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<document id="79D243E4F4055F71B483DAFA2C82E08C" ID-DOI="10.11646/zootaxa.3669.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10098002" ID-ZooBank="35D1C69A-0E41-430A-8483-BEB84F45D8D4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629862386453" checkinUser="felipe" docAuthor="Wang, Shuxia, Du, Zhaohui &amp; Li, Houhun" docDate="2013" docId="D957879E0A2AD651A099D7D0FAF924C2" docLanguage="en" docName="zt03669p455.pdf" docOrigin="Zootaxa 3669 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Promalactis spinosicostalis Wang, Du &amp; Li 2013, sp. nov." docType="treatment" docVersion="3" lastPageNumber="428" masterDocId="256EFFE60A31D64AA00ED747FFE12350" masterDocTitle="Taxonomic study of the genus Promalactis Meyrick, 1908 from Southeast Asia (Lepidoptera: Oecophoridae)" masterLastPageNumber="455" masterPageNumber="401" pageNumber="428" updateTime="1699589443423" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="034A78922E93641F0B30A9A2EF7160C5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2EFC727C9A7C3BCA50C376506310BF81">
<mods:title id="7F2200F4DEAE3F232E9D00877BFD02A1">Taxonomic study of the genus Promalactis Meyrick, 1908 from Southeast Asia (Lepidoptera: Oecophoridae)</mods:title>
</mods:titleInfo>
<mods:name id="9F7CCDCBFE728F5CA5B682970288BA1D" type="personal">
<mods:role id="69E625FF327A7E55419D13C484BA2DE5">
<mods:roleTerm id="293238A823532E551BB25DB642E19C53">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C4CF318688877F393F49C630BD12D3C2">Wang, Shuxia</mods:namePart>
<mods:affiliation id="8E0025CE8809724EB191933F24557919">These authors contributed equally to this work.</mods:affiliation>
</mods:name>
<mods:name id="FD9A430451BA69271A625D68D664FC4B" type="personal">
<mods:role id="CF9911015B1C073DCD54588437409319">
<mods:roleTerm id="9008E1894F77091ABCC4994CAFFE91A7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DAF5B8F7411F175EB2A282C756831F9F">Du, Zhaohui</mods:namePart>
<mods:affiliation id="478614EFE88776EE0535F41DC62797F9">These authors contributed equally to this work.</mods:affiliation>
</mods:name>
<mods:name id="3462F71EC65A7B07E02EA47EFD68D8B3" type="personal">
<mods:role id="71F2836DCDCAC91324A85825A92466F4">
<mods:roleTerm id="1544523AD4BDD177D5083EA4220D9373">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9487507967A65BB5FBD0D7FA8543BA28">Li, Houhun</mods:namePart>
<mods:nameIdentifier id="AC6FA8CCC711A79BFEC73329BD4F4F8F" type="email">lihouhun@nankai.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="73F0B9FCC54370595A27364E04E8AD05">text</mods:typeOfResource>
<mods:relatedItem id="722A9056F27311BCD19DE569693D750E" type="host">
<mods:titleInfo id="8076B0E0067E2AE73F524A5915A9F78C">
<mods:title id="C0D8466DBF105D324B4FB5C2BF113531">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F760E9EF79AE31337FEA8DD6595B00B8">
<mods:date id="233C50A7A924BEF76211C53C5DA343A2">2013</mods:date>
<mods:detail id="4A6A2403E62671F26EA661904C359B9D" type="pubDate">
<mods:number id="096D2851C344244020402E3AC16B5CBF">2013-06-10</mods:number>
</mods:detail>
<mods:detail id="9E65A08FBC78D8256AF3D58868EE8435" type="volume">
<mods:number id="D5658F0592E089FE6E0E1F5457024E2A">3669</mods:number>
</mods:detail>
<mods:detail id="71FC0B782580EFFA75E201ABD3DDF0E7" type="issue">
<mods:number id="7038690D6011805AC9FF852E71267687">4</mods:number>
</mods:detail>
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<mods:start id="D8EC3A9C51AEBF22715D30730B45F16E">401</mods:start>
<mods:end id="1D8812A766416A7AD6A7BA27B1E0764E">455</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="5DFCA4DBEE9BF026B97FA4E0818AE490">journal article</mods:classification>
<mods:identifier id="D6F3DC4313462F8C40F8688A984B0369" type="DOI">10.11646/zootaxa.3669.4.1</mods:identifier>
