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<document id="7F5F06C0D003B4A8D8A711BC1D5499D0" ID-CLB-Dataset="24882" ID-DOI="10.11646/zootaxa.4698.1.1" ID-GBIF-Dataset="1c94057f-64de-4f19-b047-2223e4d664e8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3542876" ID-ZooBank="0B4482B9-F89F-45CA-B929-E9B098605CD5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573811688707" checkinUser="plazi" docAuthor="Calvanese, Victor C., Brescovit, Antonio D. &amp; Bonato, Lucio" docDate="2019" docId="06130E72A519FFFCFF34FE49FC19AD52" docLanguage="en" docName="zootaxa.4698.1.1.pdf" docOrigin="Zootaxa 4698 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Aphilodon indespectus Calvanese &amp; Brescovit &amp; Bonato 2019, new species" docType="treatment" docVersion="9" lastPageNumber="31" masterDocId="FA2A760AA504FFE2FFA3FFDCFFCBAC35" masterDocTitle="Revision of the Neotropical species of Aphilodontinae (Geophilomorpha, Geophilidae), with eight new species and a first phylogenetic analysis of the subfamily" masterLastPageNumber="72" masterPageNumber="1" pageNumber="30" updateTime="1698767377376" updateUser="ExternalLinkService">
<mods:mods id="E858209966821E09E16BB988B8AE69EF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A559B6A448A67F19E9F7AEB271FE2669">
<mods:title id="7C7005E0E06FFB8E6D63096E522E94BD">Revision of the Neotropical species of Aphilodontinae (Geophilomorpha, Geophilidae), with eight new species and a first phylogenetic analysis of the subfamily</mods:title>
</mods:titleInfo>
<mods:name id="67E358DA36D5ECCE5E7BF4FDD26F6D95" type="personal">
<mods:role id="B5E4EFE429C9F46ED060133627986368">
<mods:roleTerm id="F024A60B7CB01FD2BFF44B5E87AF01CB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F064473B953911C393511F4180D94763">Calvanese, Victor C.</mods:namePart>
</mods:name>
<mods:name id="9F71D2CFD25EBAAF0846E50204262B0B" type="personal">
<mods:role id="6D0E3E287ED4DA70B00BE054CE11D35D">
<mods:roleTerm id="08B86B259B3D40BAE3B71A0985B765BE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4453EE7B1FD48388521F0135430CF882">Brescovit, Antonio D.</mods:namePart>
</mods:name>
<mods:name id="5322F257E074AD4AE52B55452BAA61E6" type="personal">
<mods:role id="C8A1F9BFF1353689C1C994CD7CE753DD">
<mods:roleTerm id="D41289A544EB794D930330FBA95772F7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="36ECC4A04255AE8825F211CF26F1DCE3">Bonato, Lucio</mods:namePart>
</mods:name>
<mods:typeOfResource id="5D6FD6B437C3703D99828BAF2A75E14F">text</mods:typeOfResource>
<mods:relatedItem id="D205A6A4E87EAD380A47C2DED3BF861F" type="host">
<mods:titleInfo id="FD4AA2BFEA3CF580AFD49DED3E31CE52">
<mods:title id="C5C8A7172C6708DFD65FD43D7F6CA0FE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2BD5EB90CE0E9B2AFC995A18DDE59C32">
<mods:date id="6F1AA0495AAD5423A2FF6801701E15C8">2019</mods:date>
<mods:detail id="A4AC5F38A13D2B1E07179F3FE4940A50" type="pubDate">
<mods:number id="E5C2E2099C3560C41A6459C569A9C0B5">2019-11-15</mods:number>
</mods:detail>
<mods:detail id="68757B904C59839C81ED07F58C0C7745" type="volume">
<mods:number id="E50FAC65FB931E3850B19AE66979852B">4698</mods:number>
</mods:detail>
<mods:detail id="A3D4CA781836E82A7C685E2AC4C6F997" type="issue">
<mods:number id="70551BB0BC539527C161503D5FA4E81C">1</mods:number>
</mods:detail>
<mods:extent id="78AEF3E40016A7E9C764FD0E7D764003" unit="page">
<mods:start id="D03F5EC9694300D0DEDC1A629A37428D">1</mods:start>
<mods:end id="E6C79E9E254C3A449AC6A9F02F4D6A78">72</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="C5F250A93EA94751E1143174D7B4DD3A">journal article</mods:classification>
<mods:identifier id="50B258DDA94C3311C887AD2B43F54CB2" type="CLB-Dataset">24882</mods:identifier>
<mods:identifier id="1108CE32D57451D1C03B3D6035C28665" type="DOI">10.11646/zootaxa.4698.1.1</mods:identifier>
<mods:identifier id="5A0E018F7332AF080A53340D2A39CB71" type="GBIF-Dataset">1c94057f-64de-4f19-b047-2223e4d664e8</mods:identifier>
<mods:identifier id="A55B6145345B058D889013AF711672C7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1C246B79A5C219A536BE44CB2E694565" type="Zenodo-Dep">3542876</mods:identifier>
