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<document id="53FC3D9BE923A407102B355E49B51149" ID-CLB-Dataset="21948" ID-DOI="10.5852/ejt.2017.376" ID-GBIF-Dataset="4298f5cf-d85e-48fb-afe8-f12c99ad65b3" ID-ISSN="2118-9773" ID-Zenodo-Dep="3838544" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590088443336" checkinUser="carolina" docAuthor="Aharon, Shlomi, Huber, Bernhard A. &amp; Gavish-Regev, Efrat" docDate="2017" docId="516F87BB1324BC5CFDBABF61FE9CEAF9" docLanguage="en" docName="ejt-2017-376.pdf.imf" docOrigin="European Journal of Taxonomy 376 (376)" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Artema magna Roewer 1960" docType="treatment" docVersion="9" lastPageNumber="44" masterDocId="AD56FFC3130CBC77FFD0BC5EFF85E94D" masterDocTitle="Daddy-long-leg giants: revision of the spider genus Artema Walckenaer, 1837 (Araneae, Pholcidae)" masterLastPageNumber="57" masterPageNumber="1" pageNumber="41" updateTime="1698840747163" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FEAA809D8EAC281B1A6DF19847DDBDBA">Daddy-long-leg giants: revision of the spider genus Artema Walckenaer, 1837 (Araneae, Pholcidae)</mods:title>
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<mods:namePart id="1B6FB140350D2D3691CBCD6FF8177B1B">Aharon, Shlomi</mods:namePart>
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<mods:affiliation id="7E8E9CFD41649921C4B0DE572E9CB828">Blaustein Institutes for Desert Research, Ben-Gurion University of the Negev, Sede Boqer Campus, Midreshet Ben-Gurion, Israel. Alexander Koenig Research Museum of Zoology, Adenauerallee 160, 53113 Bonn, Germany. The Arachnid National Natural History Collection, The Hebrew University of Jerusalem, Edmond J. Safra Campus, Givat Ram, Jerusalem, Israel.</mods:affiliation>
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<mods:namePart id="454AEE97497CE48E6C156F91D1C20F53">Huber, Bernhard A.</mods:namePart>
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<mods:affiliation id="EE57B7F57A38DB2A5798146F22A2BC4A">Alexander Koenig Research Museum of Zoology, Adenauerallee 160, 53113 Bonn, Germany. &amp; Email: b. huber @ zfmk. de &amp; urn: lsid: zoobank. org: author: 33607 F 65 - 19 BF- 4 DC 9 - 94 FD- 4 BB 88 CED 455 F</mods:affiliation>
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<mods:namePart id="94F65D4D8D1B973FE7F85587A6AF3042">Gavish-Regev, Efrat</mods:namePart>
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<mods:affiliation id="DCFAC48437A0D57645688CA338AE31F5">The Arachnid National Natural History Collection, The Hebrew University of Jerusalem, Edmond J. Safra Campus, Givat Ram, Jerusalem, Israel. &amp; urn: lsid: zoobank. org: author: FC 073 F 19 - 2202 - 4 C 89 - 8 B 43 - CEA 4 CC 5 E 2 D 50 &amp; Corresponding author: efrat. gavish-regev @ mail. huji. ac. il</mods:affiliation>
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<mods:title id="2396D8AE92710F92221221DCC7704A8F">European Journal of Taxonomy</mods:title>
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<treatment id="516F87BB1324BC5CFDBABF61FE9CEAF9" ID-DOI="http://doi.org/10.5281/zenodo.3852336" ID-GBIF-Taxon="164246584" ID-Zenodo-Dep="3852336" LSID="urn:lsid:plazi:treatment:516F87BB1324BC5CFDBABF61FE9CEAF9" httpUri="http://treatment.plazi.org/id/516F87BB1324BC5CFDBABF61FE9CEAF9" lastPageId="43" lastPageNumber="44" pageId="40" pageNumber="41">
<subSubSection id="91DC65261324BC5FFDBABF61FC4CEA15" box="[618,969,830,857]" pageId="40" pageNumber="41" type="nomenclature">
<paragraph id="D97936AD1324BC5FFDBABF61FC4CEA15" blockId="40.[618,969,830,893]" box="[618,969,830,857]" pageId="40" pageNumber="41">
<heading id="823181C11324BC5FFDBABF61FC4CEA15" box="[618,969,830,857]" centered="true" fontSize="11" level="2" pageId="40" pageNumber="41" reason="2">
<taxonomicName id="1EC64D2E1324BC5FFDBABF61FC4CEA15" ID-CoL="GVVJ" authority="Roewer, 1960" authorityName="Roewer" authorityYear="1960" box="[618,969,830,857]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="EBB2EABF1324BC5FFDBABF61FC9BEA14" bold="true" box="[618,798,831,857]" italics="true" pageId="40" pageNumber="41">Artema magna</emphasis>
<bibRefCitation id="BD574B5C1324BC5FFCF6BF61FC4CEA15" author="Roewer" box="[806,969,830,857]" firstAuthor="Roewer" pageId="40" pageNumber="41" pagination="1 - 53" refId="ref29937" refString="Roewer C. F. 1960. Solifugen und Opilioniden - Araneae Orthognathae, Haplogynae und Entelegynae (Contribution a l'etude de la faune d'Afghanistan 23). Goteborgs Kungliga Vetenskaps och Vitterhets Samhalles Handlingar 8 (7): 1 - 53." type="journal article" year="1960">Roewer, 1960</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="91DC65261324BC5FFD6BBF3CFCFCEA30" box="[699,889,866,893]" pageId="40" pageNumber="41" type="description">
<paragraph id="D97936AD1324BC5FFD6BBF3CFCFCEA30" blockId="40.[618,969,830,893]" box="[699,889,866,893]" pageId="40" pageNumber="41">
<figureCitation id="41FD2A281324BC5FFD6BBF3CFC81EA31" box="[699,772,866,892]" captionStart="Figs 12. 1" captionStartId="5.[189,241,1647,1673]" captionTargetBox="[190,1397,377,1570]" captionTargetId="figure@5.[187,1399,374,1571]" captionTargetPageId="5" captionText="Figs 12. 1. Known distributions of species of Artema except A. atlanta Walckenaer, 1837. The Japanese record of A. doriae Thorell, 1881 is not shown as it is based on a single introduced specimen. 2. Known distribution of Artema atlanta. Green dots: specimens examined; black dots: specimens not examined (data from: Banks 1898; Beatty et al. 2008; Blackwell 1958; Bösenberg &amp; Lenz 1895; Brignoli 1981; Bryant 1940, 1948; Caporiacco 1928, 1948, 1949; Chamberlin 1924; Chrysanthus 1967; Colmenares 2008; Denis 1953; Doleschall 1857; Dyal 1935; Franganillo 1926a; Gao &amp; Li 2010; Gertsch &amp; Davis 1942; Gonzalez-Sponga 2005; Karsch 1879, 1892; Leardi in Airaghi 1902; Lessert 1936; Majumder &amp; Biswas 1993; Marples 1955; Mello-Leitão 1942; Millot 1946; Moenkhaus 1898; F.O.Pickard-Cambridge 1902; Petrunkevitch 1925, 1929; Pocock 1900; Saaristo 2001; Simon 1885, 1894; Taczanowsky 1874; Thorell 1895, 1898; Vinson 1863; Yaginuma 1970). Records from Israel, Belgium, and Britain are not shown as they are based on single introduced specimens." figureDoi="http://doi.org/10.5281/zenodo.3838546" httpUri="https://zenodo.org/record/3838546/files/figure.png" pageId="40" pageNumber="41">Figs 1</figureCitation>