<mods:identifier id="72757FC9DD6514959B04FE5EA770A312" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C96D19E9430A1EF5C98708ABCAFD815C" type="Zenodo-Dep">10098002</mods:identifier>
<mods:identifier id="4FBD6DEFFB1E2A3FAFE39936E5D49C1D" type="ZooBank">35D1C69A-0E41-430A-8483-BEB84F45D8D4</mods:identifier>
</mods:mods>
<treatment id="D957879E0A2AD651A099D7D0FAF924C2" LSID="urn:lsid:plazi:treatment:D957879E0A2AD651A099D7D0FAF924C2" httpUri="http://treatment.plazi.org/id/D957879E0A2AD651A099D7D0FAF924C2" lastPageNumber="428" pageId="27" pageNumber="428">
<subSubSection id="19E465030A2AD651A099D7D0FCF123E1" box="[151,784,151,177]" pageId="27" pageNumber="428" type="nomenclature">
<paragraph id="514136880A2AD651A099D7D0FCF123E1" blockId="27.[151,784,151,210]" box="[151,784,151,177]" pageId="27" pageNumber="428">
<heading id="0A0981E40A2AD651A099D7D0FCF123E1" bold="true" box="[151,784,151,177]" fontSize="11" level="1" pageId="27" pageNumber="428" reason="1">
<emphasis id="638AEA9A0A2AD651A099D7D0FCF123E1" bold="true" box="[151,784,151,177]" pageId="27" pageNumber="428">
<taxonomicName id="96FE4D0B0A2AD651A099D7D0FD4723E1" authority="Wang, Du &amp; Li" authorityName="Wang, Du &amp; Li" authorityYear="2013" box="[151,678,151,177]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="spinosicostalis" status="sp. nov.">
<emphasis id="638AEA9A0A2AD651A099D7D0FE3C23E1" bold="true" box="[151,477,151,177]" italics="true" pageId="27" pageNumber="428">Promalactis spinosicostalis</emphasis>
Wang, Du &amp; Li
</taxonomicName>
,
<taxonomicNameLabel id="78B957E10A2AD651A2BDD7D0FCF123E1" box="[691,784,151,177]" pageId="27" pageNumber="428" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="19E465030A2AD651A099D7FDFEB02382" box="[151,337,186,210]" pageId="27" pageNumber="428" type="description">
<paragraph id="514136880A2AD651A099D7FDFEB02382" blockId="27.[151,784,151,210]" box="[151,337,186,210]" pageId="27" pageNumber="428">
(
<figureCitation id="C9C52A0D0A2AD651A091D7FDFF142382" box="[159,245,186,210]" captionStart="FIGURES 17–24" captionStartId="25.[151,263,1942,1965]" captionTargetBox="[206,1381,193,1921]" captionTargetId="figure-87@25.[206,1381,193,1921]" captionTargetPageId="25" captionText="FIGURES 17–24. Adults of Promalactis spp. 17, P. medispinata sp. nov., holotype; 18, P.palmifolia sp. nov., holotype; 19, P. pectinifera sp. nov., holotype; 20, P. sectoralis sp. nov., holotype; 21, P.semiovata sp. nov., holotype; 22, P.sinualis sp. nov., holotype; 23, P. spinosicostalis sp. nov., holotype; 24, P.strena sp. nov., holotype (17–21, 23, 24: ♂, 22: ♀)." figureDoi="http://doi.org/10.5281/zenodo.10098008" httpUri="https://zenodo.org/record/10098008/files/figure.png" pageId="27" pageNumber="428">Figs 23</figureCitation>
,
<figureCitation id="C9C52A0D0A2AD651A10FD7FDFEC12382" box="[257,288,186,210]" captionStart="FIGURES 53–58" captionStartId="40.[151,263,1892,1915]" captionTargetBox="[232,1343,183,1856]" captionTargetId="figure-100@40.[206,1381,143,1871]" captionTargetPageId="40" captionText="FIGURES 53–58. Male genitalia of Promalactis spp. 53, P. semiovata sp. nov., holotype, slide No. MNHU-NK006; 54, P. spinosicostalis sp. nov., paratype, slide No. MNHU-NK022; 55, P. strena sp. nov., holotype, 55a, enlarged apical spines of cornutus, slide No. BMNH-33543; 56, P. superimposita sp. nov., holotype, 56a, enlarged apical process of gnathos, slide No. ZMUC-NK002; 57, P. truncatapicalis sp. nov., holotype, slide No. MNHU-NK023; 58, P. undulata sp. nov., holotype, slide No. MNHU-NK002." figureDoi="http://doi.org/10.5281/zenodo.10098021" httpUri="https://zenodo.org/record/10098021/files/figure.png" pageId="27" pageNumber="428">54</figureCitation>
,
