374 lines
39 KiB
XML
374 lines
39 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.699.12031" ID-GBIF-Dataset="dbf6bb9f-7ea4-4fa3-83ee-f58263427e39" ID-PMC="PMC5673864" ID-Pensoft-Pub="1313-2970-699-1" ID-PubMed="29200927" ID-ZBK="0F92401F3F7C4896AD9D72BC84348C7D" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-699-1" ModsDocOrigin="ZooKeys 699" ModsDocTitle="A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae)" checkinTime="1505836577638" checkinUser="pensoft" docAuthor="Cherman, Mariana Alejandra, Mise, Kleber Makoto, Moron †, Miguel Angel, Vaz-de-Mello, Fernando Z. & Almeida, Lucia Massutti de" docDate="2017" docId="AA297075A9704FA336524936BC022CBC" docLanguage="en" docName="ZooKeys 699: 1-120" docOrigin="ZooKeys 699" docSource="http://dx.doi.org/10.3897/zookeys.699.12031" docTitle="Liogenys fusca Blanchard 1851" docType="treatment" docVersion="4" lastPageNumber="37" masterDocId="FFD9636FB945C41DFF8AB133FFFA9D76" masterDocTitle="A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae)" masterLastPageNumber="120" masterPageNumber="1" pageNumber="34" updateTime="1668164813467" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Cherman, Mariana Alejandra</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Mise, Kleber Makoto</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Moron †, Miguel Angel</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Vaz-de-Mello, Fernando Z.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Almeida, Lucia Massutti de</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2017</mods:date>
|
||
<mods:detail type="volume">
|
||
<mods:number>699</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>1</mods:start>
|
||
<mods:end>120</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/zookeys.699.12031</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.699.12031</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1313-2970-699-1</mods:identifier>
|
||
<mods:identifier type="ZBK">0F92401F3F7C4896AD9D72BC84348C7D</mods:identifier>
|
||
<mods:identifier type="ZooBank">0F92401F3F7C4896AD9D72BC84348C7D</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="134311176" LSID="urn:lsid:plazi:treatment:AA297075A9704FA336524936BC022CBC" httpUri="http://treatment.plazi.org/id/AA297075A9704FA336524936BC022CBC" lastPageId="37" lastPageNumber="37" pageId="33" pageNumber="34">
|
||
<subSubSection pageId="33" pageNumber="34" type="nomenclature">
|
||
<paragraph pageId="33" pageNumber="34">
|
||
<taxonomicName authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys fusca" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="fusca">
|
||
<pageBreakToken pageId="33" pageNumber="34" start="start">Liogenys</pageBreakToken>
|
||
fusca Blanchard, 1851
|
||
</taxonomicName>
|
||
Figs 64, 92
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="33" pageNumber="34" type="reference_group">
|
||
<paragraph pageId="33" pageNumber="34">
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys fuscus" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="fuscus">Liogenys fuscus</taxonomicName>
|
||
Blanchard, 1851: 168 (orig. desc.),
|
||
<bibRefCitation author="Lacordaire, JT" journalOrPublisher="Librarie Encyclopedique de Roret, Paris" pageId="106" pageNumber="107" title="Histoire naturelle des Insectes. Genera des Coleopteres, ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet odre d'insectes. Tome Troiseme. Contenant les familles des Pectinicornes et Lamellicornes." year="1856">Lacordaire 1856</bibRefCitation>
|
||
: 269 (sys.);
|
||
<bibRefCitation pageId="33" pageNumber="34">Harold 1869a</bibRefCitation>
|
||
: 1140 (check.);
|
||
<bibRefCitation pageId="33" pageNumber="34">Dalla Torre 1913</bibRefCitation>
|
||
: 318 (check.);
|
||
<bibRefCitation author="Frey, G" journalOrPublisher="Frey" pageId="105" pageNumber="106" pagination="36 - 64" title="Betimmumgstabelle und Revision der Gattung Liogenys Guerrez. (Coleoptera-Melolonthinae-Macrodactylini). Entomologischen Arbeiten aus dem Museum G." volume="20" year="1969">Frey 1969</bibRefCitation>
|
||
: 47, 55 (key, red.).
