refId="ref5374" refString="Maync, W., 1959. Deux nouvelles especes Cretacees du genre Pseudocyclammina (Foraminiferes). Revue de Micropaleontologie, 1: 179 - 189." type="journal article" year="1959">Maync, 1959</bibRefCitation>
,
<tableCitation id="6F7934B10807FFB3D1B03A1E0FA7988B" box="[1183,1244,1136,1160]" captionStart="Tabel 1" captionStartId="7.[154,219,153,177]" captionTargetBox="[166,1418,253,650]" captionTargetId="graphics-1386@7.[151,1434,245,654]" captionTargetPageId="7" captionText="Tabel 1 Comparison of dimensions of Pseudotextulariella cretosa (Cushman) (late Albian–Cenomanian), P. brevicamerata sp. nov., and P. courtionensis Brönnimann (late Berriasian–Valanginian). Test height and diameter in mm." httpUri="http://table.plazi.org/id/768451820804FFB0D5B53EF70E0A9CD2" pageId="4" pageNumber="7" tableUuid="768451820804FFB0D5B53EF70E0A9CD2">tab. 1</tableCitation>
;
<bibRefCitation id="466A7CFB0807FFB3D1C33A1F08D798AA" author="Banner, F. T. &amp; Whittaker, J. E." pageId="4" pageNumber="7" pagination="41 - 59" refId="ref4251" refString="Banner, F. T. &amp; Whittaker, J. E., 1991. Redmond's &quot; new lituolid foraminifera &quot; from the Mesozoic of Saudi Arabia. Micropaleontology, 37: 41 - 59." type="journal article" year="1991">Banner &amp; Whittaker, 1991</bibRefCitation>
). For example, the Upper Jurassic-Lower Cretaceous type-species
<taxonomicName id="E5FB7A890807FFB3D1373ADC08E698E8" authority="(Yokoyama, 1890)" baseAuthorityName="Yokoyama" baseAuthorityYear="1890" class="Globothalamea" family="Cyclamminidae" genus="Pseudocyclammina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="lituus">
<emphasis id="108FDD180807FFB3D1373ADC0E5798C9" box="[1048,1324,1202,1226]" italics="true" pageId="4" pageNumber="7">Pseudocyclammina lituus</emphasis>
(
<bibRefCitation id="466A7CFB0807FFB3D0113ADC08EF98E8" author="Yokoyama, M." pageId="4" pageNumber="7" pagination="26 - 27" refId="ref6499" refString="Yokoyama, M., 1890. Foraminiferen aus dem Kalksteine von Torinosu und Kompira. Denkschriften der Kaiserlichen Akademie der Wissenschaften, Wien, Mathematisch-Naturwissenschaftliche Classe, 57: 26 - 27." type="journal article" year="1890">Yokoyama, 1890</bibRefCitation>
)
</taxonomicName>
typically has 8 to 12 chambers in the last whorl (
<bibRefCitation id="466A7CFB0807FFB3D6173A9B0F20990E" author="Banner, F. T. &amp; Whittaker, J. E." box="[824,1115,1269,1293]" pageId="4" pageNumber="7" pagination="41 - 59" refId="ref4251" refString="Banner, F. T. &amp; Whittaker, J. E., 1991. Redmond's &quot; new lituolid foraminifera &quot; from the Mesozoic of Saudi Arabia. Micropaleontology, 37: 41 - 59." type="journal article" year="1991">Banner &amp; Whittaker, 1991</bibRefCitation>
). The Lower Cretaceous (Aptian-Albian)
<taxonomicName id="E5FB7A890807FFB3D6933B780FA9992E" authority="Maync, 1953" authorityName="Maync" authorityYear="1953" box="[956,1234,1301,1326]" class="Globothalamea" family="Cyclamminidae" genus="Pseudocyclammina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="hedbergi">
<emphasis id="108FDD180807FFB3D6933B780F40992E" box="[956,1083,1301,1325]" italics="true" pageId="4" pageNumber="7">P. hedbergi</emphasis>
<bibRefCitation id="466A7CFB0807FFB3D16B3B780FA9992E" author="Maync, W." box="[1092,1234,1301,1326]" pageId="4" pageNumber="7" pagination="101 - 102" refId="ref5335" refString="Maync, W., 1953. Pseudocyclammina hedbergi n. sp. from the Urgo-Aptian and Albian of Venezuela. Contributions from the Cushman Foundation for Foraminiferal Research. 4 (3): 101 - 102." type="journal article" year="1953">Maync, 1953</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="E5FB7A890807FFB3D0243B780EEE992E" authorityName="Maync" authorityYear="1959" box="[1291,1429,1302,1325]" class="Globothalamea" family="Cyclamminidae" genus="Pseudocyclammina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="vasconica">
<emphasis id="108FDD180807FFB3D0243B780EEE992E" box="[1291,1429,1302,1325]" italics="true" pageId="4" pageNumber="7">P. vasconica</emphasis>
</taxonomicName>
, 1959 have 4-5 and 7-8 chambers in the final whorl.
