treatments-xml/data/5D/2A/C7/5D2AC71CFF83FFF0FF4EF487F653FB57.xml

<document id="2F3AAC3AFCFCC6331304F48EC7A72483" ID-CLB-Dataset="71410" ID-DOI="10.11646/zootaxa.5150.4.6" ID-GBIF-Dataset="04240c5e-3517-479f-95bd-512bf8fc087c" ID-ISSN="1175-5326" ID-Zenodo-Dep="6626882" ID-ZooBank="D2F51EDC-7929-4112-93DF-A69983BFF4A4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1654761210179" checkinUser="plazi" docAuthor="Perger, Robert &amp; Pett, Brogan L." docDate="2022" docId="5D2AC71CFF83FFF0FF4EF487F653FB57" docLanguage="en" docName="zootaxa.5150.4.6.pdf" docOrigin="Zootaxa 5150 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mazax akephaloi Perger &amp; Pett 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="586" masterDocId="A113BF64FF80FFF7FFD9F410F23BFFEE" masterDocTitle="Mazax akephaloi sp. nov. - a new Neotropical spider species resembling ‘ headless’ Ectatomma ants (Araneae: Corinnidae: Castianeirinae)" masterLastPageNumber="590" masterPageNumber="579" pageNumber="582" updateTime="1699339233048" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="851B25D52168B8E0251E9A538EFA8B6B">Mazax akephaloi sp. nov. - a new Neotropical spider species resembling ‘ headless’ Ectatomma ants (Araneae: Corinnidae: Castianeirinae)</mods:title>
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<mods:name id="5967587E06B8EF40A9030336AA0D7207" type="personal">
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<mods:namePart id="FDDE68012361CFFACBE5BDAF0DBA3892">Perger, Robert</mods:namePart>
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<mods:affiliation id="87A916F8C0B75F437F70B8060ECBA1B9">Colección Boliviana de Fauna, La Paz, Bolivia &amp; robertperger @ hotmail. com; https: // orcid. org / 0000 - 0001 - 9930 - 9638</mods:affiliation>
<mods:nameIdentifier id="805862CEAF15D6C2C9B0825B84E2A26B" type="email">robertperger@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="8BADAFA71AAE7E9C938572DB114A4089">Pett, Brogan L.</mods:namePart>
<mods:nameIdentifier id="6E8C2CA2B05699074D64AE48D033EF16" type="ORCID">0000-0002-0461-3715</mods:nameIdentifier>
<mods:affiliation id="DF97557C308C31A5D8D25FB4AA9D1F9C">SpiDivErse, Biodiversity Inventory for Conservation (BINCO), Walmersumstraat 44, 3380 Glabbeek, Belgium &amp; brogan. pett @ outlook. com; https: // orcid. org / 0000 - 0002 - 0461 - 3715 &amp; Centre for Ecology and Conservation, College of Life and Environmental Sciences, University of Exeter, Penryn Campus, Penryn, Cornwall, TR 10 9 FE, U. K. &amp; brogan. pett @ outlook. com; https: // orcid. org / 0000 - 0002 - 0461 - 3715</mods:affiliation>
<mods:nameIdentifier id="FAC02AB6F56775420A984D90A22B69DE" type="email">brogan.pett@outlook.com</mods:nameIdentifier>
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<mods:date id="4313B2CEE4568E7E848505BCB64D1D6E">2022</mods:date>
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<treatment id="5D2AC71CFF83FFF0FF4EF487F653FB57" ID-DOI="http://doi.org/10.5281/zenodo.6638462" ID-GBIF-Taxon="195865206" ID-Zenodo-Dep="6638462" LSID="urn:lsid:plazi:treatment:5D2AC71CFF83FFF0FF4EF487F653FB57" httpUri="http://treatment.plazi.org/id/5D2AC71CFF83FFF0FF4EF487F653FB57" lastPageId="7" lastPageNumber="586" pageId="3" pageNumber="582">
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<taxonomicName id="12830D89FF83FFF4FF4EF487F35CFF5C" ID-CoL="999MD" authority="Perger &amp; Pett, 2022" authorityName="Perger &amp; Pett" authorityYear="2022" box="[151,359,151,178]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="582" phylum="Arthropoda" rank="species" species="akephaloi" status="sp. nov.">
<emphasis id="E7F7AA18FF83FFF4FF4EF487F35CFF5C" bold="true" box="[151,359,151,178]" italics="true" pageId="3" pageNumber="582">Mazax akephaloi</emphasis>
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urn:lsid:zoobank.org:act: 
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<paragraph id="D53C760AFF83FFF4FF4EF4EEF2C2FEF7" blockId="3.[151,914,219,281]" box="[151,249,254,281]" pageId="3" pageNumber="582">
<figureCitation id="4DB86A8FFF83FFF4FF4EF4EEF2C2FEF7" box="[151,249,254,281]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="3.[151,250,1694,1718]" captionStartId-1="4.[151,250,1918,1942]" captionStartId-2="5.[151,250,1633,1657]" captionStartId-3="6.[151,250,1386,1410]" captionTargetBox-0="[173,1418,649,1676]" captionTargetBox-1="[170,1417,669,1890]" captionTargetBox-2="[170,1405,921,1610]" captionTargetBox-3="[187,1408,831,1341]" captionTargetId-0="figure-137@3.[170,1418,649,1677]" captionTargetId-1="figure-286@4.[170,1418,669,1894]" captionTargetId-2="figure-334@5.[170,1418,921,1610]" captionTargetId-3="figure-292@6.[170,1418,813,1363]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 2. Mazax akephaloi sp. nov., dorsal habitus: A, holotype male (ZMH-A0015362); B, female allotype (CBF). Scale bars 0.5 mm." captionText-1="FIGURE 3. Lateral views of Mazax akephaloi sp. nov.: A, holotype male (ZMH-A0015362); B, female allotype (CBF). Scale bars 0.5 mm." captionText-2="FIGURE 4. Mazax akephaloi sp. nov., palp male holotype (ZMH-A0015362), A, C, ventral view; B, D, retrolateral view (Em = embolus, SD = sperm duct). Scale bars 0.1 mm" captionText-3="FIGURE 5. Mazax akephaloi sp. nov., epigyne female allotype (CBF), A, dorsal view; B, ventral view (CO = copulatory opening; CD = copulatory duct; ST = spermatheca; FD = fertilization duct). Scale bar 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6626886" figureDoi-1="http://doi.org/10.5281/zenodo.6626888" figureDoi-2="http://doi.org/10.5281/zenodo.6626890" figureDoi-3="http://doi.org/10.5281/zenodo.6626892" httpUri-0="https://zenodo.org/record/6626886/files/figure.png" httpUri-1="https://zenodo.org/record/6626888/files/figure.png" httpUri-2="https://zenodo.org/record/6626890/files/figure.png" httpUri-3="https://zenodo.org/record/6626892/files/figure.png" pageId="3" pageNumber="582">Figs 2–5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="9D992581FF83FFF4FF4EF556F7B3F8A8" pageId="3" pageNumber="582" type="materials_examined">
<paragraph id="D53C760AFF83FFF4FF4EF556F6C0FE6B" blockId="3.[151,1436,325,640]" pageId="3" pageNumber="582">
<emphasis id="E7F7AA18FF83FFF4FF4EF556F37CFEB1" bold="true" box="[151,327,325,352]" pageId="3" pageNumber="582">Type material.</emphasis>
<materialsCitation id="65EB7C57FF83FFF4FE89F556F6C0FE6B" ID-GBIF-Occurrence="3807271302" collectingDate="2019-12-21" collectingDateMax="2019-12-28" collectingDateMin="2019-12-21" collectorName="de la Colina &amp; R. Perger" country="Bolivia" elevation="432" latitude="-17.760834" location="Chiquitano forest" longLatPrecision="529" longitude="-63.24" municipality="Urubo" pageId="3" pageNumber="582" specimenCount="1" specimenCount-male="1" stateProvince="Santa Cruz" typeStatus="holotype">
<typeStatus id="0A38C8A8FF83FFF4FE89F556F383FE8E" box="[336,440,326,352]" pageId="3" pageNumber="582" type="holotype">Holotype</typeStatus>
<specimenCount id="C385BD83FF83FFF4FE1BF556F3E1FE8E" box="[450,474,326,352]" pageId="3" pageNumber="582" type="male">♂</specimenCount>
: 
<emphasis id="E7F7AA18FF83FFF4FE31F555F051FEB1" bold="true" box="[488,618,325,351]" pageId="3" pageNumber="582">
<collectingCountry id="AD94369AFF83FFF4FE31F555F05DFEB1" box="[488,614,325,351]" name="Bolivia" pageId="3" pageNumber="582">BOLIVIA</collectingCountry>
:
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<collectingRegion id="1747B8E8FF83FFF4FDADF556F147FE8E" box="[628,892,326,352]" country="Bolivia" name="Santa Cruz" pageId="3" pageNumber="582">Santa Cruz department</collectingRegion>
, 
<collectingRegion id="1747B8E8FF83FFF4FC50F556F63DFE8E" box="[905,1030,326,352]" country="Bolivia" name="Santa Cruz" pageId="3" pageNumber="582">Santa Cruz</collectingRegion>
<collectorName id="787613DCFF83FFF4FBC9F556F6A5FE8E" box="[1040,1182,326,352]" pageId="3" pageNumber="582">de la Colina</collectorName>
, 
<collectingMunicipality id="3558EC70FF83FFF4FB75F556F6CDFE8E" box="[1196,1270,326,352]" pageId="3" pageNumber="582">Urubo</collectingMunicipality>
, 
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, 
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, 
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<quantity id="127BDBEFFF83FFF4FEDEF57AF375FE6A" box="[263,334,362,388]" metricMagnitude="2" metricUnit="m" metricValue="4.32" pageId="3" pageNumber="582" unit="m" value="432.0">432 m</quantity>
a.s.l.
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, 
<date id="A13D50CAFF83FFF4FE4EF57AF076FE6A" box="[407,589,362,388]" pageId="3" pageNumber="582" value="2019-12-21" valueMax="2019-12-28" valueMin="2019-12-21">
<collectingDate id="B179A922FF83FFF4FE4EF57AF076FE6A" box="[407,589,362,388]" pageId="3" pageNumber="582" value="2019-12-21" valueMax="2019-12-28" valueMin="2019-12-21">21–28.XII.2019</collectingDate>
</date>
, leg. 
<collectorName id="787613DCFF83FFF4FD51F57AF0C8FE6A" box="[648,755,362,388]" pageId="3" pageNumber="582">R. Perger</collectorName>
, edge of 
<location id="D05C20D1FF83FFF4FC80F57AF627FE6A" LSID="urn:lsid:plazi:treatment:5D2AC71CFF83FFF0FF4EF487F653FB57:D05C20D1FF83FFF4FC80F57AF627FE6A" box="[857,1052,362,388]" country="Bolivia" latitude="-17.760834" longLatPrecision="529" longitude="-63.24" municipality="Urubo" name="Chiquitano forest" pageId="3" pageNumber="582" stateProvince="Santa Cruz">Chiquitano forest</location>
(ZMH-A0015362).