,
<figureCitation id="41FD2A281324BC5FFCC2BF3CFCFCEA30" box="[786,889,866,893]" captionStart-0="Figs 155161" captionStart-1="Figs 162169" captionStartId-0="41.[189,241,1894,1920]" captionStartId-1="42.[189,241,1858,1884]" captionTargetBox-0="[195,1393,1050,1844]" captionTargetBox-1="[201,1389,775,1815]" captionTargetId-0="figure@41.[189,1399,1040,1850]" captionTargetId-1="figure@42.[189,1400,762,1826]" captionTargetPageId-0="41" captionTargetPageId-1="42" captionText-0="Figs 155161. Artema magna Roewer, 1960. 155. Female prosoma (NHMG A 196, Afghanistan, Konar Valley). 156158. Male left pedipalp (ZMMU, Pakistan, Baluchistan Province, Quetta): prolateral, dorsal, and retrolateral views. 159161. Male left bulb and procursus: prolateral, dorsal, and retrolateral views. Scale lines: 155158 = 1 mm; 159161 = 0.5 mm." captionText-1="Figs 162169. Artema magna Roewer, 1960. 162164. Male chelicerae (ZMMU, Pakistan, Baluchistan Province, Quetta), lateral and frontal views (arrow: distinctive median projection). 165166, 168. Female abdomen (NHMG A 196, Afghanistan, Konar Valley): lateral, dorsal, and ventral views. 167, 169. Epigynum and anterior epigynal projections (AEP): lateral and ventral views. Scale lines: 162164, 167, 169 = 0.5 mm; 165166, 168 = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3838588" figureDoi-1="http://doi.org/10.5281/zenodo.3838590" httpUri-0="https://zenodo.org/record/3838588/files/figure.png" httpUri-1="https://zenodo.org/record/3838590/files/figure.png" pageId="40" pageNumber="41">155169</figureCitation>
</paragraph>
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<subSubSection id="91DC65261324BC5FFF6DBFFBFC4EEAF2" box="[189,971,932,959]" pageId="40" pageNumber="41" type="reference_group">
<paragraph id="D97936AD1324BC5FFF6DBFFBFC4EEAF2" blockId="40.[189,971,932,959]" box="[189,971,932,959]" pageId="40" pageNumber="41">
<treatmentCitationGroup id="F9D611831324BC5FFF6DBFFBFC4EEAF2" box="[189,971,932,959]" pageId="40" pageNumber="41">
<treatmentCitation id="586710BC1324BC5FFF6DBFFBFDF4EAF2" author="Roewer C. F." box="[189,625,932,959]" page="45" pageId="40" pageNumber="41" year="1960">
<taxonomicName id="1EC64D2E1324BC5FFF6DBFFBFDF4EAF2" ID-CoL="GVVJ" authority="Roewer, 1960: 43 - 45" authorityName="Roewer" authorityPageNumber="43 - 45" authorityYear="1960" box="[189,625,932,959]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="EBB2EABF1324BC5FFF6DBFFBFEEEEAF3" box="[189,363,933,959]" italics="true" pageId="40" pageNumber="41">Artema magna</emphasis>
<bibRefCitation id="BD574B5C1324BC5FFEA2BFFBFDF4EAF2" author="Roewer" box="[370,625,932,959]" firstAuthor="Roewer" pageId="40" pageNumber="41" pagination="1 - 53" refId="ref29937" refString="Roewer C. F. 1960. Solifugen und Opilioniden - Araneae Orthognathae, Haplogynae und Entelegynae (Contribution a l'etude de la faune d'Afghanistan 23). Goteborgs Kungliga Vetenskaps och Vitterhets Samhalles Handlingar 8 (7): 1 - 53." type="journal article" year="1960">Roewer, 1960: 4345</bibRefCitation>
</taxonomicName>
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, figs 14ah (
<specimenCount id="CFC0FD241324BC5FFCDABFFAFC9BEAF3" box="[778,798,932,958]" pageId="40" pageNumber="41" type="female"></specimenCount>
,
<collectingCountry id="A1D1763D1324BC5FFCFABFFAFC3AEAF2" box="[810,959,932,959]" name="Afghanistan" pageId="40" pageNumber="41">Afghanistan</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="91DC65261324BC5FFF6DBFB8FB97EDF3" pageId="40" pageNumber="41" type="diagnosis">
<paragraph id="D97936AD1324BC5FFF6DBFB8FEB3ED4D" blockId="40.[189,310,998,1024]" box="[189,310,998,1024]" pageId="40" pageNumber="41">
<heading id="823181C11324BC5FFF6DBFB8FEB3ED4D" bold="true" box="[189,310,998,1024]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="EBB2EABF1324BC5FFF6DBFB8FEB3ED4D" bold="true" box="[189,310,998,1024]" pageId="40" pageNumber="41">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="D97936AD1324BC5FFF6DB84BFB97EDF3" blockId="40.[189,1398,1045,1214]" pageId="40" pageNumber="41">
The male specimen we tentatively assign to this species differs from all known congeners by the cheliceral armature: each male cheliceral process with unique median projection ending with two modified hairs (arrow in
<figureCitation id="41FD2A281324BC5FFEE0B802FE16ED3A" box="[304,403,1116,1143]" captionStart="Figs 162169" captionStartId="42.[189,241,1858,1884]" captionTargetBox="[201,1389,775,1815]" captionTargetId="figure@42.[189,1400,762,1826]" captionTargetPageId="42" captionText="Figs 162169. Artema magna Roewer, 1960. 162164. Male chelicerae (ZMMU, Pakistan, Baluchistan Province, Quetta), lateral and frontal views (arrow: distinctive median projection). 165166, 168. Female abdomen (NHMG A 196, Afghanistan, Konar Valley): lateral, dorsal, and ventral views. 167, 169. Epigynum and anterior epigynal projections (AEP): lateral and ventral views. Scale lines: 162164, 167, 169 = 0.5 mm; 165166, 168 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838590" httpUri="https://zenodo.org/record/3838590/files/figure.png" pageId="40" pageNumber="41">Fig. 164</figureCitation>