<mods:identifier id="2DA71A18ABF3DECF743AC3D62B8C7AA0" type="ZooBank">0B4482B9-F89F-45CA-B929-E9B098605CD5</mods:identifier>
</mods:mods>
<treatment id="06130E72A519FFFCFF34FE49FC19AD52" ID-DOI="http://doi.org/10.5281/zenodo.5623807" ID-GBIF-Taxon="160406606" ID-Zenodo-Dep="5623807" LSID="urn:lsid:plazi:treatment:06130E72A519FFFCFF34FE49FC19AD52" httpUri="http://treatment.plazi.org/id/06130E72A519FFFCFF34FE49FC19AD52" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<subSubSection id="C6A0ECEFA519FFFFFF34FE49FD8AAD9A" box="[151,577,405,432]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="8E05BF64A519FFFFFF34FE49FD8AAD9A" blockId="29.[151,577,405,467]" box="[151,577,405,432]" pageId="29" pageNumber="30">
<heading id="D54D0808A519FFFFFF34FE49FD8AAD9A" bold="true" box="[151,577,405,432]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
<emphasis id="BCCE6376A519FFFFFF34FE49FD8AAD9A" bold="true" box="[151,577,405,432]" pageId="29" pageNumber="30">
<taxonomicName id="49BAC4E7A519FFFFFF34FE49FE61AD85" authority="Calvanese &amp; Brescovit &amp; Bonato, 2019" authorityName="Calvanese &amp; Brescovit &amp; Bonato" authorityYear="2019" box="[151,426,405,432]" class="Chilopoda" family="Aphilodontidae" genus="Aphilodon" kingdom="Animalia" order="Geophilomorpha" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="indespectus" status="sp. nov.">
<emphasis id="BCCE6376A519FFFFFF34FE49FE61AD85" bold="true" box="[151,426,405,432]" italics="true" pageId="29" pageNumber="30">Aphilodon indespectus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A7FDDE0DA519FFFFFE12FE49FD8AAD9A" box="[433,577,405,431]" pageId="29" pageNumber="30" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C6A0ECEFA519FFFFFF34FE65FDE8ADE6" box="[151,547,440,467]" pageId="29" pageNumber="30" type="description">
<paragraph id="8E05BF64A519FFFFFF34FE65FDE8ADE6" blockId="29.[151,577,405,467]" box="[151,547,440,467]" pageId="29" pageNumber="30">
(
<figureCitation id="1681A3E1A519FFFFFF3CFE65FF2DADE6" box="[159,230,441,467]" captionStart="FIGURE 4" captionStartId="12.[151,250,1691,1716]" captionTargetBox="[157,1436,184,1637]" captionTargetId="figure@12.[152,1438,174,1644]" captionTargetPageId="12" captionText="FIGURE 4. A–H, Aphilodontinae antenna. Aphilodon pereirai n. sp. (IBSP 3784): A, left antenna, ventral. Mairata butantan n. sp. (IBSP 3663): B, left antenna, ventral. Aphilodon silvestrii n. sp. (IBSP 3665): C, last antennal article of the left antenna,,ventral. Aphilodon indespectus n. sp. (IBSP 3653): D, terminal part of the left antenna, lateral; E, terminal part of the last antennal article of the left antenna, lateral; F, claviform sensilla, ventral; G, specialized setae of type “a”, ventral; H, specialized setae of type “b”, ventral. Abbreviations: CSS: claviform sensilla; SSA: specialized setae of type “a”. SSB: specialized setae of type “b”. Scale bars: 0.5 (A–B), 0.1 (C–D), 0.025 (E), 0.005 (F–H) mm." figureDoi="http://doi.org/10.5281/zenodo.3542884" httpUri="https://zenodo.org/record/3542884/files/figure.png" pageId="29" pageNumber="30">Figs 4</figureCitation>
D–H, 5A, 21, 22, 39E, 42D)
</paragraph>
</subSubSection>
<subSubSection id="C6A0ECEFA519FFFFFF34FE26FC29AEFD" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="8E05BF64A519FFFFFF34FE26FAA4AE0D" blockId="29.[151,1437,505,2045]" pageId="29" pageNumber="30">
<emphasis id="BCCE6376A519FFFFFF34FE26FE92AE21" bold="true" box="[151,345,505,532]" pageId="29" pageNumber="30">Type specimens.</emphasis>
<materialsCitation id="3ED2B539A519FFFFFEFCFE26FDB8AE0D" ID-GBIF-Occurrence="2451575467" collectorName="V. Calvanese" country="Brazil" location="Parque Ecologico da Mata da Camara" pageId="29" pageNumber="30" specimenCount="1" specimenCount-female="1" stateProvince="Sao Paulo" typeStatus="holotype">
<typeStatus id="510101C6A519FFFFFEFCFE26FE07AE21" box="[351,460,506,532]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
:
<specimenCount id="98BC74EDA519FFFFFE75FE27FE20AE20" box="[470,491,507,533]" pageId="29" pageNumber="30" type="female">♀</specimenCount>