</materialsCitation>
</paragraph>
<paragraph id="D53C760AFF83FFF4FF4EF59EF09EFE46" blockId="3.[151,1436,325,640]" box="[151,677,398,424]" pageId="3" pageNumber="582">
<materialsCitation id="65EB7C57FF83FFF4FF4EF59EF09EFE46" ID-GBIF-Occurrence="3807271307" box="[151,677,398,424]" collectionCode="CBF" pageId="3" pageNumber="582" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
<typeStatus id="0A38C8A8FF83FFF4FF4EF59EF2C2FE46" box="[151,249,398,424]" pageId="3" pageNumber="582" type="allotype">Allotype</typeStatus>
<specimenCount id="C385BD83FF83FFF4FED9F59EF32CFE46" box="[256,279,398,424]" pageId="3" pageNumber="582" type="female">♀</specimenCount>
: same data as for preceding (
<collectionCode id="B392EECFFF83FFF4FD87F59EF0A3FE46" box="[606,664,398,424]" country="Bolivia" httpUri="http://grbio.org/cool/ky5m-b42n" name="Coleccion Boliviana de Fauna" pageId="3" pageNumber="582">CBF</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="D53C760AFF83FFF4FF1EF5A2F0E6FD6E" blockId="3.[151,1436,325,640]" pageId="3" pageNumber="582">
<materialsCitation id="65EB7C57FF83FFF4FF1EF5A2F1B4FE23" ID-GBIF-Occurrence="3807271306" box="[199,911,434,461]" pageId="3" pageNumber="582" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="0A38C8A8FF83FFF4FF1EF5A2F303FE22" box="[199,312,434,460]" pageId="3" pageNumber="582" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="C385BD83FF83FFF4FE9FF5A2F351FE22" box="[326,362,434,460]" pageId="3" pageNumber="582" type="male">1♂</specimenCount>
, same data as for preceding (ZMH-A0015360)
</materialsCitation>
; 
<materialsCitation id="65EB7C57FF83FFF4FC44F5A2F7ACFE22" ID-GBIF-Occurrence="3807271305" box="[925,1431,434,460]" collectionCode="CBF" pageId="3" pageNumber="582" specimenCount="16" specimenCount-female="10" specimenCount-male="6" typeStatus="paratype">
<specimenCount id="C385BD83FF83FFF4FC44F5A2F1FBFE22" box="[925,960,434,460]" pageId="3" pageNumber="582" type="male">6♂</specimenCount>
<specimenCount id="C385BD83FF83FFF4FC13F5A2F1C6FE22" box="[970,1021,434,460]" pageId="3" pageNumber="582" type="female">10♀</specimenCount>
, same data as for preceding (
<collectionCode id="B392EECFFF83FFF4FA8CF5A2F7B4FE22" box="[1365,1423,434,460]" country="Bolivia" httpUri="http://grbio.org/cool/ky5m-b42n" name="Coleccion Boliviana de Fauna" pageId="3" pageNumber="582">CBF</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="65EB7C57FF83FFF4FF4EF5C6F083FDFD" ID-GBIF-Occurrence="3807271303" collectingDate="2015-09-09" collectionCode="CBF" collectorName="La Guardia &amp; R. Perger" country="Paraguay" elevation="480" latitude="-17.883" location="Chiquitano forest" longLatPrecision="7" longitude="-63.3177" municipality="Santa Cruz" pageId="3" pageNumber="582" specimenCount="7" specimenCount-female="4" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="C385BD83FF83FFF4FF4EF5C6F280FE1E" box="[151,187,470,496]" pageId="3" pageNumber="582" type="male">3♂</specimenCount>
<specimenCount id="C385BD83FF83FFF4FF1EF5C6F2D0FE1E" box="[199,235,470,496]" pageId="3" pageNumber="582" type="female">4♀</specimenCount>
, 
<collectingMunicipality id="3558EC70FF83FFF4FF24F5C6F346FE1E" box="[253,381,470,496]" pageId="3" pageNumber="582">Santa Cruz</collectingMunicipality>
department, 
<collectorName id="787613DCFF83FFF4FDC5F5C6F099FE1E" box="[540,674,470,496]" pageId="3" pageNumber="582">La Guardia</collectorName>
, 
<geoCoordinate id="B0B710CDFF83FFF4FD6AF5C6F110FE1E" box="[691,811,470,496]" degrees="17.8830" direction="south" orientation="latitude" pageId="3" pageNumber="582" precision="5" value="-17.883">17.8830°S</geoCoordinate>
, 
<geoCoordinate id="B0B710CDFF83FFF4FCE5F5C6F186FE1E" box="[828,957,470,496]" degrees="63.3177" direction="west" orientation="longitude" pageId="3" pageNumber="582" precision="5" value="-63.3177">63.3177°W</geoCoordinate>
, 
<elevation id="5EAE9139FF83FFF4FC14F5C6F667FE1E" box="[973,1116,470,496]" metricMagnitude="2" metricUnit="m" metricValue="4.8" pageId="3" pageNumber="582" unit="m" value="480.0">
<quantity id="127BDBEFFF83FFF4FC14F5C6F621FE1E" box="[973,1050,470,496]" metricMagnitude="2" metricUnit="m" metricValue="4.8" pageId="3" pageNumber="582" unit="m" value="480.0">480 m</quantity>
a.s.l.
</elevation>
, 
<date id="A13D50CAFF83FFF4FBB4F5C6F6DAFE1F" box="[1133,1249,470,497]" pageId="3" pageNumber="582" value="2015-09-09">
<collectingDate id="B179A922FF83FFF4FBB4F5C6F6DAFE1F" box="[1133,1249,470,497]" pageId="3" pageNumber="582" value="2015-09-09">9.IX.2015</collectingDate>
</date>
, leg. 
<collectorName id="787613DCFF83FFF4FAF1F5C6F7A3FE1E" box="[1320,1432,470,496]" pageId="3" pageNumber="582">R. Perger</collectorName>
, edge of 
<location id="D05C20D1FF83FFF4FF2CF5EAF381FDFA" LSID="urn:lsid:plazi:treatment:5D2AC71CFF83FFF0FF4EF487F653FB57:D05C20D1FF83FFF4FF2CF5EAF381FDFA" box="[245,442,506,532]" country="Paraguay" latitude="-17.883" longLatPrecision="7" longitude="-63.3177" municipality="Santa Cruz" name="Chiquitano forest" pageId="3" pageNumber="582">Chiquitano forest</location>
(
<collectionCode id="B392EECFFF83FFF4FE12F5EAF03EFDFA" box="[459,517,506,532]" country="Bolivia" httpUri="http://grbio.org/cool/ky5m-b42n" name="Coleccion Boliviana de Fauna" pageId="3" pageNumber="582">CBF</collectionCode>
). 
<collectingCountry id="AD94369AFF83FFF4FDC3F5E9F083FDFD" box="[538,696,505,531]" name="Paraguay" pageId="3" pageNumber="582">
<emphasis id="E7F7AA18FF83FFF4FDC3F5E9F083FDFD" bold="true" box="[538,696,505,531]" pageId="3" pageNumber="582">PARAGUAY</emphasis>
</collectingCountry>
</materialsCitation>
<emphasis id="E7F7AA18FF83FFF4FD6EF5E9F087FDFD" bold="true" box="[695,700,505,531]" pageId="3" pageNumber="582">:</emphasis>
<materialsCitation id="65EB7C57FF83FFF4FD1CF5EAF35AFDB2" ID-GBIF-Occurrence="3807271308" collectingDate="1982-09-06" collectionCode="MNHNP" collectorName="J. A. Kochalka" country="Paraguay" county="Parque Nacional Defensores del Chaco" latitude="-20.1227" location="Tribu Nueva" longLatPrecision="7" longitude="-60.3246" municipality="Mision Cue" pageId="3" pageNumber="582" specimenCount="1" specimenCount-male="1" stateProvince="Alto Paraguay" typeStatus="paratype">
<specimenCount id="C385BD83FF83FFF4FD1CF5EAF0D2FDFA" box="[709,745,506,532]" pageId="3" pageNumber="582" type="male">1♂</specimenCount>
, 
<collectingRegion id="1747B8E8FF83FFF4FD2FF5EAF619FDFA" box="[758,1058,506,532]" country="Paraguay" name="Alto Paraguay" pageId="3" pageNumber="582">Alto Paraguay department</collectingRegion>
, 
<collectingCounty id="3C5D0E86FF83FFF4FBF6F5EAF2DAFDD6" pageId="3" pageNumber="582">Parque Nacional Defensores del Chaco</collectingCounty>
, 
<collectingMunicipality id="3558EC70FF83FFF4FF36F60EF34CFDD6" box="[239,375,542,568]" pageId="3" pageNumber="582">Misión Cué</collectingMunicipality>
, 
<location id="D05C20D1FF83FFF4FE5DF60EF02CFDD6" LSID="urn:lsid:plazi:treatment:5D2AC71CFF83FFF0FF4EF487F653FB57:D05C20D1FF83FFF4FE5DF60EF02CFDD6" box="[388,535,542,568]" country="Paraguay" county="Parque Nacional Defensores del Chaco" latitude="-20.1227" longLatPrecision="7" longitude="-60.3246" municipality="Mision Cue" name="Tribu Nueva" pageId="3" pageNumber="582" stateProvince="Alto Paraguay">Tribu Nueva</location>
, 
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, 
<geoCoordinate id="B0B710CDFF83FFF4FD73F60EF110FDD6" box="[682,811,542,568]" degrees="60.3246" direction="west" orientation="longitude" pageId="3" pageNumber="582" precision="5" value="-60.3246">60.3246°W</geoCoordinate>
, 
<date id="A13D50CAFF83FFF4FCE1F60EF197FDD6" box="[824,940,542,568]" pageId="3" pageNumber="582" value="1982-09-06">
<collectingDate id="B179A922FF83FFF4FCE1F60EF197FDD6" box="[824,940,542,568]" pageId="3" pageNumber="582" value="1982-09-06">6.IX.1982</collectingDate>
</date>
, leg. 
<collectorName id="787613DCFF83FFF4FC35F60EF6B5FDD6" box="[1004,1166,542,568]" pageId="3" pageNumber="582">J.A. Kochalka</collectorName>
(
<collectionCode id="B392EECFFF83FFF4FB46F60EF731FDD6" box="[1183,1290,542,568]" country="0" httpUri="http://grbio.org/cool/bmqw-2x87" name="Museo Nacional de Historia Natural del Paraguay" pageId="3" pageNumber="582">MNHNP</collectionCode>
: IBNP-JAK-CR 000.00.2.717)
</materialsCitation>
; 
<materialsCitation id="65EB7C57FF83FFF4FEABF652F0E2FD6E" ID-GBIF-Occurrence="3807271304" collectingDate="1983-12-11" collectingDateMax="1983-12-12" collectingDateMin="1983-12-11" collectionCode="MNHNP" collectorName="J. A. Kochalka" country="Paraguay" latitude="-22.924" location="Laguas" longLatPrecision="75" longitude="-59.486" pageId="3" pageNumber="582" specimenCount="1" specimenCount-male="1" stateProvince="Presidente Hayes" typeStatus="paratype">
<specimenCount id="C385BD83FF83FFF4FEABF652F3ACFDB2" box="[370,407,578,604]" pageId="3" pageNumber="582" type="male">1♂</specimenCount>
; 
<collectingRegion id="1747B8E8FF83FFF4FE7FF652F0C8FDB2" box="[422,755,578,604]" country="Paraguay" name="Presidente Hayes" pageId="3" pageNumber="582">Presidente Hayes department</collectingRegion>
, 25 
<location id="D05C20D1FF83FFF4FCFDF652F143FDB2" LSID="urn:lsid:plazi:treatment:5D2AC71CFF83FFF0FF4EF487F653FB57:D05C20D1FF83FFF4FCFDF652F143FDB2" box="[804,888,578,604]" country="Paraguay" latitude="-22.924" longLatPrecision="75" longitude="-59.486" name="Laguas" pageId="3" pageNumber="582" stateProvince="Presidente Hayes">Laguas</location>
, 
<geoCoordinate id="B0B710CDFF83FFF4FC5DF652F1D4FDB2" box="[900,1007,578,604]" degrees="22.924" direction="south" orientation="latitude" pageId="3" pageNumber="582" precision="55" value="-22.924">22.924°S</geoCoordinate>
, 
<geoCoordinate id="B0B710CDFF83FFF4FC25F652F654FDB3" box="[1020,1135,578,605]" degrees="59.486" direction="west" orientation="longitude" pageId="3" pageNumber="582" precision="55" value="-59.486">59.486°W</geoCoordinate>
, 
<date id="A13D50CAFF83FFF4FBA5F652F70BFDB2" box="[1148,1328,578,604]" pageId="3" pageNumber="582" value="1983-12-11" valueMax="1983-12-12" valueMin="1983-12-11">
<collectingDate id="B179A922FF83FFF4FBA5F652F70BFDB2" box="[1148,1328,578,604]" pageId="3" pageNumber="582" value="1983-12-11" valueMax="1983-12-12" valueMin="1983-12-11">11–12.XII.1983</collectingDate>
</date>
, leg. 
<collectorName id="787613DCFF83FFF4FAB6F652F339FD6E" pageId="3" pageNumber="582">J.A. Kochalka</collectorName>
(
<collectionCode id="B392EECFFF83FFF4FEC8F676F347FD6E" box="[273,380,614,640]" country="0" httpUri="http://grbio.org/cool/bmqw-2x87" name="Museo Nacional de Historia Natural del Paraguay" pageId="3" pageNumber="582">MNHNP</collectionCode>
: IBNP-JAK-CR 000.00.2.719)
</materialsCitation>
.
</paragraph>
<caption id="81FC2682FF83FFF4FF4EF28EF327F937" ID-DOI="http://doi.org/10.5281/zenodo.6626886" ID-Zenodo-Dep="6626886" httpUri="https://zenodo.org/record/6626886/files/figure.png" pageId="3" pageNumber="582" startId="3.[151,250,1694,1718]" targetBox="[173,1418,649,1676]" targetPageId="3">
<paragraph id="D53C760AFF83FFF4FF4EF28EF327F937" blockId="3.[151,1436,1694,1754]" pageId="3" pageNumber="582">
<emphasis id="E7F7AA18FF83FFF4FF4EF28EF32FF958" bold="true" box="[151,276,1694,1718]" pageId="3" pageNumber="582">FIGURE 2.</emphasis>
<taxonomicName id="12830D89FF83FFF4FEC2F28FF3F1F958" authority="Perger &amp; Pett, 2022" authorityName="Perger &amp; Pett" authorityYear="2022" box="[283,458,1694,1718]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="582" phylum="Arthropoda" rank="species" species="akephaloi" status="sp. nov.">
<emphasis id="E7F7AA18FF83FFF4FEC2F28FF3F1F958" box="[283,458,1694,1718]" italics="true" pageId="3" pageNumber="582">Mazax akephaloi</emphasis>
</taxonomicName>
<emphasis id="E7F7AA18FF83FFF4FE08F28EF018F958" bold="true" box="[465,547,1694,1718]" pageId="3" pageNumber="582">
<taxonomicNameLabel id="FCC41763FF83FFF4FE08F28EF018F958" box="[465,547,1694,1718]" pageId="3" pageNumber="582" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, dorsal habitus: A, holotype male (
<emphasis id="E7F7AA18FF83FFF4FC54F28EF1E9F958" bold="true" box="[909,978,1694,1718]" pageId="3" pageNumber="582">ZMH-</emphasis>
A0015362); B, female allotype (CBF). Scale bars 0.5 mm.