<figureCitation id="C9C52A0D0A2AD651A125D7FDFEA92382" box="[299,328,186,210]" captionStart="FIGURES 70–73" captionStartId="44.[151,264,1916,1939]" captionTargetBox="[242,1361,188,1869]" captionTargetId="figure-77@44.[206,1381,167,1895]" captionTargetPageId="44" captionText="FIGURES 70–73. Female genitalia of Promalactis spp. 70, P. spinosicostalis sp. nov., 70a, enlarged strumae of olivary signum, slide No. MNHU-NK019; 71, P. superimposita sp. nov., 71a, enlarged signum, slide No. ZMUC-NK001; 72, P. diehli Lvovsky, 72a, enlarged spines of signum, slide No. MNHU-NK014; 73, P. semantris (Meyrick), 73a, enlarged signum, slide No. ZMUC-NK014." figureDoi="http://doi.org/10.5281/zenodo.10098029" httpUri="https://zenodo.org/record/10098029/files/figure.png" pageId="27" pageNumber="428">70</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="19E465030A2AD651A099D644FC8522FA" pageId="27" pageNumber="428" type="materials_examined">
<paragraph id="514136880A2AD651A099D644FC8522FA" blockId="27.[151,1436,258,1939]" pageId="27" pageNumber="428">
<emphasis id="638AEA9A0A2AD651A099D644FEAC224B" bold="true" box="[151,333,258,283]" pageId="27" pageNumber="428">Type material:</emphasis>
<materialsCitation id="E1963CD50A2AD651A154D645FB7F226E" collectingDate="2006-11-10" collectingDateMax="2006-11-13" collectingDateMin="2006-11-10" collectionCode="MNHU" collectorName="W. Mey &amp; K. Ebert &amp; No." country="Malaysia" elevation="1600" latitude="6.0666666" location="Mt. Kinabalu Headquarter" longLatPrecision="1305" longitude="116.6" municipality="Borneo" pageId="27" pageNumber="428" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="8E45882A0A2AD651A154D645FE23224A" box="[346,450,258,282]" pageId="27" pageNumber="428" type="holotype">Holotype</typeStatus>
<specimenCount id="47F8FD010A2AD651A1C1D644FE05224A" box="[463,484,259,282]" pageId="27" pageNumber="428" type="male">♂</specimenCount>
,
<emphasis id="638AEA9A0A2AD651A1F6D645FD6F224B" bold="true" box="[504,654,258,283]" pageId="27" pageNumber="428">
<collectingCountry id="29E976180A2AD651A1F6D645FD6F224B" box="[504,654,258,283]" name="Malaysia" pageId="27" pageNumber="428">MALAYSIA</collectingCountry>
</emphasis>
,
<location id="542160530A2AD651A2ACD644FC35224A" LSID="urn:lsid:plazi:treatment:D957879E0A2AD651A099D7D0FAF924C2:542160530A2AD651A2ACD644FC35224A" box="[674,980,258,282]" country="Malaysia" latitude="6.0666666" longLatPrecision="1305" longitude="116.6" municipality="Borneo" name="Mt. Kinabalu Headquarter" pageId="27" pageNumber="428">Mt. Kinabalu Headquarter</location>
(
<geoCoordinate id="34CA504F0A2AD651A3E4D645FBDD224A" box="[1002,1084,258,283]" degrees="6" direction="north" minutes="04" orientation="latitude" pageId="27" pageNumber="428" precision="925" value="6.0666666">6°04'N</geoCoordinate>
,
<geoCoordinate id="34CA504F0A2AD651A441D645FB58224A" box="[1103,1209,258,283]" degrees="116" direction="east" minutes="36" orientation="longitude" pageId="27" pageNumber="428" precision="925" value="116.6">116°36'E</geoCoordinate>
),
<collectingMunicipality id="B125ACF20A2AD651A4DAD644FAC8224A" box="[1236,1321,259,282]" pageId="27" pageNumber="428">Borneo</collectingMunicipality>
,
<quantity id="96069B6D0A2AD651A534D645FA76224A" box="[1338,1431,258,282]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="27" pageNumber="428" unit="m" value="1600.0">
<elevation id="DAD3D1BB0A2AD651A534D645FA76224A" box="[1338,1431,258,282]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="27" pageNumber="428" unit="m" value="1600.0">1600 m</elevation>
</quantity>
,
<date id="254010480A2AD651A099D661FEDD226E" box="[151,316,294,318]" pageId="27" pageNumber="428" value="2006-11-10" valueMax="2006-11-13" valueMin="2006-11-10">
<collectingDate id="3504E9A00A2AD651A099D661FEDD226E" box="[151,316,294,318]" pageId="27" pageNumber="428" value="2006-11-10" valueMax="2006-11-13" valueMin="2006-11-10">10−13.xi.2006</collectingDate>
</date>
, leg.