|
||
</paragraph>
|
||
<paragraph pageId="33" pageNumber="34">
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys fusca" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="fusca">Liogenys fusca</taxonomicName>
|
||
:
|
||
<bibRefCitation author="Blackwelder, RE" journalOrPublisher="United States National Museum Bulletin" pageId="103" pageNumber="104" pagination="189 - 341" title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
|
||
: 227 (check.);
|
||
<bibRefCitation author="Evans, AV" journalOrPublisher="Zootaxa" pageId="104" pageNumber="105" pagination="1 - 458" title="A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae)." url="https://doi.org/10.11646/zootaxa.211.1.1" volume="211" year="2003">Evans 2003</bibRefCitation>
|
||
: 209 (check.);
|
||
<bibRefCitation author="Evans, AV" journalOrPublisher="Zootaxa" pageId="104" pageNumber="105" url="http://digitalcommons.unl.edu/entomologypapers/2" year="2005">Evans and Smith 2005</bibRefCitation>
|
||
: 174 (check.);
|
||
<bibRefCitation author="Evans, AV" journalOrPublisher="Zootaxa" pageId="104" pageNumber="105" url="http://museum.unl.edu/research/entomology/SSSA/nwmelos.htm" year="2009">Evans and Smith 2009</bibRefCitation>
|
||
: 178 (check.).
|
||
</paragraph>
|
||
<paragraph pageId="33" pageNumber="34">
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys argentinus" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="argentinus">Liogenys argentinus</taxonomicName>
|
||
Moser, 1918: 97 (orig. desc.);
|
||
<bibRefCitation author="Frey, G" journalOrPublisher="Frey" pageId="105" pageNumber="106" pagination="36 - 64" title="Betimmumgstabelle und Revision der Gattung Liogenys Guerrez. (Coleoptera-Melolonthinae-Macrodactylini). Entomologischen Arbeiten aus dem Museum G." volume="20" year="1969">Frey 1969</bibRefCitation>
|
||
: 55 (syn.).
|
||
</paragraph>
|
||
<paragraph pageId="33" pageNumber="34">
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys argentina" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="argentina">Liogenys argentina</taxonomicName>
|
||
:
|
||
<bibRefCitation author="Blackwelder, RE" journalOrPublisher="United States National Museum Bulletin" pageId="103" pageNumber="104" pagination="189 - 341" title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
|
||
: 227 (check.).
|
||
</paragraph>
|
||
<paragraph pageId="33" pageNumber="34">
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys cuyabanus" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="cuyabanus">Liogenys cuyabanus</taxonomicName>
|
||
Moser, 1919: 12 (orig. desc.);
|
||
<bibRefCitation author="Frey, G" journalOrPublisher="Frey" pageId="105" pageNumber="106" pagination="36 - 64" title="Betimmumgstabelle und Revision der Gattung Liogenys Guerrez. (Coleoptera-Melolonthinae-Macrodactylini). Entomologischen Arbeiten aus dem Museum G." volume="20" year="1969">Frey 1969</bibRefCitation>
|
||
: 55 (syn.).
|
||
</paragraph>
|
||
<paragraph pageId="33" pageNumber="34">
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys cuyabana" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="cuyabana">Liogenys cuyabana</taxonomicName>
|
||
:
|
||
<bibRefCitation author="Blackwelder, RE" journalOrPublisher="United States National Museum Bulletin" pageId="103" pageNumber="104" pagination="189 - 341" title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
|
||
: 227 (check.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="33" pageNumber="34" type="type material">
|
||
<paragraph pageId="33" pageNumber="34">Type material.</paragraph>
|
||
<paragraph pageId="33" pageNumber="34">
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys fuscus" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="fuscus">Liogenys fuscus</taxonomicName>
|
||
female syntype (MNHN): [white handwritten] "De
|
||
<normalizedToken originalValue="l’emb">l'emb</normalizedToken>
|
||
./de
|
||
<normalizedToken originalValue="l’Uruguay/jusqu’aux/missions”">l'Uruguay/jusqu'aux/missions"</normalizedToken>
|
||
, [green handwritten] "
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys fuscus" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="fuscus">Liogenys fuscus</taxonomicName>
|
||
/Cat. Mus./Rives de
|
||
<normalizedToken originalValue="l’Uruguay">l'Uruguay</normalizedToken>
|
||
/ M. A. St Hilaire", [red printed]
|
||
<normalizedToken originalValue="“SYNTYPE”">"SYNTYPE"</normalizedToken>
|
||
, [light green printed]
|
||
<normalizedToken originalValue="“MUSÉUM">"MUSEUM</normalizedToken>
|
||
PARIS". This type is here designated the lectotype [white, outlined in red, printed], "LECTOTYPE/
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys fuscus" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="fuscus">Liogenys fuscus</taxonomicName>
|
||
/Blanchard, 1851/ des. M. A. Cherman 2012". Female syntype (MNHN): [white handwritten]
|
||
<normalizedToken originalValue="“6449/34”">"6449/34"</normalizedToken>
|
||
, [white handwritten]
|
||
<normalizedToken originalValue="“1444”">"1444"</normalizedToken>
|
||
, [light green printed]
|
||
<normalizedToken originalValue="“MUSÉUM">"MUSEUM</normalizedToken>
|
||
PARIS/ [handwritten] Santa Cruz/
|
||
<normalizedToken originalValue="D’Orbigny”">D'Orbigny"</normalizedToken>
|
||
. This type is here designated the paralectotype: [white, outlined in red, printed] "PARALECTOTYPE/
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys fuscus" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="fuscus">Liogenys fuscus</taxonomicName>
|
||
/Blanchard, 1851/ des. M. A. Cherman 2012".