</paragraph>
</caption>
<paragraph id="D53C760AFF83FFF4FF1EF318F7B3F8A8" blockId="3.[151,1437,1800,2043]" pageId="3" pageNumber="582">
<emphasis id="E7F7AA18FF83FFF4FF1EF318F3C7F8CC" bold="true" box="[199,508,1800,1826]" pageId="3" pageNumber="582">Other material examined.</emphasis>
<materialsCitation id="65EB7C57FF83FFF4FDDAF318F7BFF8A8" ID-GBIF-Occurrence="3807271301" collectingDate="1982-09-01" collectingDateMax="1982-09-06" collectingDateMin="1982-09-01" collectionCode="MNHNP" collectorName="J. A. Kochalka" country="Paraguay" county="Tribu Nueva" latitude="-20.1227" location="Parque Nacional Defensores del Chaco" longLatPrecision="7" longitude="-60.3246" municipality="Mision Cue" pageId="3" pageNumber="582" specimenCount="2" specimenCount-male="1" specimenCount-subadult="1">
<specimenCount id="C385BD83FF83FFF4FDDAF318F04CF8CC" box="[515,631,1800,1826]" pageId="3" pageNumber="582" type="subadult">1 subadult</specimenCount>
<specimenCount id="C385BD83FF83FFF4FDA7F319F0ADF8CD" box="[638,662,1801,1827]" pageId="3" pageNumber="582" type="male">♂</specimenCount>
: 
<emphasis id="E7F7AA18FF83FFF4FD7BF318F17EF8CC" bold="true" box="[674,837,1800,1826]" pageId="3" pageNumber="582">
<collectingCountry id="AD94369AFF83FFF4FD7BF318F17BF8CC" box="[674,832,1800,1826]" name="Paraguay" pageId="3" pageNumber="582">PARAGUAY</collectingCountry>
:
</emphasis>
<location id="D05C20D1FF83FFF4FC95F319F73DF8CC" LSID="urn:lsid:plazi:treatment:5D2AC71CFF83FFF0FF4EF487F653FB57:D05C20D1FF83FFF4FC95F319F73DF8CC" box="[844,1286,1800,1827]" country="Paraguay" county="Tribu Nueva" latitude="-20.1227" longLatPrecision="7" longitude="-60.3246" municipality="Mision Cue" name="Parque Nacional Defensores del Chaco" pageId="3" pageNumber="582">Parque Nacional Defensores del Chaco</location>
, 
<collectingMunicipality id="3558EC70FF83FFF4FACBF318F7A3F8CC" box="[1298,1432,1800,1826]" pageId="3" pageNumber="582">Misión Cué</collectingMunicipality>
, 
<collectingCounty id="3C5D0E86FF83FFF4FF4EF33CF31CF8A9" box="[151,295,1836,1863]" pageId="3" pageNumber="582">Tribu Nueva</collectingCounty>
, 
<geoCoordinate id="B0B710CDFF83FFF4FEEBF33CF390F8A8" box="[306,427,1836,1862]" degrees="20.1227" direction="south" orientation="latitude" pageId="3" pageNumber="582" precision="5" value="-20.1227">20.1227°S</geoCoordinate>
, 
<geoCoordinate id="B0B710CDFF83FFF4FE6FF33CF00CF8A8" box="[438,567,1836,1862]" degrees="60.3246" direction="west" orientation="longitude" pageId="3" pageNumber="582" precision="5" value="-60.3246">60.3246°W</geoCoordinate>
, 
<date id="A13D50CAFF83FFF4FD9BF33CF0E9F8A8" box="[578,722,1836,1862]" pageId="3" pageNumber="582" value="1982-09-01" valueMax="1982-09-06" valueMin="1982-09-01">
<collectingDate id="B179A922FF83FFF4FD9BF33CF0E9F8A8" box="[578,722,1836,1862]" pageId="3" pageNumber="582" value="1982-09-01" valueMax="1982-09-06" valueMin="1982-09-01">1–6.IX.1982</collectingDate>
</date>
, leg. 
<collectorName id="787613DCFF83FFF4FCD7F33CF196F8A8" box="[782,941,1836,1862]" pageId="3" pageNumber="582">J.A. Kochalka</collectorName>
(
<collectionCode id="B392EECFFF83FFF4FC65F33CF61CF8A8" box="[956,1063,1836,1862]" country="0" httpUri="http://grbio.org/cool/bmqw-2x87" name="Museo Nacional de Historia Natural del Paraguay" pageId="3" pageNumber="582">MNHNP</collectionCode>
: IBNP-JAK-CR 000.00.2.718)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9D992581FF83FFF3FF1EF340F702FEAF" lastPageId="4" lastPageNumber="583" pageId="3" pageNumber="582" type="diagnosis">
<paragraph id="D53C760AFF83FFF4FF1EF340F091F814" blockId="3.[151,1437,1800,2043]" pageId="3" pageNumber="582">
<emphasis id="E7F7AA18FF83FFF4FF1EF340F379F884" bold="true" box="[199,322,1872,1898]" pageId="3" pageNumber="582">Diagnosis.</emphasis>
<taxonomicName id="12830D89FF83FFF4FE95F341F034F884" authority="Perger &amp; Pett, 2022" authorityName="Perger &amp; Pett" authorityYear="2022" box="[332,527,1872,1898]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="582" phylum="Arthropoda" rank="species" species="akephaloi" status="sp. nov.">
<emphasis id="E7F7AA18FF83FFF4FE95F341F034F884" box="[332,527,1872,1898]" italics="true" pageId="3" pageNumber="582">Mazax akephaloi</emphasis>
</taxonomicName>
<emphasis id="E7F7AA18FF83FFF4FDC3F340F04DF884" bold="true" box="[538,630,1872,1898]" pageId="3" pageNumber="582">
<taxonomicNameLabel id="FCC41763FF83FFF4FDC3F340F04DF884" box="[538,630,1872,1898]" pageId="3" pageNumber="582" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be separated from all known congeners by a combination of the following characters: tibia I ventral spination 5– 
<quantity id="127BDBEFFF83FFF4FD19F364F0D5F860" box="[704,750,1908,1935]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="3" pageNumber="582" unit="in" value="4.0">4 in</quantity>
males (5 at prolateral margin) and 5– 
<quantity id="127BDBEFFF83FFF4FB7DF364F6EBF860" box="[1188,1232,1908,1935]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="3" pageNumber="582" unit="in" value="5.0">5 in</quantity>
females, feathery setae on carapace, and embolus sub-apical with spatulate extension (males of all known congeners have the embolus tapering evenly, either twisted or straight, see 
<bibRefCitation id="B1120BFBFF83FFF4FD75F3ACF14BF839" author="Reiskind, J." box="[684,880,1980,2007]" pageId="3" pageNumber="582" pagination="163 - 325" refId="ref7181" refString="Reiskind, J. (1969) The spider subfamily Castianeirinae of North and Central America (Araneae, Clubionidae). Bulletin of the Museum of Comparative Zoology, 138, 163 - 325." type="journal article" year="1969">Reiskind (1969))</bibRefCitation>
. Females can be diagnosed by a combination of lung-shaped ST II and slightly undulating CDs.
</paragraph>
<paragraph id="D53C760AFF84FFF3FF1EF487F702FEAF" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="583">
White feathery setae on the carapace are only shared with 
<taxonomicName id="12830D89FF84FFF3FC8FF487F19EFF5E" authorityName="Reiskind" authorityYear="1969" box="[854,933,151,176]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="pax">
<emphasis id="E7F7AA18FF84FFF3FC8FF487F19EFF5E" box="[854,933,151,176]" italics="true" pageId="4" pageNumber="583">M. pax</emphasis>
</taxonomicName>
(tibia I spination 3–3) and 
<taxonomicName id="12830D89FF84FFF3FB01F487F761FF5E" authorityName="Rubio &amp; Danisman" authorityYear="2014" box="[1240,1370,151,176]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="ramirezi">
<emphasis id="E7F7AA18FF84FFF3FB01F487F761FF5E" box="[1240,1370,151,176]" italics="true" pageId="4" pageNumber="583">M. ramirezi</emphasis>
</taxonomicName>
(tibia I spination 4–4) (
<bibRefCitation id="B1120BFBFF84FFF3FE8FF4ABF3C7FF38" author="Reiskind, J." box="[342,508,187,214]" pageId="4" pageNumber="583" pagination="163 - 325" refId="ref7181" refString="Reiskind, J. (1969) The spider subfamily Castianeirinae of North and Central America (Araneae, Clubionidae). Bulletin of the Museum of Comparative Zoology, 138, 163 - 325." type="journal article" year="1969">Reiskind 1969</bibRefCitation>
). 
<taxonomicName id="12830D89FF84FFF3FDC9F4ABF167FF3B" authority="(Simon, 1898)" baseAuthorityName="Simon" baseAuthorityYear="1898" box="[528,860,187,213]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis id="E7F7AA18FF84FFF3FDC9F4ABF08EFF3A" box="[528,693,187,212]" italics="true" pageId="4" pageNumber="583">Mazax spinosa</emphasis>
(Simon, 1898)
</taxonomicName>
and 
<taxonomicName id="12830D89FF84FFF3FC4DF4ABF694FF3B" authority="Reiskind, 1969" authorityName="Reiskind" authorityYear="1969" box="[916,1199,187,213]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="xerxes">
<emphasis id="E7F7AA18FF84FFF3FC4DF4ABF63BFF3A" box="[916,1024,187,212]" italics="true" pageId="4" pageNumber="583">M. xerxes</emphasis>
<bibRefCitation id="B1120BFBFF84FFF3FBD1F4ABF694FF3B" author="Reiskind, J." box="[1032,1199,187,213]" pageId="4" pageNumber="583" pagination="163 - 325" refId="ref7181" refString="Reiskind, J. (1969) The spider subfamily Castianeirinae of North and Central America (Araneae, Clubionidae). Bulletin of the Museum of Comparative Zoology, 138, 163 - 325." type="journal article" year="1969">Reiskind, 1969</bibRefCitation>
</taxonomicName>
have a tibia I ventral spination of 5–5, but no feathery setae on the carapace (
<bibRefCitation id="B1120BFBFF84FFF3FD05F4CFF145FF14" author="Reiskind, J." box="[732,894,223,250]" pageId="4" pageNumber="583" pagination="163 - 325" refId="ref7181" refString="Reiskind, J. (1969) The spider subfamily Castianeirinae of North and Central America (Araneae, Clubionidae). Bulletin of the Museum of Comparative Zoology, 138, 163 - 325." type="journal article" year="1969">Reiskind 1969</bibRefCitation>
).The lung-shaped ST II of females of 
<taxonomicName id="12830D89FF84FFF3FAD7F4CFF7A7FF17" authority="Perger &amp; Pett, 2022" authorityName="Perger &amp; Pett" authorityYear="2022" box="[1294,1436,223,249]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="akephaloi" status="sp. nov.">
<emphasis id="E7F7AA18FF84FFF3FAD7F4CFF7A7FF17" box="[1294,1436,223,249]" italics="true" pageId="4" pageNumber="583">M.akephaloi</emphasis>
</taxonomicName>
<emphasis id="E7F7AA18FF84FFF3FF4EF513F2D6FEF3" bold="true" box="[151,237,259,285]" pageId="4" pageNumber="583">
<taxonomicNameLabel id="FCC41763FF84FFF3FF4EF513F2D6FEF3" box="[151,237,259,285]" pageId="4" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are only shared with 
<taxonomicName id="12830D89FF84FFF3FE0BF513F06AFEF2" authorityName="Rubio &amp; Danisman" authorityYear="2014" box="[466,593,259,284]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="ramirezi">
<emphasis id="E7F7AA18FF84FFF3FE0BF513F06AFEF2" box="[466,593,259,284]" italics="true" pageId="4" pageNumber="583">M. ramirezi</emphasis>
</taxonomicName>
(CDs more twisted) and the nearly straight CDs with 
<taxonomicName id="12830D89FF84FFF3FB48F513F2F5FEAF" authority="Reiskind, 1969" authorityName="Reiskind" authorityYear="1969" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="chickeringi">
<emphasis id="E7F7AA18FF84FFF3FB48F513F714FEF3" box="[1169,1327,259,285]" italics="true" pageId="4" pageNumber="583">M. chickeringi</emphasis>
<bibRefCitation id="B1120BFBFF84FFF3FAEDF513F2F5FEAF" author="Reiskind, J." pageId="4" pageNumber="583" pagination="163 - 325" refId="ref7181" refString="Reiskind, J. (1969) The spider subfamily Castianeirinae of North and Central America (Araneae, Clubionidae). Bulletin of the Museum of Comparative Zoology, 138, 163 - 325." type="journal article" year="1969">Reiskind, 1969</bibRefCitation>
</taxonomicName>
(ST II globose), but neither have both characters combined (
<bibRefCitation id="B1120BFBFF84FFF3FCBFF537F631FEAF" author="Reiskind, J." box="[870,1034,295,321]" pageId="4" pageNumber="583" pagination="163 - 325" refId="ref7181" refString="Reiskind, J. (1969) The spider subfamily Castianeirinae of North and Central America (Araneae, Clubionidae). Bulletin of the Museum of Comparative Zoology, 138, 163 - 325." type="journal article" year="1969">Reiskind 1969</bibRefCitation>
; Rubio &amp; Danişman 2014).