<collectorName id="FC0B535E0A2AD651A18BD660FE05226F" box="[389,484,295,319]" pageId="27" pageNumber="428">W. Mey</collectorName>
&amp;
<collectorName id="FC0B535E0A2AD651A21AD660FD9D226E" box="[532,636,294,318]" pageId="27" pageNumber="428">K. Ebert</collectorName>
, genitalia slide
<collectorName id="FC0B535E0A2AD651A333D660FC87226E" box="[829,870,295,318]" pageId="27" pageNumber="428">No.</collectorName>
MNHU-NK012 (
<collectionCode id="37EFAE4D0A2AD651A434D661FB77226E" box="[1082,1174,294,318]" pageId="27" pageNumber="428">MNHU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E1963CD50A2AD651A4A0D660FEBD22D6" collectingDate="2005-08-09" collectingDateMax="2005-08-13" collectingDateMin="2005-08-09" collectionCode="MNHU-, MNHU" collectorName="W. Mey &amp; K. Ebert &amp; No." country="Malaysia" elevation="1400" location="Borneo" municipality="Mt. Kinabalu" pageId="27" pageNumber="428" specimenCode="NK019" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="8E45882A0A2AD651A4A0D660FAFE226F" box="[1198,1311,295,319]" pageId="27" pageNumber="428" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="47F8FD010A2AD651A53ED661FA81226E" box="[1328,1376,294,318]" pageId="27" pageNumber="428" type="female">1 ♀</specimenCount>
,
<collectingMunicipality id="B125ACF20A2AD651A57AD660FEE02232" pageId="27" pageNumber="428">Mt. Kinabalu</collectingMunicipality>
, kalanggaan,
<location id="542160530A2AD651A1ACD60CFE162232" LSID="urn:lsid:plazi:treatment:D957879E0A2AD651A099D7D0FAF924C2:542160530A2AD651A1ACD60CFE162232" box="[418,503,331,354]" country="Malaysia" municipality="Mt. Kinabalu" name="Borneo" pageId="27" pageNumber="428">Borneo</location>
,
<quantity id="96069B6D0A2AD651A208D60DFD832232" box="[518,610,330,354]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="27" pageNumber="428" unit="m" value="1400.0">
<elevation id="DAD3D1BB0A2AD651A208D60DFD832232" box="[518,610,330,354]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="27" pageNumber="428" unit="m" value="1400.0">1400 m</elevation>
</quantity>
,
<date id="254010480A2AD651A27CD60DFCF82232" box="[626,793,330,354]" pageId="27" pageNumber="428" value="2005-08-09" valueMax="2005-08-13" valueMin="2005-08-09">
<collectingDate id="3504E9A00A2AD651A27CD60DFCF82232" box="[626,793,330,354]" pageId="27" pageNumber="428" value="2005-08-09" valueMax="2005-08-13" valueMin="2005-08-09">9−13.viii.2005</collectingDate>
</date>
, leg.