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys argentinus" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="argentinus">Liogenys argentinus</taxonomicName>
|
||
male syntype ZMHB: [grey printed] "Argentinien/Santiago del Estero", [white handwritten] "
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
|
||
/
|
||
<taxonomicName lsidName="argentinus" pageId="33" pageNumber="34" rank="species" species="argentinus">argentinus</taxonomicName>
|
||
/Mos./Typen m# ", [red printed]
|
||
<normalizedToken originalValue="“Typus”">"Typus"</normalizedToken>
|
||
, [red printed] "SYNTYPUS/
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
|
||
/
|
||
<taxonomicName lsidName="argentinus" pageId="33" pageNumber="34" rank="species" species="argentinus">argentinus</taxonomicName>
|
||
Moser, 1918/labelled by MNHUB 2011". Genitalia mounted. This type is here designated the lectotype [white, outlined in red, printed] "LECTOTYPE/
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys argentinus" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="argentinus">Liogenys argentinus</taxonomicName>
|
||
/Moser, 1918/ des. M. A. Cherman 2012". Seven male and seven female syntypes (ZMHB): [grey printed] "Argentinien/ Santiago del Estero", [white handwritten] "
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
|
||
/
|
||
<taxonomicName lsidName="argentinus" pageId="33" pageNumber="34" rank="species" species="argentinus">argentinus</taxonomicName>
|
||
/Mos./ Typen", [red printed]
|
||
<normalizedToken originalValue="“Typus”">"Typus"</normalizedToken>
|
||
, [red printed] "SYNTYPUS/
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
|
||
/
|
||
<taxonomicName lsidName="argentinus" pageId="33" pageNumber="34" rank="species" species="argentinus">argentinus</taxonomicName>
|
||
Moser, 1918/ labelled by MNHUB 2011". These fourteen syntypes are here designated as paralectotypes, each one with the label: [white, outlined in red, printed] "PARALECTOTYPE/
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys argentinus" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="argentinus">Liogenys argentinus</taxonomicName>
|
||
/Moser, 1918/ des. M. A. Cherman 2012".
|
||
</paragraph>
|
||
<paragraph pageId="33" pageNumber="34">
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys cuyabanus" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="cuyabanus">Liogenys cuyabanus</taxonomicName>
|
||
male syntype (ZMHB): [white handwritten] "Cuyaba/Mato Grosso", [white handwritten] "
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
|
||
/
|
||
<taxonomicName lsidName="cuyabanus" pageId="33" pageNumber="34" rank="species" species="cuyabanus">cuyabanus</taxonomicName>
|
||
/Mos./Type m#", [light red printed]
|
||
<normalizedToken originalValue="“Typus”">"Typus"</normalizedToken>
|
||
, [red printed] "SYNTYPUS/
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
|
||
/
|
||
<taxonomicName lsidName="cuyabana" pageId="33" pageNumber="34" rank="species" species="cuyabana">cuyabana</taxonomicName>
|
||
Moser, 1919/labelled by MNHUB 2011". Genitalia mounted. This type is here designated the lectotype [white, outlined in red, printed] "LECTOTYPE/
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys cuyabanus" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="cuyabanus">Liogenys cuyabanus</taxonomicName>
|
||
/Moser, 1919/ des. M. A. Cherman 2012". Four female syntypes (ZMHB): [white handwritten] "Cuyaba/Mato Grosso", [white handwritten] "
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
|
||
/
|
||
<taxonomicName lsidName="cuyabanus" pageId="33" pageNumber="34" rank="species" species="cuyabanus">cuyabanus</taxonomicName>
|
||
/Mos./Typen f#", [light red printed]
|
||
<normalizedToken originalValue="“Typus”">"Typus"</normalizedToken>
|
||
, [red printed] "SYNTYPUS/
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
|
||
/
|
||
<taxonomicName lsidName="cuyabana" pageId="33" pageNumber="34" rank="species" species="cuyabana">cuyabana</taxonomicName>
|
||
Moser, 1919/labelled by MNHUB 2011". These four syntypes are here designated as paralectotypes, each one with the label: "PARALECTOTYPE/
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys cuyabanus" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="cuyabanus">Liogenys cuyabanus</taxonomicName>
|
||
/Moser, 1919/ des. M. A. Cherman 2012".