</paragraph>
</subSubSection>
<subSubSection id="9D992581FF84FFF3FF1EF55BF6BEFD6B" pageId="4" pageNumber="583" type="discussion">
<paragraph id="D53C760AFF84FFF3FF1EF55BF6BEFD6B" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="583">
<emphasis id="E7F7AA18FF84FFF3FF1EF55BF300FE8B" bold="true" box="[199,315,331,357]" pageId="4" pageNumber="583">Remarks.</emphasis>
<taxonomicName id="12830D89FF84FFF3FE9AF55BF049FE8B" authorityName="Simon" authorityYear="1896" box="[323,626,331,357]" class="Arachnida" family="Corinnidae" genus="Apochinomma" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="acanthaspis">
<emphasis id="E7F7AA18FF84FFF3FE9AF55BF049FE8B" box="[323,626,331,357]" italics="true" pageId="4" pageNumber="583">Apochinomma acanthaspis</emphasis>
</taxonomicName>
and 
<taxonomicName id="12830D89FF84FFF3FD73F55BF114FE8A" authorityName="Mello-Leitao" authorityYear="1922" box="[682,815,331,357]" class="Arachnida" family="Corinnidae" genus="Apochinomma" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="armatum">
<emphasis id="E7F7AA18FF84FFF3FD73F55BF114FE8A" box="[682,815,331,357]" italics="true" pageId="4" pageNumber="583">A. armatum</emphasis>
</taxonomicName>
possibly belong to 
<taxonomicName id="12830D89FF84FFF3FBD4F55BF663FE8A" authorityName="O. Pickard-Cambridge" authorityYear="1898" box="[1037,1112,331,356]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="583" phylum="Arthropoda" rank="genus">
<emphasis id="E7F7AA18FF84FFF3FBD4F55BF663FE8A" box="[1037,1112,331,356]" italics="true" pageId="4" pageNumber="583">Mazax</emphasis>
</taxonomicName>
but were not included in the most recent taxonomic works on this genus (
<bibRefCitation id="B1120BFBFF84FFF3FD4BF57FF100FE67" author="Reiskind, J." box="[658,827,367,393]" pageId="4" pageNumber="583" pagination="163 - 325" refId="ref7181" refString="Reiskind, J. (1969) The spider subfamily Castianeirinae of North and Central America (Araneae, Clubionidae). Bulletin of the Museum of Comparative Zoology, 138, 163 - 325." type="journal article" year="1969">Reiskind 1969</bibRefCitation>
; Rubio &amp; Danişman 2014). The types of both species were not available for study. The type of 
<taxonomicName id="12830D89FF84FFF3FDB6F583F0CEFE42" authorityName="Mello-Leitao" authorityYear="1922" box="[623,757,403,429]" class="Arachnida" family="Corinnidae" genus="Apochinomma" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="armatum">
<emphasis id="E7F7AA18FF84FFF3FDB6F583F0CEFE42" box="[623,757,403,429]" italics="true" pageId="4" pageNumber="583">A. armatum</emphasis>
</taxonomicName>
was likely destroyed in a recent fire (A. Kury, unpublished data) and the type of 
<taxonomicName id="12830D89FF84FFF3FE5EF5A7F016FE3F" authorityName="Simon" authorityYear="1896" box="[391,557,439,465]" class="Arachnida" family="Corinnidae" genus="Apochinomma" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="acanthaspis">
<emphasis id="E7F7AA18FF84FFF3FE5EF5A7F016FE3F" box="[391,557,439,465]" italics="true" pageId="4" pageNumber="583">A. acanthaspis</emphasis>
</taxonomicName>
is likely lost (C. Rollard, 
<emphasis id="E7F7AA18FF84FFF3FC8AF5A7F651FE3E" box="[851,1130,439,465]" italics="true" pageId="4" pageNumber="583">personal communication</emphasis>
). According to the original descriptions and illustration by 
<bibRefCitation id="B1120BFBFF84FFF3FDD5F5CBF095FE18" author="Simon, E." box="[524,686,475,502]" pageId="4" pageNumber="583" pagination="400 - 422" refId="ref7384" refString="Simon, E. (1896) Descriptions d'arachnides nouveaux de la famille des Clubionidae. Annales de la Societe Entomologique de Belgique, 40, 400 - 422." type="journal article" year="1896">Simon (1896)</bibRefCitation>
, the 
<typeStatus id="0A38C8A8FF84FFF3FD37F5CBF16BFE1B" box="[750,848,475,501]" pageId="4" pageNumber="583" type="holotype">holotype</typeStatus>
female of 
<taxonomicName id="12830D89FF84FFF3FC0EF5CBF6BAFE1B" authorityName="Simon" authorityYear="1896" box="[983,1153,475,501]" class="Arachnida" family="Corinnidae" genus="Apochinomma" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="acanthaspis">
<emphasis id="E7F7AA18FF84FFF3FC0EF5CBF6BAFE1B" box="[983,1153,475,501]" italics="true" pageId="4" pageNumber="583">A. acanthaspis</emphasis>
</taxonomicName>
has a considerably less pronounced abdominal constriction and flatter ventral sclerite, lacks the first pair of abdominal setae (present in 
<taxonomicName id="12830D89FF84FFF3FF4EF633F314FDD3" authority="Perger &amp; Pett, 2022" authorityName="Perger &amp; Pett" authorityYear="2022" box="[151,303,547,573]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="akephaloi" status="sp. nov.">
<emphasis id="E7F7AA18FF84FFF3FF4EF633F314FDD3" box="[151,303,547,573]" italics="true" pageId="4" pageNumber="583">M. akephaloi</emphasis>
</taxonomicName>
<emphasis id="E7F7AA18FF84FFF3FEE3F633F3A3FDD3" bold="true" box="[314,408,547,573]" pageId="4" pageNumber="583">
<taxonomicNameLabel id="FCC41763FF84FFF3FEE3F633F3A3FDD3" box="[314,408,547,573]" pageId="4" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), and has the metatarsus I with spine formula 3–3 (2– 
<quantity id="127BDBEFFF84FFF3FBC5F633F670FDD3" box="[1052,1099,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="583" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="12830D89FF84FFF3FB8FF633F6D6FDD3" authority="Perger &amp; Pett, 2022" authorityName="Perger &amp; Pett" authorityYear="2022" box="[1110,1261,547,573]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="akephaloi" status="sp. nov.">
<emphasis id="E7F7AA18FF84FFF3FB8FF633F6D6FDD3" box="[1110,1261,547,573]" italics="true" pageId="4" pageNumber="583">M. akephaloi</emphasis>
</taxonomicName>
<emphasis id="E7F7AA18FF84FFF3FB21F633F76DFDD3" bold="true" box="[1272,1366,547,573]" pageId="4" pageNumber="583">
<taxonomicNameLabel id="FCC41763FF84FFF3FB21F633F76DFDD3" box="[1272,1366,547,573]" pageId="4" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). The female 
<typeStatus id="0A38C8A8FF84FFF3FF33F657F377FD8F" box="[234,332,583,609]" pageId="4" pageNumber="583" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="12830D89FF84FFF3FEA8F657F3CEFD8E" authorityName="Mello-Leitao" authorityYear="1922" box="[369,501,583,609]" class="Arachnida" family="Corinnidae" genus="Apochinomma" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="armatum">
<emphasis id="E7F7AA18FF84FFF3FEA8F657F3CEFD8E" box="[369,501,583,609]" italics="true" pageId="4" pageNumber="583">A. armatum</emphasis>
</taxonomicName>
has a tibia I spine formula of 2–2 and whitish coxae II–III (
<bibRefCitation id="B1120BFBFF84FFF3FB49F657F75EFD8F" author="Mello-Leitao, C. F. de" box="[1168,1381,583,609]" pageId="4" pageNumber="583" pagination="17 - 56" refId="ref6287" refString="Mello-Leitao, C. F. de (1922) Novas clubionidas do Brasil. Archivos da Escola Superior de Agricultura e Medicina Veterinaria, Rio de Janeiro, 6, 17 - 56." type="journal article" year="1922">Mello-Leitão 1922</bibRefCitation>
) (
<taxonomicName id="12830D89FF84FFF3FAA7F657F33EFD6B" authority="Perger &amp; Pett, 2022" authorityName="Perger &amp; Pett" authorityYear="2022" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="akephaloi" status="sp. nov.">
<emphasis id="E7F7AA18FF84FFF3FAA7F657F33EFD6B" italics="true" pageId="4" pageNumber="583">M. akephaloi</emphasis>
</taxonomicName>
<emphasis id="E7F7AA18FF84FFF3FED4F67BF35DFD6B" bold="true" box="[269,358,619,645]" pageId="4" pageNumber="583">
<taxonomicNameLabel id="FCC41763FF84FFF3FED4F67BF35DFD6B" box="[269,358,619,645]" pageId="4" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
with tibia I with spine formula of 5– 
<quantity id="127BDBEFFF84FFF3FCDCF67AF114FD6B" box="[773,815,618,645]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="4" pageNumber="583" unit="in" value="5.0">5 in</quantity>
females and brownish coxae).
</paragraph>
</subSubSection>
<caption id="81FC2682FF84FFF3FF4EF36EF327F854" ID-DOI="http://doi.org/10.5281/zenodo.6626888" ID-Zenodo-Dep="6626888" httpUri="https://zenodo.org/record/6626888/files/figure.png" pageId="4" pageNumber="583" startId="4.[151,250,1918,1942]" targetBox="[170,1417,669,1890]" targetPageId="4">
<paragraph id="D53C760AFF84FFF3FF4EF36EF327F854" blockId="4.[151,1436,1918,1978]" pageId="4" pageNumber="583">
<emphasis id="E7F7AA18FF84FFF3FF4EF36EF328F878" bold="true" box="[151,275,1918,1942]" pageId="4" pageNumber="583">FIGURE 3.</emphasis>
Lateral views of 
<taxonomicName id="12830D89FF84FFF3FE1DF36FF049F878" authority="Perger &amp; Pett, 2022" authorityName="Perger &amp; Pett" authorityYear="2022" box="[452,626,1918,1942]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="akephaloi" status="sp. nov.">
<emphasis id="E7F7AA18FF84FFF3FE1DF36FF049F878" box="[452,626,1918,1942]" italics="true" pageId="4" pageNumber="583">Mazax akephaloi</emphasis>
</taxonomicName>
<emphasis id="E7F7AA18FF84FFF3FDA1F36EF0F3F878" bold="true" box="[632,712,1918,1942]" pageId="4" pageNumber="583">
<taxonomicNameLabel id="FCC41763FF84FFF3FDA1F36EF0F3F878" box="[632,712,1918,1942]" pageId="4" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A, holotype male (
<emphasis id="E7F7AA18FF84FFF3FC4BF36EF1ECF878" bold="true" box="[914,983,1918,1942]" pageId="4" pageNumber="583">ZMH-</emphasis>
A0015362); B, female allotype (CBF). Scale bars 0.5 mm.