, from
<location id="8B65E9BFA519FFFFFD95FE27FC15AE21" LSID="urn:lsid:plazi:treatment:06130E72A519FFFCFF34FE49FC19AD52:8B65E9BFA519FFFFFD95FE27FC15AE21" box="[566,990,506,533]" country="Brazil" name="Parque Ecologico da Mata da Camara" pageId="29" pageNumber="30" stateProvince="Sao Paulo">Parque Ecológico da Mata da Câmara</location>
, São Roque,
<collectingRegion id="4C7E7186A519FFFFFBD3FE26FB2FAE21" box="[1136,1252,506,532]" country="Brazil" name="Sao Paulo" pageId="29" pageNumber="30">São Paulo</collectingRegion>
,
<collectingCountry id="F6ADFFF4A519FFFFFB4DFE26FAF9AE21" box="[1262,1330,506,532]" name="Brazil" pageId="29" pageNumber="30">Brazil</collectingCountry>
06/2014,
<collectorName id="234FDAB2A519FFFFFF34FDC3FEE2AE0D" box="[151,297,542,569]" pageId="29" pageNumber="30">V. Calvanese</collectorName>
col., deposited in IBSP 3653
</materialsCitation>
.
<materialsCitation id="3ED2B539A519FFFFFDDEFDC3FAA7AE0D" ID-GBIF-Occurrence="2451575464" box="[637,1388,541,569]" collectorName="V. Calvanese" country="Brazil" location="Parque Ecologico da Mata da Camara" pageId="29" pageNumber="30" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Sao Paulo" typeStatus="paratype">
<typeStatus id="510101C6A519FFFFFDDEFDC3FD25AE0C" box="[637,750,543,569]" pageId="29" pageNumber="30" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="98BC74EDA519FFFFFD5AFDC2FCD6AE0D" box="[761,797,542,568]" pageId="29" pageNumber="30" type="female">3♀</specimenCount>
,
<specimenCount id="98BC74EDA519FFFFFC89FDC2FC84AE0D" box="[810,847,542,568]" pageId="29" pageNumber="30" type="male">4♂</specimenCount>
, same data as holotype, deposited in IBSP 3653
</materialsCitation>
.
</paragraph>
<paragraph id="8E05BF64A519FFFFFF64FD9EFA5CAE69" blockId="29.[151,1437,505,2045]" box="[199,1431,577,605]" pageId="29" pageNumber="30">
<materialsCitation id="3ED2B539A519FFFFFF64FD9EFA58AE69" ID-GBIF-Occurrence="2451575469" box="[199,1427,577,605]" country="Brazil" latitude="-23.528055" location="Parque Ecologico da Mata da Camara" longLatPrecision="21" longitude="-47.114166" municipality="Sao Roque" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Sao Paulo" typeStatus="holotype">
<emphasis id="BCCE6376A519FFFFFF64FD9EFEADAE69" bold="true" box="[199,358,577,604]" pageId="29" pageNumber="30">
<typeStatus id="510101C6A519FFFFFF64FD9EFEC9AE69" box="[199,258,578,604]" pageId="29" pageNumber="30">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F6ADFFF4A519FFFFFECFFD9EFE7EAE69" box="[364,437,578,604]" name="Brazil" pageId="29" pageNumber="30">Brazil</collectingCountry>
:
<collectingRegion id="4C7E7186A519FFFFFE63FD9EFDFEAE69" box="[448,565,578,604]" country="Brazil" name="Sao Paulo" pageId="29" pageNumber="30">São Paulo</collectingRegion>
:
<collectingMunicipality id="6E61251EA519FFFFFD9CFD9EFD75AE68" box="[575,702,578,605]" pageId="29" pageNumber="30">São Roque</collectingMunicipality>
:
<location id="8B65E9BFA519FFFFFD6BFD9FFBBEAE69" LSID="urn:lsid:plazi:treatment:06130E72A519FFFCFF34FE49FC19AD52:8B65E9BFA519FFFFFD6BFD9FFBBEAE69" box="[712,1141,578,605]" country="Brazil" latitude="-23.528055" longLatPrecision="21" longitude="-47.114166" municipality="Sao Roque" name="Parque Ecologico da Mata da Camara" pageId="29" pageNumber="30" stateProvince="Sao Paulo">Parque Ecológico da Mata da Câmara</location>
:
<geoCoordinate id="EB8ED9A3A519FFFFFBDCFD9EFACEAE69" box="[1151,1285,578,604]" degrees="23" direction="south" minutes="31" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="41" value="-23.528055">23°31’41”S</geoCoordinate>
,
<geoCoordinate id="EB8ED9A3A519FFFFFAB2FD9DFA58AE69" box="[1297,1427,577,604]" degrees="47" direction="west" minutes="6" orientation="longitude" pageId="29" pageNumber="30" precision="15" seconds="51" value="-47.114166">47°6’51”W</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph id="8E05BF64A519FFFFFF64FDB9FC29AEFD" blockId="29.[151,1437,505,2045]" pageId="29" pageNumber="30">