<collectorName id="FC0B535E0A2AD651A353D60CFC5A2233" box="[861,955,331,355]" pageId="27" pageNumber="428">W. Mey</collectorName>
&amp;
<collectorName id="FC0B535E0A2AD651A3E9D60CFBAC2232" box="[999,1101,330,354]" pageId="27" pageNumber="428">K. Ebert</collectorName>
, genitalia slide
<collectorName id="FC0B535E0A2AD651A507D60CFAD32232" box="[1289,1330,331,354]" pageId="27" pageNumber="428">No.</collectorName>
<collectionCode id="37EFAE4D0A2AD651A533D60CFA7D2232" box="[1341,1436,331,354]" pageId="27" pageNumber="428">MNHU-</collectionCode>
<specimenCode id="01589EF30A2AD651A099D629FF0822D6" box="[151,233,366,390]" collectionCode="MNHU-" pageId="27" pageNumber="428">NK019</specimenCode>
(
<collectionCode id="37EFAE4D0A2AD651A0F9D629FEB222D6" box="[247,339,366,390]" pageId="27" pageNumber="428">MNHU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E1963CD50A2AD651A168D629FC8022FA" collectingDate="2005-08-10" collectingDateMax="2005-08-13" collectingDateMin="2005-08-10" collectionCode="MNHU" collectorName="Mt. Kinabalu NP &amp; Mey &amp; Ebert &amp; No." country="Malaysia" county="Liwagu River Trail" elevation="1500" location="River Trail" municipality="Liwagu River" pageId="27" pageNumber="428" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
<specimenCount id="47F8FD010A2AD651A168D629FE6E22D6" box="[358,399,366,390]" pageId="27" pageNumber="428" type="male">1 ♂</specimenCount>
, light tower,
<collectingCounty id="B8204E040A2AD651A22CD629FD1A22D6" box="[546,763,366,390]" pageId="27" pageNumber="428">
<collectingMunicipality id="B125ACF20A2AD651A22CD629FD5D22D6" box="[546,700,366,390]" pageId="27" pageNumber="428">
Liwagu
<location id="542160530A2AD651A270D629FD5D22D6" LSID="urn:lsid:plazi:treatment:D957879E0A2AD651A099D7D0FAF924C2:542160530A2AD651A270D629FD5D22D6" box="[638,700,366,390]" country="Malaysia" county="Liwagu River Trail" municipality="Liwagu River" name="River Trail" pageId="27" pageNumber="428" stateProvince="Sabah">River</location>
</collectingMunicipality>
Trail
</collectingCounty>
,
<collectorName id="FC0B535E0A2AD651A308D628FC2422D6" box="[774,965,366,390]" pageId="27" pageNumber="428">
<location id="542160530A2AD651A308D628FC7B22D6" LSID="urn:lsid:plazi:treatment:D957879E0A2AD651A099D7D0FAF924C2:542160530A2AD651A308D628FC7B22D6" box="[774,922,366,390]" country="Malaysia" county="Liwagu River Trail" municipality="Liwagu River" name="Mt. Kinabalu" pageId="27" pageNumber="428" stateProvince="Sabah">Mt. Kinabalu</location>
NP
</collectorName>
,
<collectingRegion id="933AF86A0A2AD651A3C1D629FBF722D6" box="[975,1046,366,390]" country="Malaysia" name="Sabah" pageId="27" pageNumber="428">Sabah</collectingRegion>
,
<location id="542160530A2AD651A42ED628FB9422D6" LSID="urn:lsid:plazi:treatment:D957879E0A2AD651A099D7D0FAF924C2:542160530A2AD651A42ED628FB9422D6" box="[1056,1141,367,390]" country="Malaysia" county="Liwagu River Trail" municipality="Liwagu River" name="Borneo" pageId="27" pageNumber="428" stateProvince="Sabah">Borneo</location>
,
<quantity id="96069B6D0A2AD651A48ED629FB3622D6" box="[1152,1239,366,390]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="428" unit="m" value="1500.0">
<elevation id="DAD3D1BB0A2AD651A48ED629FB3622D6" box="[1152,1239,366,390]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="428" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<date id="254010480A2AD651A4ECD629FA7622D6" box="[1250,1431,366,390]" pageId="27" pageNumber="428" value="2005-08-10" valueMax="2005-08-13" valueMin="2005-08-10">
<collectingDate id="3504E9A00A2AD651A4ECD629FA7622D6" box="[1250,1431,366,390]" pageId="27" pageNumber="428" value="2005-08-10" valueMax="2005-08-13" valueMin="2005-08-10">10−13.viii.2005</collectingDate>
</date>
, leg.