</paragraph>
</caption>
<subSubSection id="9D992581FF85FFF1FF1EF487F092FCFB" lastPageId="6" lastPageNumber="585" pageId="5" pageNumber="584" type="etymology">
<paragraph id="D53C760AFF85FFF2FF1EF487F3E6FE8B" blockId="5.[151,1437,151,897]" pageId="5" pageNumber="584">
<emphasis id="E7F7AA18FF85FFF2FF1EF487F375FF5F" bold="true" box="[199,334,151,177]" pageId="5" pageNumber="584">Etymology.</emphasis>
The specific epithet, 
<taxonomicName id="12830D89FF85FFF2FD90F487F08CFF5F" authorityName="Perger &amp; Pett" authorityYear="2022" box="[585,695,151,177]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="584" phylum="Arthropoda" rank="species" species="akephaloi">
<emphasis id="E7F7AA18FF85FFF2FD90F487F08CFF5F" box="[585,695,151,177]" italics="true" pageId="5" pageNumber="584">akephaloi</emphasis>
</taxonomicName>
, means &quot;headless ones&quot; (ἀκέφαλοι) in Greek, mythical headless men who were rumored, in antiquity and later, to inhabit remote parts of the world (
<bibRefCitation id="B1120BFBFF85FFF2FB53F4ABF76EFF3B" author="Syropoulos, S." box="[1162,1365,187,213]" pageId="5" pageNumber="584" refId="ref7442" refString="Syropoulos, S. (2018) A bestiary of monsters in Greek mythology. Oxford University Press, Oxford, 139 pp." type="book" year="2018">Syropoulos 2018</bibRefCitation>
). One hypothesis for the origin of the myth of the 
<taxonomicName id="12830D89FF85FFF2FD51F4CFF0CDFF17" authorityName="Perger &amp; Pett" authorityYear="2022" box="[648,758,223,249]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="584" phylum="Arthropoda" rank="species" species="akephaloi">
<emphasis id="E7F7AA18FF85FFF2FD51F4CFF0CDFF17" box="[648,758,223,249]" italics="true" pageId="5" pageNumber="584">akephaloi</emphasis>
</taxonomicName>
is the observation of the combat tactic of the North African Blemmyae tribe in which they keep their heads pressed close to the chest (
<bibRefCitation id="B1120BFBFF85FFF2FC05F513F646FEF3" author="Dijkstra, J. H. F." box="[988,1149,259,285]" pageId="5" pageNumber="584" pagination="1145 - 1146" refId="ref5965" refString="Dijkstra, J. H. F. (2013) Blemmyes. In: Erskine, A., Bagnall, R. S., Brodersen, K., Champion, C. B. &amp; Huebner, S. R. (Eds.), The Encyclopedia of Ancient History. Wiley, pp. 1145 - 1146. https: // doi. org / 10.1002 / 9781444338386" type="book chapter" year="2013">Dijkstra 2013</bibRefCitation>
). The epithet refers to the observation that 
<taxonomicName id="12830D89FF85FFF2FE8DF537F3D3FEAF" authority="Perger &amp; Pett, 2022" authorityName="Perger &amp; Pett" authorityYear="2022" box="[340,488,295,321]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="584" phylum="Arthropoda" rank="species" species="akephaloi" status="sp. nov.">
<emphasis id="E7F7AA18FF85FFF2FE8DF537F3D3FEAF" box="[340,488,295,321]" italics="true" pageId="5" pageNumber="584">M. akephaloi</emphasis>
</taxonomicName>
<emphasis id="E7F7AA18FF85FFF2FE29F537F070FEAF" bold="true" box="[496,587,295,321]" pageId="5" pageNumber="584">
<taxonomicNameLabel id="FCC41763FF85FFF2FE29F537F070FEAF" box="[496,587,295,321]" pageId="5" pageNumber="584" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
lack a structure resembling the head of their ant model 
<taxonomicName id="12830D89FF85FFF2FB1FF537F757FEAE" authorityName="Forel" authorityYear="1908" box="[1222,1388,295,320]" class="Insecta" family="Formicidae" genus="Ectatomma" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="584" phylum="Arthropoda" rank="species" species="permagnum">
<emphasis id="E7F7AA18FF85FFF2FB1FF537F757FEAE" box="[1222,1388,295,320]" italics="true" pageId="5" pageNumber="584">E. permagnum</emphasis>
</taxonomicName>
and have a skull-shaped sternum.
</paragraph>
<paragraph id="D53C760AFF85FFF2FF1EF57FF652FE1B" blockId="5.[151,1437,151,897]" pageId="5" pageNumber="584">
<emphasis id="E7F7AA18FF85FFF2FF1EF57FF341FE67" bold="true" box="[199,378,367,393]" pageId="5" pageNumber="584">
Male 
<typeStatus id="0A38C8A8FF85FFF2FED5F57FF34EFE67" box="[268,373,367,393]" pageId="5" pageNumber="584" type="holotype">holotype</typeStatus>
.
</emphasis>
Body length 6.79; carapace length 2.96, width 1.5, carapace index 50.6; cephalic width 0.84, cephalic index 55.85; abdomen length 3.32, maximum width anterior part 1.21, maximum width posterior part 1.39, abdominal index 41.8; dorsal sclerite length 2.66, maximum width same as maximum abdomen width. Eyes: AER 0.55; AME–AME 0.09; AME–ALE 0.02; PER 0.64; PME–PME 0.14; PME–PLE 0.07.
</paragraph>
<paragraph id="D53C760AFF85FFF2FF1EF5EEF3E8FD1F" blockId="5.[151,1437,151,897]" pageId="5" pageNumber="584">
<emphasis id="E7F7AA18FF85FFF2FF1EF5EEF3C5FDF6" bold="true" box="[199,510,510,537]" pageId="5" pageNumber="584">Color and microsculpture.</emphasis>
Dorsum dark blackish-brown in life, with purplish tinge when seen in sunlight (color faded to reddish-brown in ethanol; 
<figureCitation id="4DB86A8FFF85FFF2FDF5F633F0B2FDD3" box="[556,649,547,573]" captionStart="FIGURE 2" captionStartId="3.[151,250,1694,1718]" captionTargetBox="[173,1418,649,1676]" captionTargetId="figure-137@3.[170,1418,649,1677]" captionTargetPageId="3" captionText="FIGURE 2. Mazax akephaloi sp. nov., dorsal habitus: A, holotype male (ZMH-A0015362); B, female allotype (CBF). Scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6626886" httpUri="https://zenodo.org/record/6626886/files/figure.png" pageId="5" pageNumber="584">Figs 2A</figureCitation>
, 
<figureCitation id="4DB86A8FFF85FFF2FD4EF633F086FDD3" box="[663,701,547,573]" captionStart="FIGURE 3" captionStartId="4.[151,250,1918,1942]" captionTargetBox="[170,1417,669,1890]" captionTargetId="figure-286@4.[170,1418,669,1894]" captionTargetPageId="4" captionText="FIGURE 3. Lateral views of Mazax akephaloi sp. nov.: A, holotype male (ZMH-A0015362); B, female allotype (CBF). Scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6626888" httpUri="https://zenodo.org/record/6626888/files/figure.png" pageId="5" pageNumber="584">3A</figureCitation>
); carapace and posterior part of sclerite posterior of constriction weakly shiny, smooth, microsculpture finely reticulate, with evenly distributed, fine pits; petiole and anterior part of sclerite heavily wrinkled and shiny, wrinkles on petiole transverse and on anterior sclerite longitudinal, abdomen posterior of dorsal sclerite glabrous, shiny; legs glabrous, shiny, with regularly arranged narrow, transverse ridges from which emerge setae, dark brown; femora I–II translucent, yellowish to white prolateral to ventrally; tarsi I–IV cream with dark brown tips.
</paragraph>
<paragraph id="D53C760AFF85FFF2FF1EF6EAF64FFC6F" blockId="5.[151,1437,151,897]" pageId="5" pageNumber="584">
<emphasis id="E7F7AA18FF85FFF2FF1EF6EAF30BFCFA" bold="true" box="[199,304,762,788]" pageId="5" pageNumber="584">Setation.</emphasis>
Dorsum with separate white feathery setae, forming dense transverse band in abdominal constriction; separate, erect, simple, moderately long yellowish-brassy setae all over dorsum, denser and longer on posterior part of abdomen, similar simple and feathery setae on legs. First pair of abdominal setae simple, indistinct, second pair of abdominal spines heavily sclerotized, emerging from two distinct tubercles (
<figureCitation id="4DB86A8FFF85FFF2FBD4F777F653FC6F" box="[1037,1128,871,897]" captionStart="FIGURE 4" captionStartId="5.[151,250,1633,1657]" captionTargetBox="[170,1405,921,1610]" captionTargetId="figure-334@5.[170,1418,921,1610]" captionTargetPageId="5" captionText="FIGURE 4. Mazax akephaloi sp. nov., palp male holotype (ZMH-A0015362), A, C, ventral view; B, D, retrolateral view (Em = embolus, SD = sperm duct). Scale bars 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.6626890" httpUri="https://zenodo.org/record/6626890/files/figure.png" pageId="5" pageNumber="584">Fig. 4A</figureCitation>
).
</paragraph>
<caption id="81FC2682FF85FFF2FF4EF271F0B2F973" ID-DOI="http://doi.org/10.5281/zenodo.6626890" ID-Zenodo-Dep="6626890" httpUri="https://zenodo.org/record/6626890/files/figure.png" pageId="5" pageNumber="584" startId="5.[151,250,1633,1657]" targetBox="[170,1405,921,1610]" targetPageId="5">
<paragraph id="D53C760AFF85FFF2FF4EF271F0B2F973" blockId="5.[151,1436,1633,1694]" pageId="5" pageNumber="584">
<emphasis id="E7F7AA18FF85FFF2FF4EF271F328F994" bold="true" box="[151,275,1633,1658]" pageId="5" pageNumber="584">FIGURE 4.</emphasis>
<taxonomicName id="12830D89FF85FFF2FEC0F272F3FCF994" authority="Perger &amp; Pett, 2022" authorityName="Perger &amp; Pett" authorityYear="2022" box="[281,455,1634,1658]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="584" phylum="Arthropoda" rank="species" species="akephaloi" status="sp. nov.">
<emphasis id="E7F7AA18FF85FFF2FEC0F272F3FCF994" box="[281,455,1634,1658]" italics="true" pageId="5" pageNumber="584">Mazax akephaloi</emphasis>
</taxonomicName>
<emphasis id="E7F7AA18FF85FFF2FE14F272F025F994" bold="true" box="[461,542,1634,1658]" pageId="5" pageNumber="584">
<taxonomicNameLabel id="FCC41763FF85FFF2FE14F272F025F994" box="[461,542,1634,1658]" pageId="5" pageNumber="584" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, palp male holotype (
<emphasis id="E7F7AA18FF85FFF2FD25F272F17AF994" bold="true" box="[764,833,1634,1658]" pageId="5" pageNumber="584">ZMH-</emphasis>
A0015362), A, C, ventral view; B, D, retrolateral view (Em = embolus, SD = sperm duct). Scale bars 0.1 mm
</paragraph>
</caption>
<paragraph id="D53C760AFF85FFF2FF1EF2DCF6DBF8E4" blockId="5.[151,1437,1740,2018]" pageId="5" pageNumber="584">
<emphasis id="E7F7AA18FF85FFF2FF1EF2DCF37AF908" bold="true" box="[199,321,1740,1766]" pageId="5" pageNumber="584">Carapace.</emphasis>
Long pyriform, front slightly convex, cephalic area laterally somewhat narrowed, broadening towards middle, widest in middle, narrowing posteriorly, posterior margin truncate (
<figureCitation id="4DB86A8FFF85FFF2FB9DF2E0F69AF8E4" box="[1092,1185,1776,1802]" captionStart="FIGURE 2" captionStartId="3.[151,250,1694,1718]" captionTargetBox="[173,1418,649,1676]" captionTargetId="figure-137@3.[170,1418,649,1677]" captionTargetPageId="3" captionText="FIGURE 2. Mazax akephaloi sp. nov., dorsal habitus: A, holotype male (ZMH-A0015362); B, female allotype (CBF). Scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6626886" httpUri="https://zenodo.org/record/6626886/files/figure.png" pageId="5" pageNumber="584">Figs 2A</figureCitation>
, 
<figureCitation id="4DB86A8FFF85FFF2FB74F2E0F6E8F8E4" box="[1197,1235,1776,1802]" captionStart="FIGURE 3" captionStartId="4.[151,250,1918,1942]" captionTargetBox="[170,1417,669,1890]" captionTargetId="figure-286@4.[170,1418,669,1894]" captionTargetPageId="4" captionText="FIGURE 3. Lateral views of Mazax akephaloi sp. nov.: A, holotype male (ZMH-A0015362); B, female allotype (CBF). Scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6626888" httpUri="https://zenodo.org/record/6626888/files/figure.png" pageId="5" pageNumber="584">3A</figureCitation>
).
</paragraph>
<paragraph id="D53C760AFF85FFF2FF1EF304F03EF8BC" blockId="5.[151,1437,1740,2018]" pageId="5" pageNumber="584">
<emphasis id="E7F7AA18FF85FFF2FF1EF304F33DF8C0" bold="true" box="[199,262,1812,1838]" pageId="5" pageNumber="584">Eyes.</emphasis>
Both eye rows slightly recurved, eyes approximately equal, with narrow, slightly darker rings, remaining characters as in genus diagnosis.
</paragraph>
<paragraph id="D53C760AFF85FFF2FF1EF34CF6E2F874" blockId="5.[151,1437,1740,2018]" pageId="5" pageNumber="584">
<emphasis id="E7F7AA18FF85FFF2FF1EF34CF375F898" bold="true" box="[199,334,1884,1910]" pageId="5" pageNumber="584">Chelicerae.</emphasis>
Two teeth on both the pro- and retromargins. Promargin with distal tooth about twice the size of basal tooth. Retromargin with two subequal teeth, slightly smaller than largest promarginal tooth.