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34" type="type material">
|
||
<paragraph pageId="33" pageNumber="34">Non-type material.</paragraph>
|
||
<paragraph lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34">
|
||
PARAGUAY. "Estancia
|
||
<normalizedToken originalValue="Postillón">Postillon</normalizedToken>
|
||
, Puerto
|
||
<taxonomicName lsidName="max" pageId="33" pageNumber="34" rank="species" species="max">max</taxonomicName>
|
||
. A. Rio Paraguay", without date, Louis des Arts Jr. Col., 1 ex. (ZMHB); PH: Puerto Pinasco, IX/1916, without collector, 2 ex. (AMNH); Campo
|
||
<normalizedToken originalValue="León">Leon</normalizedToken>
|
||
, X/1979, A. Martinez col., 1 ex. (CMNC); BQ: Filadelfia, X/1979, A. Martinez col., 2 ex. (CMNC). BOLIVIA. SC: Santa Cruz de la Sierra, without date, M.
|
||
<normalizedToken originalValue="D’Orbigny">D'Orbigny</normalizedToken>
|
||
col., 2 ex. (MNHN); Camiri, 9/XI/1946, Maldonado col., 2 ex. (CMNC); Comarapa, "9 km SW",
|
||
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-17.9821">17°58.926'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-64.57275">64°34.365'W</geoCoordinate>
|
||
, XII/2008, Edmonds and Vidaurre cols., 1 ex. (CEMT); San Ramon, "Laguna Brava", 15/IX/1987, A. Castillo col., 1 ex. (AMNH); TA: Tatarenda, without date, 618 m, Erland Nordenskj col., 1 ex. (AMHB). BRAZIL. TO: Porto Nacional, XI/1953, R. Aires col., 1 ex. (DZUP); BA: Encruzilhada,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-15.576388">15°34'35"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-40.947502">40°56'51"W</geoCoordinate>
|
||
, 15/XII/2012, 850 m, Rafael and Grossi col., 2 ex. (INPA); MG: Arinos, 6/XI/1964, Exp. Dep. Zool. Col., 1 ex. (MZSP);
|
||
<normalizedToken originalValue="Águas">Aguas</normalizedToken>
|
||
Vermelhas, "Faz. Faccino",12/XII/2012, P. Grossi col., 1 ex. (EPGC); DF:
|
||
<normalizedToken originalValue="Brasília">Brasilia</normalizedToken>
|
||
,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-15.736195">15°44'10.30"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-47.93517">47°56'6.6"W</geoCoordinate>
|
||
, XII/2009, 1070 m, Carvalho col., 1 ex. (CEMT); Planaltina, "Embrapa Cerrados",
|
||
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-15.6044445">15°36'16"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-47.737778">47°44'16"W</geoCoordinate>
|
||
, 18/XI/2009, C. Oliveira col., 1 ex. (CEMT); GO:
|
||
<normalizedToken originalValue="Aragarças">Aragarcas</normalizedToken>
|
||
, X/1959,
|
||
<pageBreakToken pageId="34" pageNumber="35" start="start">M</pageBreakToken>
|
||
. Alvarenga, 2 ex. (DZUP); MT:
|
||
<normalizedToken originalValue="Barão">Barao</normalizedToken>
|
||
de
|
||
<normalizedToken originalValue="Melgaço">Melgaco</normalizedToken>
|
||
, 21/V/2011, Sol col., 1 ex. (CEMT); Barra do
|
||
<normalizedToken originalValue="Garças">Garcas</normalizedToken>
|
||
, 1990, Serrano col., 1 ex.; 15/X/1988, Capucho col., 1 ex.; 4/X/1988, Pereira col., 1ex. (CEMT); Boa
|
||
<normalizedToken originalValue="Esperança">Esperanca</normalizedToken>
|
||
3/XI/1991, Oliveira col., 1 ex. (CEMT);
|
||
<normalizedToken originalValue="Cáceres">Caceres</normalizedToken>
|
||
, 9/XI/1984, C. Elias col., 1 ex. (DZUP); Chapada dos
|
||
<normalizedToken originalValue="Guimarães">Guimaraes</normalizedToken>
|
||
, 27-30/IX/2005, Marques col., 5 ex.; 5/X/2005, Marques col., 1 ex.; 30/V/2011, Sol col., 1 ex.; 27/V/2011, Klemp col., 1 ex.; "EE Buriti", 6/X/1984, Dal Ponte col., 1 ex.;
|
||
<normalizedToken originalValue="“Água">"Agua</normalizedToken>
|
||
Fria", 8/X/1989, without collector, 1 ex.; (CEMT);
|
||
<normalizedToken originalValue="Cuiabá">Cuiaba</normalizedToken>
|
||