) that appear isolated and not linked (by further hairs) to main modified hairs ridge as in
<taxonomicName id="1EC64D2E1324BC5FFF0BB8DEFED0EDD7" authority="Aharon &amp; Huber &amp; Gavish-Regev, 2017" authorityName="Aharon &amp; Huber &amp; Gavish-Regev" authorityYear="2017" box="[219,341,1152,1178]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="nephilit" status="sp. nov.">
<emphasis id="EBB2EABF1324BC5FFF0BB8DEFED0EDD7" box="[219,341,1152,1178]" italics="true" pageId="40" pageNumber="41">A. nephilit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F08157C41324BC5FFE8BB8DEFE30EDD7" box="[347,437,1152,1178]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
Female differs from all congeners by wide epigynal plate (~2.5 × as wide as long) and by distinctively protruding median posterior margin (
<figureCitation id="41FD2A281324BC5FFC8EB8FDFB80EDF3" box="[862,1029,1187,1214]" captionStart="Figs 162169" captionStartId="42.[189,241,1858,1884]" captionTargetBox="[201,1389,775,1815]" captionTargetId="figure@42.[189,1400,762,1826]" captionTargetPageId="42" captionText="Figs 162169. Artema magna Roewer, 1960. 162164. Male chelicerae (ZMMU, Pakistan, Baluchistan Province, Quetta), lateral and frontal views (arrow: distinctive median projection). 165166, 168. Female abdomen (NHMG A 196, Afghanistan, Konar Valley): lateral, dorsal, and ventral views. 167, 169. Epigynum and anterior epigynal projections (AEP): lateral and ventral views. Scale lines: 162164, 167, 169 = 0.5 mm; 165166, 168 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838590" httpUri="https://zenodo.org/record/3838590/files/figure.png" pageId="40" pageNumber="41">Figs 168169</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="91DC65261324BC5FFF6DB8BBFC6EECB2" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="D97936AD1324BC5FFF6DB8BBFE2FEDB2" blockId="40.[189,426,1253,1279]" box="[189,426,1253,1279]" pageId="40" pageNumber="41">
<heading id="823181C11324BC5FFF6DB8BBFE2FEDB2" bold="true" box="[189,426,1253,1279]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="EBB2EABF1324BC5FFF6DB8BBFE2FEDB2" bold="true" box="[189,426,1253,1279]" pageId="40" pageNumber="41">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="69AE3CF01324BC5FFF3AB94BFCE5EC3B" ID-GBIF-Occurrence="2621730302" collectingDate="1958-02-06" collectionCode="NHMG" collectorName="K. Lindberg" country="Afghanistan" latitude="34.61" location="Qachgar cave" longLatPrecision="719" longitude="70.76" municipality="Qachgar cave" pageId="40" pageNumber="41" specimenCode="NHMG A 196" specimenCount="2" specimenCount-female="1" specimenCount-juvenile="1" stateProvince="Konar valley" typeStatus="holotype">
<paragraph id="D97936AD1324BC5FFF3AB94BFED9EC62" blockId="40.[189,1399,1301,1398]" box="[234,348,1301,1327]" pageId="40" pageNumber="41">
<emphasis id="EBB2EABF1324BC5FFF3AB94BFED9EC62" bold="true" box="[234,348,1301,1327]" pageId="40" pageNumber="41">
<typeStatus id="067D880F1324BC5FFF3AB94BFED9EC62" box="[234,348,1301,1327]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="D97936AD1324BC5FFF6DB966FCE5EC3B" blockId="40.[189,1399,1301,1398]" pageId="40" pageNumber="41">
<collectingCountry id="A1D1763D1324BC5FFF6DB966FE15EC1F" box="[189,400,1336,1362]" name="Afghanistan" pageId="40" pageNumber="41">AFGHANISTAN</collectingCountry>
:
<specimenCount id="CFC0FD241324BC5FFE71B967FE32EC1E" box="[417,439,1337,1363]" pageId="40" pageNumber="41" type="female"></specimenCount>
(together with
<specimenCount id="CFC0FD241324BC5FFDAFB966FD4EEC1E" box="[639,715,1336,1363]" pageId="40" pageNumber="41" type="juvenile">1 juv.</specimenCount>
),
<collectingRegion id="1B02F84F1324BC5FFD36B967FC03EC1F" box="[742,902,1336,1363]" pageId="40" pageNumber="41">Konar valley</collectingRegion>
,
<location id="DC1960761324BC5FFC49B966FBC6EC1E" LSID="urn:lsid:plazi:treatment:516F87BB1324BC5CFDBABF61FE9CEAF9:DC1960761324BC5FFC49B966FBC6EC1E" box="[921,1091,1336,1363]" country="Afghanistan" latitude="34.61" longLatPrecision="719" longitude="70.76" municipality="Qachgar cave" name="Qachgar cave" pageId="40" pageNumber="41" stateProvince="Konar valley">Qachgar cave</location>
,
<location id="DC1960761324BC5FFB85B967FB63EC1F" LSID="urn:lsid:plazi:treatment:516F87BB1324BC5CFDBABF61FE9CEAF9:DC1960761324BC5FFB85B967FB63EC1F" box="[1109,1254,1336,1363]" country="Afghanistan" latitude="34.61" longLatPrecision="719" longitude="70.76" municipality="Qachgar cave" name="near Nurgal" pageId="40" pageNumber="41" stateProvince="Konar valley">near Nurgal</location>
[
<geoCoordinate id="BCF2506A1324BC5FFB2CB966FAF7EC1E" box="[1276,1394,1336,1363]" degrees="34.61" direction="north" orientation="latitude" pageId="40" pageNumber="41" precision="555" value="34.61">34.61° N</geoCoordinate>
,
<geoCoordinate id="BCF2506A1324BC5FFF6DB905FEADEC3B" box="[189,296,1371,1398]" degrees="70.76" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="555" value="70.76">70.76° E</geoCoordinate>
],
<date id="AD78106D1324BC5FFEECB905FE4FEC3B" box="[316,458,1371,1398]" pageId="40" pageNumber="41" value="1958-02-06">
<collectingDate id="BD3CE9851324BC5FFEECB905FE4FEC3B" box="[316,458,1371,1398]" pageId="40" pageNumber="41" value="1958-02-06">6 Feb. 1958</collectingDate>
</date>
,
<collectorName id="7433537B1324BC5FFE06B902FDE0EC3B" box="[470,613,1372,1398]" pageId="40" pageNumber="41">K. Lindberg</collectorName>
leg. (
<collectionCode id="BFD7AE681324BC5FFD77B902FC86EC3B" box="[679,771,1372,1398]" pageId="40" pageNumber="41">NHMG</collectionCode>
A 196).