</paragraph>
<paragraph id="D53C760AFF85FFF2FF1EF3B4F70AF850" blockId="5.[151,1437,1740,2018]" box="[199,1329,1956,1982]" pageId="5" pageNumber="584">
<emphasis id="E7F7AA18FF85FFF2FF1EF3B4F30DF850" bold="true" box="[199,310,1956,1982]" pageId="5" pageNumber="584">Sternum.</emphasis>
Skull-shaped, narrowing posteriorly with conspicuous indentation just anterior to coxa III.
</paragraph>
<paragraph id="D53C760AFF85FFF1FF1EF3D8F18FFEF3" blockId="5.[151,1437,1740,2018]" lastBlockId="6.[151,1437,151,789]" lastPageId="6" lastPageNumber="585" pageId="5" pageNumber="584">
<emphasis id="E7F7AA18FF85FFF2FF1EF3D8F378F80C" bold="true" box="[199,323,1992,2018]" pageId="5" pageNumber="584">Abdomen.</emphasis>
Petiole conspicuous, elongated, with anterior margin concave; abdomen roughly obovate, distinctly constricted dorsally and laterally at about middle, anterior part dorsally bulged out to transverse elliptic bead, posterior part orbicular, broader than anterior part; dorsal sclerite completely covering abdomen laterally, sclerite length 80% of abdomen length, slightly convex posteriorly; ventral sclerite not reaching to level of inframamillary sclerite, latter narrow, subrectangular, broader than long (
<figureCitation id="4DB86A8FFF86FFF1FCC1F513F14EFEF3" box="[792,885,259,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1694,1718]" captionTargetBox="[173,1418,649,1676]" captionTargetId="figure-137@3.[170,1418,649,1677]" captionTargetPageId="3" captionText="FIGURE 2. Mazax akephaloi sp. nov., dorsal habitus: A, holotype male (ZMH-A0015362); B, female allotype (CBF). Scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6626886" httpUri="https://zenodo.org/record/6626886/files/figure.png" pageId="6" pageNumber="585">Figs 2A</figureCitation>
, 
<figureCitation id="4DB86A8FFF86FFF1FC58F513F19CFEF3" box="[897,935,259,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1918,1942]" captionTargetBox="[170,1417,669,1890]" captionTargetId="figure-286@4.[170,1418,669,1894]" captionTargetPageId="4" captionText="FIGURE 3. Lateral views of Mazax akephaloi sp. nov.: A, holotype male (ZMH-A0015362); B, female allotype (CBF). Scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6626888" httpUri="https://zenodo.org/record/6626888/files/figure.png" pageId="6" pageNumber="585">3A</figureCitation>
).
</paragraph>
<paragraph id="D53C760AFF86FFF1FF1EF537F7A0FE43" blockId="6.[151,1437,151,789]" pageId="6" pageNumber="585">
<emphasis id="E7F7AA18FF86FFF1FF1EF537F33FFEAF" bold="true" box="[199,260,295,321]" pageId="6" pageNumber="585">Palp.</emphasis>
Tibia with two distinct, long setae and several shorter setae, margin distinctly edged, RTA absent; maximum width of tibia 58% of maximum width of bulb when viewed retrolaterally; tarsus with globose genital bulb drawn out into moderately long, broad neck, with short, sclerotized embolus, sub-apical with spatulate extension, embolus ending in pointed tip with apical twist; sperm ducts with two loops, one retrolateral and one median to embolus tube (
<figureCitation id="4DB86A8FFF86FFF1FA93F583F7B5FE43" box="[1354,1422,403,429]" captionStart="FIGURE 4" captionStartId="5.[151,250,1633,1657]" captionTargetBox="[170,1405,921,1610]" captionTargetId="figure-334@5.[170,1418,921,1610]" captionTargetPageId="5" captionText="FIGURE 4. Mazax akephaloi sp. nov., palp male holotype (ZMH-A0015362), A, C, ventral view; B, D, retrolateral view (Em = embolus, SD = sperm duct). Scale bars 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.6626890" httpUri="https://zenodo.org/record/6626890/files/figure.png" pageId="6" pageNumber="585">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="D53C760AFF86FFF1FF1EF5A7F3B4FDD3" blockId="6.[151,1437,151,789]" pageId="6" pageNumber="585">
<emphasis id="E7F7AA18FF86FFF1FF1EF5A7F3B6FE3F" bold="true" box="[199,397,439,465]" pageId="6" pageNumber="585">
Female 
<typeStatus id="0A38C8A8FF86FFF1FEFEF5A7F3B3FE3F" box="[295,392,439,465]" pageId="6" pageNumber="585" type="allotype">allotype</typeStatus>
.
</emphasis>
Body length 6.23; carapace length 3.05, width 1.59, carapace index 52.13; cephalic width 0.89, cephalic index 56; abdomen length 2.58, width 1.58, abdominal index 61.24; dorsal sclerite length (width agrees with abdominal width) 1.26; Eyes: AER 0.58, AME–AME 0.09, AME–ALE 0.04, PER 0.69, PME–PME 0.13, PME–PLE 0.14.
</paragraph>
<paragraph id="D53C760AFF86FFF1FF1EF657F099FD47" blockId="6.[151,1437,151,789]" pageId="6" pageNumber="585">
Color, microsculpture, setation, carapace shape and eye characteristics as in male. Abdomen larger, lateral constriction of abdomen much less pronounced as in male (cf. 
<figureCitation id="4DB86A8FFF86FFF1FC8FF67BF1A5FD6B" box="[854,926,619,645]" captionStart="FIGURE 2" captionStartId="3.[151,250,1694,1718]" captionTargetBox="[173,1418,649,1676]" captionTargetId="figure-137@3.[170,1418,649,1677]" captionTargetPageId="3" captionText="FIGURE 2. Mazax akephaloi sp. nov., dorsal habitus: A, holotype male (ZMH-A0015362); B, female allotype (CBF). Scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6626886" httpUri="https://zenodo.org/record/6626886/files/figure.png" pageId="6" pageNumber="585">Figs 2</figureCitation>
, 
<figureCitation id="4DB86A8FFF86FFF1FC73F67BF182FD6B" box="[938,953,619,645]" captionStart="FIGURE 3" captionStartId="4.[151,250,1918,1942]" captionTargetBox="[170,1417,669,1890]" captionTargetId="figure-286@4.[170,1418,669,1894]" captionTargetPageId="4" captionText="FIGURE 3. Lateral views of Mazax akephaloi sp. nov.: A, holotype male (ZMH-A0015362); B, female allotype (CBF). Scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6626888" httpUri="https://zenodo.org/record/6626888/files/figure.png" pageId="6" pageNumber="585">3</figureCitation>
), dorsal sclerite suboval, only extending to abdominal constriction, ventral sclerite absent.
</paragraph>
<paragraph id="D53C760AFF86FFF1FF1EF6A2F092FCFB" blockId="6.[151,1437,151,789]" pageId="6" pageNumber="585">
<emphasis id="E7F7AA18FF86FFF1FF1EF6A2F30BFD22" bold="true" box="[199,304,690,716]" pageId="6" pageNumber="585">Epigyne.</emphasis>
ST II large, lung-shaped, about four to five times the diameter of very small circular ST I. CD joins posterior end of ST II, almost straight and projected laterally leading to small oval CO that are posterolateral to ST II. FD arises at anterior margin of ST I (
<figureCitation id="4DB86A8FFF86FFF1FD8EF6EBF0A0FCFB" box="[599,667,762,789]" captionStart="FIGURE 5" captionStartId="6.[151,250,1386,1410]" captionTargetBox="[187,1408,831,1341]" captionTargetId="figure-292@6.[170,1418,813,1363]" captionTargetPageId="6" captionText="FIGURE 5. Mazax akephaloi sp. nov., epigyne female allotype (CBF), A, dorsal view; B, ventral view (CO = copulatory opening; CD = copulatory duct; ST = spermatheca; FD = fertilization duct). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6626892" httpUri="https://zenodo.org/record/6626892/files/figure.png" pageId="6" pageNumber="585">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="81FC2682FF86FFF1FF4EF17AF66EFA48" ID-DOI="http://doi.org/10.5281/zenodo.6626892" ID-Zenodo-Dep="6626892" httpUri="https://zenodo.org/record/6626892/files/figure.png" pageId="6" pageNumber="585" startId="6.[151,250,1386,1410]" targetBox="[187,1408,831,1341]" targetPageId="6">
<paragraph id="D53C760AFF86FFF1FF4EF17AF66EFA48" blockId="6.[151,1436,1386,1446]" pageId="6" pageNumber="585">
<emphasis id="E7F7AA18FF86FFF1FF4EF17AF32DFA6C" bold="true" box="[151,278,1386,1410]" pageId="6" pageNumber="585">FIGURE 5.</emphasis>
<taxonomicName id="12830D89FF86FFF1FEC6F17BF3EBFA6C" authority="Perger &amp; Pett, 2022" authorityName="Perger &amp; Pett" authorityYear="2022" box="[287,464,1386,1410]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="akephaloi" status="sp. nov.">
<emphasis id="E7F7AA18FF86FFF1FEC6F17BF3EBFA6C" box="[287,464,1386,1410]" italics="true" pageId="6" pageNumber="585">Mazax akephaloi</emphasis>
</taxonomicName>
<emphasis id="E7F7AA18FF86FFF1FE00F17AF016FA6C" bold="true" box="[473,557,1386,1410]" pageId="6" pageNumber="585">
<taxonomicNameLabel id="FCC41763FF86FFF1FE00F17AF016FA6C" box="[473,557,1386,1410]" pageId="6" pageNumber="585" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, epigyne female allotype (CBF), A, dorsal view; B, ventral view (CO = copulatory opening; CD = copulatory duct; ST = spermatheca; FD = fertilization duct). Scale bar 0.5 mm.
</paragraph>
</caption>
<subSubSection id="9D992581FF86FFF0FF1EF1C4F653FB57" lastPageId="7" lastPageNumber="586" pageId="6" pageNumber="585" type="description">
<paragraph id="D53C760AFF86FFF1FF1EF1C4F1A1F9FD" blockId="6.[151,1437,1492,2023]" pageId="6" pageNumber="585">
<emphasis id="E7F7AA18FF86FFF1FF1EF1C4F304FA00" bold="true" box="[199,319,1492,1518]" pageId="6" pageNumber="585">Variation.</emphasis>
While the abdominal constriction of the male is determined by the shape of the large dorsal sclerite, it varied in females according to the nutritional or reproductive state.