, without date and collector, 8 ex.; 7/IX/1988, Loverde col. 1 ex.; 6/X/1988, Oliveira, Alves and Macieski cols., 4 ex.; 7/X/1988, Resende col., 1 ex.; 10/XI/1990, Aicio col., 3 ex.; 13/X/1991, Souza col., 1 ex.; 8/XI/1991, Aquino col., 1 ex.; 14/II/1992, Abido col., 1 ex.; 10/VIII/1992, Edenir col., 1 ex.; 8/IV/1993, Almindo Filho col., 1 ex.; 6/V/1993, Serrano col., 1 ex.; 20-25/VII/1993, Serrano col., 2 ex.; 17/VIII/1993, Serrano col., 1 ex.; 19/VIII/1993, Romio col., 1 ex.; 24/VIII/1993, Pinto col., 4 ex.; 8/XI/1993, Godoy col., 1 ex.; 21/XI/1993, Silva col., 4 ex.; 10/IV/1994, silveira col., 1 ex.; 12/III/1994, Ormond col., 1 ex.; 9/IX/1994, Montori col., 1 ex.; 1, 29/X/1994, Montori col. 2 ex.; 16-18/X/1994, Solange col., 2 ex.; 20/VI/2009, Silva col., 1 ex.; 11/IV/2010, Marcondes col., 1 ex.; "campus UFMT" 24/X/1988, Oliveira col., 1 ex.; 4/X/1990, Marques col., 1 ex.; 8/XI/1990, Alves Costa col., 1 ex.; 14/IX/1991, Souza col., 1 ex.; X/2008, Vaz-de-Mello col., 12 ex.; 13/IX/1988, Marques col., 1 ex.; 10/XI/1988, Leni col., 1 ex.; 13/IX/1988, Marques col. 1 ex; "Fazenda Birvana", X-XI/1988, Serrano col., 2 ex.; "Duque de Caxias, 15/III/2011, Cardodo col., 2 ex.; "
|
||
<normalizedToken originalValue="Rodoviária”">Rodoviaria"</normalizedToken>
|
||
, 15/IV/2011, Vaz-de-Mello col., 14 ex.; "Santa Cruz", 30/XI/2010, Kramer col., 1 ex.; "Res.
|
||
<normalizedToken originalValue="Coxipó”">Coxipo"</normalizedToken>
|
||
, 12/IV/1989, Gianni col., 1 ex.; 22/XI/2010, F. Vaz-de-Mello col., 1 ex.; "Boa
|
||
<normalizedToken originalValue="Esperança”">Esperanca"</normalizedToken>
|
||
X/2010, Vaz-de-Mello col., 1 ex.; 5/V/2010, Rabello col., 1ex.; "Jardim
|
||
<normalizedToken originalValue="Vitória”">Vitoria"</normalizedToken>
|
||
22/VIII/2009,
|
||
<normalizedToken originalValue="Galvão">Galvao</normalizedToken>
|
||
col., 2 ex.; "Recanto do Sol" 23/X/2009, Dal Mas col., 1 ex.;
|
||
<normalizedToken originalValue="“Cophamil”">"Cophamil"</normalizedToken>
|
||
27/VIII/1990, Regina col., 1 ex.; "Santa Amalia", 22/V/2011, Klemp col., 1 ex.;
|
||
<normalizedToken originalValue="“Verdão”">"Verdao"</normalizedToken>
|
||
,25/X/1989, Roseni col., 1 ex.; "Jardim das
|
||
<normalizedToken originalValue="Américas”">Americas"</normalizedToken>
|
||
11/VI/2010, Amado col., 1 ex.; Lucas do Rio Verde, 3/I/2012, Camera col., 2 ex. (CEMT); Nova Mutum, "Alto da Colina", 20/VII/2010, Amhold col., 1 ex. (CEMT); Nossa Senhora do Livramento, 15/XI/1992, 2 ex.; 12/XII/1992, 1 ex. (CEMT); Pari, 6/X/1988, Auxiliadora col., 1 ex. (CEMT); Pocone, "SESC pantanal", 10/IV/2009, Abreu col., 1 ex.; 14/V/2002, without collector, 2 ex.; 6/IX/2002, Myazaki col., 4 ex.; 7/IX/2002, without collector, 5 ex.; 12/X/2002; without collector, 1 ex.; "Fazenda Santa Maria", 23/XII/2011, Semedo col., 1 ex.; "Fazenda Bom Jesus", 24/XII/2010, Santos col., 1 ex.; "Transpantaneira, Porto Jofre", 5/V/1984, Silva col., 1 ex. (CEMT); Porto
|
||
<normalizedToken originalValue="Esperidião">Esperidiao</normalizedToken>
|
||
, XI/1984, Magno, Alvarenga cols., 3 ex. (MNRJ);
|
||
<normalizedToken originalValue="Rondonópolis">Rondonopolis</normalizedToken>
|
||
, 6/IX/1991, Viegas col., 1 ex. (CEMT); Rosario Oeste, XI/1976, A. Maller col., 2 ex. (MNRJ); Santo Antonio do Leverger, 16/XI/2008, Becker col., 1 ex. (CEMT);
|
||
<normalizedToken originalValue="Tangará">Tangara</normalizedToken>
|
||
da serra, "Chacara Asa Branca",
|
||
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-14.545">14°32'42"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-58.674446">58°40'28"W</geoCoordinate>
|
||
, XI/2011, Meurer col., 8 ex. (CEMT).
|
||
<normalizedToken originalValue="Várzea">Varzea</normalizedToken>
|
||
Grande, 8/VI/2011, Uchaki col. 1 ex.; 27/X/1991, Bento col., 1 ex.; 5/XI/2008, Silva col., 1 ex. (CEMT); MS:
|
||
<normalizedToken originalValue="Corumbá">Corumba</normalizedToken>
|
||
,
|
||
