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<document id="E566572C1DA1971BC3171DF047753BCB" ID-CLB-Dataset="28412" ID-DOI="10.11646/zootaxa.4565.1.1" ID-GBIF-Dataset="e9ffcfc7-9f7c-4212-ae6e-b842b31de6e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="2589489" ID-ZooBank="B9919693-2478-4089-B82E-C25EDFDB2A96" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552246080949" checkinUser="plazi" docAuthor="Chen, Hao, Zhang, Hui, Chen, Yongxia &amp; Freyhof, Jörg" docDate="2019" docId="5212F95FFFD2AA75B5CAF8CBFE613081" docLanguage="en" docName="zootaxa.4565.1.1.pdf" docOrigin="Zootaxa 4565 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Barbatula pechiliensis" docType="treatment" docVersion="9" lastPageNumber="19" masterDocId="AE2B8127FFDCAA67B55DFFF6FF9E324D" masterDocTitle="A review of the Barbatula loaches (Teleostei: Nemacheilidae) from north-eastern China, with the description of four new species" masterLastPageNumber="36" masterPageNumber="1" pageNumber="15" updateTime="1698701226448" updateUser="ExternalLinkService">
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<mods:title id="D10D06E23ECCFBCFF402A801E52F5957">A review of the Barbatula loaches (Teleostei: Nemacheilidae) from north-eastern China, with the description of four new species</mods:title>
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<mods:namePart id="C83F8035AC81AEB0EBEE45B787A62470">Chen, Hao</mods:namePart>
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<mods:namePart id="ADA013AC4FD5CFE96F9B02E6FFC74E4A">Chen, Yongxia</mods:namePart>
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<treatment id="5212F95FFFD2AA75B5CAF8CBFE613081" ID-DOI="http://doi.org/10.5281/zenodo.5923827" ID-GBIF-Taxon="154366722" ID-Zenodo-Dep="5923827" LSID="urn:lsid:plazi:treatment:5212F95FFFD2AA75B5CAF8CBFE613081" httpUri="http://treatment.plazi.org/id/5212F95FFFD2AA75B5CAF8CBFE613081" lastPageId="18" lastPageNumber="19" pageId="14" pageNumber="15">
<subSubSection id="92A11BC2FFD2AA69B5CAF8CBFDC13515" box="[151,607,1853,1880]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="DA044849FFD2AA69B5CAF8CBFDC13515" blockId="14.[151,607,1853,1913]" box="[151,607,1853,1880]" pageId="14" pageNumber="15">
<heading id="814CFF25FFD2AA69B5CAF8CBFDC13515" bold="true" box="[151,607,1853,1880]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<taxonomicName id="1DBB33CAFFD2AA69B5CAF8CBFDC13515" ID-CoL="46657" authority="(Fowler, 1899)" baseAuthorityName="Fowler" baseAuthorityYear="1899" box="[151,607,1853,1880]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="pechiliensis">
<emphasis id="E8CF945BFFD2AA69B5CAF8CBFDC13515" bold="true" box="[151,607,1853,1880]" pageId="14" pageNumber="15">
<emphasis id="E8CF945BFFD2AA69B5CAF8CBFE3B351A" bold="true" box="[151,421,1853,1879]" italics="true" pageId="14" pageNumber="15">Barbatula pechiliensis</emphasis>
(
<bibRefCitation id="BE2A35B8FFD2AA69B4E8F8C8FDC93515" author="Fowler, H. W." box="[437,599,1854,1880]" pageId="14" pageNumber="15" pagination="179 - 182" refId="ref16114" refString="Fowler, H. W. (1899) Notes on a small collection of Chinese fishes. Proceedings of the Academy of Natural Sciences of Philadelphia, 51, 179 - 182." type="journal article" year="1899">Fowler, 1899</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="92A11BC2FFD2AA69B5CAF896FCEA35B8" pageId="14" pageNumber="15" type="description">
<paragraph id="DA044849FFD2AA69B5CAF896FEBD3534" blockId="14.[151,607,1853,1913]" box="[151,291,1888,1913]" pageId="14" pageNumber="15">
(
<figureCitation id="428054CCFFD2AA69B5C2F896FE853534" box="[159,283,1888,1913]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStart-2="FIGURE 18" captionStartId-0="15.[151,250,1653,1675]" captionStartId-1="16.[151,250,1476,1498]" captionStartId-2="18.[151,250,1144,1166]" captionTargetBox-0="[233,1312,681,1627]" captionTargetBox-1="[151,1436,225,1405]" captionTargetBox-2="[151,1399,781,1123]" captionTargetId-0="figure@15.[151,1436,661,1631]" captionTargetId-1="figure@16.[151,1436,193,1454]" captionTargetId-2="figure@18.[151,1436,772,1123]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="18" captionText-0="FIGURE 16. Barbatula pechiliensis, from the top: HU1608897, 69.8 mm SL; HU 1805102, 90.77 mm SL; HU1600026, 63.6 mm SL; China: Hebei prov.: Luan River drainage; IHB200608002042, 69.4mm SL; China: Liaoning prov.: Daling River drainage." captionText-1="FIGURE 17. Barbatula pechiliensis, from the top: HU 1805102, 90.8 mm SL; China: Hebei prov.: Luan River drainage; HU 1804062, 76.8 mm SL; China: Inner Mongolia: Luan River drainage; HU 1608042, 47.9 mm SL; China: Liaoning prov.: Xiaoling River drainage; IHB 200608002042, 69.4mm SL; China: Liaoning prov.: Daling River drainage." captionText-2="FIGURE 18. Barbatula pechiliensis, ab: HU1805102, 90.77 mm SL; c: HU1600024, 79.4 mm SL; China: Hebei prov.: Luan River drainage. Scale bars: ab = 1 mm, c = 0.1 mm. Designations: m.i, median incision; m.le,; m.i, mental lobe lateral expansion; l.le, lower lip lateral expansion." httpUri-0="https://zenodo.org/record/2589521/files/figure.png" httpUri-1="https://zenodo.org/record/2589523/files/figure.png" httpUri-2="https://zenodo.org/record/2589525/files/figure.png" pageId="14" pageNumber="15">Fig. 1618</figureCitation>
)
</paragraph>
<paragraph id="DA044849FFD2AA69B5CAF854FCEA35B8" blockId="14.[151,1436,1954,2037]" pageId="14" pageNumber="15">
<emphasis id="E8CF945BFFD2AA69B5CAF854FEED35F5" box="[151,371,1954,1976]" italics="true" pageId="14" pageNumber="15">Nemchilus pechiliens</emphasis>
<bibRefCitation id="BE2A35B8FFD2AA69B421F854FD9835F5" author="Fowler, H. W." box="[380,518,1954,1976]" pageId="14" pageNumber="15" pagination="179 - 182" refId="ref16114" refString="Fowler, H. W. (1899) Notes on a small collection of Chinese fishes. Proceedings of the Academy of Natural Sciences of Philadelphia, 51, 179 - 182." type="journal article" year="1899">Fowler, 1899</bibRefCitation>
(
<collectingCountry id="A2AC08D9FFD2AA69B74AF854FDC735F5" box="[535,601,1954,1976]" name="China" pageId="14" pageNumber="15">China</collectingCountry>
:
<collectingRegion id="187F86ABFFD2AA69B73BF855FC9535F5" box="[614,779,1954,1976]" country="China" name="Nei Mongol" pageId="14" pageNumber="15">Inner Mongolia</collectingRegion>
: Tan lan Ho River about
<quantity id="1D43E5ACFFD2AA69B143F854FBE735F5" box="[1054,1145,1954,1976]" metricMagnitude="4" metricUnit="m" metricValue="4.828032" pageId="14" pageNumber="15" unit="mi" value="30.0">30 miles</quantity>
northeast of Lama-miau or Dolon-nor)
<taxonomicName id="1DBB33CAFFD2AA69B5CAF829FDB735B8" authority=": Nichols, 1925: 3" authorityName=": Nichols" authorityPageNumber="3" authorityYear="1925" box="[151,553,2015,2037]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="subSpecies" species="toni" subSpecies="fowleri">
<emphasis id="E8CF945BFFD2AA69B5CAF829FEE435B8" box="[151,378,2015,2037]" italics="true" pageId="14" pageNumber="15">Barbatula toni fowleri</emphasis>
:
<bibRefCitation id="BE2A35B8FFD2AA69B4DAF829FD8735B8" author="Nichols, J T" box="[391,537,2015,2037]" pageId="14" pageNumber="15" pagination="1 - 7" refId="ref16500" refString="Nichols, J T. (1925) Nemacheilus and related loaches in China. American Museum Novitates, 171, 1 - 7." type="journal article" year="1925">Nichols, 1925</bibRefCitation>
:3
</taxonomicName>
(
<collectingCountry id="A2AC08D9FFD2AA69B76BF829FDE635B8" box="[566,632,2015,2037]" name="China" pageId="14" pageNumber="15">China</collectingCountry>
: Chihli: Eastern Tombs).
</paragraph>
</subSubSection>
<subSubSection id="92A11BC2FFD3AA68B5CAFF61FCEA35AD" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="DA044849FFD3AA68B5CAFF61FCEF302D" blockId="15.[151,1437,151,608]" pageId="15" pageNumber="16">
<emphasis id="E8CF945BFFD3AA68B5CAFF61FE1C32FD" bold="true" box="[151,386,151,176]" pageId="15" pageNumber="16">Material examined.</emphasis>
HU 180406264, 1805100107, 11,
<quantity id="1D43E5ACFFD3AA68B672FF61FC7C32E2" box="[815,994,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="8.125" metricValueMax="10.69" metricValueMin="5.56" pageId="15" pageNumber="16" unit="mm" value="81.25" valueMax="106.9" valueMin="55.6">55.6106.9 mm</quantity>
SL; FSJF 4045, 4,
<quantity id="1D43E5ACFFD3AA68B19BFF61FAF532E2" box="[1222,1387,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="9.84" metricValueMin="7.159999999999999" pageId="15" pageNumber="16" unit="mm" value="85.0" valueMax="98.4" valueMin="71.6">71.698.4 mm</quantity>
SL;
<materialsCitation id="6AD34214FFD3AA68B5CAFF4AFEF53351" ID-GBIF-Occurrence="2012822334" collectingDate="2018-05" country="China" county="Tuligen River" latitude="42.683334" location="upper Luan River" municipality="Dolon-nor" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Nei Mongol">
<collectingCountry id="A2AC08D9FFD3AA68B5CAFF4AFF403298" box="[151,222,188,213]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
:
<collectingRegion id="187F86ABFFD3AA68B5B2FF4BFE363298" box="[239,424,188,213]" country="China" name="Nei Mongol" pageId="15" pageNumber="16">Inner Mongolia</collectingRegion>
:
<collectingCounty id="336530C5FFD3AA68B4E4FF4AFDC43298" box="[441,602,188,213]" pageId="15" pageNumber="16">Tuligen River</collectingCounty>
<quantity id="1D43E5ACFFD3AA68B73AFF4AFD2B3298" box="[615,693,188,213]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="15" pageNumber="16" unit="km" value="45.0">45 km</quantity>
north-east of
<collectingMunicipality id="3A60D233FFD3AA68B63EFF4AFC443298" box="[867,986,188,213]" pageId="15" pageNumber="16">Dolon-nor</collectingMunicipality>
,
<location id="DF641E92FFD3AA68B6B6FF4BFB233298" LSID="urn:lsid:plazi:treatment:5212F95FFFD2AA75B5CAF8CBFE613081:DF641E92FFD3AA68B6B6FF4BFB233298" box="[1003,1213,188,213]" country="China" county="Tuligen River" latitude="42.683334" municipality="Dolon-nor" name="upper Luan River" pageId="15" pageNumber="16" stateProvince="Nei Mongol">upper Luan River</location>
drainage,
<geoCoordinate id="BF8F2E8EFFD3AA68B062FF4AFA023298" box="[1343,1436,188,213]" direction="north" orientation="latitude" pageId="15" pageNumber="16" precision="925" value="42.683334">42°41N</geoCoordinate>
116°93E (4);
<location id="DF641E92FFD3AA68B46BFF29FE4732B5" LSID="urn:lsid:plazi:treatment:5212F95FFFD2AA75B5CAF8CBFE613081:DF641E92FFD3AA68B46BFF29FE4732B5" box="[310,473,223,248]" country="China" county="Tuligen River" latitude="42.683334" municipality="Dolon-nor" name="Yongxia Chen" pageId="15" pageNumber="16" stateProvince="Nei Mongol">Yongxia Chen</location>
,
<date id="AE056E89FFD3AA68B4B8FF16FDC732B5" box="[485,601,224,248]" pageId="15" pageNumber="16" value="2018-05">
<collectingDate id="BE419761FFD3AA68B4B8FF16FDC732B5" box="[485,601,224,248]" pageId="15" pageNumber="16" value="2018-05">May 2018</collectingDate>
</date>
.—HU 160002430, 1608881, 1608993, 1608897, 1608899, 1608900, 12,
<quantity id="1D43E5ACFFD3AA68B5CAFEF2FEA33351" box="[151,317,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="7.695" metricValueMax="9.44" metricValueMin="5.95" pageId="15" pageNumber="16" unit="mm" value="76.95" valueMax="94.4" valueMin="59.5">59.594.4 mm</quantity>
SL
</materialsCitation>
;
<materialsCitation id="6AD34214FFD3AA68B427FEF2FAD8330D" ID-GBIF-Occurrence="2012822335" collectingDate="2015-09" country="China" county="Xiaoluan River" location="upper Luan River" longitude="117.01667" municipality="Guojiatunzhen" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Hebei">
<collectingCountry id="A2AC08D9FFD3AA68B427FEF2FE5F3350" box="[378,449,260,285]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
:
<collectingRegion id="187F86ABFFD3AA68B492FEF2FD8C3350" box="[463,530,260,285]" country="China" name="Hebei" pageId="15" pageNumber="16">Hebei</collectingRegion>
prov.:
<collectingCounty id="336530C5FFD3AA68B73AFEF2FC883350" box="[615,790,260,285]" pageId="15" pageNumber="16">Xiaoluan River</collectingCounty>
<quantity id="1D43E5ACFFD3AA68B67CFEF2FCF53350" box="[801,875,260,285]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="15" pageNumber="16" unit="km" value="18.0">18 km</quantity>
north-west of
<collectingMunicipality id="3A60D233FFD3AA68B147FEF2FB5C3350" box="[1050,1218,260,285]" pageId="15" pageNumber="16">Guojiatunzhen</collectingMunicipality>
,
<location id="DF641E92FFD3AA68B18DFEF3FA023350" LSID="urn:lsid:plazi:treatment:5212F95FFFD2AA75B5CAF8CBFE613081:DF641E92FFD3AA68B18DFEF3FA023350" box="[1232,1436,260,285]" country="China" county="Xiaoluan River" longitude="117.01667" municipality="Guojiatunzhen" name="upper Luan River" pageId="15" pageNumber="16" stateProvince="Hebei">upper Luan River</location>
drainage, 41°69N
<geoCoordinate id="BF8F2E8EFFD3AA68B434FED1FE4E330D" box="[361,464,295,320]" direction="east" orientation="longitude" pageId="15" pageNumber="16" precision="925" value="117.01667">117°01E</geoCoordinate>
(2);
<location id="DF641E92FFD3AA68B75AFED1FD34330D" LSID="urn:lsid:plazi:treatment:5212F95FFFD2AA75B5CAF8CBFE613081:DF641E92FFD3AA68B75AFED1FD34330D" box="[519,682,295,320]" country="China" county="Xiaoluan River" longitude="117.01667" municipality="Guojiatunzhen" name="Yongxia Chen" pageId="15" pageNumber="16" stateProvince="Hebei">Yongxia Chen</location>
,
<date id="AE056E89FFD3AA68B7E8FEDEFCB3330D" box="[693,813,295,320]" pageId="15" pageNumber="16" value="2015-09">
<collectingDate id="BE419761FFD3AA68B7E8FEDEFCB3330D" box="[693,813,295,320]" pageId="15" pageNumber="16" value="2015-09">Sept. 2015</collectingDate>
</date>
.—HU 1608292, 1608311, 2,
<quantity id="1D43E5ACFFD3AA68B124FED1FA823372" box="[1145,1308,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="6.49" metricValueMax="6.75" metricValueMin="6.2299999999999995" pageId="15" pageNumber="16" unit="mm" value="64.9" valueMax="67.5" valueMin="62.3">62.367.5 mm</quantity>
SL
</materialsCitation>
;
<materialsCitation id="6AD34214FFD3AA68B00FFED1FC6933C5" ID-GBIF-Occurrence="2012822320" collectingDate="2015-09" country="China" county="Shandian River" location="upper Luan River" municipality="Shandian River Reservoir" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Hebei">
<collectingCountry id="A2AC08D9FFD3AA68B00FFED1FA07330D" box="[1362,1433,295,320]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
:
<collectingRegion id="187F86ABFFD3AA68B5CAFEBAFF443328" box="[151,218,332,357]" country="China" name="Hebei" pageId="15" pageNumber="16">Hebei</collectingRegion>
prov.:
<collectingCounty id="336530C5FFD3AA68B469FEBAFE793328" box="[308,487,332,357]" pageId="15" pageNumber="16">Shandian River</collectingCounty>
<quantity id="1D43E5ACFFD3AA68B4A9FEBAFDAD3328" box="[500,563,332,357]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="16" unit="km" value="4.0">4 km</quantity>
north of
<collectingMunicipality id="3A60D233FFD3AA68B7F1FEBAFC453328" box="[684,987,332,357]" pageId="15" pageNumber="16">Shandian River Reservoir</collectingMunicipality>
,
<location id="DF641E92FFD3AA68B6B1FEBBFB233328" LSID="urn:lsid:plazi:treatment:5212F95FFFD2AA75B5CAF8CBFE613081:DF641E92FFD3AA68B6B1FEBBFB233328" box="[1004,1213,332,357]" country="China" county="Shandian River" municipality="Shandian River Reservoir" name="upper Luan River" pageId="15" pageNumber="16" stateProvince="Hebei">upper Luan River</location>
drainage, 41°68N 115°79E (3);
<date id="AE056E89FFD3AA68B462FE86FE2333C5" box="[319,445,367,392]" pageId="15" pageNumber="16" value="2015-09">
<collectingDate id="BE419761FFD3AA68B462FE86FE2333C5" box="[319,445,367,392]" pageId="15" pageNumber="16" value="2015-09">Sept. 2015</collectingDate>
</date>
.—HU 1608309, 1608314, 2,
<quantity id="1D43E5ACFFD3AA68B642FE99FC5933CA" box="[799,967,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="7.375" metricValueMax="7.62" metricValueMin="7.13" pageId="15" pageNumber="16" unit="mm" value="73.75" valueMax="76.2" valueMin="71.3">71.376.2 mm</quantity>
SL
</materialsCitation>
;
<materialsCitation id="6AD34214FFD3AA68B15AFE99FCBB339D" ID-GBIF-Occurrence="2012822314" collectingDate="2015-09" country="China" county="Beiwanzi of Fengning Manchu Autonomous County" latitude="41.05" location="upper Hai River" longLatPrecision="1159" longitude="116.78333" municipality="Tang River" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Hebei">
<collectingCountry id="A2AC08D9FFD3AA68B15AFE99FBD033C5" box="[1031,1102,367,392]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
:
<collectingRegion id="187F86ABFFD3AA68B103FE99FB3F33C5" box="[1118,1185,367,392]" country="China" name="Hebei" pageId="15" pageNumber="16">Hebei</collectingRegion>
prov.:
<collectingMunicipality id="3A60D233FFD3AA68B1A7FE86FAE233C5" box="[1274,1404,367,392]" pageId="15" pageNumber="16">Tang River</collectingMunicipality>
at
<collectingCounty id="336530C5FFD3AA68B5CAFE62FC9F33E1" box="[151,769,404,429]" pageId="15" pageNumber="16">Beiwanzi of Fengning Manchu Autonomous County</collectingCounty>
,
<location id="DF641E92FFD3AA68B64CFE63FC4E33E0" LSID="urn:lsid:plazi:treatment:5212F95FFFD2AA75B5CAF8CBFE613081:DF641E92FFD3AA68B64CFE63FC4E33E0" box="[785,976,404,429]" country="China" county="Beiwanzi of Fengning Manchu Autonomous County" latitude="41.05" longLatPrecision="1159" longitude="116.78333" municipality="Tang River" name="upper Hai River" pageId="15" pageNumber="16" stateProvince="Hebei">upper Hai River</location>
drainage,
<geoCoordinate id="BF8F2E8EFFD3AA68B10CFE62FB3033E0" box="[1105,1198,404,429]" direction="north" orientation="latitude" pageId="15" pageNumber="16" precision="925" value="41.05">41°03N</geoCoordinate>
<geoCoordinate id="BF8F2E8EFFD3AA68B1E6FE62FABC33E0" box="[1211,1314,404,429]" direction="east" orientation="longitude" pageId="15" pageNumber="16" precision="925" value="116.78333">116°47E</geoCoordinate>
(1);
<date id="AE056E89FFD3AA68B03EFE62FF4E339D" pageId="15" pageNumber="16" value="2015-09">
<collectingDate id="BE419761FFD3AA68B03EFE62FF4E339D" pageId="15" pageNumber="16" value="2015-09">Sept. 2015</collectingDate>
</date>
.—HU 160918587, 1608914, 4, 70.0
<quantity id="1D43E5ACFFD3AA68B7CFFE41FD663382" box="[658,760,439,464]" metricMagnitude="-2" metricUnit="m" metricValue="9.05" pageId="15" pageNumber="16" unit="mm" value="90.5">90.5 mm</quantity>
SL
</materialsCitation>
;
<materialsCitation id="6AD34214FFD3AA68B669FE41FDE83055" ID-GBIF-Occurrence="2012822327" collectingDate="2017-08" country="China" county="Yin River" latitude="42.566666" location="upper Liao River" municipality="Chutoulangzhen" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Nei Mongol">
<collectingCountry id="A2AC08D9FFD3AA68B669FE41FCE2339D" box="[820,892,439,464]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
:
<collectingRegion id="187F86ABFFD3AA68B6D5FE4EFBA0339D" box="[904,1086,439,464]" country="China" name="Nei Mongol" pageId="15" pageNumber="16">Inner Mongolia</collectingRegion>
:
<collectingCounty id="336530C5FFD3AA68B116FE41FB22339D" box="[1099,1212,439,464]" pageId="15" pageNumber="16">Yin River</collectingCounty>
<quantity id="1D43E5ACFFD3AA68B19BFE41FA9F339D" box="[1222,1281,439,464]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="16" unit="km" value="5.0">5 km</quantity>
north-east of
<collectingMunicipality id="3A60D233FFD3AA68B5CAFE2AFECC33B8" box="[151,338,476,501]" pageId="15" pageNumber="16">Chutoulangzhen</collectingMunicipality>
,
<location id="DF641E92FFD3AA68B43BFE2BFDA633B8" LSID="urn:lsid:plazi:treatment:5212F95FFFD2AA75B5CAF8CBFE613081:DF641E92FFD3AA68B43BFE2BFDA633B8" box="[358,568,476,501]" country="China" county="Yin River" latitude="42.566666" municipality="Chutoulangzhen" name="upper Liao River" pageId="15" pageNumber="16" stateProvince="Nei Mongol">upper Liao River</location>
drainage,
<geoCoordinate id="BF8F2E8EFFD3AA68B79DFE2AFC8333B8" box="[704,797,476,501]" direction="north" orientation="latitude" pageId="15" pageNumber="16" precision="925" value="42.566666">42°34N</geoCoordinate>
118°72E (6);
<date id="AE056E89FFD3AA68B683FE2AFBC133B9" box="[990,1119,476,500]" pageId="15" pageNumber="16" value="2017-08">
<collectingDate id="BE419761FFD3AA68B683FE2AFBC133B9" box="[990,1119,476,500]" pageId="15" pageNumber="16" value="2017-08">Aug. 2017</collectingDate>
</date>
.—HU 1608038, 1608040, 1608042, 1608044, 4,
<quantity id="1D43E5ACFFD3AA68B4FDFE09FDD9305A" box="[416,583,511,536]" metricMagnitude="-2" metricUnit="m" metricValue="5.04" metricValueMax="5.32" metricValueMin="4.76" pageId="15" pageNumber="16" unit="mm" value="50.4" valueMax="53.2" valueMin="47.6">47.653.2 mm</quantity>
SL
</materialsCitation>
;
<materialsCitation id="6AD34214FFD3AA68B7DBFE09FEBB302D" ID-GBIF-Occurrence="2012822326" collectingDate="2016-08" collectionCode="IHB" country="China" county="Nu'er River" location="upper Xiaoling River" longitude="120.6" municipality="Xintaimenzhen" pageId="15" pageNumber="16" specimenCode="IHB 2006082037-49, 2006082051-55, 18, 50.1, 88.3" specimenCount="1" stateProvince="Liaoning">
<collectingCountry id="A2AC08D9FFD3AA68B7DBFE09FD503055" box="[646,718,511,536]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
:
<collectingRegion id="187F86ABFFD3AA68B781FE09FCDF3055" box="[732,833,511,536]" country="China" name="Liaoning" pageId="15" pageNumber="16">Liaoning</collectingRegion>
prov.:
<collectingCounty id="336530C5FFD3AA68B6C5FDF6FBBB3055" box="[920,1061,511,536]" pageId="15" pageNumber="16">Nuer River</collectingCounty>
, south of
<collectingMunicipality id="3A60D233FFD3AA68B1C2FE09FAD33055" box="[1183,1357,511,536]" pageId="15" pageNumber="16">Xintaimenzhen</collectingMunicipality>
,
<location id="DF641E92FFD3AA68B001FDF6FEDF3070" LSID="urn:lsid:plazi:treatment:5212F95FFFD2AA75B5CAF8CBFE613081:DF641E92FFD3AA68B001FDF6FEDF3070" country="China" county="Nu'er River" longitude="120.6" municipality="Xintaimenzhen" name="upper Xiaoling River" pageId="15" pageNumber="16" stateProvince="Liaoning">upper Xiaoling River</location>
drainage, 40°84N
<geoCoordinate id="BF8F2E8EFFD3AA68B778FDD2FD133070" box="[549,653,548,573]" direction="east" orientation="longitude" pageId="15" pageNumber="16" precision="925" value="120.6">120°36E</geoCoordinate>
(9);
<date id="AE056E89FFD3AA68B797FDD2FCD83070" box="[714,838,548,573]" pageId="15" pageNumber="16" value="2016-08">
<collectingDate id="BE419761FFD3AA68B797FDD2FCD83070" box="[714,838,548,573]" pageId="15" pageNumber="16" value="2016-08">Aug. 2016</collectingDate>
</date>
.—
<specimenCode id="8A1DE032FFD3AA68B63AFDD2FBC43070" box="[871,1114,548,573]" collectionCode="IHB" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35160" lsid="urn:lsid:biocol.org:col:35160" name="Institute of Hydrobiology, Chinese Academy of Sciences" pageId="15" pageNumber="16">IHB 200608203749</specimenCode>
,
<specimenCode id="8A1DE032FFD3AA68B135FDD2FABC3070" box="[1128,1314,548,573]" collectionCode="IHB" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35160" lsid="urn:lsid:biocol.org:col:35160" name="Institute of Hydrobiology, Chinese Academy of Sciences" pageId="15" pageNumber="16">200608205155</specimenCode>
,
<specimenCode id="8A1DE032FFD3AA68B06DFDD2FAD03071" box="[1328,1358,548,572]" collectionCode="IHB" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35160" lsid="urn:lsid:biocol.org:col:35160" name="Institute of Hydrobiology, Chinese Academy of Sciences" pageId="15" pageNumber="16">18</specimenCode>
,
<quantity id="1D43E5ACFFD3AA68B000FDD2FF653012" metricMagnitude="-2" metricUnit="m" metricValue="6.92" metricValueMax="8.83" metricValueMin="5.01" pageId="15" pageNumber="16" unit="mm" value="69.2" valueMax="88.3" valueMin="50.1">
<specimenCode id="8A1DE032FFD3AA68B000FDD2FA133070" box="[1373,1421,548,573]" collectionCode="IHB" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35160" lsid="urn:lsid:biocol.org:col:35160" name="Institute of Hydrobiology, Chinese Academy of Sciences" pageId="15" pageNumber="16">50.1</specimenCode>
<specimenCode id="8A1DE032FFD3AA68B5CAFDBEFF56302D" box="[151,200,584,608]" collectionCode="IHB" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35160" lsid="urn:lsid:biocol.org:col:35160" name="Institute of Hydrobiology, Chinese Academy of Sciences" pageId="15" pageNumber="16">88.3</specimenCode>
mm
</quantity>
SL
</materialsCitation>
;
<materialsCitation id="6AD34214FFD3AA68B46CFDB1FCF3302D" ID-GBIF-Occurrence="2012822321" box="[305,877,583,608]" country="China" location="Daling River" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Liaoning">
<collectingCountry id="A2AC08D9FFD3AA68B46CFDB1FEE6302D" box="[305,376,583,608]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
:
<collectingRegion id="187F86ABFFD3AA68B4DEFDB1FE76302D" box="[387,488,583,608]" country="China" name="Liaoning" pageId="15" pageNumber="16">Liaoning</collectingRegion>
prov.:
<location id="DF641E92FFD3AA68B76BFDB1FD59302D" LSID="urn:lsid:plazi:treatment:5212F95FFFD2AA75B5CAF8CBFE613081:DF641E92FFD3AA68B76BFDB1FD59302D" box="[566,711,583,608]" country="China" name="Daling River" pageId="15" pageNumber="16" stateProvince="Liaoning">Daling River</location>
at Yixian (10)
</materialsCitation>
.
</paragraph>
<caption id="8EC418C1FFD3AA68B5CAF983FF6B3485" httpUri="https://zenodo.org/record/2589521/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,250,1653,1675]" targetBox="[233,1312,681,1627]" targetPageId="15">
<paragraph id="DA044849FFD3AA68B5CAF983FF6B3485" blockId="15.[151,1436,1652,1736]" pageId="15" pageNumber="16">
<emphasis id="E8CF945BFFD3AA68B5CAF983FEBE34C7" bold="true" box="[151,288,1653,1675]" pageId="15" pageNumber="16">FIGURE 16.</emphasis>
<taxonomicName id="1DBB33CAFFD3AA68B47BF982FD9534C7" baseAuthorityName="Fowler" baseAuthorityYear="1899" box="[294,523,1652,1674]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="pechiliensis">
<emphasis id="E8CF945BFFD3AA68B47BF982FD9534C7" box="[294,523,1652,1674]" italics="true" pageId="15" pageNumber="16">Barbatula pechiliensis</emphasis>
</taxonomicName>
, from the top: HU1608897, 69.8 mm SL; HU 1805102, 90.77 mm SL; HU1600026, 63.6 mm SL; China: Hebei prov.: Luan River drainage; IHB200608002042, 69.4mm SL; China: Liaoning prov.: Daling River drainage.
</paragraph>
</caption>
<paragraph id="DA044849FFD3AA68B59AF919FCEA35AD" blockId="15.[151,1437,1775,2016]" pageId="15" pageNumber="16">
<emphasis id="E8CF945BFFD3AA68B59AF919FCAF3545" bold="true" box="[199,817,1775,1800]" pageId="15" pageNumber="16">Material used in the molecular genetic analysis.</emphasis>
HU1608314;
<collectingCountry id="A2AC08D9FFD3AA68B6BEF919FBB53545" box="[995,1067,1775,1800]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
:
<collectingRegion id="187F86ABFFD3AA68B160F919FB1E3545" box="[1085,1152,1775,1800]" country="China" name="Hebei" pageId="15" pageNumber="16">Hebei</collectingRegion>
prov.: Tang River (1). (GenBank accession number:
<accessionNumber id="C5E8D5AAFFD3AA68B4B1F8E5FDF23561" box="[492,620,1811,1836]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY451917" pageId="15" pageNumber="16">KY451917</accessionNumber>
).—HU1600026;
<collectingCountry id="A2AC08D9FFD3AA68B66DF8E5FCE93561" box="[816,887,1811,1836]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
:
<collectingRegion id="187F86ABFFD3AA68B6DEF8E5FC583561" box="[899,966,1811,1836]" country="China" name="Hebei" pageId="15" pageNumber="16">Hebei</collectingRegion>
prov.: Xiaoluan River, upper Luan River drainage, 41°69N
<geoCoordinate id="BF8F2E8EFFD3AA68B42FF8C1FE47351D" box="[370,473,1847,1872]" direction="east" orientation="longitude" pageId="15" pageNumber="16" precision="925" value="117.01667">117°01E</geoCoordinate>
(2). (GenBank accession number:
<accessionNumber id="C5E8D5AAFFD3AA68B62EF8C1FC6B351D" box="[883,1013,1847,1872]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY451914" pageId="15" pageNumber="16">KY451914</accessionNumber>
).—HU1608311;
<collectingCountry id="A2AC08D9FFD3AA68B1E5F8C1FB61351D" box="[1208,1279,1847,1872]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
:
<collectingRegion id="187F86ABFFD3AA68B053F8C1FACF351D" box="[1294,1361,1847,1872]" country="China" name="Hebei" pageId="15" pageNumber="16">Hebei</collectingRegion>
prov.: Shandian River, upper Luan River drainage, 41°68N 115°79E (3). (GenBank accession number:
<accessionNumber id="C5E8D5AAFFD3AA68B1AEF8ADFAED3539" box="[1267,1395,1883,1908]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY451919" pageId="15" pageNumber="16">KY451919</accessionNumber>
).— HU1608040;
<collectingCountry id="A2AC08D9FFD3AA68B46DF889FEE935D5" box="[304,375,1919,1944]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
:
<collectingRegion id="187F86ABFFD3AA68B4DCF889FE7835D5" box="[385,486,1919,1944]" country="China" name="Liaoning" pageId="15" pageNumber="16">Liaoning</collectingRegion>
prov.: Nuer River, upper Xiaoling River drainage, 40°84N
<geoCoordinate id="BF8F2E8EFFD3AA68B1D1F889FB6A35D5" box="[1164,1268,1919,1944]" direction="east" orientation="longitude" pageId="15" pageNumber="16" precision="925" value="120.6">120°36E</geoCoordinate>
(9). (GenBank accession number:
<accessionNumber id="C5E8D5AAFFD3AA68B433F855FE7135F1" box="[366,495,1955,1980]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF770517" pageId="15" pageNumber="16">MF770517</accessionNumber>
).—HU1804062;
<collectingCountry id="A2AC08D9FFD3AA68B7EDF855FD6935F1" box="[688,759,1955,1980]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
:
<collectingRegion id="187F86ABFFD3AA68B65CF852FC2B35F1" box="[769,949,1955,1980]" country="China" name="Nei Mongol" pageId="15" pageNumber="16">Inner Mongolia</collectingRegion>
: Tuligen River, upper Luan River drainage,
<geoCoordinate id="BF8F2E8EFFD3AA68B5CAF831FF6B35AD" box="[151,245,1991,2016]" direction="north" orientation="latitude" pageId="15" pageNumber="16" precision="925" value="42.683334">42°41N</geoCoordinate>
116°93E (4) (GenBank accession number:
<accessionNumber id="C5E8D5AAFFD3AA68B7BFF831FCF735AD" box="[738,873,1991,2016]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH316129" pageId="15" pageNumber="16">MH316129</accessionNumber>
).
</paragraph>
</subSubSection>
<caption id="8EC418C1FFCCAA77B5CAFA32FB54345A" httpUri="https://zenodo.org/record/2589523/files/figure.png" pageId="16" pageNumber="17" startId="16.[151,250,1476,1498]" targetBox="[151,1436,225,1405]" targetPageId="16">
<paragraph id="DA044849FFCCAA77B5CAFA32FB54345A" blockId="16.[151,1436,1476,1559]" pageId="16" pageNumber="17">
<emphasis id="E8CF945BFFCCAA77B5CAFA32FEBE3794" bold="true" box="[151,288,1476,1498]" pageId="16" pageNumber="17">FIGURE 17.</emphasis>
<taxonomicName id="1DBB33CAFFCCAA77B47AFA32FD933797" baseAuthorityName="Fowler" baseAuthorityYear="1899" box="[295,525,1476,1498]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="pechiliensis">
<emphasis id="E8CF945BFFCCAA77B47AFA32FD933797" box="[295,525,1476,1498]" italics="true" pageId="16" pageNumber="17">Barbatula pechiliensis</emphasis>
</taxonomicName>
, from the top: HU 1805102, 90.8 mm SL; China: Hebei prov.: Luan River drainage; HU 1804062, 76.8 mm SL; China: Inner Mongolia: Luan River drainage; HU 1608042, 47.9 mm SL; China: Liaoning prov.: Xiaoling River drainage; IHB 200608002042, 69.4mm SL; China: Liaoning prov.: Daling River drainage.
</paragraph>
</caption>
<subSubSection id="92A11BC2FFCCAA77B59AF9C8FA0E35AE" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="DA044849FFCCAA77B59AF9C8FBA435F2" blockId="16.[151,1436,1598,2019]" pageId="16" pageNumber="17">
<emphasis id="E8CF945BFFCCAA77B59AF9C8FEDF341A" bold="true" box="[199,321,1598,1623]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
<taxonomicName id="1DBB33CAFFCCAA77B414F9C8FDD8341A" baseAuthorityName="Fowler" baseAuthorityYear="1899" box="[329,582,1598,1623]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="pechiliensis">
<emphasis id="E8CF945BFFCCAA77B414F9C8FDD8341A" box="[329,582,1598,1623]" italics="true" pageId="16" pageNumber="17">Barbatula pechiliensis</emphasis>
</taxonomicName>
is distinguished from the other species of
<taxonomicName id="1DBB33CAFFCCAA77B17AF9C8FB07341A" box="[1063,1177,1598,1623]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="E8CF945BFFCCAA77B17AF9C8FB07341A" box="[1063,1177,1598,1623]" italics="true" pageId="16" pageNumber="17">Barbatula</emphasis>
</taxonomicName>
in north-eastern
<collectingCountry id="A2AC08D9FFCCAA77B004F9C8FA02341A" box="[1369,1436,1598,1623]" name="China" pageId="16" pageNumber="17">China</collectingCountry>
by a combination of characters, none of them unique: a shallow median incision in the upper lip, its depth is 10- 40% of the width of the upper-lip (vs. deep, 40-60% in
<taxonomicName id="1DBB33CAFFCCAA77B654F97EFCFE34D2" baseAuthorityName="Bleeker" baseAuthorityYear="1865" box="[777,864,1670,1695]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="nuda">
<emphasis id="E8CF945BFFCCAA77B654F97EFCFE34D2" box="[777,864,1670,1695]" italics="true" pageId="16" pageNumber="17">B. nuda</emphasis>
</taxonomicName>
and
<taxonomicName id="1DBB33CAFFCCAA77B6C5F97EFC6934D2" authorityName="Cao, Causse &amp; Zhang" authorityYear="2012" box="[920,1015,1670,1695]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="gibba">
<emphasis id="E8CF945BFFCCAA77B6C5F97EFC6934D2" box="[920,1015,1670,1695]" italics="true" pageId="16" pageNumber="17">B. gibba</emphasis>
</taxonomicName>
), a blade-shaped lateral expansion of the mental lobe (vs. without expansion in
<taxonomicName id="1DBB33CAFFCCAA77B727F95AFD4D348E" baseAuthorityName="Bleeker" baseAuthorityYear="1865" box="[634,723,1706,1731]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="nuda">
<emphasis id="E8CF945BFFCCAA77B727F95AFD4D348E" box="[634,723,1706,1731]" italics="true" pageId="16" pageNumber="17">B. nuda</emphasis>
</taxonomicName>
,
<emphasis id="E8CF945BFFCCAA77B7BEF95AFC07348E" box="[739,921,1706,1731]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="1DBB33CAFFCCAA77B7BEF95AFCAE348E" baseAuthorityName="Dybowski" baseAuthorityYear="1869" box="[739,816,1708,1731]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="toni">B. toni</taxonomicName>
<taxonomicName id="1DBB33CAFFCCAA77B665F95AFC07348E" authorityName="Cao, Causse &amp; Zhang" authorityYear="2012" box="[824,921,1706,1731]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="gibba">B. gibba</taxonomicName>
</emphasis>
and
<taxonomicName id="1DBB33CAFFCCAA77B68EF95AFBFF348E" authorityName="Tchang" authorityYear="1932" box="[979,1121,1706,1731]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="kirinensis">
<emphasis id="E8CF945BFFCCAA77B68EF95AFBFF348E" box="[979,1121,1706,1731]" italics="true" pageId="16" pageNumber="17">B. kirinensis</emphasis>
</taxonomicName>
), the upper jaw completely covered by the upper lip (vs. partly covered in
<taxonomicName id="1DBB33CAFFCCAA77B7ECF926FC9734AA" baseAuthorityName="Bleeker" baseAuthorityYear="1865" box="[689,777,1742,1767]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="nuda">
<emphasis id="E8CF945BFFCCAA77B7ECF926FC9734AA" box="[689,777,1742,1767]" italics="true" pageId="16" pageNumber="17">B. nuda</emphasis>
</taxonomicName>
), a short lateral expansion of the lower lip, its length 60 90% of the width of the upper lip (vs. without expansion in
<taxonomicName id="1DBB33CAFFCCAA77B66EF902FC143546" baseAuthorityName="Bleeker" baseAuthorityYear="1865" box="[819,906,1778,1803]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="nuda">
<emphasis id="E8CF945BFFCCAA77B66EF902FC143546" box="[819,906,1778,1803]" italics="true" pageId="16" pageNumber="17">B. nuda</emphasis>
</taxonomicName>
), having the nostrils widely spaced (vs. closely set in
<taxonomicName id="1DBB33CAFFCCAA77B587F8EEFEAC3562" baseAuthorityName="Bleeker" baseAuthorityYear="1865" box="[218,306,1814,1839]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="nuda">
<emphasis id="E8CF945BFFCCAA77B587F8EEFEAC3562" box="[218,306,1814,1839]" italics="true" pageId="16" pageNumber="17">B. nuda</emphasis>
</taxonomicName>
), snout shorter than the postorbital head length (vs. equal to in
<taxonomicName id="1DBB33CAFFCCAA77B6A6F8EEFBCC3562" baseAuthorityName="Bleeker" baseAuthorityYear="1865" box="[1019,1106,1814,1839]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="nuda">
<emphasis id="E8CF945BFFCCAA77B6A6F8EEFBCC3562" box="[1019,1106,1814,1839]" italics="true" pageId="16" pageNumber="17">B. nuda</emphasis>
</taxonomicName>
and
<taxonomicName id="1DBB33CAFFCCAA77B1D4F8EEFB4A3562" baseAuthorityName="Dybowski" baseAuthorityYear="1869" box="[1161,1236,1816,1839]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="toni">
<emphasis id="E8CF945BFFCCAA77B1D4F8EEFB4A3562" box="[1161,1236,1816,1839]" italics="true" pageId="16" pageNumber="17">B. toni</emphasis>
</taxonomicName>
), a convex dorsal profile (caudal-peduncle depth 4358% of body depth vs. slightly convex, caudal-peduncle depth 6284% of body depth in
<taxonomicName id="1DBB33CAFFCCAA77B5A5F896FED1353A" baseAuthorityName="Bleeker" baseAuthorityYear="1865" box="[248,335,1886,1911]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="nuda">
<emphasis id="E8CF945BFFCCAA77B5A5F896FED1353A" box="[248,335,1886,1911]" italics="true" pageId="16" pageNumber="17">B. nuda</emphasis>
</taxonomicName>
and
<taxonomicName id="1DBB33CAFFCCAA77B4D8F896FD8F353A" authorityName="Tchang" authorityYear="1932" box="[389,529,1886,1911]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="kirinensis">
<emphasis id="E8CF945BFFCCAA77B4D8F896FD8F353A" box="[389,529,1886,1911]" italics="true" pageId="16" pageNumber="17">B. kirinensis</emphasis>
</taxonomicName>
), 7½ branched dorsal-fin rays (vs. 6½ in
<taxonomicName id="1DBB33CAFFCCAA77B682F896FB10353A" authorityName="Prokofiev" authorityYear="2007" box="[991,1166,1888,1911]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="potaninorum">
<emphasis id="E8CF945BFFCCAA77B682F896FB10353A" box="[991,1166,1888,1911]" italics="true" pageId="16" pageNumber="17">B. potaninorum</emphasis>
</taxonomicName>
) and scales restricted to the caudal peduncle (vs. sparsely set scales in front of the dorsal-fin origin in
<taxonomicName id="1DBB33CAFFCCAA77B171F872FBE535D6" baseAuthorityName="Dybowski" baseAuthorityYear="1869" box="[1068,1147,1924,1947]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="toni">
<emphasis id="E8CF945BFFCCAA77B171F872FBE535D6" box="[1068,1147,1924,1947]" italics="true" pageId="16" pageNumber="17">B. toni</emphasis>
</taxonomicName>
and
<taxonomicName id="1DBB33CAFFCCAA77B1E5F872FAD635D6" authorityName="Tchang" authorityYear="1932" box="[1208,1352,1922,1947]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="kirinensis">
<emphasis id="E8CF945BFFCCAA77B1E5F872FAD635D6" box="[1208,1352,1922,1947]" italics="true" pageId="16" pageNumber="17">B. kirinensis</emphasis>
</taxonomicName>
, scales restricted to the back and flank behind the dorsal-fin base in
<taxonomicName id="1DBB33CAFFCCAA77B660F85EFC0B35F2" baseAuthorityName="Bleeker" baseAuthorityYear="1865" box="[829,917,1958,1983]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="nuda">
<emphasis id="E8CF945BFFCCAA77B660F85EFC0B35F2" box="[829,917,1958,1983]" italics="true" pageId="16" pageNumber="17">B. nuda</emphasis>
</taxonomicName>
and
<taxonomicName id="1DBB33CAFFCCAA77B696F85EFBB435F2" authorityName="Cao, Causse &amp; Zhang" authorityYear="2012" box="[971,1066,1958,1983]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="gibba">
<emphasis id="E8CF945BFFCCAA77B696F85EFBB435F2" box="[971,1066,1958,1983]" italics="true" pageId="16" pageNumber="17">B. gibba</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="DA044849FFCCAA77B59AF83DFA0E35AE" blockId="16.[151,1436,1598,2019]" box="[199,1424,1994,2019]" pageId="16" pageNumber="17">
See
<tableCitation id="97397DF2FFCCAA77B5ABF83CFED735AE" box="[246,329,1994,2019]" captionStart="TABLE 3" captionStartId="31.[147,216,807,825]" captionText="TABLE 3. (Continued)" pageId="16" pageNumber="17">Table 3</tableCitation>
for the character states shown by the different species of
<taxonomicName id="1DBB33CAFFCCAA77B690F83CFBA135AE" box="[973,1087,1994,2019]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="E8CF945BFFCCAA77B690F83CFBA135AE" box="[973,1087,1994,2019]" italics="true" pageId="16" pageNumber="17">Barbatula</emphasis>
</taxonomicName>
found in north-eastern
<collectingCountry id="A2AC08D9FFCCAA77B018F83CFA1235AE" box="[1349,1420,1994,2019]" name="China" pageId="16" pageNumber="17">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="92A11BC2FFCDAA76B59AFF61FA95377D" pageId="17" pageNumber="18" type="description">
<paragraph id="DA044849FFCDAA76B59AFF61FC983055" blockId="17.[151,1437,151,2018]" pageId="17" pageNumber="18">
<emphasis id="E8CF945BFFCDAA76B59AFF61FEC732FD" bold="true" box="[199,345,151,176]" pageId="17" pageNumber="18">Description.</emphasis>
See
<figureCitation id="428054CCFFCDAA76B4CEFF61FDA932FD" box="[403,567,151,176]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStartId-0="15.[151,250,1653,1675]" captionStartId-1="16.[151,250,1476,1498]" captionTargetBox-0="[233,1312,681,1627]" captionTargetBox-1="[151,1436,225,1405]" captionTargetId-0="figure@15.[151,1436,661,1631]" captionTargetId-1="figure@16.[151,1436,193,1454]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIGURE 16. Barbatula pechiliensis, from the top: HU1608897, 69.8 mm SL; HU 1805102, 90.77 mm SL; HU1600026, 63.6 mm SL; China: Hebei prov.: Luan River drainage; IHB200608002042, 69.4mm SL; China: Liaoning prov.: Daling River drainage." captionText-1="FIGURE 17. Barbatula pechiliensis, from the top: HU 1805102, 90.8 mm SL; China: Hebei prov.: Luan River drainage; HU 1804062, 76.8 mm SL; China: Inner Mongolia: Luan River drainage; HU 1608042, 47.9 mm SL; China: Liaoning prov.: Xiaoling River drainage; IHB 200608002042, 69.4mm SL; China: Liaoning prov.: Daling River drainage." httpUri-0="https://zenodo.org/record/2589521/files/figure.png" httpUri-1="https://zenodo.org/record/2589523/files/figure.png" pageId="17" pageNumber="18">Figures 1617</figureCitation>
for general appearance and
<tableCitation id="97397DF2FFCDAA76B622FF61FC4B32FD" box="[895,981,151,176]" captionStart="TABLE 2" captionStartId="28.[147,222,518,538]" captionText="TABLE 2. Morphometric characters of eight species of Barbatula. Ho = holotype; SD = standard deviation." pageId="17" pageNumber="18">Table 2</tableCitation>
for morphometric data. Body elongate, roundish; caudal peduncle compressed. Dorsal profile convex; caudal-peduncle depth 4358% of body depth at dorsal-fin origin. Ventral profile between anal-fin base and caudal-fin origin concave. Head wider than deep, head depth 7692% of its width at posterior margin of operculum. Mouth inferior and arched (
<figureCitation id="428054CCFFCDAA76B120FEF2FB433351" box="[1149,1245,260,285]" captionStart="FIGURE 18" captionStartId="18.[151,250,1144,1166]" captionTargetBox="[151,1399,781,1123]" captionTargetId="figure@18.[151,1436,772,1123]" captionTargetPageId="18" captionText="FIGURE 18. Barbatula pechiliensis, ab: HU1805102, 90.77 mm SL; c: HU1600024, 79.4 mm SL; China: Hebei prov.: Luan River drainage. Scale bars: ab = 1 mm, c = 0.1 mm. Designations: m.i, median incision; m.le,; m.i, mental lobe lateral expansion; l.le, lower lip lateral expansion." httpUri="https://zenodo.org/record/2589525/files/figure.png" pageId="17" pageNumber="18">Fig. 18a</figureCitation>
). Snout rounded, short and blunt; head width at trailing edge of operculum 1.31.4 times in its width at anterior nostril. Snout shorter than postorbital head length. Eye small, set close to dorsal profile. Anterior and posterior nostril widely spaced (
<figureCitation id="428054CCFFCDAA76B5C2FE99FE9833C5" box="[159,262,367,392]" captionStart="FIGURE 18" captionStartId="18.[151,250,1144,1166]" captionTargetBox="[151,1399,781,1123]" captionTargetId="figure@18.[151,1436,772,1123]" captionTargetPageId="18" captionText="FIGURE 18. Barbatula pechiliensis, ab: HU1805102, 90.77 mm SL; c: HU1600024, 79.4 mm SL; China: Hebei prov.: Luan River drainage. Scale bars: ab = 1 mm, c = 0.1 mm. Designations: m.i, median incision; m.le,; m.i, mental lobe lateral expansion; l.le, lower lip lateral expansion." httpUri="https://zenodo.org/record/2589525/files/figure.png" pageId="17" pageNumber="18">Fig. 18b</figureCitation>
), gap between nostrils equal or larger than width of posterior nostril. Caudal-fin truncate. Caudalpeduncle length 72100% HL, caudal-peduncle width 3966% its depth. Distance between anus and anal-fin origin shorter than or equal to eye diameter. Dorsal-fin origin situated at middle between posterior nostril and caudal-fin base. Pelvic-fin origin situated below or in front of the vertical through dorsal-fin origin. Anal-fin origin anterior to midpoint between pelvic-fin origin and caudal-fin base.
</paragraph>
<paragraph id="DA044849FFCDAA76B59AFDD2FBF631CD" blockId="17.[151,1437,151,2018]" pageId="17" pageNumber="18">
Lateral line complete, reaching to caudal-fin base. Scales restricted to caudal peduncle, densely set. Scales on caudal peduncle small, oval, with a large focal zone and 2527 radial grooves (
<figureCitation id="428054CCFFCDAA76B157FDB1FBF4302D" box="[1034,1130,583,608]" captionStart="FIGURE 18" captionStartId="18.[151,250,1144,1166]" captionTargetBox="[151,1399,781,1123]" captionTargetId="figure@18.[151,1436,772,1123]" captionTargetPageId="18" captionText="FIGURE 18. Barbatula pechiliensis, ab: HU1805102, 90.77 mm SL; c: HU1600024, 79.4 mm SL; China: Hebei prov.: Luan River drainage. Scale bars: ab = 1 mm, c = 0.1 mm. Designations: m.i, median incision; m.le,; m.i, mental lobe lateral expansion; l.le, lower lip lateral expansion." httpUri="https://zenodo.org/record/2589525/files/figure.png" pageId="17" pageNumber="18">Fig. 18c</figureCitation>
). Anterior nostril with long tube, posterior tube short or absent. Width of anterior nostril equal to width of posterior nostril (
<figureCitation id="428054CCFFCDAA76B183FD9AFADF30C8" box="[1246,1345,620,645]" captionStart="FIGURE 18" captionStartId="18.[151,250,1144,1166]" captionTargetBox="[151,1399,781,1123]" captionTargetId="figure@18.[151,1436,772,1123]" captionTargetPageId="18" captionText="FIGURE 18. Barbatula pechiliensis, ab: HU1805102, 90.77 mm SL; c: HU1600024, 79.4 mm SL; China: Hebei prov.: Luan River drainage. Scale bars: ab = 1 mm, c = 0.1 mm. Designations: m.i, median incision; m.le,; m.i, mental lobe lateral expansion; l.le, lower lip lateral expansion." httpUri="https://zenodo.org/record/2589525/files/figure.png" pageId="17" pageNumber="18">Fig. 18b</figureCitation>
). Upper jaw completely covered by upper lip. Upper lip with shallow median incision, its depth 1040% of width of upper lip. Lower lip widely separate in middle. Mental lobe with blade-shaped lateral expansion. Interspace between mental lobes narrow, exposing small part of lower jaw. In few individuals lower jaw with elongated, club-shaped protrusion, formed by produced antero-ventral angles of symphyseal tip of dentary. Lower lip with short lateral expansion, its length 6090% of width of upper lip (
<figureCitation id="428054CCFFCDAA76B7B5FCE9FCD73175" box="[744,841,799,824]" captionStart="FIGURE 18" captionStartId="18.[151,250,1144,1166]" captionTargetBox="[151,1399,781,1123]" captionTargetId="figure@18.[151,1436,772,1123]" captionTargetPageId="18" captionText="FIGURE 18. Barbatula pechiliensis, ab: HU1805102, 90.77 mm SL; c: HU1600024, 79.4 mm SL; China: Hebei prov.: Luan River drainage. Scale bars: ab = 1 mm, c = 0.1 mm. Designations: m.i, median incision; m.le,; m.i, mental lobe lateral expansion; l.le, lower lip lateral expansion." httpUri="https://zenodo.org/record/2589525/files/figure.png" pageId="17" pageNumber="18">Fig. 18a</figureCitation>
). Maxillary barbel usually reaching to posterior eyemargin, inner rostral barbel usually reaching to anterior margin of posterior nostril, outer rostral barbel usually reaching to anterior half or middle of snout. In few individuals, barbels slightly shorter.
</paragraph>
<paragraph id="DA044849FFCDAA76B59AFC7AFF623615" blockId="17.[151,1437,151,2018]" pageId="17" pageNumber="18">Dorsal fin with 4 unbranched and 7½ branched rays. Anal fin with 2 unbranched and 5½ branched rays. Caudal-fin with 78+8 branched rays. Pectoral fin with 1 unbranched and 1112 branched rays. Pelvic fin with 1 unbranched and 67 branched rays. Infraorbital canal confluent with occipital canal, separated from supraorbital canal. Infraorbital canal with 1213, supraorbital canal with 78, occipital canal with 3, and mandibular canal and preopercular canal with 1012 pores. Intestine Z-shaped, anterior margin of zigzag loop not touching U-shaped stomach.</paragraph>
<paragraph id="DA044849FFCDAA76B59AFB92FACA36A5" blockId="17.[151,1437,151,2018]" pageId="17" pageNumber="18">
<emphasis id="E8CF945BFFCDAA76B59AFB92FD863630" bold="true" box="[199,536,1124,1149]" pageId="17" pageNumber="18">Colouration in preservative.</emphasis>
Background colour pale yellow with dark-brown pattern. Back with 912 large, dark-brown bars. Interspaces between bars equal to or wider than width of them. Flank with large cloud-like mottling. Caudal fin hyaline with 35 dark-brown bands formed by spots. Dorsal fin hyaline with 35 dark-brown bands formed by spots. Pectoral and pelvic fins hyaline without pigmentation or with few dark-brown spots.
</paragraph>
<paragraph id="DA044849FFCDAA76B59AFB02FA95377D" blockId="17.[151,1437,151,2018]" pageId="17" pageNumber="18">
<emphasis id="E8CF945BFFCDAA76B59AFB02FE263740" bold="true" box="[199,440,1268,1293]" pageId="17" pageNumber="18">Sexual dimorphism.</emphasis>
Males have a longer pectoral-fin than females (pectoral-fin length 1619% SL vs. 13 15% in female) and the 2nd7rd branched pectoral-fin rays are thickened in males (vs. not in female).
</paragraph>
</subSubSection>
<subSubSection id="92A11BC2FFCDAA76B59AFACAFE9137D0" pageId="17" pageNumber="18" type="distribution">
<paragraph id="DA044849FFCDAA76B59AFACAFE9137D0" blockId="17.[151,1437,151,2018]" pageId="17" pageNumber="18">
<emphasis id="E8CF945BFFCDAA76B59AFACAFEFE3718" bold="true" box="[199,352,1340,1365]" pageId="17" pageNumber="18">Distribution.</emphasis>
Known from the Luan and Hai Rivers, the Yin River, which is a tributary of the West Liao River, the Dali-Nur Lake in Inner
<collectingCountry id="A2AC08D9FFCDAA76B491FAA9FDA73735" box="[460,569,1375,1400]" name="Mongolia" pageId="17" pageNumber="18">Mongolia</collectingCountry>
and the Nuer River, which is a tributary of the Xiaoling River. See
<figureCitation id="428054CCFFCDAA76B063FAA9FA023735" box="[1342,1436,1375,1400]" captionStart="FIGURE 2" captionStartId="3.[151,250,1882,1904]" captionTargetBox="[173,1389,888,1818]" captionTargetId="figure@3.[151,1436,876,1861]" captionTargetPageId="3" captionText="FIGURE 2. Map showing the collection sites. Details for each number are listed in material examined." httpUri="https://zenodo.org/record/2589493/files/figure.png" pageId="17" pageNumber="18">Figure 2</figureCitation>
for details.
</paragraph>
</subSubSection>
<subSubSection id="92A11BC2FFCDAA76B59AFA51FC8535AF" pageId="17" pageNumber="18" type="discussion">
<paragraph id="DA044849FFCDAA76B59AFA51FD993563" blockId="17.[151,1437,151,2018]" pageId="17" pageNumber="18">
<emphasis id="E8CF945BFFCDAA76B59AFA51FEA5378D" bold="true" box="[199,315,1447,1472]" pageId="17" pageNumber="18">Remarks.</emphasis>
<taxonomicName id="1DBB33CAFFCDAA76B41EFA51FDDF378D" baseAuthorityName="Fowler" baseAuthorityYear="1899" box="[323,577,1447,1472]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="pechiliensis">
<emphasis id="E8CF945BFFCDAA76B41EFA51FDDF378D" box="[323,577,1447,1472]" italics="true" pageId="17" pageNumber="18">Barbatula pechiliensis</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="BE2A35B8FFCDAA76B645FA51FC25378D" author="Fowler, H. W." box="[792,955,1447,1472]" pageId="17" pageNumber="18" pagination="179 - 182" refId="ref16114" refString="Fowler, H. W. (1899) Notes on a small collection of Chinese fishes. Proceedings of the Academy of Natural Sciences of Philadelphia, 51, 179 - 182." type="journal article" year="1899">Fowler (1899)</bibRefCitation>
from the Tan Lan Ho River
<quantity id="1D43E5ACFFCDAA76B05BFA51FAD1378D" box="[1286,1359,1447,1472]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="18" unit="km" value="50.0">50 km</quantity>
northeast of Dolon-nor in the Chinese Pechili province. The 19
<superScript id="2DCEE501FFCDAA76B669FA3DFCDE3794" attach="left" box="[820,832,1483,1497]" fontSize="6" pageId="17" pageNumber="18">th</superScript>
century Pechili province is situated in todays Inner
<collectingCountry id="A2AC08D9FFCDAA76B5CAFA03FE963443" box="[151,264,1525,1550]" name="Mongolia" pageId="17" pageNumber="18">Mongolia</collectingCountry>
. The Tan Lan Ho River of the 19
<superScript id="2DCEE501FFCDAA76B729FA04FD1E344D" attach="left" box="[628,640,1522,1536]" fontSize="6" pageId="17" pageNumber="18">th</superScript>
century is modern day Tuligen River in the upper Luan River drainage. We found only one species of
<taxonomicName id="1DBB33CAFFCDAA76B4B1F9EFFDC0347F" box="[492,606,1561,1586]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="E8CF945BFFCDAA76B4B1F9EFFDC0347F" box="[492,606,1561,1586]" italics="true" pageId="17" pageNumber="18">Barbatula</emphasis>
</taxonomicName>
in the Tuligen River north-east of Dolon-nor. The
<typeStatus id="0500F6EBFFCDAA76B1C6F9ECFB55347F" box="[1179,1227,1562,1586]" pageId="17" pageNumber="18">type</typeStatus>
of
<taxonomicName id="1DBB33CAFFCDAA76B1ACF9ECFA0B347F" baseAuthorityName="Fowler" baseAuthorityYear="1899" box="[1265,1429,1561,1586]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="pechiliensis">
<emphasis id="E8CF945BFFCDAA76B1ACF9ECFA0B347F" box="[1265,1429,1561,1586]" italics="true" pageId="17" pageNumber="18">B. pechiliensis</emphasis>
</taxonomicName>
, the only individual examined by
<bibRefCitation id="BE2A35B8FFCDAA76B756F9CBFD31341B" author="Fowler, H. W." box="[523,687,1597,1622]" pageId="17" pageNumber="18" pagination="179 - 182" refId="ref16114" refString="Fowler, H. W. (1899) Notes on a small collection of Chinese fishes. Proceedings of the Academy of Natural Sciences of Philadelphia, 51, 179 - 182." type="journal article" year="1899">Fowler (1899)</bibRefCitation>
, was noted to be lost (
<bibRefCitation id="BE2A35B8FFCDAA76B6F6F9C8FBC1341B" author="Cao, L. &amp; Causse, R. &amp; Zhang, E." box="[939,1119,1597,1622]" pageId="17" pageNumber="18" pagination="236 - 248" refId="ref16027" refString="Cao, L., Causse, R. &amp; Zhang, E. (2012) Revision of the loach species Barbatula nuda (Bleeker 1865) (Pisces: Balitoridae) from North China, with a description of a new species from Inner Mongolia. Zootaxa, 3586, 236 - 248." type="journal article" year="2012">
Cao
<emphasis id="E8CF945BFFCDAA76B6BFF9C8FB8D341B" box="[994,1043,1597,1622]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. 2012
</bibRefCitation>
). The description by
<bibRefCitation id="BE2A35B8FFCDAA76B010F9CBFF7F3437" author="Fowler, H. W." pageId="17" pageNumber="18" pagination="179 - 182" refId="ref16114" refString="Fowler, H. W. (1899) Notes on a small collection of Chinese fishes. Proceedings of the Academy of Natural Sciences of Philadelphia, 51, 179 - 182." type="journal article" year="1899">Fowler (1899)</bibRefCitation>
is uninformative except the position of the dorsal-fin origin, which is situated slightly behind the vertical through the pelvic-fin origin in
<taxonomicName id="1DBB33CAFFCDAA76B756F970FD2C34D3" baseAuthorityName="Fowler" baseAuthorityYear="1899" box="[523,690,1669,1694]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="pechiliensis">
<emphasis id="E8CF945BFFCDAA76B756F970FD2C34D3" box="[523,690,1669,1694]" italics="true" pageId="17" pageNumber="18">B. pechiliensis</emphasis>
</taxonomicName>
, as in our material examined. Therefore, we are convinced that loaches collected at the
<typeStatus id="0500F6EBFFCDAA76B4F2F95CFE41348F" box="[431,479,1706,1730]" pageId="17" pageNumber="18">type</typeStatus>
locality of
<taxonomicName id="1DBB33CAFFCDAA76B734F95CFC91348F" baseAuthorityName="Fowler" baseAuthorityYear="1899" box="[617,783,1705,1730]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="pechiliensis">
<emphasis id="E8CF945BFFCDAA76B734F95CFC91348F" box="[617,783,1705,1730]" italics="true" pageId="17" pageNumber="18">B. pechiliensis</emphasis>
</taxonomicName>
indeed represent this species.
<bibRefCitation id="BE2A35B8FFCDAA76B129F95FFABD348F" author="Kottelat, M." box="[1140,1315,1705,1730]" pageId="17" pageNumber="18" pagination="1 - 199" refId="ref16350" refString="Kottelat, M. (2012) Conspectus cobitidum: An inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidei). The Raffles Bulletin of Zoology, 26, 1 - 199." type="journal article" year="2012">Kottelat (2012)</bibRefCitation>
treated
<taxonomicName id="1DBB33CAFFCDAA76B0D9F95CFE8534AB" baseAuthorityName="Fowler" baseAuthorityYear="1899" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="pechiliensis">
<emphasis id="E8CF945BFFCDAA76B0D9F95CFE8534AB" italics="true" pageId="17" pageNumber="18">B. pechiliensis</emphasis>
</taxonomicName>
as a synonym of
<taxonomicName id="1DBB33CAFFCDAA76B4A6F938FDC934AB" baseAuthorityName="Bleeker" baseAuthorityYear="1865" box="[507,599,1741,1766]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="nuda">
<emphasis id="E8CF945BFFCDAA76B4A6F938FDC934AB" box="[507,599,1741,1766]" italics="true" pageId="17" pageNumber="18">B. nuda</emphasis>
</taxonomicName>
and
<bibRefCitation id="BE2A35B8FFCDAA76B7C5F938FCF934AB" author="Cao, L. &amp; Causse, R. &amp; Zhang, E." box="[664,871,1741,1766]" pageId="17" pageNumber="18" pagination="236 - 248" refId="ref16027" refString="Cao, L., Causse, R. &amp; Zhang, E. (2012) Revision of the loach species Barbatula nuda (Bleeker 1865) (Pisces: Balitoridae) from North China, with a description of a new species from Inner Mongolia. Zootaxa, 3586, 236 - 248." type="journal article" year="2012">
Cao
<emphasis id="E8CF945BFFCDAA76B78FF938FC9634AB" box="[722,776,1741,1766]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. (2012)
</bibRefCitation>
considered it to be a synonym of
<taxonomicName id="1DBB33CAFFCDAA76B050F938FAC334A8" baseAuthorityName="Dybowski" baseAuthorityYear="1869" box="[1293,1373,1742,1765]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="toni">
<emphasis id="E8CF945BFFCDAA76B050F938FAC334A8" box="[1293,1373,1742,1765]" italics="true" pageId="17" pageNumber="18">B. toni</emphasis>
</taxonomicName>
. Our morphological and molecular data demonstrate that
<taxonomicName id="1DBB33CAFFCDAA76B7B9F904FC163547" baseAuthorityName="Fowler" baseAuthorityYear="1899" box="[740,904,1777,1802]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="pechiliensis">
<emphasis id="E8CF945BFFCDAA76B7B9F904FC163547" box="[740,904,1777,1802]" italics="true" pageId="17" pageNumber="18">B. pechiliensis</emphasis>
</taxonomicName>
is well distinguished from
<taxonomicName id="1DBB33CAFFCDAA76B1E2F904FA863547" baseAuthorityName="Bleeker" baseAuthorityYear="1865" box="[1215,1304,1777,1802]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="nuda">
<emphasis id="E8CF945BFFCDAA76B1E2F904FA863547" box="[1215,1304,1777,1802]" italics="true" pageId="17" pageNumber="18">B. nuda</emphasis>
</taxonomicName>
and
<taxonomicName id="1DBB33CAFFCDAA76B00DF904FA023544" baseAuthorityName="Dybowski" baseAuthorityYear="1869" box="[1360,1436,1778,1801]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="toni">
<emphasis id="E8CF945BFFCDAA76B00DF904FA023544" box="[1360,1436,1778,1801]" italics="true" pageId="17" pageNumber="18">B. toni</emphasis>
</taxonomicName>
and we treat it as a valid species.
</paragraph>
<paragraph id="DA044849FFCDAA76B59AF8CFFC8535AF" blockId="17.[151,1437,151,2018]" pageId="17" pageNumber="18">
<bibRefCitation id="BE2A35B8FFCDAA76B59AF8CFFE17351F" author="Prokofiev, A. M." box="[199,393,1849,1874]" pageId="17" pageNumber="18" pagination="457 - 464" refId="ref16679" refString="Prokofiev, A. M. (2015) A new species of Barbatula from the Russian Altai (Teleostei: Nemacheilidae). Zootaxa, 4052 (4), 457 - 464. https: // doi. org / 10.11646 / zootaxa. 4052.4.3" type="journal article" year="2015">Prokofiev (2015)</bibRefCitation>
lists
<taxonomicName id="1DBB33CAFFCDAA76B495F8CCFDF0351F" baseAuthorityName="Fowler" baseAuthorityYear="1899" box="[456,622,1849,1874]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="pechiliensis">
<emphasis id="E8CF945BFFCDAA76B495F8CCFDF0351F" box="[456,622,1849,1874]" italics="true" pageId="17" pageNumber="18">B. pechiliensis</emphasis>
</taxonomicName>
as a synonym of
<taxonomicName id="1DBB33CAFFCDAA76B661F8CCFC70351C" authorityName="Prokofiev" authorityYear="2007" box="[828,1006,1850,1873]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="potaninorum">
<emphasis id="E8CF945BFFCDAA76B661F8CCFC70351C" box="[828,1006,1850,1873]" italics="true" pageId="17" pageNumber="18">B. potaninorum</emphasis>
</taxonomicName>
. Unfortunately, we have no access to material of
<taxonomicName id="1DBB33CAFFCDAA76B446F8A9FE533538" authorityName="Prokofiev" authorityYear="2007" box="[283,461,1886,1910]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="potaninorum">
<emphasis id="E8CF945BFFCDAA76B446F8A9FE533538" box="[283,461,1886,1910]" italics="true" pageId="17" pageNumber="18">B. potaninorum</emphasis>
</taxonomicName>
.
<taxonomicName id="1DBB33CAFFCDAA76B481F8ABFD47353B" baseAuthorityName="Fowler" baseAuthorityYear="1899" box="[476,729,1885,1910]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="pechiliensis">
<emphasis id="E8CF945BFFCDAA76B481F8ABFD47353B" box="[476,729,1885,1910]" italics="true" pageId="17" pageNumber="18">Barbatula pechiliensis</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="1DBB33CAFFCDAA76B686F8A9FB123538" authorityName="Prokofiev" authorityYear="2007" box="[987,1164,1886,1910]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="potaninorum">
<emphasis id="E8CF945BFFCDAA76B686F8A9FB123538" box="[987,1164,1886,1910]" italics="true" pageId="17" pageNumber="18">B. potaninorum</emphasis>
</taxonomicName>
by the pelvic-fin origin being situated below the vertical through the dorsal-fin origin (vs. in front), having 7½ branched dorsal-fin rays (vs. 6½), the mental lobe with a blade-shaped lateral expansion (vs. without) and the scales restricted to the caudal peduncle (vs. scales present on complete back and flank).
</paragraph>
</subSubSection>
<subSubSection id="92A11BC2FFCEAA75B59AFF61FE613081" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="DA044849FFCEAA75B59AFF61FE613081" blockId="18.[151,1436,151,717]" pageId="18" pageNumber="19">
<taxonomicName id="1DBB33CAFFCEAA75B59AFF61FE5B32FD" ID-CoL="7JDFV" authorityName=": Nichols" authorityYear="1925" box="[199,453,151,176]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="subSpecies" species="toni" subSpecies="fowleri">
<emphasis id="E8CF945BFFCEAA75B59AFF61FE5B32FD" box="[199,453,151,176]" italics="true" pageId="18" pageNumber="19">Barbatula toni fowleri</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="BE2A35B8FFCEAA75B7C1FF61FCD832FD" author="Nichols, J T" box="[668,838,151,176]" pageId="18" pageNumber="19" pagination="1 - 7" refId="ref16500" refString="Nichols, J T. (1925) Nemacheilus and related loaches in China. American Museum Novitates, 171, 1 - 7." type="journal article" year="1925">Nichols (1925)</bibRefCitation>
from Eastern Tombs in Zunhua City in the Chinese Chihli (todays
<collectingRegion id="187F86ABFFCEAA75B415FF4AFE133298" box="[328,397,188,213]" country="China" name="Hebei" pageId="18" pageNumber="19">Hebei</collectingRegion>
) province (see
<figureCitation id="428054CCFFCEAA75B760FF4AFD1C3299" box="[573,642,188,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1882,1904]" captionTargetBox="[173,1389,888,1818]" captionTargetId="figure@3.[151,1436,876,1861]" captionTargetPageId="3" captionText="FIGURE 2. Map showing the collection sites. Details for each number are listed in material examined." httpUri="https://zenodo.org/record/2589493/files/figure.png" pageId="18" pageNumber="19">Fig. 2</figureCitation>
). As is the
<typeStatus id="0500F6EBFFCEAA75B65AFF4BFCA93298" box="[775,823,189,213]" pageId="18" pageNumber="19">type</typeStatus>
locality of
<taxonomicName id="1DBB33CAFFCEAA75B6E6FF4BFBFE3298" ID-CoL="46657" baseAuthorityName="Fowler" baseAuthorityYear="1899" box="[955,1120,188,213]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="pechiliensis">
<emphasis id="E8CF945BFFCEAA75B6E6FF4BFBFE3298" box="[955,1120,188,213]" italics="true" pageId="18" pageNumber="19">B. pechiliensis</emphasis>
</taxonomicName>
, the
<typeStatus id="0500F6EBFFCEAA75B1C6FF4BFB553298" box="[1179,1227,189,213]" pageId="18" pageNumber="19">type</typeStatus>
locality of
<taxonomicName id="1DBB33CAFFCEAA75B00DFF4BFF7932B5" ID-CoL="7JDFV" authorityName=": Nichols" authorityYear="1925" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="subSpecies" species="toni" subSpecies="fowleri">
<emphasis id="E8CF945BFFCEAA75B00DFF4BFF7932B5" italics="true" pageId="18" pageNumber="19">B. toni fowleri</emphasis>
</taxonomicName>
is situated in the Luan River drainage. Unfortunately, we were unable to find
<taxonomicName id="1DBB33CAFFCEAA75B10EFF29FB5B32B5" ID-CoL="37Z7" box="[1107,1221,223,248]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="E8CF945BFFCEAA75B10EFF29FB5B32B5" box="[1107,1221,223,248]" italics="true" pageId="18" pageNumber="19">Barbatula</emphasis>
</taxonomicName>
loaches at the
<typeStatus id="0500F6EBFFCEAA75B031FF16FA0232B5" box="[1388,1436,224,248]" pageId="18" pageNumber="19">type</typeStatus>
locality of
<emphasis id="E8CF945BFFCEAA75B44AFEF3FE563350" box="[279,456,260,285]" italics="true" pageId="18" pageNumber="19">
<taxonomicName id="1DBB33CAFFCEAA75B44AFEF3FE5D3350" ID-CoL="7JDFV" authorityName=": Nichols" authorityYear="1925" box="[279,451,260,285]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="subSpecies" species="toni" subSpecies="fowleri">B. toni fowleri</taxonomicName>
.
</emphasis>
We found
<taxonomicName id="1DBB33CAFFCEAA75B710FEF2FD213350" ID-CoL="37Z7" box="[589,703,260,285]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="E8CF945BFFCEAA75B710FEF2FD213350" box="[589,703,260,285]" italics="true" pageId="18" pageNumber="19">Barbatula</emphasis>
</taxonomicName>
in the Shandian and Xiaoluan Rivers in the upper Luan River drainage which are identified as
<emphasis id="E8CF945BFFCEAA75B740FEDFFD53330D" box="[541,717,295,320]" italics="true" pageId="18" pageNumber="19">
<taxonomicName id="1DBB33CAFFCEAA75B740FEDFFD57330D" ID-CoL="46657" baseAuthorityName="Fowler" baseAuthorityYear="1899" box="[541,713,295,320]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="pechiliensis">B. pechiliensis</taxonomicName>
.
</emphasis>
We found only one species of
<taxonomicName id="1DBB33CAFFCEAA75B117FED1FB22330D" ID-CoL="37Z7" box="[1098,1212,295,320]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="E8CF945BFFCEAA75B117FED1FB22330D" box="[1098,1212,295,320]" italics="true" pageId="18" pageNumber="19">Barbatula</emphasis>
</taxonomicName>
in the Luan River drainage, but cannot rule out the possibility that a second species might occur there.
<bibRefCitation id="BE2A35B8FFCEAA75B10AFEBAFA9E3328" author="Nichols, J T" box="[1111,1280,332,357]" pageId="18" pageNumber="19" pagination="1 - 7" refId="ref16500" refString="Nichols, J T. (1925) Nemacheilus and related loaches in China. American Museum Novitates, 171, 1 - 7." type="journal article" year="1925">Nichols (1925)</bibRefCitation>
stated that
<taxonomicName id="1DBB33CAFFCEAA75B0D9FEBBFE8533C5" ID-CoL="7JDFV" authorityName=": Nichols" authorityYear="1925" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="subSpecies" species="toni" subSpecies="fowleri">
<emphasis id="E8CF945BFFCEAA75B0D9FEBBFE8533C5" italics="true" pageId="18" pageNumber="19">B. toni fowleri</emphasis>
</taxonomicName>
has closely-set nostrils and the scales were only evident on the caudal peduncle, character states in agreement with
<taxonomicName id="1DBB33CAFFCEAA75B416FE63FE7333E0" ID-CoL="46657" baseAuthorityName="Fowler" baseAuthorityYear="1899" box="[331,493,404,429]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="pechiliensis">
<emphasis id="E8CF945BFFCEAA75B416FE63FE7333E0" box="[331,493,404,429]" italics="true" pageId="18" pageNumber="19">B. pechiliensis</emphasis>
</taxonomicName>
. We examined photos of the
<typeStatus id="0500F6EBFFCEAA75B66FFE63FCFC33E0" box="[818,866,405,429]" pageId="18" pageNumber="19">type</typeStatus>
of
<taxonomicName id="1DBB33CAFFCEAA75B6DAFE63FBB933E0" ID-CoL="7JDFV" authorityName=": Nichols" authorityYear="1925" box="[903,1063,404,429]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="subSpecies" species="toni" subSpecies="fowleri">
<emphasis id="E8CF945BFFCEAA75B6DAFE63FBB933E0" box="[903,1063,404,429]" italics="true" pageId="18" pageNumber="19">B. toni fowleri</emphasis>
</taxonomicName>
(AMNH 8409,
<figureCitation id="428054CCFFCEAA75B184FE62FABC33E0" box="[1241,1314,404,429]" captionStart="FIGURE 16" captionStartId="15.[151,250,1653,1675]" captionTargetBox="[233,1312,681,1627]" captionTargetId="figure@15.[151,1436,661,1631]" captionTargetPageId="15" captionText="FIGURE 16. Barbatula pechiliensis, from the top: HU1608897, 69.8 mm SL; HU 1805102, 90.77 mm SL; HU1600026, 63.6 mm SL; China: Hebei prov.: Luan River drainage; IHB200608002042, 69.4mm SL; China: Liaoning prov.: Daling River drainage." httpUri="https://zenodo.org/record/2589521/files/figure.png" pageId="18" pageNumber="19">Fig 16</figureCitation>
) and it has a blunt and short snout, widely spaced nostrils, a shallow median incision in the upper lip, a short lateral expansion at lower lip and the pelvic-fin origin situated below the vertical through the dorsal-fin origin. The
<typeStatus id="0500F6EBFFCEAA75B1A5FE2BFAB633B8" box="[1272,1320,477,501]" pageId="18" pageNumber="19">type</typeStatus>
of
<taxonomicName id="1DBB33CAFFCEAA75B00DFE2BFF793055" ID-CoL="7JDFV" authorityName=": Nichols" authorityYear="1925" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="subSpecies" species="toni" subSpecies="fowleri">
<emphasis id="E8CF945BFFCEAA75B00DFE2BFF793055" italics="true" pageId="18" pageNumber="19">B. toni fowleri</emphasis>
</taxonomicName>
has few scales on the back anterior to the dorsal-fin origin but no scales on the flank anterior to the dorsalfin origin. It is also different from
<taxonomicName id="1DBB33CAFFCEAA75B774FDD3FD513070" ID-CoL="46657" baseAuthorityName="Fowler" baseAuthorityYear="1899" box="[553,719,548,573]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="pechiliensis">
<emphasis id="E8CF945BFFCEAA75B774FDD3FD513070" box="[553,719,548,573]" italics="true" pageId="18" pageNumber="19">B. pechiliensis</emphasis>
</taxonomicName>
by having the scales densely set on the caudal peduncle only, while in
<taxonomicName id="1DBB33CAFFCEAA75B5A7FDBFFE0C302D" box="[250,402,583,608]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="pechilienis">
<emphasis id="E8CF945BFFCEAA75B5A7FDBFFE0C302D" box="[250,402,583,608]" italics="true" pageId="18" pageNumber="19">B. pechilienis</emphasis>
</taxonomicName>
, the scales are densely set below the dorsal-fin base. We cannot exclude that there might be some variation between different populations of
<taxonomicName id="1DBB33CAFFCEAA75B7E7FD9BFCC930C8" box="[698,855,620,645]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="pechilensis">
<emphasis id="E8CF945BFFCEAA75B7E7FD9BFCC930C8" box="[698,855,620,645]" italics="true" pageId="18" pageNumber="19">B. pechilensis</emphasis>
</taxonomicName>
in the scale patterns and more field work is needed to resolve this question. Until it can be demonstrated that
<taxonomicName id="1DBB33CAFFCEAA75B64AFD67FC1A30E5" ID-CoL="7JDFV" box="[791,900,655,680]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="fowleri">
<emphasis id="E8CF945BFFCEAA75B64AFD67FC1A30E5" box="[791,900,655,680]" italics="true" pageId="18" pageNumber="19">B. fowleri</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="1DBB33CAFFCEAA75B120FD67FA8130E5" ID-CoL="46657" baseAuthorityName="Fowler" baseAuthorityYear="1899" box="[1149,1311,655,680]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="pechiliensis">
<emphasis id="E8CF945BFFCEAA75B120FD67FA8130E5" box="[1149,1311,655,680]" italics="true" pageId="18" pageNumber="19">B. pechiliensis</emphasis>
</taxonomicName>
, we treat it as a synonym of
<taxonomicName id="1DBB33CAFFCEAA75B409FD43FE663080" ID-CoL="46657" baseAuthorityName="Fowler" baseAuthorityYear="1899" box="[340,504,692,717]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="pechiliensis">
<emphasis id="E8CF945BFFCEAA75B409FD43FE663080" box="[340,504,692,717]" italics="true" pageId="18" pageNumber="19">B. pechiliensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>