<collectorName id="FC0B535E0A2AD651A0C9D6D4FF1B22FB" box="[199,250,403,427]" pageId="27" pageNumber="428">Mey</collectorName>
&amp;
<collectorName id="FC0B535E0A2AD651A110D6D5FEBC22FA" box="[286,349,402,426]" pageId="27" pageNumber="428">Ebert</collectorName>
, genitalia slide
<collectorName id="FC0B535E0A2AD651A202D6D4FDD422FA" box="[524,565,403,426]" pageId="27" pageNumber="428">No.</collectorName>
MNHU-NK022 (
<collectionCode id="37EFAE4D0A2AD651A2F2D6D5FCB922FA" box="[764,856,402,426]" pageId="27" pageNumber="428">MNHU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="19E465030A2AD651A0C9D6F1FD3325C6" pageId="27" pageNumber="428" type="description">
<paragraph id="514136880A2AD651A0C9D6F1FC28275E" blockId="27.[151,1436,258,1939]" pageId="27" pageNumber="428">
<emphasis id="638AEA9A0A2AD651A0C9D6F1FEB9229F" bold="true" box="[199,344,438,463]" inLineHeading="true" pageId="27" pageNumber="428" reason="1">Description.</emphasis>
Imago (
<figureCitation id="C9C52A0D0A2AD651A1B7D6F1FDED229E" box="[441,524,438,462]" captionStart="FIGURES 17–24" captionStartId="25.[151,263,1942,1965]" captionTargetBox="[206,1381,193,1921]" captionTargetId="figure-87@25.[206,1381,193,1921]" captionTargetPageId="25" captionText="FIGURES 17–24. Adults of Promalactis spp. 17, P. medispinata sp. nov., holotype; 18, P.palmifolia sp. nov., holotype; 19, P. pectinifera sp. nov., holotype; 20, P. sectoralis sp. nov., holotype; 21, P.semiovata sp. nov., holotype; 22, P.sinualis sp. nov., holotype; 23, P. spinosicostalis sp. nov., holotype; 24, P.strena sp. nov., holotype (17–21, 23, 24: ♂, 22: ♀)." figureDoi="http://doi.org/10.5281/zenodo.10098008" httpUri="https://zenodo.org/record/10098008/files/figure.png" pageId="27" pageNumber="428">Fig. 23</figureCitation>
) with wingspan 10.0−11.0 mm. Head with vertex and frons shining milk-white, occiput dark brown. Labial palpus with basal and second segments dark brown on outer surface, first segment white on inner surface, second segment dark ochreous yellow on inner surface; third segment with basal 1/4 white, distal 3/4 dark brown, apex white, longer than second. Antenna with scape white, black at apex; flagellum white and black on dorsal surface, dark brown on ventral surface. Thorax and tegula dark ochreous brown. Forewing ochreous brown, markings white edged with black scales: first fascia from base of costal margin to basal 1/5 of dorsal margin, second fascia extending obliquely from 1/4 of costal margin to middle of dorsal margin, slightly arched outward, broader anteriorly, with some black scales on outside anteriorly; costal margin with a subtriangular white patch at 2/3, extending down across midwing, tinged with dark-gray scales on posterior and outer margins, with very small triangular dark-gray spot antero-laterally; an irregularly quadrangular, dark-gray spot before tornus, extending to posterior margin of costal patch, its inner edge with black scales, outside with a small white spot at tornus; apex with a white spot, extending downward along termen, edged with black scales on its inner margin; cilia orange yellow mixed with black, dark gray along distal part of costal margin, gray along distal part of dorsal margin. Foreleg black, tibia with three equational white spots on dorsal surface, tarsus with white spots on dorsal surface; midleg black on dorsal surface, gray on ventral surface, tibia with a white spot near base on dorsal surface, with a tuft of long white scales at apex, tarsus with white spots on dorsal surface; hindleg yellow on ventral surface, gray on dorsal surface, tarsus with white spots on dorsal surface.