<normalizedToken originalValue="“Nhecolandia”">"Nhecolandia"</normalizedToken>
|
||
, 17/X/1953, C. R.
|
||
<normalizedToken originalValue="Gonçalves">Goncalves</normalizedToken>
|
||
col., 11 ex. (DZUP); 11 ex. (MNRJ); "Fazenda
|
||
<normalizedToken originalValue="São">Sao</normalizedToken>
|
||
Bento
|
||
<normalizedToken originalValue="próxima">proxima</normalizedToken>
|
||
a Porto Jofre", 6/XII/2010, Tortato col., 1 ex. (CEMT); Salobra, 29/VIII/1940, Com
|
||
<pageBreakToken pageId="35" pageNumber="36" start="start">IOC</pageBreakToken>
|
||
col., 1 ex.; XI/1941, Com IOC col., 2 ex. (CEIOC); Porto Murtinho, "Faz. Campo Florido", 11/XII/2012, M Savaris and S Lampert col., 2 ex. (DZUP). Rio Verde [Rio Verde do Mato Grosso], XI/1963, 400 m, A. Maller, 2 ex (DZUP); "Zona da NOB Oswaldo Cruz Correntes" [Dois
|
||
<normalizedToken originalValue="Irmãos">Irmaos</normalizedToken>
|
||
do Burito], 17/X/1938, without collector, 4 ex. (CEIOC); SP:
|
||
<normalizedToken originalValue="Jundiaí">Jundiai</normalizedToken>
|
||
, 10/II/2012, Castro col., 1 ex.; 30/XII/2011, Castro col., 1 ex. (CEMT); Campinas, without date and collector, 1 ex. (ZMHB); RS:
|
||
<normalizedToken originalValue="São">Sao</normalizedToken>
|
||
Francisco de Paula,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-29.422888">29°25'22.4"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-50.386448">50°23'11.2"W</geoCoordinate>
|
||
, 10/XII/2009, P. Grossi col., 1 ex. (CEMT). ARGENTINA. SA: Aguaray, XII/1959, A. Martinez col., 1 ex. (MZSP); FO: Laguna Yema,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-24.35582">24°21'20,95"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-61.315342">61°18'55,23"W</geoCoordinate>
|
||
, 11/XII/2008, Ocampo, San Blas, Campon cols., 5 ex. (IADIZA); Rio Pilcomayo, 9/X/1906, Hermann J. col., 1 ex. (ZMHB); SE: without date, Wagner col., 4 ex. (MLPA); TU, IX/1900, C. Bruch col., 1 ex. (MACN); CH: Asustado, without date, Richter col., 1 ex. (MLPA), Resistencia, XII/1961, A. Martinez col., 2 ex. (CMNC); CR: Bella Vista, IV/1931, without collector, 1 ex. (MLPA); CO:
|
||
<normalizedToken originalValue="“Córdoba">"Cordoba</normalizedToken>
|
||
, Casa Hernandez", XI/1965, A. Martinez col., 2 ex. (CMNC); Rio Cuarto, without date, Bremer col., 1 ex. (ZMHB).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="35" pageNumber="36" type="diagnosis">
|
||
<paragraph pageId="35" pageNumber="36">Diagnosis.</paragraph>
|
||
<paragraph pageId="35" pageNumber="36">Body, pronotum and elytra purplish brown or dark brown; elongate; distance between eyes slightly more than twice the width of one eye; frons slightly longer than clypeus; clypeal lateral margin convex, with a projection that varies from rounded to sharp; pronotal convexity at sides distinct (Fig. 64D); pronotal posterior corners sharp, obtuse-angled; prothorax scaly posteriorly (Fig. 64A); pro-, meso- and metasternum, pro- and metacoxae scaly (Figs 64B, E); male mesotibia sub-quadrate in cross section; inner margin of male metatibia abruptly sub-basally or medially produced towards apex; protarsi, male claws unequal, symmetrical in females; basal metatarsomere one-half the length of tarsomere II and wider; pygidium noticeable convex and wide, pygidial width exceeding distance between spiracles of propygidium; total length of parameres more than five times the length of their apex; inner margins slightly convergent; apex harpoon-like with lateral angle projecting straight downward (Fig. 64G). In lateral view parameres concave (Fig. 64G).</paragraph>
|
||
<caption pageId="35" pageNumber="36">
|
||
<paragraph pageId="35" pageNumber="36">
|
||
Figure 64.
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys fusca" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="fusca">Liogenys fusca</taxonomicName>
|
||