<emphasis id="BCCE6376A519FFFFFF64FDB9FE7AAEB5" bold="true" box="[199,433,613,640]" pageId="29" pageNumber="30">Material examined.</emphasis>
<materialsCitation id="3ED2B539A519FFFFFE1BFDB9FE9AAE91" ID-GBIF-Occurrence="2451575466" collectionCode="IBSP" collectorName="J. M. Pereira" country="Brazil" latitude="-22.68361" location="Bananal" longLatPrecision="21" longitude="-44.323334" pageId="29" pageNumber="30" specimenCode="IBSP 3654" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Sao Paulo">
<emphasis id="BCCE6376A519FFFFFE1BFDB9FD63AEB5" bold="true" box="[440,680,613,640]" pageId="29" pageNumber="30">
<collectingCountry id="F6ADFFF4A519FFFFFE1BFDB9FDEEAEB5" box="[440,549,613,640]" name="Brazil" pageId="29" pageNumber="30">BRAZIL</collectingCountry>
.
<emphasis id="BCCE6376A519FFFFFD91FDBAFD63AEB5" bold="true" box="[562,680,613,640]" italics="true" pageId="29" pageNumber="30">
<collectingRegion id="4C7E7186A519FFFFFD91FDBAFD63AEB5" box="[562,680,613,640]" country="Brazil" name="Sao Paulo" pageId="29" pageNumber="30">São Paulo</collectingRegion>
</emphasis>
</emphasis>
:
<location id="8B65E9BFA519FFFFFD1BFDBAFCDFAEB5" LSID="urn:lsid:plazi:treatment:06130E72A519FFFCFF34FE49FC19AD52:8B65E9BFA519FFFFFD1BFDBAFCDFAEB5" box="[696,788,614,640]" country="Brazil" latitude="-22.68361" longLatPrecision="21" longitude="-44.323334" name="Bananal" pageId="29" pageNumber="30" stateProvince="Sao Paulo">Bananal</location>
(
<geoCoordinate id="EB8ED9A3A519FFFFFC87FDBAFC60AEB5" box="[804,939,614,640]" degrees="22" direction="south" minutes="41" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="01" value="-22.68361">22°41’01”S</geoCoordinate>
,
<geoCoordinate id="EB8ED9A3A519FFFFFC14FDBAFB81AEB5" box="[951,1098,614,640]" degrees="44" direction="west" minutes="19" orientation="longitude" pageId="29" pageNumber="30" precision="15" seconds="24" value="-44.323334">44°19’24”W</geoCoordinate>
), 2010,
<collectorName id="234FDAB2A519FFFFFB07FDBBFAF8AEB5" box="[1188,1331,614,641]" pageId="29" pageNumber="30">J. M. Pereira</collectorName>
col.,
<specimenCount id="98BC74EDA519FFFFFAD1FDBAFA5DAEB5" box="[1394,1430,614,640]" pageId="29" pageNumber="30" type="female">3♀</specimenCount>
,
<specimenCount id="98BC74EDA519FFFFFF34FD56FF70AE91" box="[151,187,650,676]" pageId="29" pageNumber="30" type="male">3♂</specimenCount>
(
<specimenCode id="DE1C171FA519FFFFFF6AFD56FE82AE91" box="[201,329,649,676]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="29" pageNumber="30">IBSP 3654</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3ED2B539A519FFFFFEFFFD56FDF0AEFD" ID-GBIF-Occurrence="2451575468" collectionCode="IBSP" collectorName="V. Calvanese" country="Brazil" latitude="-23.528055" location="Parque Ecologico da Mata da Camara" longLatPrecision="21" longitude="-47.114166" municipality="Sao Roque" pageId="29" pageNumber="30" specimenCode="IBSP 3653" specimenCount="1" specimenCount-female="1" stateProvince="Sao Paulo" typeStatus="holotype">
<collectingMunicipality id="6E61251EA519FFFFFEFFFD56FE12AE90" box="[348,473,650,677]" pageId="29" pageNumber="30">São Roque</collectingMunicipality>
,
<location id="8B65E9BFA519FFFFFE46FD57FC46AE91" LSID="urn:lsid:plazi:treatment:06130E72A519FFFCFF34FE49FC19AD52:8B65E9BFA519FFFFFE46FD57FC46AE91" box="[485,909,650,677]" country="Brazil" latitude="-23.528055" longLatPrecision="21" longitude="-47.114166" municipality="Sao Roque" name="Parque Ecologico da Mata da Camara" pageId="29" pageNumber="30" stateProvince="Sao Paulo">Parque Ecológico da Mata da Câmara</location>
(
<geoCoordinate id="EB8ED9A3A519FFFFFC3EFD56FBEEAE91" box="[925,1061,650,676]" degrees="23" direction="south" minutes="31" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="41" value="-23.528055">23°31’41”S</geoCoordinate>
,
<geoCoordinate id="EB8ED9A3A519FFFFFB93FD55FB7FAE91" box="[1072,1204,649,676]" degrees="47" direction="west" minutes="6" orientation="longitude" pageId="29" pageNumber="30" precision="15" seconds="51" value="-47.114166">47°6’51”W</geoCoordinate>
), 06/2014,
<collectorName id="234FDAB2A519FFFFFA90FD57FF02AEFC" pageId="29" pageNumber="30">V. Calvanese</collectorName>