</paragraph>
<paragraph id="514136880A2AD651A0C9D35DFE6426EE" blockId="27.[151,1436,258,1939]" pageId="27" pageNumber="428">
Male genitalia (
<figureCitation id="C9C52A0D0A2AD651A177D35DFE2F2762" box="[377,462,1050,1074]" captionStart="FIGURES 53–58" captionStartId="40.[151,263,1892,1915]" captionTargetBox="[232,1343,183,1856]" captionTargetId="figure-100@40.[206,1381,143,1871]" captionTargetPageId="40" captionText="FIGURES 53–58. Male genitalia of Promalactis spp. 53, P. semiovata sp. nov., holotype, slide No. MNHU-NK006; 54, P. spinosicostalis sp. nov., paratype, slide No. MNHU-NK022; 55, P. strena sp. nov., holotype, 55a, enlarged apical spines of cornutus, slide No. BMNH-33543; 56, P. superimposita sp. nov., holotype, 56a, enlarged apical process of gnathos, slide No. ZMUC-NK002; 57, P. truncatapicalis sp. nov., holotype, slide No. MNHU-NK023; 58, P. undulata sp. nov., holotype, slide No. MNHU-NK002." figureDoi="http://doi.org/10.5281/zenodo.10098021" httpUri="https://zenodo.org/record/10098021/files/figure.png" pageId="27" pageNumber="428">Fig. 54</figureCitation>
). Uncus broad, sclerotized laterally, basal 1/3 parallel laterally, narrowed to 3/4, distal 1/4 broadened to straight apex. Gnathos about 3/5 length of uncus, broad tongue shaped, scobinate, narrowly rounded at apex; lateral arms short, triangular, about 1/2 length of gnathos. Tegumen narrow, branched from posterior 3/5, rounded anteriorly. Valva narrow; costa about 3/4 length of valva, with dense long spines on ventral margin except at base, distal 2/5 with denser and longer spines; ventral margin rough, concave medially; apex bluntly rounded, setose, with a small notch at middle. Vinculum roundly projected postero-medially, forming a transverse, arched narrow band; saccus large, about twice length as uncus, very broad at base, gradually narrowed to 2/3, distal 1/3 thumb-like, rounded at apex. Juxta broadly leaf-like, very narrow at base, with a slender basal handle reaching 3/5 length of saccus, folded laterally from basal 1/4 to 3/5, distal 2/5 spinulose laterally, apex narrowly rounded, reaching posterior 1/5 of tegumen. Aedeagus straight, basal 2/5 thin, distal 3/5 gradually broadened, with a large densely spinulose bursa near apex, apex flower-like; cornutus very thin, indistinct, about 1/ 3 length of aedeagus.
</paragraph>
<paragraph id="514136880A2AD651A0C9D28DFD3325C6" blockId="27.[151,1436,258,1939]" pageId="27" pageNumber="428">
Female genitalia (
<figureCitation id="C9C52A0D0A2AD651A1A8D28DFDE426B2" box="[422,517,1482,1506]" captionStart="FIGURES 70–73" captionStartId="44.[151,264,1916,1939]" captionTargetBox="[242,1361,188,1869]" captionTargetId="figure-77@44.[206,1381,167,1895]" captionTargetPageId="44" captionText="FIGURES 70–73. Female genitalia of Promalactis spp. 70, P. spinosicostalis sp. nov., 70a, enlarged strumae of olivary signum, slide No. MNHU-NK019; 71, P. superimposita sp. nov., 71a, enlarged signum, slide No. ZMUC-NK001; 72, P. diehli Lvovsky, 72a, enlarged spines of signum, slide No. MNHU-NK014; 73, P. semantris (Meyrick), 73a, enlarged signum, slide No. ZMUC-NK014." figureDoi="http://doi.org/10.5281/zenodo.10098029" httpUri="https://zenodo.org/record/10098029/files/figure.png" pageId="27" pageNumber="428">Fig. 70</figureCitation>
). Apophysis anterioris about 2/3 length of apophysis posterioris. Lamella postvaginalis heavily sclerotized, nearly oval, with two equational, sclerotized longitudinal folds at middle. Ductus bursae short, about 1/3 length of corpus bursae, sclerotized near base, with two elongately triangular, weakly sclerotized plates extending to posterior 3/10 of corpus bursae. Corpus bursae elongately ovate, membranous; with three signa: one large, olivary, bearing two small strumae at middle; two small, nearly rounded, with small teeth on right margin, placed at both sides of the larger one.
