278 lines
38 KiB
XML
278 lines
38 KiB
XML
<document id="686FAE4546295E5C40A81E1BA774F3D9" ID-DOI="10.1649/072.067.0mo4.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="4907452" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623090705760" checkinUser="carolina" docAuthor="Lord, Nathan P. & McHugh, Joseph V." docDate="2013" docId="1F35BA507D65FA29ACDA85D0FCFDF0AD" docLanguage="en" docName="TheColeopBull.67.mo12.1-107.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 12) 67" docStyle="DocumentStyle:4CAA047AD53AEBA766D4E826C2986B94.5:TheColeopBull.2010-2016.monograph" docStyleId="4CAA047AD53AEBA766D4E826C2986B94" docStyleName="TheColeopBull.2010-2016.monograph" docStyleVersion="5" docTitle="Deretaphrus parviceps Lea 1898" docType="treatment" docVersion="3" lastPageNumber="45" masterDocId="E30CC2287D4EFA05AC3E8037FFB8F13C" masterDocTitle="A Taxonomic Revision of the Genus Deretaphrus Newman, 1842 (Coleoptera: Cucujoidea: Bothrideridae)" masterLastPageNumber="107" masterPageNumber="1" pageNumber="44" updateTime="1698955835990" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="791679D0CD10DD6159821FECD3CA7258">A Taxonomic Revision of the Genus Deretaphrus Newman, 1842 (Coleoptera: Cucujoidea: Bothrideridae)</mods:title>
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<mods:namePart id="A5B7B8486F7D0D12CEBCDA426872F979">Lord, Nathan P.</mods:namePart>
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<mods:affiliation id="A7FF45009DC75A22A07D5083217BD57E">Department of Biology, 401 WIDB, Brigham Young University, Provo, UT 84602, U. S. A.</mods:affiliation>
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<mods:namePart id="87638CD6505C9A3DF9A42D97ADE52988">McHugh, Joseph V.</mods:namePart>
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<mods:affiliation id="B8FA0B4237186DAE2BBAA5A40067825C">Department of Entomology, University of Georgia Athens, GA 30602, U. S. A.</mods:affiliation>
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<mods:title id="381FD342522520723BF919FA0F5F039B">The Coleopterists Bulletin</mods:title>
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<mods:part id="C143B8E71C6E746C22DE49AC1EB475A4">
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<mods:date id="60AAD13C2479DA5939331CD359A4C445">2013</mods:date>
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<mods:detail id="204EAA9B4DBCF2DC7DACBF4DD1BFA335" type="series">
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<mods:title id="1346069B226A5E701D6A9F96E448C822">mo 12</mods:title>
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<mods:number id="1BD0CF80D9485FA8DC44A375562B82E7">2013-12-20</mods:number>
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<subSubSection id="DF8658CD7D65FA2EACDA85D0FD89F4C1" box="[228,561,1511,1533]" pageId="43" pageNumber="44" type="nomenclature">
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<paragraph id="97230B467D65FA2EACDA85D0FD89F4C1" blockId="43.[156,633,1511,1560]" box="[228,561,1511,1533]" pageId="43" pageNumber="44">
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<heading id="CC6BBC2A7D65FA2EACDA85D0FD89F4C1" box="[228,561,1511,1533]" centered="true" fontSize="10" level="2" pageId="43" pageNumber="44" reason="2">
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<taxonomicName id="509C70C57D65FA2EACDA85D0FD89F4C1" authority="Lea, 1898" authorityName="Lea" authorityYear="1898" box="[228,561,1511,1533]" class="Insecta" family="Bothrideridae" genus="Deretaphrus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="parviceps">
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<emphasis id="A5E8D7547D65FA2EACDA85D0FD89F4C1" bold="true" box="[228,561,1511,1533]" pageId="43" pageNumber="44">
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<emphasis id="A5E8D7547D65FA2EACDA85D0FE7BF4C0" bold="true" box="[228,451,1511,1532]" italics="true" pageId="43" pageNumber="44">Deretaphrus parviceps</emphasis>
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<bibRefCitation id="F30D76B77D65FA2EADF585DFFD89F4C1" author="Lea" box="[459,561,1512,1533]" pageId="43" pageNumber="44" pagination="521 - 645" refId="ref63142" refString="Lea, A. M. 1898. Descriptions of new species of Australian Coleoptera. Part V. The Proceedings of the Linnean Society of New South Wales 23: 521 - 645." type="journal article" year="1898">Lea, 1898</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="DF8658CD7D65FA2EACA28635FDC1F72B" box="[156,633,1538,1560]" pageId="43" pageNumber="44" type="description">
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<paragraph id="97230B467D65FA2EACA28635FDC1F72B" blockId="43.[156,633,1511,1560]" box="[156,633,1538,1560]" pageId="43" pageNumber="44">
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(
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<figureCitation id="0FA717C37D65FA2EAC9D8635FF4EF72B" box="[163,246,1538,1559]" captionStart="Figs" captionStartId="71.[175,215,1597,1616]" captionTargetBox="[151,1171,194,1569]" captionTargetId="figure-1@71.[151,1172,193,1619]" captionTargetPageId="71" captionText="Figs. 35–41. Deretaphrus species, head. 35) D. bucculentus, dorsal view; 36) D. carinatus, dorsal view; 37) D. incultus, dorsal view; 38) D. parviceps, dorsal view; 39) D. rodmani, dorsal view; 40) D. ocularis, dorsal view; 41) D. ocularis, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4907486" httpUri="https://zenodo.org/record/4907486/files/figure.png" pageId="43" pageNumber="44">Figs. 38</figureCitation>