<taxonomicName id="E5FB7A890807FFB3D04F3B590E7F9973" authority="Gendrot, 1968" authorityName="Gendrot" authorityYear="1968" class="Globothalamea" family="Cyclamminidae" genus="Pseudocyclammina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="sphaeroidea">
<emphasis id="108FDD180807FFB3D04F3B590F219973" italics="true" pageId="4" pageNumber="7">Pseudocyclammina sphaeroidea</emphasis>
<bibRefCitation id="466A7CFB0807FFB3D1493B360E7F9973" author="Gendrot, C." box="[1126,1284,1368,1392]" pageId="4" pageNumber="7" pagination="657 - 694" refId="ref4986" refString="Gendrot, C., 1968. Stratigraphie et micropaleontologie du Senonien de la region es Martigues pres Marseille (Bouches-du-Rhone). Eclogae Geologicae Helvetia, 61: 657 - 694." type="journal article" year="1968">Gendrot, 1968</bibRefCitation>
</taxonomicName>
(=
<taxonomicName id="E5FB7A890807FFB3D01D3B3708CF9992" class="Globothalamea" family="Mayncinidae" genus="Mayncina" kingdom="Chromista" order="Lituolida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="sphaeroidea">
<emphasis id="108FDD180807FFB3D01D3B3708CF9992" italics="true" pageId="4" pageNumber="7">Mayncina sphaeroidea</emphasis>
</taxonomicName>
acc. to
<bibRefCitation id="466A7CFB0807FFB3D1223B170FBB9992" author="De Castro, P." box="[1037,1216,1401,1425]" pageId="4" pageNumber="7" pagination="21 - 38" refId="ref4766" refString="De Castro, P., 1991. Mesozoic. In: Barattolo, F., De Castro, P. &amp; Parente, M. (Eds.), 5 th International Symposium on Fossil Algae Capri, 7 - 12 April 1991, Field Trip Guide-Book: 21 - 38." type="book chapter" year="1991">De Castro, 1991</bibRefCitation>
) described from the upper Santonian of southeastern
<collectingCountry id="5AEC419A0807FFB3D1943BF50E7F99B1" box="[1211,1284,1435,1458]" name="France" pageId="4" pageNumber="7">France</collectingCountry>
is distinctly smaller than
<taxonomicName id="E5FB7A890807FFB3D6913BD20F2199D0" authorityName="SCHLAGINTWEIT &amp; YAZDI-MOGHADAM" authorityYear="2023" box="[958,1114,1467,1491]" class="Globothalamea" family="Cyclamminidae" genus="Pseudocyclammina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="sarvakensis">
<emphasis id="108FDD180807FFB3D6913BD20F2199D0" box="[958,1114,1467,1491]" italics="true" pageId="4" pageNumber="7">P. sarvakensis</emphasis>
</taxonomicName>
with test length of
<quantity id="E503ACEF0807FFB3D01C3BD5082299F7" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="4" pageNumber="7" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.5 to 0.6 mm</quantity>
, often uncoiling (