col.,
<specimenCount id="98BC74EDA519FFFFFEA4FD72FEE1AEFD" box="[263,298,686,712]" pageId="29" pageNumber="30" type="female">1♀</specimenCount>
[
<typeStatus id="510101C6A519FFFFFE9BFD71FE6BAEFD" box="[312,416,685,712]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
] (
<specimenCode id="DE1C171FA519FFFFFE17FD72FDF8AEFD" box="[436,563,685,712]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="29" pageNumber="30">IBSP 3653</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3ED2B539A519FFFFFDE4FD72FC15AEFD" ID-GBIF-Occurrence="2451575463" box="[583,990,685,712]" collectionCode="IBSP" collectorName="V. Calvanese" country="Brazil" isTailEnumeration="true" latitude="-23.528055" location="Parque Ecologico da Mata da Camara" longLatPrecision="21" longitude="-47.114166" municipality="Sao Roque" pageId="29" pageNumber="30" specimenCode="IBSP 3653" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Sao Paulo" typeStatus="paratype">
<specimenCount id="98BC74EDA519FFFFFDE4FD72FDA2AEFD" box="[583,617,686,712]" pageId="29" pageNumber="30" type="female">3♀</specimenCount>
and
<specimenCount id="98BC74EDA519FFFFFD03FD72FD08AEFD" box="[672,707,686,712]" pageId="29" pageNumber="30" type="male">4♂</specimenCount>
[
<typeStatus id="510101C6A519FFFFFD73FD71FC8AAEFD" box="[720,833,685,712]" pageId="29" pageNumber="30" type="paratype">paratypes</typeStatus>
] (
<specimenCode id="DE1C171FA519FFFFFCF5FD72FC1DAEFD" box="[854,982,685,712]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="29" pageNumber="30">IBSP 3653</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C6A0ECEFA519FFFFFF64FD0EFE73AF25" pageId="29" pageNumber="30" type="etymology">
<paragraph id="8E05BF64A519FFFFFF64FD0EFE73AF25" blockId="29.[151,1437,505,2045]" pageId="29" pageNumber="30">
<emphasis id="BCCE6376A519FFFFFF64FD0EFE99AED9" bold="true" box="[199,338,722,748]" pageId="29" pageNumber="30">Etymology:</emphasis>
The epithet is an adjective and means “invisible”, referring to the difficulty in finding specimens of the species in the field.
</paragraph>
</subSubSection>
<subSubSection id="C6A0ECEFA519FFFFFF64FCC5FC01A8D1" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="8E05BF64A519FFFFFF64FCC5FE41A84D" blockId="29.[151,1437,505,2045]" pageId="29" pageNumber="30">
<emphasis id="BCCE6376A519FFFFFF64FCC5FE89AF01" bold="true" box="[199,322,793,820]" pageId="29" pageNumber="30">Diagnosis.</emphasis>
A species of
<taxonomicName id="49BAC4E7A519FFFFFE40FCC6FD93AF01" box="[483,600,794,820]" class="Chilopoda" family="Aphilodontidae" genus="Aphilodon" kingdom="Animalia" order="Geophilomorpha" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="BCCE6376A519FFFFFE40FCC6FD93AF01" box="[483,600,794,820]" italics="true" pageId="29" pageNumber="30">Aphilodon</emphasis>
</taxonomicName>
with: around 39–41 leg-bearing segments; known body length in adults
<quantity id="49421281A519FFFFFF34FCE2FED8AF6C" box="[151,275,830,857]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.9" metricValueMin="1.0" pageId="29" pageNumber="30" unit="mm" value="14.5" valueMax="19.0" valueMin="10.0">10–19 mm</quantity>
; cephalic plate length/width ratio ca. 1.1; antenna ca. 2 times as long as cephalic plate; pectinate lamella of mandible with straight denticles; coxosternite of second maxillae not distinctly shorter in middle part; telopodite of second maxillae with article 3 shorter than article 2, with 1 or 2 setae in apical or subapical position and a subapical sensillum; forcipular pretergite exposed; forcipule with lateral margin posteriorly straight; only vestigial forcipular denticles; tarsungulum ca. 0.7 times as long as trochanteroprefemur+femur; all pleurites relatively small and well-separated by membranous regions; each coxopleuron with ca. 10–15 coxal pores in adults, all sparse; ultimate presternite exposed and apparently divided in middle part; metasternite of ultimate leg-bearing segment length/width ratio ca. 0.5–0.6; ultimate legs 1.8 times as long as penultimate legs; tarsus 1 of ultimate legs ca. 2.0 times as long as tibia.