</paragraph>
</subSubSection>
<subSubSection id="19E465030A2AD651A0C9D1E5FC0D2476" pageId="27" pageNumber="428" type="diagnosis">
<paragraph id="514136880A2AD651A0C9D1E5FC0D2476" blockId="27.[151,1436,258,1939]" pageId="27" pageNumber="428">
<emphasis id="638AEA9A0A2AD651A0C9D1E5FEA025EB" bold="true" box="[199,321,1698,1723]" inLineHeading="true" pageId="27" pageNumber="428" reason="1">Diagnosis.</emphasis>
This new species is similar to new species
<taxonomicName id="96FE4D0B0A2AD651A339D1E4FC4D25EA" authority="Wang &amp; Du &amp; Li, 2013" authorityName="Wang &amp; Du &amp; Li" authorityYear="2013" box="[823,940,1699,1722]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="angusta" status="sp. nov.">
<emphasis id="638AEA9A0A2AD651A339D1E4FC4D25EA" box="[823,940,1699,1722]" italics="true" pageId="27" pageNumber="428">P. angusta</emphasis>
</taxonomicName>
<emphasis id="638AEA9A0A2AD651A3BBD1E4FBF025EB" bold="true" box="[949,1041,1699,1723]" pageId="27" pageNumber="428">
<taxonomicNameLabel id="78B957E10A2AD651A3BBD1E4FBF025EB" box="[949,1041,1699,1723]" pageId="27" pageNumber="428" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but can be separated by the costa with dense spines on ventral margin, the broad juxta not bilobed and the aedeagus with a large bursa bearing spinules distally in the male genitalia. In
<taxonomicName id="96FE4D0B0A2AD651A272D1ACFD142452" authority="Wang &amp; Du &amp; Li, 2013" authorityName="Wang &amp; Du &amp; Li" authorityYear="2013" box="[636,757,1771,1794]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="angusta" status="sp. nov.">
<emphasis id="638AEA9A0A2AD651A272D1ACFD142452" box="[636,757,1771,1794]" italics="true" pageId="27" pageNumber="428">P. angusta</emphasis>
</taxonomicName>
<emphasis id="638AEA9A0A2AD651A2F1D1ACFCBD2453" bold="true" box="[767,860,1771,1795]" pageId="27" pageNumber="428">
<taxonomicNameLabel id="78B957E10A2AD651A2F1D1ACFCBD2453" box="[767,860,1771,1795]" pageId="27" pageNumber="428" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the costa has no spines, the juxta is narrow and bilobed in distal 1/5 and the aedeagus has no bursa bearing spinules distally.
</paragraph>
</subSubSection>
<subSubSection id="19E465030A2AD651A0C9D075FDA0241A" box="[199,577,1842,1867]" pageId="27" pageNumber="428" type="distribution">
<paragraph id="514136880A2AD651A0C9D075FDA0241A" blockId="27.[151,1436,258,1939]" box="[199,577,1842,1867]" pageId="27" pageNumber="428">
<emphasis id="638AEA9A0A2AD651A0C9D075FE81241B" bold="true" box="[199,352,1842,1867]" inLineHeading="true" pageId="27" pageNumber="428" reason="1">Distribution.</emphasis>
<collectingCountry id="29E976180A2AD651A169D075FE2C241A" box="[359,461,1842,1866]" name="Malaysia" pageId="27" pageNumber="428">Malaysia</collectingCountry>
(Borneo).
</paragraph>
</subSubSection>
<subSubSection id="19E465030A2AD651A0C9D010FAF924C2" pageId="27" pageNumber="428" type="etymology">
<paragraph id="514136880A2AD651A0C9D010FAF924C2" blockId="27.[151,1436,258,1939]" pageId="27" pageNumber="428">
<emphasis id="638AEA9A0A2AD651A0C9D010FEAC243F" bold="true" box="[199,333,1879,1903]" inLineHeading="true" pageId="27" pageNumber="428" reason="1">Etymology.</emphasis>
The specific name is derived from the Latin
<emphasis id="638AEA9A0A2AD651A37FD010FC30243E" box="[881,977,1879,1902]" italics="true" pageId="27" pageNumber="428">spinosus</emphasis>
(= having many spines) and the Latin
<emphasis id="638AEA9A0A2AD651A099D03CFF0C24C3" box="[151,237,1915,1939]" italics="true" pageId="27" pageNumber="428">costalis</emphasis>
(= costa), referring to the ventral margin of the costa having dense spines in the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>
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