</paragraph>
</materialsCitation>
<paragraph id="D97936AD1324BC5FFF3AB9C3FE23ECFA" blockId="40.[189,1399,1437,1536]" box="[234,422,1437,1463]" pageId="40" pageNumber="41">
<emphasis id="EBB2EABF1324BC5FFF3AB9C3FE23ECFA" bold="true" box="[234,422,1437,1463]" pageId="40" pageNumber="41">Other material</emphasis>
</paragraph>
<paragraph id="D97936AD1324BC5FFF6DB99FFC6EECB2" blockId="40.[189,1399,1437,1536]" pageId="40" pageNumber="41">
<materialsCitation id="69AE3CF01324BC5FFF6DB99FFC6EECB2" ID-GBIF-Occurrence="3763211454" collectingDate="2005-07-27" collectionCode="ZMMU" collectorName="S. V. Ovchinnikov" country="Pakistan" latitude="30.18" location="Quetta Town" longLatPrecision="734" longitude="66.99" municipality="Quetta Town" pageId="40" pageNumber="41" specimenCount="1" specimenCount-male="1" stateProvince="Baluchistan Province">
<collectingCountry id="A1D1763D1324BC5FFF6DB99FFECBEC96" box="[189,334,1473,1499]" name="Pakistan" pageId="40" pageNumber="41">PAKISTAN</collectingCountry>
:
<specimenCount id="CFC0FD241324BC5FFE87B99FFE01EC91" box="[343,388,1473,1500]" pageId="40" pageNumber="41" type="male">1 ♂</specimenCount>
, assigned tentatively (see Notes below),
<collectingRegion id="1B02F84F1324BC5FFC8AB99FFBD3EC96" box="[858,1110,1473,1499]" country="Pakistan" name="Balochistan" pageId="40" pageNumber="41">Baluchistan Province</collectingRegion>
,
<collectingMunicipality id="391DACD71324BC5FFBB0B99FFB73EC91" box="[1120,1270,1473,1500]" pageId="40" pageNumber="41">Quetta Town</collectingMunicipality>
[
<geoCoordinate id="BCF2506A1324BC5FFAD3B99FFAF7EC91" box="[1283,1394,1473,1500]" degrees="30.18" direction="north" orientation="latitude" pageId="40" pageNumber="41" precision="555" value="30.18">30.18° N</geoCoordinate>
,
<geoCoordinate id="BCF2506A1324BC5FFF6DB9BBFEADECB2" box="[189,296,1508,1535]" degrees="66.99" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="555" value="66.99">66.99° E</geoCoordinate>
], in building,
<date id="AD78106D1324BC5FFE1BB9BBFDDAEF4D" box="[459,607,1509,1536]" pageId="40" pageNumber="41" value="2005-07-27">
<collectingDate id="BD3CE9851324BC5FFE1BB9BBFDDAEF4D" box="[459,607,1509,1536]" pageId="40" pageNumber="41" value="2005-07-27">27 Jul. 2005</collectingDate>
</date>
,
<collectorName id="7433537B1324BC5FFDBAB9BBFCBEECB2" box="[618,827,1509,1535]" pageId="40" pageNumber="41">S.V. Ovchinnikov</collectorName>
leg. (
<collectionCode id="BFD7AE681324BC5FFCADB9BBFC5AECB2" box="[893,991,1509,1535]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="40" pageNumber="41" type="Museum">ZMMU</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="91DC65261324BC5DFF6DBA78FC6BEB4D" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41" type="description">
<paragraph id="D97936AD1324BC5FFF6DBA78FECBEF0D" blockId="40.[189,334,1574,1600]" box="[189,334,1574,1600]" pageId="40" pageNumber="41">
<heading id="823181C11324BC5FFF6DBA78FECBEF0D" bold="true" box="[189,334,1574,1600]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="EBB2EABF1324BC5FFF6DBA78FECBEF0D" bold="true" box="[189,334,1574,1600]" pageId="40" pageNumber="41">Description</emphasis>
</heading>
</paragraph>
<paragraph id="D97936AD1324BC5FFF3ABA08FD7BEF3C" blockId="40.[189,1399,1622,1755]" box="[234,766,1622,1649]" pageId="40" pageNumber="41">
<emphasis id="EBB2EABF1324BC5FFF3ABA08FEAFEF3D" bold="true" box="[234,298,1622,1648]" pageId="40" pageNumber="41">Male</emphasis>
(
<collectingCountry id="A1D1763D1324BC5FFEE9BA08FE26EF3D" box="[313,419,1622,1648]" name="Pakistan" pageId="40" pageNumber="41">Pakistan</collectingCountry>
: Quetta; assigned tentatively)
</paragraph>
<paragraph id="D97936AD1324BC5FFF6DBA24FB6AEF96" blockId="40.[189,1399,1622,1755]" pageId="40" pageNumber="41">MEASUREMENTS. Total body length 9.3, carapace width 4.3. Leg 1: 54.8 (14.0 + 1.8 + 15.8 + 20.3 + 2.9), tibia 2: 11.8, tibia 3: 9.0, tibia 4: 12.0; tibia 1 L/d: 39. Distance PMEPME 210 μm, diameter PME 230 μm, distance PMEALE 100 μm, distance AMEAME 30 μm, diameter AME 210 μm.</paragraph>
<paragraph id="D97936AD1324BC5FFF6DBAA5FCF4EEA6" blockId="40.[189,1399,1787,2027]" pageId="40" pageNumber="41">
COLOR. Carapace light ochre to beige with small brown spots laterally forming few small lines near median pit, with light brown median band that splits at posterior margin of ocular area to brown lines following base of ocular area and light brown band medially on ocular area; clypeus with wide ochre brown band below AME that fades towards dark brown edge of clypeus (as in
<figureCitation id="41FD2A281324BC5FFBB2BB38FB4CEECC" box="[1122,1225,1894,1921]" captionStart="Figs 155161" captionStartId="41.[189,241,1894,1920]" captionTargetBox="[195,1393,1050,1844]" captionTargetId="figure@41.[189,1399,1040,1850]" captionTargetPageId="41" captionText="Figs 155161. Artema magna Roewer, 1960. 155. Female prosoma (NHMG A 196, Afghanistan, Konar Valley). 156158. Male left pedipalp (ZMMU, Pakistan, Baluchistan Province, Quetta): prolateral, dorsal, and retrolateral views. 159161. Male left bulb and procursus: prolateral, dorsal, and retrolateral views. Scale lines: 155158 = 1 mm; 159161 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3838588" httpUri="https://zenodo.org/record/3838588/files/figure.png" pageId="40" pageNumber="41">Fig. 155</figureCitation>
); legs ochre to light brown with dark rings on femora subdistally, patellae + tibiae proximally, and tibiae subdistally, tips of femora and tibiae pale ochre; sternum ochre, with brown margins and median light brown dots forming cross-like pattern; abdomen beige without pattern.