<figureCitation id="BAC01D8F0807FFB3D1313BB20F1B99F7" box="[1054,1120,1500,1524]" captionStart="Fig" captionStartId="5.[206,241,1726,1747]" captionTargetBox="[205,1379,165,1684]" captionTargetId="figure-33@5.[205,1379,165,1684]" captionTargetPageId="5" captionText="Fig. 4 Upper Cretaceous species of Pseudocyclammina Yabe &amp; Hanzawa, 1926; note the different scales. a-g Pseudocyclammina sarvakensis sp. nov., middle Cenomanian Sarvak Formation of SW Iran, Tang-e Darbast section, thin-section RAP 181 (b, d-f), and Shahneshin section, thin-section AKHS 97 (c, g). a Equatorial section, microspheric specimen? (from Sampò,1969 illustrated as Cyclammina sp. (?) in pl. XLV, fig. 2). b Equatorial section, holotype. c, e Oblique sections. d Subaxial section. f Oblique tangential section. g Oblique equatorial section, megalospheric specimen? h Pseudocyclammina massiliensis Maync, early-middle Campanian of Brač Island, Croatia. i Pseudocylammina rugosa (Orbigny), equatorial section, Cenomanian of W France (from Maync, 1952, pl. 12, fig. 10). j-m Pseudocyclammina sphaeroidea Gendrot, Upper Cretaceous (?Coniacian) of Austria (from Schlagintweit, 1992, pl. 1, figs. 13-16). Subaxial section (j), equatorial section of a specimen with uncoiling final stage (k), equatorial section (l), equatorial section slightly oblique (m)." figureDoi="http://doi.org/10.5281/zenodo.10520933" httpUri="https://zenodo.org/record/10520933/files/figure.png" pageId="4" pageNumber="7">Fig. 4</figureCitation>
j-m). The youngest record of the genus
<taxonomicName id="E5FB7A890807FFB3D6B03B930F159A16" authorityName="Yabe &amp; Hanzawa" authorityYear="1926" box="[927,1134,1533,1557]" class="Globothalamea" family="Cyclamminidae" genus="Pseudocyclammina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="genus">
<emphasis id="108FDD180807FFB3D6B03B930F159A16" box="[927,1134,1533,1557]" italics="true" pageId="4" pageNumber="7">Pseudocyclammina</emphasis>
</taxonomicName>
seems to be from the upper Campanian (
<bibRefCitation id="466A7CFB0807FFB3D6EC38700F1D9A35" author="Parente, M." box="[963,1126,1566,1590]" pageId="4" pageNumber="7" pagination="155 - 170" refId="ref5676" refString="Parente, M., 1994. A revised stratigraphy of the Upper Cretaceous to Oligocene units from southeastern Salento (Apulia, southern Itlay). Bolletinmo della Societa Paleontologica Italiana, 33 (2): 155 - 170." type="journal article" year="1994">Parente, 1994</bibRefCitation>
, pl. 1, fig. 5,
<taxonomicName id="E5FB7A890807FFB3D001387008B89A54" class="Globothalamea" family="Cyclamminidae" genus="Pseudocyclammina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="undetermined">
<emphasis id="108FDD180807FFB3D001387008E29A54" italics="true" pageId="4" pageNumber="7">Pseudocyclammina</emphasis>
sp.
</taxonomicName>
, in our opinion referable to
<taxonomicName id="E5FB7A890807FFB3D1D3382E08049A7B" authority="Maync" authorityName="Maync" authorityYear="1959" class="Globothalamea" family="Cyclamminidae" genus="Pseudocyclammina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="massiliensis">
<emphasis id="108FDD180807FFB3D1D3382E0EE79A54" box="[1276,1436,1599,1623]" italics="true" pageId="4" pageNumber="7">P. massiliensis</emphasis>
Maync
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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