Blanchard. A Dorsal view B Lateral view C Frontal view D Clypeus and pronotum E Pygidium F Male protarsal claw G Parameres, dorsal view H. Parameres, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="redescription">
|
||
<paragraph pageId="35" pageNumber="36">Redescription.</paragraph>
|
||
<paragraph lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
|
||
Length: 12.4-14.0 mm; width: 6.5-7.6 mm. Purplish brown. Head: distance between eyes slightly more than twice the width of one eye; frons slightly longer than clypeus; clypeal emargination rounded, shallow and narrow; outer sides of anterior teeth sub-parallel or parallel; clypeal lateral margin convex, with a projection that varies from rounded to sharp; distance between clypeal lateral projection and anterior margin of eye slightly longer than one eye; distance between clypeal lateral projection and anterior tooth longer than basal width of anterior tooth; obtuse angle between outer side of anterior teeth and clypeal lateral projection; canthus not exceeding the outer margin of the eye; distal maxillary palpomere, maximum width more than twice width of apex; fovea deep and oval, extending past the transverse midline of the palpomere; labium transversely carinated, as wide as it is long; antenna 10-articulated, lamellae lighter in color than flagellum and in males lamellae longer than the flagellum. Thorax: anterior margin of pronotum slightly produced medially; maximum length of pronotum exceeding the length of tarsomeres I, II and III together; disc glabrous, punctures fine and dense, smooth at posterior longitudinal midline; pronotal
|
||
<pageBreakToken pageId="36" pageNumber="37" start="start">convexity</pageBreakToken>
|
||
at sides distinct; pronotal posterior corners sharp, obtuse-angled, in some cases almost right-angled; prothorax scaly posteriorly; proepisternum with long bristles and scales; mesepisternum scaly, as are the sides of metasternum, also with few long bristles on the anterior margin; distance between meso- and metacoxae up to twice longer than the metacoxa; scutellum ogival, finely punctured. Elytra: shiny, glabrous, uniform dark brown to purplish, elytra more than three times longer than the pronotum; elytral suture and elytron unicolored, distinctly elevated; pair of inner ridges more noticeable than the other three outer pairs. Legs: procoxa scaly on infra-carinal and outer surface; punctures visible at 12
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
magnification; three protibial teeth, middle and apical equal in size, distance between basal and middle teeth slightly longer than between middle and apical; protibial inner apical spur present; mesofemural disc setose, with a row of long bristles on anterior and posterior margins; mesotibia sub-quadrate in cross section in males, cylindrical in females; disc coarsely sculptured, two mesotibial transverse carinae, the apical one complete; basal apophysis of metacoxa produced beyond the outer margin of trochanter; metatibial apical spurs of different lengths, the longest one shorter than the diameter of the tibial apex; inner margin of male metatibia carinated and abruptly sub-basally produced towards apex, apical inner surface setose; metatibial disc finely sculptured in males, coarsely sculptured in females; two metatibial transverse carinae present posteriorly; basal metatarsomere one-half the length of tarsomere II and wider, in males protarsomere II long; pro- and mesotarsomeres I to IV enlarged, protarsomeres slightly wider than the mesotarsomeres and more than twice as wide as metatarsi; protarsal claws asymmetrical in males, symmetrically bifid in females; superior tooth longer and as wide as the inferior; distance between teeth as long as the inferior tooth. Abdomen: band of abundant scales visible at the lowest magnification beneath