</paragraph>
<paragraph id="D97936AD1325BC5EFF6DBD52FCE7E8FF" blockId="41.[189,1399,267,435]" pageId="41" pageNumber="42">
BODY. Ocular area slightly elevated; carapace with median pit and distinct posterior furrow; clypeus unmodified; sternum wider than long (3.0/2.1); chelicerae as in
<figureCitation id="41FD2A281325BC5EFC02BD70FBFCE805" box="[978,1145,302,328]" captionStart="Figs 162169" captionStartId="42.[189,241,1858,1884]" captionTargetBox="[201,1389,775,1815]" captionTargetId="figure@42.[189,1400,762,1826]" captionTargetPageId="42" captionText="Figs 162169. Artema magna Roewer, 1960. 162164. Male chelicerae (ZMMU, Pakistan, Baluchistan Province, Quetta), lateral and frontal views (arrow: distinctive median projection). 165166, 168. Female abdomen (NHMG A 196, Afghanistan, Konar Valley): lateral, dorsal, and ventral views. 167, 169. Epigynum and anterior epigynal projections (AEP): lateral and ventral views. Scale lines: 162164, 167, 169 = 0.5 mm; 165166, 168 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838590" httpUri="https://zenodo.org/record/3838590/files/figure.png" pageId="41" pageNumber="42">Figs 162164</figureCitation>
, with frontal row of ~17 modified (cone-shaped) hairs on each side with median projection ending with two modified hairs that appear isolated and not linked to main modified hairs ridge; without stridulatory ridges; Abdomen globose and high; gonopore with four epiandrous spigots.
</paragraph>
<paragraph id="D97936AD1325BC5EFF6DBD8DFE10EBAE" blockId="41.[189,1399,466,739]" pageId="41" pageNumber="42">
PALPS. As in
<figureCitation id="41FD2A281325BC5EFE81BD8CFE70E8A0" box="[337,501,466,493]" captionStart="Figs 155161" captionStartId="41.[189,241,1894,1920]" captionTargetBox="[195,1393,1050,1844]" captionTargetId="figure@41.[189,1399,1040,1850]" captionTargetPageId="41" captionText="Figs 155161. Artema magna Roewer, 1960. 155. Female prosoma (NHMG A 196, Afghanistan, Konar Valley). 156158. Male left pedipalp (ZMMU, Pakistan, Baluchistan Province, Quetta): prolateral, dorsal, and retrolateral views. 159161. Male left bulb and procursus: prolateral, dorsal, and retrolateral views. Scale lines: 155158 = 1 mm; 159161 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3838588" httpUri="https://zenodo.org/record/3838588/files/figure.png" pageId="41" pageNumber="42">Figs 156158</figureCitation>
; coxa unmodified; trochanter with short ventral projection; femur with short, somewhat pointed retrolateral process proximally; ventral large membranous area proximally bordered on both sides by sclerotized ridges; femur-patella hinges close together dorsally; patella very short; procursus with proximal dorsal process, with weakly developed ventral pocket, with three weak distal dorsal notches on prolateral margin (
<figureCitation id="41FD2A281325BC5EFDADBE01FD62EB37" box="[637,743,607,634]" captionStart="Figs 155161" captionStartId="41.[189,241,1894,1920]" captionTargetBox="[195,1393,1050,1844]" captionTargetId="figure@41.[189,1399,1040,1850]" captionTargetPageId="41" captionText="Figs 155161. Artema magna Roewer, 1960. 155. Female prosoma (NHMG A 196, Afghanistan, Konar Valley). 156158. Male left pedipalp (ZMMU, Pakistan, Baluchistan Province, Quetta): prolateral, dorsal, and retrolateral views. 159161. Male left bulb and procursus: prolateral, dorsal, and retrolateral views. Scale lines: 155158 = 1 mm; 159161 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3838588" httpUri="https://zenodo.org/record/3838588/files/figure.png" pageId="41" pageNumber="42">Fig. 160</figureCitation>
); bulb with membranous embolus rising from base of processes a and b, process a slightly spiraling with pointed tip; process b elongated and pointed, process c robust, strongly curved towards prolateral (
<figureCitation id="41FD2A281325BC5EFD18BEFBFCA8EB8D" box="[712,813,677,704]" captionStart="Figs 155161" captionStartId="41.[189,241,1894,1920]" captionTargetBox="[195,1393,1050,1844]" captionTargetId="figure@41.[189,1399,1040,1850]" captionTargetPageId="41" captionText="Figs 155161. Artema magna Roewer, 1960. 155. Female prosoma (NHMG A 196, Afghanistan, Konar Valley). 156158. Male left pedipalp (ZMMU, Pakistan, Baluchistan Province, Quetta): prolateral, dorsal, and retrolateral views. 159161. Male left bulb and procursus: prolateral, dorsal, and retrolateral views. Scale lines: 155158 = 1 mm; 159161 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3838588" httpUri="https://zenodo.org/record/3838588/files/figure.png" pageId="41" pageNumber="42">Fig. 160</figureCitation>
), process d low rounded projection on ventral side of bulb (
<figureCitation id="41FD2A281325BC5EFEF1BE96FE0DEBAE" box="[289,392,712,739]" captionStart="Figs 155161" captionStartId="41.[189,241,1894,1920]" captionTargetBox="[195,1393,1050,1844]" captionTargetId="figure@41.[189,1399,1040,1850]" captionTargetPageId="41" captionText="Figs 155161. Artema magna Roewer, 1960. 155. Female prosoma (NHMG A 196, Afghanistan, Konar Valley). 156158. Male left pedipalp (ZMMU, Pakistan, Baluchistan Province, Quetta): prolateral, dorsal, and retrolateral views. 159161. Male left bulb and procursus: prolateral, dorsal, and retrolateral views. Scale lines: 155158 = 1 mm; 159161 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3838588" httpUri="https://zenodo.org/record/3838588/files/figure.png" pageId="41" pageNumber="42">Fig. 159</figureCitation>
).
</paragraph>
<paragraph id="D97936AD1325BC5EFF6DBF5DFAB1EA72" blockId="41.[189,1398,770,832]" pageId="41" pageNumber="42">LEGS.Without spines;with long curved hairs, especially on tibiae and metatarsi; retrolateral trichobothrium on tibia 1 at 10%; prolateral trichobothrium present on all tibiae; pseudosegmentation not visible.</paragraph>
<paragraph id="D97936AD1325BC5EFF3ABF3EFE42EA37" blockId="41.[189,1399,863,996]" box="[234,455,863,890]" pageId="41" pageNumber="42">
<emphasis id="EBB2EABF1325BC5EFF3ABF3EFEC0EA37" bold="true" box="[234,325,864,890]" pageId="41" pageNumber="42">Female</emphasis>
(
<typeStatus id="067D880F1325BC5EFE85BF01FE3AEA37" box="[341,447,863,890]" pageId="41" pageNumber="42" type="holotype">holotype</typeStatus>
)
</paragraph>
<paragraph id="D97936AD1325BC5DFF6DBFDDFE36E821" blockId="41.[189,1399,863,996]" lastBlockId="42.[189,1398,267,364]" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
In general similar to male; tibia 1: 17.8; abdomen beige with lateral gray stripes with large marks dorsally (
<figureCitation id="41FD2A281325BC5EFEFCBFF8FE51EA8C" box="[300,468,934,961]" captionStart="Figs 162169" captionStartId="42.[189,241,1858,1884]" captionTargetBox="[201,1389,775,1815]" captionTargetId="figure@42.[189,1400,762,1826]" captionTargetPageId="42" captionText="Figs 162169. Artema magna Roewer, 1960. 162164. Male chelicerae (ZMMU, Pakistan, Baluchistan Province, Quetta), lateral and frontal views (arrow: distinctive median projection). 165166, 168. Female abdomen (NHMG A 196, Afghanistan, Konar Valley): lateral, dorsal, and ventral views. 167, 169. Epigynum and anterior epigynal projections (AEP): lateral and ventral views. Scale lines: 162164, 167, 169 = 0.5 mm; 165166, 168 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838590" httpUri="https://zenodo.org/record/3838590/files/figure.png" pageId="41" pageNumber="42">Figs 165166</figureCitation>
); stridulatory files laterally on chelicerae present; epigynum ~2.5 × wider than long, median posterior margin protruding (
<figureCitation id="41FD2A281325BC5EFD16BF97FCF5EAA9" box="[710,880,969,996]" captionStart="Figs 162169" captionStartId="42.[189,241,1858,1884]" captionTargetBox="[201,1389,775,1815]" captionTargetId="figure@42.[189,1400,762,1826]" captionTargetPageId="42" captionText="Figs 162169. Artema magna Roewer, 1960. 162164. Male chelicerae (ZMMU, Pakistan, Baluchistan Province, Quetta), lateral and frontal views (arrow: distinctive median projection). 165166, 168. Female abdomen (NHMG A 196, Afghanistan, Konar Valley): lateral, dorsal, and ventral views. 167, 169. Epigynum and anterior epigynal projections (AEP): lateral and ventral views. Scale lines: 162164, 167, 169 = 0.5 mm; 165166, 168 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838590" httpUri="https://zenodo.org/record/3838590/files/figure.png" pageId="41" pageNumber="42">Figs 168169</figureCitation>
); epigynal plate consisting of two swollen sclerotized lateral areas, pale median area, and dark median anterior sclerite; pair of projections anterior to epigynum (AEP) oval, not prominent; sternum with light brown dots forming cross-like pattern more distinct than in male.
</paragraph>
<caption id="8DB966251325BC5EFF6DBB38FCDBEEA7" ID-DOI="http://doi.org/10.5281/zenodo.3838588" ID-Zenodo-Dep="3838588" httpUri="https://zenodo.org/record/3838588/files/figure.png" pageId="41" pageNumber="42" startId="41.[189,241,1894,1920]" targetBox="[195,1393,1050,1844]" targetPageId="41">
<paragraph id="D97936AD1325BC5EFF6DBB38FCDBEEA7" blockId="41.[189,1399,1894,2026]" pageId="41" pageNumber="42">
<emphasis id="EBB2EABF1325BC5EFF6DBB38FEE0EECD" bold="true" box="[189,357,1894,1920]" pageId="41" pageNumber="42">Figs 155161.</emphasis>
<taxonomicName id="1EC64D2E1325BC5EFEBDBB39FD42EECD" authority="Roewer, 1960" authorityName="Roewer" authorityYear="1960" box="[365,711,1894,1921]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="EBB2EABF1325BC5EFEBDBB39FD9FEECD" box="[365,538,1895,1921]" italics="true" pageId="41" pageNumber="42">Artema magna</emphasis>
<bibRefCitation id="BD574B5C1325BC5EFDF0BB39FD42EECD" author="Roewer" box="[544,711,1894,1921]" firstAuthor="Roewer" pageId="41" pageNumber="42" pagination="1 - 53" refId="ref29937" refString="Roewer C. F. 1960. Solifugen und Opilioniden - Araneae Orthognathae, Haplogynae und Entelegynae (Contribution a l'etude de la faune d'Afghanistan 23). Goteborgs Kungliga Vetenskaps och Vitterhets Samhalles Handlingar 8 (7): 1 - 53." type="journal article" year="1960">Roewer, 1960</bibRefCitation>
</taxonomicName>
.
<emphasis id="EBB2EABF1325BC5EFD01BB38FD78EECD" bold="true" box="[721,765,1894,1920]" pageId="41" pageNumber="42">155</emphasis>
. Female prosoma (NHMG A 196, Afghanistan, Konar Valley).
<emphasis id="EBB2EABF1325BC5EFEF6BBD7FE08EEEE" bold="true" box="[294,397,1929,1955]" pageId="41" pageNumber="42">156158</emphasis>
. Male left pedipalp (ZMMU, Pakistan, Baluchistan Province, Quetta): prolateral, dorsal, and retrolateral views.
<emphasis id="EBB2EABF1325BC5EFDCEBBF2FD01EE8B" bold="true" box="[542,644,1964,1990]" pageId="41" pageNumber="42">159161</emphasis>
. Male left bulb and procursus: prolateral, dorsal, and retrolateral views. Scale lines: 155158 = 1 mm; 159161 = 0.5 mm.
</paragraph>
</caption>
<paragraph id="D97936AD1326BC5DFF6DBDCDFEFBE8E0" blockId="42.[189,382,403,429]" box="[189,382,403,429]" pageId="42" pageNumber="43">
<heading id="823181C11326BC5DFF6DBDCDFEFBE8E0" bold="true" box="[189,382,403,429]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="EBB2EABF1326BC5DFF6DBDCDFEFBE8E0" bold="true" box="[189,382,403,429]" pageId="42" pageNumber="43">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="D97936AD1326BC5DFF6DBD9CFC6BEB4D" blockId="42.[189,1398,450,512]" pageId="42" pageNumber="43">
The female
<typeStatus id="067D880F1326BC5DFE95BD9CFE29E891" box="[325,428,450,476]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
was collected in a cave. The only known male was collected inside a house, together with a male of a different, unnamed species (see
<taxonomicName id="1EC64D2E1326BC5DFD2ABDBBFCD5E8B2" authorityName="Walckenaer" authorityYear="1837" box="[762,848,485,511]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="EBB2EABF1326BC5DFD2ABDBBFCD5E8B2" box="[762,848,485,511]" italics="true" pageId="42" pageNumber="43">Artema</emphasis>
</taxonomicName>
sp. c below).
</paragraph>
</subSubSection>
<subSubSection id="91DC65261326BC5DFF6DBE78FAA3EBDF" pageId="42" pageNumber="43" type="distribution">
<paragraph id="D97936AD1326BC5DFF6DBE78FED3EB0D" blockId="42.[189,342,550,576]" box="[189,342,550,576]" pageId="42" pageNumber="43">
<heading id="823181C11326BC5DFF6DBE78FED3EB0D" bold="true" box="[189,342,550,576]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="EBB2EABF1326BC5DFF6DBE78FED3EB0D" bold="true" box="[189,342,550,576]" pageId="42" pageNumber="43">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="D97936AD1326BC5DFF6DBE0BFAA3EBDF" blockId="42.[189,1398,597,659]" pageId="42" pageNumber="43">
If the male from
<collectingCountry id="A1D1763D1326BC5DFE5DBE0BFE75EB22" box="[397,496,597,623]" name="Pakistan" pageId="42" pageNumber="43">Pakistan</collectingCountry>
is correctly assigned to this species, then
<taxonomicName id="1EC64D2E1326BC5DFC3FBE0BFB25EB22" authorityName="Roewer" authorityYear="1960" box="[1007,1184,597,623]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="EBB2EABF1326BC5DFC3FBE0BFB25EB22" box="[1007,1184,597,623]" italics="true" pageId="42" pageNumber="43">Artema magna</emphasis>
</taxonomicName>
may have a wide distribution along the mountain ridge from north-eastern
<collectingCountry id="A1D1763D1326BC5DFC8ABE26FC6CEBDF" box="[858,1001,632,658]" name="Afghanistan" pageId="42" pageNumber="43">Afghanistan</collectingCountry>
to south-western
<collectingCountry id="A1D1763D1326BC5DFB6BBE26FAA7EBDF" box="[1211,1314,632,658]" name="Pakistan" pageId="42" pageNumber="43">Pakistan</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="8DB966251326BC5DFF6DBB1CFCA8EEA7" ID-DOI="http://doi.org/10.5281/zenodo.3838590" ID-Zenodo-Dep="3838590" httpUri="https://zenodo.org/record/3838590/files/figure.png" pageId="42" pageNumber="43" startId="42.[189,241,1858,1884]" targetBox="[201,1389,775,1815]" targetPageId="42">
<paragraph id="D97936AD1326BC5DFF6DBB1CFCA8EEA7" blockId="42.[189,1399,1858,2026]" pageId="42" pageNumber="43">
<emphasis id="EBB2EABF1326BC5DFF6DBB1CFEE3EE11" bold="true" box="[189,358,1858,1884]" pageId="42" pageNumber="43">Figs 162169.</emphasis>
<taxonomicName id="1EC64D2E1326BC5DFEBDBB1DFD42EE11" authority="Roewer, 1960" authorityName="Roewer" authorityYear="1960" box="[365,711,1858,1885]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="EBB2EABF1326BC5DFEBDBB1DFD9FEE11" box="[365,538,1859,1885]" italics="true" pageId="42" pageNumber="43">Artema magna</emphasis>
<bibRefCitation id="BD574B5C1326BC5DFDF1BB1DFD42EE11" author="Roewer" box="[545,711,1858,1885]" firstAuthor="Roewer" pageId="42" pageNumber="43" pagination="1 - 53" refId="ref29937" refString="Roewer C. F. 1960. Solifugen und Opilioniden - Araneae Orthognathae, Haplogynae und Entelegynae (Contribution a l'etude de la faune d'Afghanistan 23). Goteborgs Kungliga Vetenskaps och Vitterhets Samhalles Handlingar 8 (7): 1 - 53." type="journal article" year="1960">Roewer, 1960</bibRefCitation>
</taxonomicName>
.
<emphasis id="EBB2EABF1326BC5DFD02BB1CFCBCEE11" bold="true" box="[722,825,1858,1884]" pageId="42" pageNumber="43">162164</emphasis>
. Male chelicerae (ZMMU, Pakistan, Baluchistan Province, Quetta), lateral and frontal views (arrow: distinctive median projection).
<emphasis id="EBB2EABF1326BC5DFAD8BB38FF6CEEEE" bold="true" pageId="42" pageNumber="43">165166, 168</emphasis>
. Female abdomen (NHMG A 196, Afghanistan, Konar Valley): lateral, dorsal, and ventral views.
<emphasis id="EBB2EABF1326BC5DFF6DBBF2FEA2EE8B" bold="true" box="[189,295,1964,1990]" pageId="42" pageNumber="43">167, 169</emphasis>
. Epigynum and anterior epigynal projections (AEP): lateral and ventral views. Scale lines: 162164, 167, 169 = 0.5 mm; 165166, 168 = 1 mm.
</paragraph>
</caption>
<subSubSection id="91DC65261327BC5CFF6DBD55FE9CEAF9" pageId="43" pageNumber="44" type="discussion">
<paragraph id="D97936AD1327BC5CFF6DBD55FE86E868" blockId="43.[189,259,267,293]" box="[189,259,267,293]" pageId="43" pageNumber="44">
<heading id="823181C11327BC5CFF6DBD55FE86E868" bold="true" box="[189,259,267,293]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="EBB2EABF1327BC5CFF6DBD55FE86E868" bold="true" box="[189,259,267,293]" pageId="43" pageNumber="44">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="D97936AD1327BC5CFF6DBD64FDA4E8AC" blockId="43.[189,1399,313,481]" pageId="43" pageNumber="44">
We tentatively assign the male from
<collectingCountry id="A1D1763D1327BC5CFD57BD67FD6FE81E" box="[647,746,313,339]" name="Pakistan" pageId="43" pageNumber="44">Pakistan</collectingCountry>
to
<taxonomicName id="1EC64D2E1327BC5CFCC9BD64FC0BE81E" authorityName="Roewer" authorityYear="1960" box="[793,910,314,340]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="EBB2EABF1327BC5CFCC9BD64FC0BE81E" box="[793,910,314,340]" italics="true" pageId="43" pageNumber="44">A. magna</emphasis>
</taxonomicName>
mainly because both this male and the female
<typeStatus id="067D880F1327BC5CFEC4BD03FEFEE83A" box="[276,379,349,375]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
show some specific similarities to
<taxonomicName id="1EC64D2E1327BC5CFCC6BD03FC15E83A" authority="Aharon &amp; Huber &amp; Gavish-Regev, 2017" authorityName="Aharon &amp; Huber &amp; Gavish-Regev" authorityYear="2017" box="[790,912,349,375]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="nephilit" status="sp. nov.">
<emphasis id="EBB2EABF1327BC5CFCC6BD03FC15E83A" box="[790,912,349,375]" italics="true" pageId="43" pageNumber="44">A. nephilit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F08157C41327BC5CFC47BD03FC77E83A" box="[919,1010,349,375]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
rather than to the geographically closer
<taxonomicName id="1EC64D2E1327BC5CFEDBBDDFFEFDE8D7" authorityName="Thorell" authorityYear="1881" box="[267,376,384,411]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="doriae">
<emphasis id="EBB2EABF1327BC5CFEDBBDDFFEFDE8D7" box="[267,376,384,411]" italics="true" pageId="43" pageNumber="44">A. doriae</emphasis>
</taxonomicName>
and
<taxonomicName id="1EC64D2E1327BC5CFE61BDDFFDEDE8D7" authorityName="Spassky" authorityYear="1934" box="[433,616,385,411]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="transcaspica">
<emphasis id="EBB2EABF1327BC5CFE61BDDFFDEDE8D7" box="[433,616,385,411]" italics="true" pageId="43" pageNumber="44">A. transcaspica</emphasis>
</taxonomicName>
. Males of
<taxonomicName id="1EC64D2E1327BC5CFD36BDDFFCD2E8D7" authorityName="Roewer" authorityYear="1960" box="[742,855,385,411]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="EBB2EABF1327BC5CFD36BDDFFCD2E8D7" box="[742,855,385,411]" italics="true" pageId="43" pageNumber="44">A. magna</emphasis>
</taxonomicName>
and
<taxonomicName id="1EC64D2E1327BC5CFC40BDDFFB8EE8D7" authority="Aharon &amp; Huber &amp; Gavish-Regev, 2017" authorityName="Aharon &amp; Huber &amp; Gavish-Regev" authorityYear="2017" box="[912,1035,384,411]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="nephilit" status="sp. nov.">
<emphasis id="EBB2EABF1327BC5CFC40BDDFFB8EE8D7" box="[912,1035,384,411]" italics="true" pageId="43" pageNumber="44">A. nephilit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F08157C41327BC5CFBC2BDDFFBE8E8D6" box="[1042,1133,385,411]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
share the pair of inner projections on the cheliceral process and cheliceral lateral shape, while females have wider epigyna than
<taxonomicName id="1EC64D2E1327BC5CFF6DBD99FEAEE8AC" authorityName="Thorell" authorityYear="1881" box="[189,299,455,481]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="doriae">
<emphasis id="EBB2EABF1327BC5CFF6DBD99FEAEE8AC" box="[189,299,455,481]" italics="true" pageId="43" pageNumber="44">A. doriae</emphasis>
</taxonomicName>
and
<taxonomicName id="1EC64D2E1327BC5CFEB3BD99FD9FE8AC" authorityName="Spassky" authorityYear="1934" box="[355,538,455,481]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="transcaspica">
<emphasis id="EBB2EABF1327BC5CFEB3BD99FD9FE8AC" box="[355,538,455,481]" italics="true" pageId="43" pageNumber="44">A. transcaspica</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="D97936AD1327BC5CFF6DBE56FEB1EB24" blockId="43.[189,1398,520,617]" pageId="43" pageNumber="44">
The dubious
<taxonomicName id="1EC64D2E1327BC5CFE87BE57FD67EB6E" authority="(Roewer, 1960)" baseAuthorityName="Roewer" baseAuthorityYear="1960" box="[343,738,520,547]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="ziaretana">
<emphasis id="EBB2EABF1327BC5CFE87BE57FDA6EB6F" box="[343,547,521,547]" italics="true" pageId="43" pageNumber="44">Artema ziaretana</emphasis>
(
<bibRefCitation id="BD574B5C1327BC5CFDE3BE56FD5FEB6E" author="Roewer" box="[563,730,520,547]" firstAuthor="Roewer" pageId="43" pageNumber="44" pagination="1 - 53" refId="ref29937" refString="Roewer C. F. 1960. Solifugen und Opilioniden - Araneae Orthognathae, Haplogynae und Entelegynae (Contribution a l'etude de la faune d'Afghanistan 23). Goteborgs Kungliga Vetenskaps och Vitterhets Samhalles Handlingar 8 (7): 1 - 53." type="journal article" year="1960">Roewer, 1960</bibRefCitation>
)
</taxonomicName>
is based on a single juvenile specimen originating from near Jalalabad (located only ~
<quantity id="1E3E9B481327BC5CFDCBBE75FDE0EB08" box="[539,613,555,582]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="43" pageNumber="44" unit="km" value="35.0">35 km</quantity>
from the
<typeStatus id="067D880F1327BC5CFD07BE72FC8FEB0B" box="[727,778,556,582]" pageId="43" pageNumber="44">type</typeStatus>
locality of
<taxonomicName id="1EC64D2E1327BC5CFC5FBE72FC7AEB08" authorityName="Roewer" authorityYear="1960" box="[911,1023,556,582]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="EBB2EABF1327BC5CFC5FBE72FC7AEB08" box="[911,1023,556,582]" italics="true" pageId="43" pageNumber="44">A. magna</emphasis>
</taxonomicName>
) and may well be a synonym of
<taxonomicName id="1EC64D2E1327BC5CFF6DBE11FEABEB24" authorityName="Roewer" authorityYear="1960" box="[189,302,591,617]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="EBB2EABF1327BC5CFF6DBE11FEABEB24" box="[189,302,591,617]" italics="true" pageId="43" pageNumber="44">A. magna</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="D97936AD1327BC5CFF6DBECFFDB2EBE3" blockId="43.[189,567,657,686]" box="[189,567,657,686]" pageId="43" pageNumber="44">
<emphasis id="EBB2EABF1327BC5CFF6DBECFFDB2EBE3" bold="true" box="[189,567,657,686]" italics="true" pageId="43" pageNumber="44">Undescribed putative species</emphasis>
</paragraph>
<paragraph id="D97936AD1327BC5CFF6DBE98FE9CEAF9" blockId="43.[189,1399,710,948]" pageId="43" pageNumber="44">
The following specimens are not formally described either because of small sample sizes (two putative species are represented by only one specimen each) or because of unclear species limits. Our aim here is to direct future collecting efforts while avoiding the potential taxonomic problems resulting from formal descriptions based on inadequate data. For more specific justifications see individual descriptions below. During the final editing of the manuscript we received further specimens from southern
<collectingCountry id="A1D1763D1327BC5CFB6ABF0AFB6CEA23" box="[1210,1257,852,878]" name="Iran" pageId="43" pageNumber="44">Iran</collectingCountry>
(
<collectingRegion id="1B02F84F1327BC5CFB27BF0DFAF7EA23" box="[1271,1394,851,878]" country="Iran" name="Fars" pageId="43" pageNumber="44">Fars Prov.</collectingRegion>
, Khonj, Khan Cave,
<geoCoordinate id="BCF2506A1327BC5CFE7BBF29FD8EEADC" box="[427,523,887,913]" degrees="27.7" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="5555" value="27.7">27.7° N</geoCoordinate>
,
<geoCoordinate id="BCF2506A1327BC5CFDC9BF28FDF0EADC" box="[537,629,886,913]" degrees="53.3" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5555" value="53.3">53.3° E</geoCoordinate>
, M.S. Tahami leg., in ZMSU) that seem to represent yet another species.
</paragraph>
</subSubSection>
</treatment>
</document>