</paragraph>
<paragraph id="D53C760AFF86FFF1FF1EF20CF7B6F929" blockId="6.[151,1437,1492,2023]" pageId="6" pageNumber="585">
<emphasis id="E7F7AA18FF86FFF1FF1EF20CF081F9D8" bold="true" box="[199,698,1564,1591]" pageId="6" pageNumber="585">Geographical and ecological distribution.</emphasis>
<taxonomicName id="12830D89FF86FFF1FD1AF20DF1BEF9D8" authority="Perger &amp; Pett, 2022" authorityName="Perger &amp; Pett" authorityYear="2022" box="[707,901,1564,1590]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="akephaloi" status="sp. nov.">
<emphasis id="E7F7AA18FF86FFF1FD1AF20DF1BEF9D8" box="[707,901,1564,1590]" italics="true" pageId="6" pageNumber="585">Mazax akephaloi</emphasis>
</taxonomicName>
<emphasis id="E7F7AA18FF86FFF1FC57F20DF1D1F9D9" bold="true" box="[910,1002,1565,1591]" pageId="6" pageNumber="585">
<taxonomicNameLabel id="FCC41763FF86FFF1FC57F20DF1D1F9D9" box="[910,1002,1565,1591]" pageId="6" pageNumber="585" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known from the Bolivian locations of La Guardia and Santa Cruz de la Colina in the Santa Cruz Department and the Paraguayan locations of Misión Cué, Tribu Nueva (
<collectingRegion id="1747B8E8FF86FFF1FEADF274F093F991" box="[372,680,1636,1663]" country="Paraguay" name="Alto Paraguay" pageId="6" pageNumber="585">Alto Paraguay Department</collectingRegion>
) and 25 Laguas (
<collectingRegion id="1747B8E8FF86FFF1FCA8F274F6F1F991" box="[881,1226,1636,1663]" country="Paraguay" name="Presidente Hayes" pageId="6" pageNumber="585">Presidente Hayes Department</collectingRegion>
). According to the ecoregion delineation by 
<bibRefCitation id="B1120BFBFF86FFF1FE69F298F0B8F94D" author="Olson, D. M. &amp; Dinerstein, E. &amp; Wikramanayake, E. D. &amp; Burgess, N. D. &amp; Powell, G. V. N. &amp; Underwood, E. C. &amp; D'Amico, J. A. &amp; Itoua, I. &amp; Strand, H. E. &amp; Morrison, J. C. &amp; Loucks, C. J. &amp; Allnutt, T. F. &amp; Ricketts, T. H. &amp; Kura, Y. &amp; Lamoreux, J. F. &amp; Wettengel, W. W. &amp; Hedao, P. &amp; Kassem, K. R." box="[432,643,1672,1699]" pageId="6" pageNumber="585" pagination="933 - 938" refId="ref6360" refString="Olson, D. M., Dinerstein, E., Wikramanayake, E. D., Burgess, N. D., Powell, G. V. N., Underwood, E. C., D'Amico, J. A., Itoua, I., Strand, H. E., Morrison, J. C., Loucks, C. J., Allnutt, T. F., Ricketts, T. H., Kura, Y., Lamoreux, J. F., Wettengel, W. W., Hedao, P. &amp; Kassem, K. R. (2001) Terrestrial Ecoregions of the World: A New Map of Life on Earth. BioScience, 51, 933 - 938." type="journal article" year="2001">
Olson 
<emphasis id="E7F7AA18FF86FFF1FE20F299F00BF94C" box="[505,560,1672,1698]" italics="true" pageId="6" pageNumber="585">et al.</emphasis>
(2001)
</bibRefCitation>
, the locations are situated in the Chiquitano dry forest (Santa Cruz de la Colina), the Chaco dry forest (La Guardia, Misión Cué, Tribu Nueva) and the Humid Chaco (25 Laguas) (
<figureCitation id="4DB86A8FFF86FFF1FAE3F2BCF747F929" box="[1338,1404,1708,1735]" captionStart="FIGURE 1" captionStartId="2.[151,250,1059,1083]" captionTargetBox="[177,1413,384,1029]" captionTargetId="figure-83@2.[173,1414,381,1035]" captionTargetPageId="2" captionText="FIGURE 1. Ecoregion distribution of Mazax akephaloi sp. nov., according to the regionalization by Olson et al. (2001); A, South America; country borders of Bolivia and Paraguay shown; B, ecoregions of M. akephaloi sp. nov.; Bolivia: Santa Cruz department: 1, Santa Cruz de la Colina; 2, La Guardia. Paraguay: 3, Alto Paraguay department, Parque Nacional Defensores del Chaco, Misión Cué, Tribu Nueva; 4, Presidente Hayes department, 25 Laguas; df, dry forest; map produced with QGIS (version 2.14.3, http:// www.qgis.org/en/site)." figureDoi="http://doi.org/10.5281/zenodo.6626884" httpUri="https://zenodo.org/record/6626884/files/figure.png" pageId="6" pageNumber="585">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="D53C760AFF86FFF1FF1EF2C1F6E3F809" blockId="6.[151,1437,1492,2023]" pageId="6" pageNumber="585">
In 
<collectingCountry id="AD94369AFF86FFF1FF3FF2C0F300F904" box="[230,315,1744,1770]" name="Bolivia" pageId="6" pageNumber="585">Bolivia</collectingCountry>
, individuals of 
<taxonomicName id="12830D89FF86FFF1FE30F2C1F046F904" authority="Perger &amp; Pett, 2022" authorityName="Perger &amp; Pett" authorityYear="2022" box="[489,637,1744,1770]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="akephaloi" status="sp. nov.">
<emphasis id="E7F7AA18FF86FFF1FE30F2C1F046F904" box="[489,637,1744,1770]" italics="true" pageId="6" pageNumber="585">M. akephaloi</emphasis>
</taxonomicName>
<emphasis id="E7F7AA18FF86FFF1FD5CF2C1F0E4F905" bold="true" box="[645,735,1745,1771]" pageId="6" pageNumber="585">
<taxonomicNameLabel id="FCC41763FF86FFF1FD5CF2C1F0E4F905" box="[645,735,1745,1771]" pageId="6" pageNumber="585" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
were observed foraging diurnally on the ground and leaf litter along the edges of Chiquitano forest fragments that were surrounded by Cerrado-like forests and savanna grass (
<figureCitation id="4DB86A8FFF86FFF1FAB6F2E4F286F8DD" captionStart="FIGURE 6" captionStartId="7.[151,250,609,633]" captionTargetBox="[170,1418,181,585]" captionTargetId="figure-17@7.[170,1418,181,585]" captionTargetPageId="7" captionText="FIGURE 6. Mazax akephaloi sp. nov.: A, habitat at edge of Chiquitano forest; B, female with captured moth; C, potential ant model Ectatomma permagnum Forel, 1908, worker." figureDoi="http://doi.org/10.5281/zenodo.6626894" httpUri="https://zenodo.org/record/6626894/files/figure.png" pageId="6" pageNumber="585">Fig. 6A</figureCitation>
). Despite several surveys employing beating tray sampling and manual search (
<bibRefCitation id="B1120BFBFF86FFF1FB90F309F77DF8DD" author="Perger, R. &amp; Perger, Y. N." box="[1097,1350,1817,1843]" pageId="6" pageNumber="585" pagination="403 - 408" refId="ref6666" refString="Perger, R. &amp; Perger, Y. N. (2017) A species richness hotspot of ant-mimicking sac spiders (Araneae: Corinnidae: Castianeirinae) at the Bolivian Orocline, with new country records for three genera and nine species. Annals of the Entomological Society of America, 110 (4), 403 - 408. https: // doi. org / 10.1093 / aesa / sax 040" type="journal article" year="2017">Perger &amp; Perger 2017</bibRefCitation>
; 
<bibRefCitation id="B1120BFBFF86FFF1FA8DF309F307F8B9" author="Perger, R. &amp; Rubio, G. D." pageId="6" pageNumber="585" pagination="0190582" refId="ref6741" refString="Perger, R. &amp; Rubio, G. D. (2018) A wolf in sheep's clothing: The description of a fly resembling jumping spider of the genus Scoturius Simon, 1901 (Araneae: Salticidae: Huriini). PLoS ONE, 13 (1), e 0190582. https: // doi. org / 10.1371 / journal. pone. 0190582" type="journal article" year="2018">Perger &amp; Rubio 2018</bibRefCitation>
, 
<bibRefCitation id="B1120BFBFF86FFF1FE91F32DF3B5F8B9" author="Perger, R. &amp; Rubio, G. D." box="[328,398,1853,1879]" pageId="6" pageNumber="585" pagination="151 - 164" refId="ref6809" refString="Perger, R. &amp; Rubio, G. D. (2020 a) Contributions to the knowledge of Neotropical ant-like spiders: Myrmecotypus tahyinandu sp. n. from Bolivian Chiquitano forest, a new country record for M. niger, and indirect evidence for species-specific mimicry (Araneae: Corinnidae: Castianeirinae). Zootaxa, 4790 (1), 151 - 164. https: // doi. org / 10.11646 / zootaxa. 4790.1.9" type="journal article" year="2020">2020a</bibRefCitation>
, b), the species was not observed in arboreal habitats or in other Bolivian forest ecoregions. Considering the distribution (
<figureCitation id="4DB86A8FFF86FFF1FE3AF370F01EF895" box="[483,549,1888,1915]" captionStart="FIGURE 1" captionStartId="2.[151,250,1059,1083]" captionTargetBox="[177,1413,384,1029]" captionTargetId="figure-83@2.[173,1414,381,1035]" captionTargetPageId="2" captionText="FIGURE 1. Ecoregion distribution of Mazax akephaloi sp. nov., according to the regionalization by Olson et al. (2001); A, South America; country borders of Bolivia and Paraguay shown; B, ecoregions of M. akephaloi sp. nov.; Bolivia: Santa Cruz department: 1, Santa Cruz de la Colina; 2, La Guardia. Paraguay: 3, Alto Paraguay department, Parque Nacional Defensores del Chaco, Misión Cué, Tribu Nueva; 4, Presidente Hayes department, 25 Laguas; df, dry forest; map produced with QGIS (version 2.14.3, http:// www.qgis.org/en/site)." figureDoi="http://doi.org/10.5281/zenodo.6626884" httpUri="https://zenodo.org/record/6626884/files/figure.png" pageId="6" pageNumber="585">Fig. 1</figureCitation>
) and observations in open habitats, this species is likely typical for Chaco dry forests. 
<taxonomicName id="12830D89FF86FFF1FF29F395F38BF870" authority="Perger &amp; Pett, 2022" authorityName="Perger &amp; Pett" authorityYear="2022" box="[240,432,1924,1950]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="akephaloi" status="sp. nov.">
<emphasis id="E7F7AA18FF86FFF1FF29F395F38BF870" box="[240,432,1924,1950]" italics="true" pageId="6" pageNumber="585">Mazax akephaloi</emphasis>
</taxonomicName>
<emphasis id="E7F7AA18FF86FFF1FE61F395F02AF871" bold="true" box="[440,529,1925,1951]" pageId="6" pageNumber="585">
<taxonomicNameLabel id="FCC41763FF86FFF1FE61F395F02AF871" box="[440,529,1925,1951]" pageId="6" pageNumber="585" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the only species of 
<taxonomicName id="12830D89FF86FFF1FCD2F395F16DF870" authorityName="O. Pickard-Cambridge" authorityYear="1898" box="[779,854,1925,1950]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="585" phylum="Arthropoda" rank="genus">
<emphasis id="E7F7AA18FF86FFF1FCD2F395F16DF870" box="[779,854,1925,1950]" italics="true" pageId="6" pageNumber="585">Mazax</emphasis>
</taxonomicName>
that is currently known from 
<collectingCountry id="AD94369AFF86FFF1FB7FF394F6C3F870" box="[1190,1272,1924,1950]" name="Bolivia" pageId="6" pageNumber="585">Bolivia</collectingCountry>
and 
<collectingCountry id="AD94369AFF86FFF1FAF6F395F7A2F871" box="[1327,1433,1925,1951]" name="Paraguay" pageId="6" pageNumber="585">Paraguay</collectingCountry>
. This species is possibly replaced by 
<taxonomicName id="12830D89FF86FFF1FDE4F3B9F0F8F82C" authorityName="Rubio &amp; Danisman" authorityYear="2014" box="[573,707,1961,1986]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="ramirezi">
<emphasis id="E7F7AA18FF86FFF1FDE4F3B9F0F8F82C" box="[573,707,1961,1986]" italics="true" pageId="6" pageNumber="585">M. ramirezi</emphasis>
</taxonomicName>
south of the 
<collectingRegion id="1747B8E8FF86FFF1FC86F3B8F19CF82C" box="[863,935,1960,1986]" country="Argentina" name="Chaco" pageId="6" pageNumber="585">Chaco</collectingRegion>
area in 
<collectingCountry id="AD94369AFF86FFF1FBDFF3B8F64EF82C" box="[1030,1141,1960,1986]" name="Argentina" pageId="6" pageNumber="585">Argentina</collectingCountry>
(
<collectingRegion id="1747B8E8FF86FFF1FB5FF3B9F7ABF82D" box="[1158,1424,1960,1987]" country="Argentina" name="Buenos Aires" pageId="6" pageNumber="585">Buenos Aires province</collectingRegion>
), making it unlikely that the latter species occurs in 
<collectingCountry id="AD94369AFF86FFF1FD12F3DCF126F808" box="[715,797,1996,2022]" name="Bolivia" pageId="6" pageNumber="585">Bolivia</collectingCountry>
(as reported by 
<bibRefCitation id="B1120BFBFF86FFF1FC0AF3DDF6F7F809" author="Perger, R. &amp; Perger, Y. N." box="[979,1228,1997,2023]" pageId="6" pageNumber="585" pagination="403 - 408" refId="ref6666" refString="Perger, R. &amp; Perger, Y. N. (2017) A species richness hotspot of ant-mimicking sac spiders (Araneae: Corinnidae: Castianeirinae) at the Bolivian Orocline, with new country records for three genera and nine species. Annals of the Entomological Society of America, 110 (4), 403 - 408. https: // doi. org / 10.1093 / aesa / sax 040" type="journal article" year="2017">Perger &amp; Perger 2017</bibRefCitation>
).