</paragraph>
<paragraph id="8E05BF64A519FFFFFF64FB5FFAFFA8F5" blockId="29.[151,1437,505,2045]" pageId="29" pageNumber="30">
<taxonomicName id="49BAC4E7A519FFFFFF64FB5FFEA0A8A9" authority="Calvanese &amp; Brescovit &amp; Bonato, 2019" authorityName="Calvanese &amp; Brescovit &amp; Bonato" authorityYear="2019" box="[199,363,1154,1181]" class="Chilopoda" family="Aphilodontidae" genus="Aphilodon" kingdom="Animalia" order="Geophilomorpha" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="indespectus" status="sp. nov.">
<emphasis id="BCCE6376A519FFFFFF64FB5FFEA0A8A9" box="[199,363,1154,1181]" italics="true" pageId="29" pageNumber="30">A. indespectus</emphasis>
</taxonomicName>
<emphasis id="BCCE6376A519FFFFFED0FB5EFE78A8A9" bold="true" box="[371,435,1154,1180]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A7FDDE0DA519FFFFFED0FB5EFE78A8A9" box="[371,435,1154,1180]" pageId="29" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from all other species of
<taxonomicName id="49BAC4E7A519FFFFFC89FB5EFC54A8A9" box="[810,927,1154,1180]" class="Chilopoda" family="Aphilodontidae" genus="Aphilodon" kingdom="Animalia" order="Geophilomorpha" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="BCCE6376A519FFFFFC89FB5EFC54A8A9" box="[810,927,1154,1180]" italics="true" pageId="29" pageNumber="30">Aphilodon</emphasis>
</taxonomicName>
by having only vestigial forcipular denticles (
<figureCitation id="1681A3E1A519FFFFFF3CFB7AFEC0A8F5" box="[159,267,1190,1216]" captionStart="FIGURE 21" captionStartId="30.[151,250,814,839]" captionTargetBox="[159,1429,378,791]" captionTargetId="figure@30.[151,1437,373,794]" captionTargetPageId="30" captionText="FIGURE 21. A–C, Aphilodon indespectus n. sp. Female paratype (IBSP 3653): A, forcipular segment, ventral; B, intermediate part of trunk, left side, lateral; C, terminal part of body, ventral. Color legend: red, scutellum; green, stigmatopleurite; lilac, posterior sclerite “I”; yellow, anterior sclerite “II”; light blue, anterior sclerite “III”; dark blue, anterior sclerite “III”. Abbreviations: TSU, tarsus. Scale bars: 0.3 mm (A–C)." figureDoi="http://doi.org/10.5281/zenodo.3542921" httpUri="https://zenodo.org/record/3542921/files/figure.png" pageId="29" pageNumber="30">Figs 21A</figureCitation>
,
<figureCitation id="1681A3E1A519FFFFFEB5FB7AFE82A8F5" box="[278,329,1190,1216]" captionStart="FIGURE 22" captionStartId="30.[151,250,1640,1665]" captionTargetBox="[157,1428,1002,1615]" captionTargetId="figure@30.[151,1436,997,1621]" captionTargetPageId="30" captionText="FIGURE 22. A–E, Aphilodon indespectus n. sp. Female paratype (IBSP 3653): A, habitus, ventral; B, anterior part of body, dorsal; C, same, ventral; D, posterior part of body, dorsal; E same, ventral. Scale bars: 1 (A), 0.5 (B–E) mm." figureDoi="http://doi.org/10.5281/zenodo.3542923" httpUri="https://zenodo.org/record/3542923/files/figure.png" pageId="29" pageNumber="30">22C</figureCitation>
), and all pleurites relatively small and well-separated by membranous regions (
<figureCitation id="1681A3E1A519FFFFFB63FB7AFAECA8F5" box="[1216,1319,1190,1216]" captionStart="FIGURE 21" captionStartId="30.[151,250,814,839]" captionTargetBox="[159,1429,378,791]" captionTargetId="figure@30.[151,1437,373,794]" captionTargetPageId="30" captionText="FIGURE 21. A–C, Aphilodon indespectus n. sp. Female paratype (IBSP 3653): A, forcipular segment, ventral; B, intermediate part of trunk, left side, lateral; C, terminal part of body, ventral. Color legend: red, scutellum; green, stigmatopleurite; lilac, posterior sclerite “I”; yellow, anterior sclerite “II”; light blue, anterior sclerite “III”; dark blue, anterior sclerite “III”. Abbreviations: TSU, tarsus. Scale bars: 0.3 mm (A–C)." figureDoi="http://doi.org/10.5281/zenodo.3542921" httpUri="https://zenodo.org/record/3542921/files/figure.png" pageId="29" pageNumber="30">Fig. 21B</figureCitation>
).