the outer margin of elytra; ventrites bristled on disc; propygidium visible, glabrous; pygidium noticeable convex, sub-quadrate, wide, pygidial width exceeding distance between spiracles of propygidium; pygidial disc bristled only on apex, coarsely punctured, pygidial apex quadrate. Parameres: width of basal region equal to the parameres together at its transverse midline, parameral split at 2/3; total length of parameres more than five times the length of their apex; inner margins slightly convergent; apex harpoon-like with lateral angle projecting straight downward (Fig. 64G). In lateral view parameres concave (Fig. 64H).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="36" pageNumber="37" type="type-locality">
|
||
<paragraph pageId="36" pageNumber="37">Type-locality.</paragraph>
|
||
<paragraph pageId="36" pageNumber="37">
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys fuscus" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="fuscus">Liogenys fuscus</taxonomicName>
|
||
: Rives de
|
||
<normalizedToken originalValue="l’Uruguay">l'Uruguay</normalizedToken>
|
||
[today Rio Grande do Sul, Brazil];
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys argentinus" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="argentinus">Liogenys argentinus</taxonomicName>
|
||
: ARGENTINA. Santiago del Estero (syn.);
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys cuyabanus" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="cuyabanus">Liogenys cuyabanus</taxonomicName>
|
||
: BRAZIL.
|
||
<normalizedToken originalValue="Cuiabá">Cuiaba</normalizedToken>
|
||
, Mato Grosso (syn.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="36" pageNumber="37" type="distribution">
|
||
<paragraph pageId="36" pageNumber="37">Geographical distribution.</paragraph>
|
||
<paragraph pageId="36" pageNumber="37">BRAZIL (TO, BA, MG, DF, GO, MT, MS, SP, RS); PARAGUAY (BQ, PH); BOLIVIA (Santa Cruz, Tarija); ARGENTINA (SA, TU, FO, CH, SE, CR, CO).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="remarks">
|
||
<paragraph pageId="36" pageNumber="37">Remarks.</paragraph>
|
||
<paragraph lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys fusca" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="fusca">Liogenys fusca</taxonomicName>
|
||
resembles
|
||
<taxonomicName lsidName="L. bidentata" pageId="36" pageNumber="37" rank="species" species="bidentata">L. bidentata</taxonomicName>
|
||
(Fig. 57) and
|
||
<taxonomicName lsidName="L. pallidicornis" pageId="36" pageNumber="37" rank="species" species="pallidicornis">L. pallidicornis</taxonomicName>
|
||
(Fig. 67).
|
||
<bibRefCitation author="Cherman, MA" journalOrPublisher="Systematic Entomology" pageId="104" pageNumber="105" pagination="744 - 770" title="Phylogenetic relationships within Diplotaxini Kirby (Coleoptera: Melolonthidae: Melolonthinae) with emphasis on Liogenys Guerin-Meneville." url="https://doi.org/10.1111/syen.12188" volume="41" year="2016">Cherman et al. (2016)</bibRefCitation>
|
||
support that those species are closely related.
|
||
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys fusca" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="fusca">Liogenys fusca</taxonomicName>
|
||
differs from the other two in the quadridentate clypeus, being more evident in males; clypeal emargination rounded; elytral suture distinctly elevated; elytral pair of inner ridges more noticeable than the outers; pygidial disc swollen; basal metatar
|
||
<pageBreakToken pageId="37" pageNumber="38" start="start">somere</pageBreakToken>
|
||
one-half the length of tarsomere II and wider; inner margin of male metatibia abruptly sub-basally produced towards apex, claws of protarsi asymmetrical, one bifid and one simple (Fig. 64F) and parameres concave.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |