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<document id="C761CF13F3CB697AA49CA99F54E6E440" ID-DOI="10.5281/zenodo.5347271" ID-ISSN="2345-7600" ID-Zenodo-Dep="5347271" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630203671219" checkinUser="tatiana" docAuthor="Nguyen, Anh D." docDate="2012" docId="191D7C72CD2BE812FF7A9279FCC7F830" docLanguage="en" docName="60rbz289-311.pdf" docOrigin="Raffles Bulletin of Zoology 60 (2)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Tylopus golovatchi Nguyen 2012, new species" docType="treatment" docVersion="4" lastPageNumber="298" masterDocId="E524040ACD22E81BFF859465FF88FFC5" masterDocTitle="Tylopus Millipedes In Vietnam (Diplopoda: Polydesmida: Paradoxosomatidae: Sulciferini), With Descriptions Of Five New Species" masterLastPageNumber="311" masterPageNumber="289" pageNumber="298" updateTime="1699183855207" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="508C8B2DF7F6088AB755F4A0310BC7EC">Tylopus Millipedes In Vietnam (Diplopoda: Polydesmida: Paradoxosomatidae: Sulciferini), With Descriptions Of Five New Species</mods:title>
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<mods:namePart id="207C47F5DE3852B5F9EBD51FFBA4C3DD">Nguyen, Anh D.</mods:namePart>
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<mods:date id="233B94E546659C3FB37D52B20F6D3DD9">2012</mods:date>
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<heading id="CA437A08CD2BE812FF7A9279FDEBF9F0" bold="true" box="[255,611,1564,1589]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="298" reason="2">
<emphasis id="A3C01176CD2BE812FF7A9279FDEBF9F0" bold="true" box="[255,611,1564,1589]" pageId="9" pageNumber="298">
<taxonomicName id="56B4B6E7CD2BE812FF7A9279FE47F9F0" ID-CoL="7CZMQ" authority="Nguyen, 2012" authorityName="Nguyen" authorityYear="2012" box="[255,463,1564,1589]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="298" phylum="Arthropoda" rank="species" species="golovatchi" status="sp. nov.">
<emphasis id="A3C01176CD2BE812FF7A9279FE47F9F0" bold="true" box="[255,463,1564,1589]" italics="true" pageId="9" pageNumber="298">Tylopus golovatchi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="B8F3AC0DCD2BE812FE5B9278FDEBF9F0" box="[478,611,1565,1589]" pageId="9" pageNumber="298" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D9AE9EEFCD2BE812FEE59258FD89F990" box="[352,513,1597,1621]" pageId="9" pageNumber="298" type="description">
<paragraph id="910BCD64CD2BE812FEE59258FD89F990" blockId="9.[255,611,1564,1621]" box="[352,513,1597,1621]" pageId="9" pageNumber="298">
(
<figureCitation id="098FD1E1CD2BE812FEED9258FE3CF991" box="[360,436,1597,1621]" captionStart="Fig" captionStartId="10.[136,171,2062,2084]" captionTargetBox="[149,1468,336,1927]" captionTargetId="figure-8@10.[118,1505,318,1949]" captionTargetPageId="10" captionText="Fig. 7. Tylopus golovatchi, new species, holotype: hypoproct, ventral view (A); sternite 5, posterioventral view (B); left gonopod, lateral view (C), mesal view (D); postfemoral region, dorsal view (E). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5347285" httpUri="https://zenodo.org/record/5347285/files/figure.png" pageId="9" pageNumber="298">Figs. 7</figureCitation>
,
<figureCitation id="098FD1E1CD2BE812FE449258FE58F990" box="[449,464,1597,1621]" captionStart="Fig" captionStartId="11.[113,148,2062,2084]" captionTargetBox="[170,1393,189,2024]" captionTargetId="figure-7@11.[169,1393,189,2024]" captionTargetPageId="11" captionText="Fig. 8. Tylopus golovatchi, new species, holotype: left gonopod, mesal view (A), ventral view (B), and lateral view (C); postfemoral region, lateral view (D)." figureDoi="http://doi.org/10.5281/zenodo.5347287" httpUri="https://zenodo.org/record/5347287/files/figure.png" pageId="9" pageNumber="298">8</figureCitation>
,
<figureCitation id="098FD1E1CD2BE812FE589258FE70F990" box="[477,504,1597,1621]" captionStart="Fig" captionStartId="21.[113,147,1802,1824]" captionTargetBox="[113,753,1083,1781]" captionTargetId="figure-736@21.[113,753,1083,1779]" captionTargetPageId="21" captionText="Fig. 18.Distribution map ofthe millipede genus Tylopus Jeekel, 1968 in Vietnam. 1: Sa Pa, Lao Cai Prov. (T. crassipes, T. procurvus, T. nodulipes, T. sigma, T. sapaensis, T. maculatus, T. magnicus); 2: Van Ban, Lao Cai Prov. (T. crassipes, T. nodulipes); 3: Xuan Son NP (T. strongylosomoides, T. golovatchi); 4: Tam Dao NP (T. tamdaoensis, T. roseiparaterga, T. strongylosomoides); 5: Cuc Phuong NP (T. granulatus, T. hilaroides, T. topali); 6: Pu Mat NP (T. nodulipes); 7: Huong Son, Ha Tinh prov. (T. nodulipes); 8: Bach MaBa Na NP (T. hilaris, T. multilatus); 9: Ngoc Linh Mts., Kon Tum prov. (T. hilaris, T. phanluongi); 10: Bi DoupNui Ba NP (T. spinisterna)." figureDoi="http://doi.org/10.5281/zenodo.5347309" httpUri="https://zenodo.org/record/5347309/files/figure.png" pageId="9" pageNumber="298">18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D9AE9EEFCD2BE812FFF4921EFF44F8E7" pageId="9" pageNumber="298" type="materials_examined">
<paragraph id="910BCD64CD2BE812FFF4921EFECDF92D" blockId="9.[113,752,1659,1826]" pageId="9" pageNumber="298">
<emphasis id="A3C01176CD2BE812FFF4921EFEBFF954" bold="true" box="[113,311,1659,1681]" italics="true" pageId="9" pageNumber="298">Material examined.</emphasis>
<materialsCitation id="21DCC739CD2BE812FEE49219FECDF92D" collectingDate="2006-01-15" collectorName="Nguyen Van Quang" latitude="21.2" location="Male" longLatPrecision="1264" longitude="105.01667" pageId="9" pageNumber="298" specimenCount="1" stateProvince="Phu Tho" typeStatus="holotype">
<typeStatus id="4E0F73C6CD2BE812FEE49219FE37F957" box="[353,447,1660,1682]" pageId="9" pageNumber="298" type="holotype">Holotype</typeStatus>
:
<location id="946B9BBFCD2BE812FE4E9219FE75F957" LSID="urn:lsid:plazi:treatment:191D7C72CD2BE812FF7A9279FCC7F830:946B9BBFCD2BE812FE4E9219FE75F957" box="[459,509,1660,1682]" latitude="21.2" longLatPrecision="1264" longitude="105.01667" name="Male" pageId="9" pageNumber="298" stateProvince="Phu Tho">Male</location>
in a bottle
<emphasis id="A3C01176CD2BE812FDF3921EFD78F955" bold="true" box="[630,752,1659,1680]" pageId="9" pageNumber="298">IEBR-141H</emphasis>
with a label “
<collectingRegion id="53700386CD2BE812FF7792FDFE2AF96B" box="[242,418,1688,1710]" country="Vietnam" name="Phu Tho" pageId="9" pageNumber="298">Phu Tho province</collectingRegion>
, Xuan Son National Park (21°03'
<geoCoordinate id="F480ABA3CD2BE812FFF492D0FF4DF90F" box="[113,197,1717,1738]" degrees="21" direction="north" minutes="12" orientation="latitude" pageId="9" pageNumber="298" precision="925" value="21.2">21°12'N</geoCoordinate>
, 104°51'
<geoCoordinate id="F480ABA3CD2BE812FEA892D0FE01F90E" box="[301,393,1717,1739]" degrees="105" direction="east" minutes="01" orientation="longitude" pageId="9" pageNumber="298" precision="925" value="105.01667">105°01'E</geoCoordinate>
), primary forest, coll.
<collectorName id="3C41A8B2CD2BE812FDF692D3FF3BF922" pageId="9" pageNumber="298">Nguyen Van Quang</collectorName>
,
<date id="E50AEBA4CD2BE812FF3B92B7FEBFF92D" box="[190,311,1746,1768]" pageId="9" pageNumber="298" value="2006-01-15">
<collectingDate id="F54E124CCD2BE812FF3B92B7FEBFF92D" box="[190,311,1746,1768]" pageId="9" pageNumber="298" value="2006-01-15">15 Jan.2006</collectingDate>
</date>
”.
</materialsCitation>
</paragraph>
<paragraph id="910BCD64CD2BE812FFF4928AFF44F8E7" blockId="9.[113,752,1659,1826]" pageId="9" pageNumber="298">
<materialsCitation id="21DCC739CD2BE812FFF4928AFF40F8E7" collectingDate="2006-01-15" collectorName="Nguyen Van Quang" latitude="21.2" location="Male" longLatPrecision="1264" longitude="105.01667" pageId="9" pageNumber="298" specimenCount="1" specimenCount-female="1" stateProvince="Phu Tho" typeStatus="paratype">
<typeStatus id="4E0F73C6CD2BE812FFF4928AFF5BF8C1" box="[113,211,1775,1796]" pageId="9" pageNumber="298" type="paratype">Paratype</typeStatus>
:
<specimenCount id="87B206EDCD2BE812FF67928AFEC9F8C0" box="[226,321,1775,1797]" pageId="9" pageNumber="298" type="female">1 female</specimenCount>
(
<emphasis id="A3C01176CD2BE812FED3928AFE5DF8C1" bold="true" box="[342,469,1775,1796]" pageId="9" pageNumber="298">IEBR-141P</emphasis>
), collected together with holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D9AE9EEFCD2BE812FFF4932CFDDAF867" pageId="9" pageNumber="298" type="etymology">
<paragraph id="910BCD64CD2BE812FFF4932CFDDAF867" blockId="9.[113,751,1865,1954]" pageId="9" pageNumber="298">
<emphasis id="A3C01176CD2BE812FFF4932CFF65F8A7" bold="true" box="[113,237,1865,1890]" italics="true" pageId="9" pageNumber="298">Etymology.</emphasis>
— “
<taxonomicName id="56B4B6E7CD2BE812FEAC932EFE13F8A7" authorityName="Nguyen" authorityYear="2012" box="[297,411,1867,1890]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="298" phylum="Arthropoda" rank="species" species="golovatchi">
<emphasis id="A3C01176CD2BE812FEAC932EFE13F8A7" box="[297,411,1867,1890]" italics="true" pageId="9" pageNumber="298">golovatchi</emphasis>
</taxonomicName>
”, named in honor of Dr. S. I. Golovatch, an outstanding specialist of millipedes for his major contribution to global myriapodology.
</paragraph>
</subSubSection>
<subSubSection id="D9AE9EEFCD2BE812FFF493AFFE18F787" pageId="9" pageNumber="298" type="diagnosis">
<paragraph id="910BCD64CD2BE812FFF493AFFE18F787" blockId="9.[113,752,1994,2114]" pageId="9" pageNumber="298">
<emphasis id="A3C01176CD2BE812FFF493AFFF78F827" bold="true" box="[113,240,1994,2018]" italics="true" pageId="9" pageNumber="298">Diagnosis.</emphasis>
— The species is very similar to
<taxonomicName id="56B4B6E7CD2BE812FD1293AEFEB0F7C7" baseAuthorityName="Korsos &amp; Golovatch" baseAuthorityYear="1989" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="298" phylum="Arthropoda" rank="species" species="strongylosomoides">
<emphasis id="A3C01176CD2BE812FD1293AEFEB0F7C7" italics="true" pageId="9" pageNumber="298">Tylopus strongylosomoides</emphasis>
</taxonomicName>
, but it can be easily distinguished from the latter by the presence of a long, slender, finger-shaped process
<emphasis id="A3C01176CD2BE812FF4C9C4FFF50F787" bold="true" box="[201,216,2090,2114]" pageId="9" pageNumber="298">h</emphasis>
on the gonopod.
</paragraph>
</subSubSection>
<subSubSection id="D9AE9EEFCD2BE812FCA994D8FCCAF851" pageId="9" pageNumber="298" type="description">
<paragraph id="910BCD64CD2BE812FCA994D8FC61FEF0" blockId="9.[812,1452,189,1941]" pageId="9" pageNumber="298">
<emphasis id="A3C01176CD2BE812FCA994D8FC3AFF10" bold="true" box="[812,946,189,213]" italics="true" pageId="9" pageNumber="298">Description.</emphasis>
— Size: Body length about [17.9] mm (male),
<quantity id="564C6081CD2BE812FCA994B8FC18FF31" box="[812,912,221,245]" metricMagnitude="-2" metricUnit="m" metricValue="2.13" pageId="9" pageNumber="298" unit="mm" value="21.3">21.3 mm</quantity>
(female). Width of midbody prozona [1.6] mm (male),
<quantity id="564C6081CD2BE812FC019498FC52FED1" box="[900,986,253,277]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="9" pageNumber="298" unit="mm" value="2.2">2.2 mm</quantity>
(female) and metazona [1.9] mm (male),
<quantity id="564C6081CD2BE812FCA99578FCF7FEF1" box="[812,895,285,309]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="9" pageNumber="298" unit="mm" value="2.6">2.6 mm</quantity>
(female).
</paragraph>
<paragraph id="910BCD64CD2BE812FCA99558FB2BFE70" blockId="9.[812,1452,189,1941]" pageId="9" pageNumber="298">Colouration: Body generally yellowish brown except legs and sterna paler; whole antennomere 6 and 7 blackish brown. Prozona and metaterga with several small blackish brown areas. Female generally yellowish.</paragraph>
<paragraph id="910BCD64CD2BE812FCA995D8FAC0FDD0" blockId="9.[812,1452,189,1941]" pageId="9" pageNumber="298">
Head: Slightly broader than collum. Epicranial suture obviously distinct. Antenna claviform, reaching body ring 4 laterally. Antennomere 2=3&gt;4=5&gt;6&gt;
<quantity id="564C6081CD2BE812FB499598FB7EFDD0" box="[1228,1270,509,533]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="9" pageNumber="298" unit="in" value="7.0">7 in</quantity>
length.
</paragraph>
<paragraph id="910BCD64CD2BE812FCA99678FC03FD50" blockId="9.[812,1452,189,1941]" pageId="9" pageNumber="298">Collum: Slightly narrower than body ring 2, subtrapeziform; surface shining, finely rugulose, with traces of row of 2+2 setae near front margin. Paratergum poorly developed, rounded.</paragraph>
<paragraph id="910BCD64CD2BE812FCA996F8FB5CFBD0" blockId="9.[812,1452,189,1941]" pageId="9" pageNumber="298">
Body ring 3&lt;4&lt;2=
<quantity id="564C6081CD2BE812FC7C96F8FBACFD70" box="[1017,1060,669,693]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="9" pageNumber="298" unit="in" value="5.0">5 in</quantity>
width, parallel-sided on body rings 517, thereafter gradually tapering towards telson. Prozona shining, somewhat smooth. Metatergal surface shining, finely rugose, with rows of 2+2 and 3+3 small knobs in front of and behind transverse sulcus, respectively. Transverse sulcus poorly-developed on body rings 24, but more distinct and well developed (reaching base of paraterga) on body rings 519. Pleura with dense microgranules. Pleurosternal keels well developed on body rings 24, beak-shaped on body rings 57, reduced on body rings 811, and completely absent on subsequent body rings. Waist between pro- and metazona broad, finely striate. Axial line evident.
</paragraph>
<paragraph id="910BCD64CD2BE812FCA99078FAE5FB10" blockId="9.[812,1452,189,1941]" pageId="9" pageNumber="298">Paraterga: Poorly developed, somewhat rounded, slightly lower than metatergal surface, not surpassing posterior contour of metaterga 217. Caudal corner rounded, neither pointed nor acute. Calluses minute, almost reduced, without any incision laterally. Ozopores located near caudal corner on lateral side of paraterga 5, 7, 910, 1213, 1519.</paragraph>
<paragraph id="910BCD64CD2BE812FCA990B8FC10FA90" blockId="9.[812,1452,189,1941]" pageId="9" pageNumber="298">
Telson: Epiproct short, broadly truncated without terminal tubercles. Tip with four spinnerets. Hypoproct somewhat parabolical, rounded with two distolateral setiferous knobs (
<figureCitation id="098FD1E1CD2BE812FCB19158FC03FA90" box="[820,907,1341,1365]" captionStart="Fig" captionStartId="10.[136,171,2062,2084]" captionTargetBox="[149,1468,336,1927]" captionTargetId="figure-8@10.[118,1505,318,1949]" captionTargetPageId="10" captionText="Fig. 7. Tylopus golovatchi, new species, holotype: hypoproct, ventral view (A); sternite 5, posterioventral view (B); left gonopod, lateral view (C), mesal view (D); postfemoral region, dorsal view (E). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5347285" httpUri="https://zenodo.org/record/5347285/files/figure.png" pageId="9" pageNumber="298">Fig. 7A</figureCitation>
).
</paragraph>
<paragraph id="910BCD64CD2BE812FCA99138FC7AFA10" blockId="9.[812,1452,189,1941]" pageId="9" pageNumber="298">
Sterna: Sparsely setose, without any modifications; longitudinal and transverse sutures obvious. Male sternum 5 with two tooth-shaped laminae and a pair of cones between coxae 4 (
<figureCitation id="098FD1E1CD2BE812FC1591D8FC6DFA10" box="[912,997,1469,1493]" captionStart="Fig" captionStartId="10.[136,171,2062,2084]" captionTargetBox="[149,1468,336,1927]" captionTargetId="figure-8@10.[118,1505,318,1949]" captionTargetPageId="10" captionText="Fig. 7. Tylopus golovatchi, new species, holotype: hypoproct, ventral view (A); sternite 5, posterioventral view (B); left gonopod, lateral view (C), mesal view (D); postfemoral region, dorsal view (E). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5347285" httpUri="https://zenodo.org/record/5347285/files/figure.png" pageId="9" pageNumber="298">Fig. 7B</figureCitation>
).
</paragraph>
<paragraph id="910BCD64CD2BE812FCA991B8FBA0F990" blockId="9.[812,1452,189,1941]" pageId="9" pageNumber="298">Legs: Strong (male), thin (female), about 1.4 (male) and 1.3 (female) times as long as midbody height. Prefemora normal, not swollen. Ventral part with large tubercles. Tarsi without setose brushes.</paragraph>
<paragraph id="910BCD64CD2BE812FCA99238FCCAF851" blockId="9.[812,1452,189,1941]" pageId="9" pageNumber="298">
Gonopod (
<figureCitation id="098FD1E1CD2BE812FC249238FBA8F9B0" box="[929,1056,1629,1653]" captionStart="Fig" captionStartId="10.[136,171,2062,2084]" captionTargetBox="[149,1468,336,1927]" captionTargetId="figure-8@10.[118,1505,318,1949]" captionTargetPageId="10" captionText="Fig. 7. Tylopus golovatchi, new species, holotype: hypoproct, ventral view (A); sternite 5, posterioventral view (B); left gonopod, lateral view (C), mesal view (D); postfemoral region, dorsal view (E). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5347285" httpUri="https://zenodo.org/record/5347285/files/figure.png" pageId="9" pageNumber="298">Figs. 7CE</figureCitation>
,
<figureCitation id="098FD1E1CD2BE812FBAB9238FBFCF9B0" box="[1070,1140,1629,1653]" captionStart="Fig" captionStartId="11.[113,148,2062,2084]" captionTargetBox="[170,1393,189,2024]" captionTargetId="figure-7@11.[169,1393,189,2024]" captionTargetPageId="11" captionText="Fig. 8. Tylopus golovatchi, new species, holotype: left gonopod, mesal view (A), ventral view (B), and lateral view (C); postfemoral region, lateral view (D)." figureDoi="http://doi.org/10.5281/zenodo.5347287" httpUri="https://zenodo.org/record/5347287/files/figure.png" pageId="9" pageNumber="298">8AD</figureCitation>
): coxite short, with sparsely setose part distoventrally. Prefemur densely setose, set off from femorite by an oblique sulcus laterally. Femorite enlarged distally, slightly twisted and grooved mesally, demarcated from postfemoral region by a V-shaped sulcus laterally. Lamina
<emphasis id="A3C01176CD2BE812FB859298FB8FF8D0" bold="true" box="[1024,1031,1789,1813]" pageId="9" pageNumber="298">l</emphasis>
subrectangular, concealing a long, slender, finger-shaped process
<emphasis id="A3C01176CD2BE812FBFF9378FB01F8F0" bold="true" box="[1146,1161,1821,1845]" pageId="9" pageNumber="298">h</emphasis>
. Process
<emphasis id="A3C01176CD2BE812FB779378FB76F8F0" bold="true" box="[1266,1278,1821,1845]" pageId="9" pageNumber="298">z</emphasis>
, lobes
<emphasis id="A3C01176CD2BE812FACB9378FAECF8F0" bold="true" box="[1358,1380,1821,1845]" pageId="9" pageNumber="298">m</emphasis>
and
<emphasis id="A3C01176CD2BE812FA199378FA23F8F0" bold="true" box="[1436,1451,1821,1845]" pageId="9" pageNumber="298">n</emphasis>
absent. Solenophore strongly coiled, completely sheathing flagelliform solenomere. Tip of solenophore with process
<emphasis id="A3C01176CD2BE812FCA99318FCB3F850" bold="true" box="[812,827,1917,1941]" pageId="9" pageNumber="298">k</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="D9AE9EEFCD2BE812FCA993D9FCC7F830" pageId="9" pageNumber="298" type="distribution">
<paragraph id="910BCD64CD2BE812FCA993D9FCC7F830" blockId="9.[812,1452,1980,2037]" pageId="9" pageNumber="298">
<emphasis id="A3C01176CD2BE812FCA993D9FC31F810" bold="true" box="[812,953,1980,2005]" italics="true" pageId="9" pageNumber="298">Distribution.</emphasis>
— Known only from the
<typeStatus id="4E0F73C6CD2BE812FB6193D8FA9AF810" box="[1252,1298,1981,2005]" pageId="9" pageNumber="298">type</typeStatus>
locality (
<figureCitation id="098FD1E1CD2BE812FA0593D8FCCFF830" captionStart="Fig" captionStartId="21.[113,147,1802,1824]" captionTargetBox="[113,753,1083,1781]" captionTargetId="figure-736@21.[113,753,1083,1779]" captionTargetPageId="21" captionText="Fig. 18.Distribution map ofthe millipede genus Tylopus Jeekel, 1968 in Vietnam. 1: Sa Pa, Lao Cai Prov. (T. crassipes, T. procurvus, T. nodulipes, T. sigma, T. sapaensis, T. maculatus, T. magnicus); 2: Van Ban, Lao Cai Prov. (T. crassipes, T. nodulipes); 3: Xuan Son NP (T. strongylosomoides, T. golovatchi); 4: Tam Dao NP (T. tamdaoensis, T. roseiparaterga, T. strongylosomoides); 5: Cuc Phuong NP (T. granulatus, T. hilaroides, T. topali); 6: Pu Mat NP (T. nodulipes); 7: Huong Son, Ha Tinh prov. (T. nodulipes); 8: Bach MaBa Na NP (T. hilaris, T. multilatus); 9: Ngoc Linh Mts., Kon Tum prov. (T. hilaris, T. phanluongi); 10: Bi DoupNui Ba NP (T. spinisterna)." figureDoi="http://doi.org/10.5281/zenodo.5347309" httpUri="https://zenodo.org/record/5347309/files/figure.png" pageId="9" pageNumber="298">Fig. 18</figureCitation>
)
</paragraph>
</subSubSection>
</treatment>
</document>