</paragraph>
<paragraph id="8E05BF64A519FFFFFF64FB16FC01A8D1" blockId="29.[151,1437,505,2045]" box="[199,970,1225,1253]" pageId="29" pageNumber="30">
<emphasis id="BCCE6376A519FFFFFF64FB16FE71A8D1" bold="true" box="[199,442,1225,1252]" pageId="29" pageNumber="30">Recorded specimens</emphasis>
: 14, including
<specimenCount id="98BC74EDA519FFFFFDC7FB17FD43A8D0" box="[612,648,1227,1253]" pageId="29" pageNumber="30" type="female">7♀</specimenCount>
and
<specimenCount id="98BC74EDA519FFFFFD1DFB17FD28A8D0" box="[702,739,1227,1253]" pageId="29" pageNumber="30" type="male">7♂</specimenCount>
, from two localities.
</paragraph>
</subSubSection>
<subSubSection id="C6A0ECEFA519FFFFFF64FB31FCFFA93C" box="[199,820,1261,1289]" pageId="29" pageNumber="30" type="distribution">
<paragraph id="8E05BF64A519FFFFFF64FB31FCFFA93C" blockId="29.[151,1437,505,2045]" box="[199,820,1261,1289]" pageId="29" pageNumber="30">
<emphasis id="BCCE6376A519FFFFFF64FB31FEAAA93D" bold="true" box="[199,353,1261,1288]" pageId="29" pageNumber="30">Distribution.</emphasis>
Brazil—São Paulo: Bananal; São Roque.
</paragraph>
</subSubSection>
<subSubSection id="C6A0ECEFA519FFFCFF64FACDFC19AD52" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="description">
<paragraph id="8E05BF64A519FFFFFF64FACDFD13A919" blockId="29.[151,1437,505,2045]" box="[199,728,1297,1324]" pageId="29" pageNumber="30">
<emphasis id="BCCE6376A519FFFFFF64FACDFE2EA919" bold="true" box="[199,485,1297,1324]" pageId="29" pageNumber="30">
Description of
<typeStatus id="510101C6A519FFFFFED4FACEFE2BA919" box="[375,480,1298,1324]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
.
</emphasis>
Female (IBSP 3653).
</paragraph>
<paragraph id="8E05BF64A519FFFFFF64FAEBFC3AA9AC" blockId="29.[151,1437,505,2045]" pageId="29" pageNumber="30">
<emphasis id="BCCE6376A519FFFFFF64FAEBFE87A965" box="[199,332,1334,1360]" italics="true" pageId="29" pageNumber="30">Entire body</emphasis>
: length 17, maximum width 0.5.
<emphasis id="BCCE6376A519FFFFFD70FAEAFCB1A965" box="[723,890,1334,1360]" italics="true" pageId="29" pageNumber="30">Cephalic plate</emphasis>
: more or less oval, length 0.49 and width 0.45 (length/width ratio 1.08), with relative long and sparse SE.
<emphasis id="BCCE6376A519FFFFFC8DFA87FC40A940" box="[814,907,1371,1397]" italics="true" pageId="29" pageNumber="30">Antenna</emphasis>
: length 0.9, 2.0 times as long as cephalic plate; AA II–XIII length/width ratio 0.58–0.91; LAA 2.0 times as long as AA XIII.
</paragraph>
<paragraph id="8E05BF64A519FFFFFF64FA7EFF2DAA79" blockId="29.[151,1437,505,2045]" pageId="29" pageNumber="30">
<emphasis id="BCCE6376A519FFFFFF64FA7EFEE1A989" box="[199,298,1442,1468]" italics="true" pageId="29" pageNumber="30">Clypeus:</emphasis>
4+4 LS, 1+1 PS, 9+9 SCS.
<emphasis id="BCCE6376A519FFFFFDC4FA7EFD19A989" box="[615,722,1442,1468]" italics="true" pageId="29" pageNumber="30">Mandible</emphasis>
: PL with 14 denticles.
<emphasis id="BCCE6376A519FFFFFC76FA7FFBB8A989" box="[981,1139,1442,1468]" italics="true" pageId="29" pageNumber="30">First maxillae</emphasis>
: left TFM length 0.05 and width 0.04; DA with 2 lateral apical SE; MPC with 2 sub-lateral SE and 1 or 2 little sub-apical SL.