</paragraph>
<caption id="81FC2682FF87FFF0FF4EF671F09EFD73" ID-DOI="http://doi.org/10.5281/zenodo.6626894" ID-Zenodo-Dep="6626894" httpUri="https://zenodo.org/record/6626894/files/figure.png" pageId="7" pageNumber="586" startId="7.[151,250,609,633]" targetBox="[170,1418,181,585]" targetPageId="7">
<paragraph id="D53C760AFF87FFF0FF4EF671F09EFD73" blockId="7.[151,1436,609,669]" pageId="7" pageNumber="586">
<emphasis id="E7F7AA18FF87FFF0FF4EF671F328FD97" bold="true" box="[151,275,609,633]" pageId="7" pageNumber="586">FIGURE 6.</emphasis>
<taxonomicName id="12830D89FF87FFF0FEC3F672F3F2FD97" authority="Perger &amp; Pett, 2022" authorityName="Perger &amp; Pett" authorityYear="2022" box="[282,457,609,633]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="akephaloi" status="sp. nov.">
<emphasis id="E7F7AA18FF87FFF0FEC3F672F3F2FD97" box="[282,457,609,633]" italics="true" pageId="7" pageNumber="586">Mazax akephaloi</emphasis>
</taxonomicName>
<emphasis id="E7F7AA18FF87FFF0FE16F671F01BFD97" bold="true" box="[463,544,609,633]" pageId="7" pageNumber="586">
<taxonomicNameLabel id="FCC41763FF87FFF0FE16F671F01BFD97" box="[463,544,609,633]" pageId="7" pageNumber="586" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A, habitat at edge of Chiquitano forest; B, female with captured moth; C, potential ant model 
<taxonomicName id="12830D89FF87FFF0FF04F696F076FD73" authority="Forel, 1908" authorityName="Forel" authorityYear="1908" box="[221,589,645,669]" class="Insecta" family="Formicidae" genus="Ectatomma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="permagnum">
<emphasis id="E7F7AA18FF87FFF0FF04F696F3EBFD73" box="[221,464,646,669]" italics="true" pageId="7" pageNumber="586">Ectatomma permagnum</emphasis>
Forel, 1908
</taxonomicName>
, worker.
</paragraph>
</caption>
<paragraph id="D53C760AFF87FFF0FF1EF6DBF745FC98" blockId="7.[151,1437,715,1210]" pageId="7" pageNumber="586">
<emphasis id="E7F7AA18FF87FFF0FF1EF6DBF353FD0B" bold="true" box="[199,360,715,741]" pageId="7" pageNumber="586">Ant mimicry.</emphasis>
Seven ant species - 
<taxonomicName id="12830D89FF87FFF0FD8DF6DCF1CBFD0B" authority="Forel, 1908" authorityName="Forel" authorityYear="1908" box="[596,1008,715,741]" class="Insecta" family="Formicidae" genus="Ectatomma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="permagnum">
<emphasis id="E7F7AA18FF87FFF0FD8DF6DCF159FD0B" box="[596,866,716,741]" italics="true" pageId="7" pageNumber="586">Ectatomma permagnum</emphasis>
Forel, 1908
</taxonomicName>
, 
<taxonomicName id="12830D89FF87FFF0FC24F6DBF68EFD08" box="[1021,1205,715,742]" class="Insecta" family="Formicidae" genus="Acromyrmex" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="E7F7AA18FF87FFF0FC24F6DBF6B0FD0B" box="[1021,1163,715,741]" italics="true" pageId="7" pageNumber="586">Acromyrmex</emphasis>
sp.
</taxonomicName>
, 
<taxonomicName id="12830D89FF87FFF0FB1AF6DBF7ACFD08" box="[1219,1431,715,742]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="E7F7AA18FF87FFF0FB1AF6DBF756FD0B" box="[1219,1389,715,741]" italics="true" pageId="7" pageNumber="586">Odontomachus</emphasis>
sp.
</taxonomicName>
, 
<taxonomicName id="12830D89FF87FFF0FF4EF6FFF030FCE7" authority="Mayr, 1862" authorityName="Mayr" authorityYear="1862" box="[151,523,751,778]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="E7F7AA18FF87FFF0FF4EF6FFF3BBFCE7" box="[151,384,751,777]" italics="true" pageId="7" pageNumber="586">Camponotus crassus</emphasis>
Mayr, 1862
</taxonomicName>
, 
<taxonomicName id="12830D89FF87FFF0FDCFF6FFF1CCFCE7" authority="(Guerin-Meneville, 1838)" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1838" box="[534,1015,751,777]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="sericeiventris">
<emphasis id="E7F7AA18FF87FFF0FDCFF6FFF0F6FCE7" box="[534,717,751,777]" italics="true" pageId="7" pageNumber="586">C. sericeiventris</emphasis>
(Guérin-Méneville, 1838)
</taxonomicName>
, 
<taxonomicName id="12830D89FF87FFF0FBDBF6E0F7A7FCE7" authority="(Latreille, 1802)" baseAuthorityName="Latreille" baseAuthorityYear="1802" box="[1026,1436,751,777]" class="Insecta" family="Formicidae" genus="Neoponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="apicalis">
<emphasis id="E7F7AA18FF87FFF0FBDBF6E0F6E4FCE7" box="[1026,1247,751,777]" italics="true" pageId="7" pageNumber="586">Neoponera apicalis</emphasis>
(Latreille, 1802)
</taxonomicName>
and 
<taxonomicName id="12830D89FF87FFF0FF11F704F03BFCC3" authority="(Fabricius, 1804)" baseAuthorityName="Fabricius" baseAuthorityYear="1804" box="[200,512,787,813]" class="Insecta" family="Formicidae" genus="Neoponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="villosa">
<emphasis id="E7F7AA18FF87FFF0FF11F704F30EFCC3" box="[200,309,787,813]" italics="true" pageId="7" pageNumber="586">N. villosa</emphasis>
(Fabricius, 1804)
</taxonomicName>
- with a similar or larger body length than adults of 
<taxonomicName id="12830D89FF87FFF0FB83F704F6D4FCC3" authority="Perger &amp; Pett, 2022" authorityName="Perger &amp; Pett" authorityYear="2022" box="[1114,1263,787,813]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="akephaloi" status="sp. nov.">
<emphasis id="E7F7AA18FF87FFF0FB83F704F6D4FCC3" box="[1114,1263,787,813]" italics="true" pageId="7" pageNumber="586">M. akephaloi</emphasis>
</taxonomicName>
<emphasis id="E7F7AA18FF87FFF0FB21F703F768FCC3" bold="true" box="[1272,1363,787,813]" pageId="7" pageNumber="586">
<taxonomicNameLabel id="FCC41763FF87FFF0FB21F703F768FCC3" box="[1272,1363,787,813]" pageId="7" pageNumber="586" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(body length 6.23–7.24) were found in the investigated ground habitats in the two Bolivian locations. Among the grounddwelling ants, only the two 
<taxonomicName id="12830D89FF87FFF0FE17F74CF0B9FC98" box="[462,642,860,886]" class="Insecta" family="Formicidae" genus="Neoponera" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="E7F7AA18FF87FFF0FE17F74CF071FC9B" box="[462,586,860,885]" italics="true" pageId="7" pageNumber="586">Neoponera</emphasis>
spp.
</taxonomicName>
, 
<taxonomicName id="12830D89FF87FFF0FD57F74BF165FC98" box="[654,862,859,886]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="E7F7AA18FF87FFF0FD57F74BF103FC9B" box="[654,824,859,885]" italics="true" pageId="7" pageNumber="586">Odontomachus</emphasis>
sp.
</taxonomicName>
and 
<taxonomicName id="12830D89FF87FFF0FC4DF74CF601FC9B" authorityName="Forel" authorityYear="1908" box="[916,1082,860,885]" class="Insecta" family="Formicidae" genus="Ectatomma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="permagnum">
<emphasis id="E7F7AA18FF87FFF0FC4DF74CF601FC9B" box="[916,1082,860,885]" italics="true" pageId="7" pageNumber="586">E. permagnum</emphasis>
</taxonomicName>
had an elongated metasoma.
</paragraph>
<paragraph id="D53C760AFF87FFF0FF1EF76FF042FBC4" blockId="7.[151,1437,715,1210]" pageId="7" pageNumber="586">
Characters that increased the ant resemblance in 
<taxonomicName id="12830D89FF87FFF0FD36F790F1B8FC77" authority="Perger &amp; Pett, 2022" authorityName="Perger &amp; Pett" authorityYear="2022" box="[751,899,895,921]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="akephaloi" status="sp. nov.">
<emphasis id="E7F7AA18FF87FFF0FD36F790F1B8FC77" box="[751,899,895,921]" italics="true" pageId="7" pageNumber="586">M. akephaloi</emphasis>
</taxonomicName>
<emphasis id="E7F7AA18FF87FFF0FC52F76FF1DEFC77" bold="true" box="[907,997,895,921]" pageId="7" pageNumber="586">
<taxonomicNameLabel id="FCC41763FF87FFF0FC52F76FF1DEFC77" box="[907,997,895,921]" pageId="7" pageNumber="586" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but were not specific for this mimetic species pair (e.g., also found in the mimetic pair 
<taxonomicName id="12830D89FF87FFF0FD1EF7B4F10FFC53" baseAuthorityName="Fabricius" baseAuthorityYear="1804" box="[711,820,931,957]" class="Insecta" family="Formicidae" genus="Neoponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="villosa">
<emphasis id="E7F7AA18FF87FFF0FD1EF7B4F10FFC53" box="[711,820,931,957]" italics="true" pageId="7" pageNumber="586">N. villosa</emphasis>
</taxonomicName>
and 
<taxonomicName id="12830D89FF87FFF0FCB7F7B3F6D5FC53" authority="(Perger 2021))" authorityName="" baseAuthorityName="Perger" baseAuthorityYear="2021" box="[878,1262,931,957]" class="Arachnida" family="Corinnidae" genus="Sphecotypus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="E7F7AA18FF87FFF0FCB7F7B3F606FC53" box="[878,1085,931,957]" italics="true" pageId="7" pageNumber="586">Sphecotypus niger</emphasis>
(Perger 2021))
</taxonomicName>
, included long, erected yellowish-brassy setae on the abdomen, a horizontal band of hairs increasing the illusion of an abdominal segmentation between the ant post-petiole and tergite IV, and several transversal bulges posterior to the dorsal sclerite resembling the ant tergites (
<figureCitation id="4DB86A8FFF87FFF0FDFFF01FF051FBC4" box="[550,618,1039,1066]" captionStart="FIGURE 7" captionStartId="8.[151,250,1352,1376]" captionTargetBox="[202,1383,184,1315]" captionTargetId="figure-26@8.[193,1394,181,1328]" captionTargetPageId="8" captionText="FIGURE 7. Morphological resemblance between possible ant model and spider mimic: A, C, Ectatomma permagnum Forel, 1908, worker (head semitransparent because it is not imitated by the spider); B, D, Mazax akephaloi sp. nov. (B, D), male, paratype (CBF); Bu, transversal bulges; P, process; Co, constriction; T, tergites. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6626898" httpUri="https://zenodo.org/record/6626898/files/figure.png" pageId="7" pageNumber="586">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="D53C760AFF87FFF0FF1EF023F653FB57" blockId="7.[151,1437,715,1210]" pageId="7" pageNumber="586">
Ethological similarities between 
<taxonomicName id="12830D89FF87FFF0FDE6F024F0DDFBA3" authorityName="Forel" authorityYear="1908" box="[575,742,1076,1101]" class="Insecta" family="Formicidae" genus="Ectatomma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="permagnum">
<emphasis id="E7F7AA18FF87FFF0FDE6F024F0DDFBA3" box="[575,742,1076,1101]" italics="true" pageId="7" pageNumber="586">E. permagnum</emphasis>
</taxonomicName>
and 
<taxonomicName id="12830D89FF87FFF0FCFBF024F18CFBA3" authority="Perger &amp; Pett, 2022" authorityName="Perger &amp; Pett" authorityYear="2022" box="[802,951,1075,1101]" class="Arachnida" family="Corinnidae" genus="Mazax" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="akephaloi" status="sp. nov.">
<emphasis id="E7F7AA18FF87FFF0FCFBF024F18CFBA3" box="[802,951,1075,1101]" italics="true" pageId="7" pageNumber="586">M. akephaloi</emphasis>
</taxonomicName>
<emphasis id="E7F7AA18FF87FFF0FC18F023F626FBA3" bold="true" box="[961,1053,1075,1101]" pageId="7" pageNumber="586">
<taxonomicNameLabel id="FCC41763FF87FFF0FC18F023F626FBA3" box="[961,1053,1075,1101]" pageId="7" pageNumber="586" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, such as relatively slow foraging speed, with frequent stops in which the ants lifted their heads and conspicuously moved their antennae (imitated by the spiders by moving the prolegs in a similar fashion), were also observed in the mimetic pair 
<taxonomicName id="12830D89FF87FFF0FB09F06CF706FB7B" baseAuthorityName="Fabricius" baseAuthorityYear="1804" box="[1232,1341,1147,1173]" class="Insecta" family="Formicidae" genus="Neoponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="villosa">
<emphasis id="E7F7AA18FF87FFF0FB09F06CF706FB7B" box="[1232,1341,1147,1173]" italics="true" pageId="7" pageNumber="586">N. villosa</emphasis>
</taxonomicName>
/ 
<taxonomicName id="12830D89FF87FFF0FA9DF06BF309FB57" authority="(Perger 2021)" authorityName="" baseAuthorityName="Perger" baseAuthorityYear="2021" class="Arachnida" family="Corinnidae" genus="Sphecotypus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="E7F7AA18FF87FFF0FA9DF06BF7A7FB7B" box="[1348,1436,1147,1173]" italics="true" pageId="7" pageNumber="586">S. niger</emphasis>
(Perger 2021)
</taxonomicName>
and appear to be typical for poneromorph ants and their mimetic spiders.
</paragraph>
</subSubSection>
</treatment>
</document>