<emphasis id="BCCE6376A519FFFFFB7FFA1AFA57A9D5" box="[1244,1436,1478,1504]" italics="true" pageId="29" pageNumber="30">Second maxillae:</emphasis>
8 little SE distributed in a row along posterior margin of CSM, between TSM; left TSM length 0.08 (1.86 times as long as TFM) and maximum width (basal margin) 0.04; both TSM with 2 subapical SE and one little SL in apical region.
</paragraph>
<paragraph id="8E05BF64A519FFFFFF64F98AFBC5AB35" blockId="29.[151,1437,505,2045]" pageId="29" pageNumber="30">
<emphasis id="BCCE6376A519FFFFFF64F98AFE6FAA45" box="[199,420,1622,1648]" italics="true" pageId="29" pageNumber="30">Forcipular segment</emphasis>
: FTE sub-trapezoidal to rectangular, length 0.32 (0.65 times as long as cephalic plate) and maximum width 0.85, with scattered SE; FPT length 0.06 and width 0.69; FCX length 0.28 and width 0.32 (length/ width ratio 0.87); FRL length 0.24, not reaching anterior margin of cephalic plate, with 4 vestigial denticles and with 1 SE in position corresponding to FD1, 5 SE in position corresponding to FD2 and 3 SE in position corresponding to FD3; TA 0.75 times as long as TFF; DT very short, detected only with SEM.
</paragraph>
<paragraph id="8E05BF64A519FFFFFF64F8D6FEEDABC9" blockId="29.[151,1437,505,2045]" pageId="29" pageNumber="30">
<emphasis id="BCCE6376A519FFFFFF64F8D6FE9CAB11" box="[199,343,1802,1828]" italics="true" pageId="29" pageNumber="30">Walking legs</emphasis>
: 41 pairs; FL 0.70 times as long as second leg, with procoxae extended to body midline; leg 25 length 0.75.
<emphasis id="BCCE6376A519FFFFFE81F8F3FC54AB7D" box="[290,927,1838,1864]" italics="true" pageId="29" pageNumber="30">Tergum from the first to penultimate leg-bearing segment</emphasis>
: TE rectangular, with slightly rounded edges; TE 25 length 0.28 and width 0.56 (length/width ratio 0.5), with relatively large and scattered SE; PT also elongated, 0.6 times as long as TE, with relatively large and scattered SE.
<emphasis id="BCCE6376A519FFFFFCCFF8ABFF38AB81" italics="true" pageId="29" pageNumber="30">Sternum from the first to penultimate leg-bearing segment</emphasis>
: STF oval; other ST elongated; ST 25 length 0.32 and width 0.26 (length/width ratio 1.23); PST visible in most anterior segments, apparently divided up to segment 7, entirely and broadly exposed in middle and terminal part of body.
</paragraph>
<paragraph id="8E05BF64A51AFFFCFF64FF45FE52AD2A" blockId="30.[151,1437,153,359]" pageId="30" pageNumber="31">
<emphasis id="BCCE6376A51AFFFCFF64FF45FDD1AC86" box="[199,538,153,179]" italics="true" pageId="30" pageNumber="31">Ultimate leg-bearing segment</emphasis>
: MTL longer than wide, trapezoidal, length 0.11 and width 0.21 (length/width ratio 0.52); UPS entire.
<emphasis id="BCCE6376A51AFFFCFE0FFF61FD85ACE2" box="[428,590,189,215]" italics="true" pageId="30" pageNumber="31">Ultimate legs:</emphasis>
thin, length 1.1, ca. 1.8 times as long as penultimate leg; each CXP with 12–13 COP, scattered, smallest pore the closest to MTL; tarsus I ca. 2.0 times as long as tibia.
<emphasis id="BCCE6376A51AFFFCFB16FF3DFA57ACCE" box="[1205,1436,225,251]" italics="true" pageId="30" pageNumber="31">Postpedal segments:</emphasis>
GF with 2+2 distal SE.
</paragraph>
<paragraph id="8E05BF64A51AFFFCFF64FEF5FC19AD52" blockId="30.[151,1437,153,359]" pageId="30" pageNumber="31">
<emphasis id="BCCE6376A51AFFFCFF64FEF5FE51AD76" bold="true" box="[199,410,296,323]" pageId="30" pageNumber="31">Known variation.</emphasis>
<emphasis id="BCCE6376A51AFFFCFE01FEF5FE17AD76" box="[418,476,297,323]" italics="true" pageId="30" pageNumber="31">Male</emphasis>
: ultimate legs thicker and hairy, with tarsus I ca. 1.6 times as long as tibia; GM biarticulated; GM1 with 5–6 SE arranged in 2 rows; GM2 with 6–8 distal SE.
</paragraph>
</subSubSection>
</treatment>
</document>
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