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,
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<figureCitation id="0FA717C37D65FA2EAD3C8635FEA4F72B" box="[258,284,1538,1559]" captionStart="Figs" captionStartId="78.[197,237,1618,1637]" captionTargetBox="[173,1193,193,1590]" captionTargetId="figure-1@78.[173,1194,193,1623]" captionTargetPageId="78" captionText="Figs. 75–81. Deretaphrus species, pronotum. 75) D. parviceps; 76) D. piceus; 77) D. puncticollis; 78) D. rodmani;" figureDoi="http://doi.org/10.5281/zenodo.4907500" httpUri="https://zenodo.org/record/4907500/files/figure.png" pageId="43" pageNumber="44">75</figureCitation>
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,
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<figureCitation id="0FA717C37D65FA2EAD168635FEC4F72B" box="[296,380,1538,1560]" captionStart="Figs" captionStartId="85.[175,215,1605,1624]" captionTargetBox="[151,1171,193,1577]" captionTargetId="figure-1@85.[151,1172,193,1612]" captionTargetPageId="85" captionText="Figs. 118–123. Deretaphrus species, elytra, dorsal and lateral views. 118–119) D. parviceps; 120–121) D. piceus;" figureDoi="http://doi.org/10.5281/zenodo.4907516" httpUri="https://zenodo.org/record/4907516/files/figure.png" pageId="43" pageNumber="44">118–119</figureCitation>
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,
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<figureCitation id="0FA717C37D65FA2EADB68635FE16F72B" box="[392,430,1538,1559]" captionStart="Figs" captionStartId="91.[175,215,1597,1616]" captionTargetBox="[170,1163,195,1569]" captionTargetId="figure-1@91.[151,1173,183,1616]" captionTargetPageId="91" captionText="Figs. 158–164. Deretaphrus species, phallobase. 158) D. parviceps; 159) D. piceus; 160) D. puncticollis;" figureDoi="http://doi.org/10.5281/zenodo.4907528" httpUri="https://zenodo.org/record/4907528/files/figure.png" pageId="43" pageNumber="44">158</figureCitation>
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,
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<figureCitation id="0FA717C37D65FA2EAD848635FE58F72B" box="[442,480,1538,1559]" captionStart="Figs" captionStartId="92.[197,237,1321,1340]" captionTargetBox="[186,1185,196,1294]" captionTargetId="figure-1@92.[173,1195,183,1358]" captionTargetPageId="92" captionText="Figs. 165–172. Deretaphrus species, penis (left: dorsal view; right: lateral view; phallobase, to scale with penis). 165) D. boliviensis; 166) D. bucculentus; 167) D. gracilis; 168) D. hoplites; 169) D. ignarus; 170) D. oregonensis; 171) D. parviceps; 172) D. puncticollis." figureDoi="http://doi.org/10.5281/zenodo.4907530" httpUri="https://zenodo.org/record/4907530/files/figure.png" pageId="43" pageNumber="44">171</figureCitation>
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,
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<figureCitation id="0FA717C37D65FA2EADD28635FDAAF72B" box="[492,530,1538,1559]" captionStart="Figs" captionStartId="93.[175,215,1480,1499]" captionTargetBox="[252,1071,238,1451]" captionTargetId="figure-1@93.[151,1173,183,1504]" captionTargetPageId="93" captionText="Figs. 173–174. Distribution of Deretaphrus species. 173) D. alveolatus, D. antennatus, D. parviceps, and D. rodmani;" figureDoi="http://doi.org/10.5281/zenodo.4907532" httpUri="https://zenodo.org/record/4907532/files/figure.png" pageId="43" pageNumber="44">173</figureCitation>
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,
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<figureCitation id="0FA717C37D65FA2EAE238635FDCAF72B" box="[541,626,1538,1560]" captionStart="Figs" captionStartId="102.[197,237,1618,1637]" captionTargetBox="[173,1193,193,1590]" captionTargetId="figure-1@102.[173,1194,193,1623]" captionTargetPageId="102" captionText="Figs. 219–227. Primary type specimens and specimen labels of Deretaphrus species. 219) D. iridescens, lectotype labels; 220–221) D. lateropunctatus, holotype, BMNH; 222–223) D. ocularis, holotype, SAMA; 224–225) D. oregonensis, lectotype, MCZ; 226–227) D. parviceps, lectotype, SAMA." figureDoi="http://doi.org/10.5281/zenodo.4907552" httpUri="https://zenodo.org/record/4907552/files/figure.png" pageId="43" pageNumber="44">226–227</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="DF8658CD7D65FA2EACA9861AFEA1F74B" pageId="43" pageNumber="44" type="reference_group">
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<paragraph id="97230B467D65FA2EACA9861AFEA1F74B" blockId="43.[151,638,1581,1655]" pageId="43" pageNumber="44">
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<treatmentCitation id="163D2D577D65FA2EACA9861AFDA8F77E" author="Lea" box="[151,528,1581,1602]" page="548" pageId="43" pageNumber="44" year="1898">
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<taxonomicName id="509C70C57D65FA2EACA9861AFDA8F77E" authority="Lea, 1898: 548" authorityName="Lea" authorityPageNumber="548" authorityYear="1898" box="[151,528,1581,1602]" class="Insecta" family="Bothrideridae" genus="Deretaphrus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="parviceps">
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<emphasis id="A5E8D7547D65FA2EACA9861AFECAF77E" box="[151,370,1581,1602]" italics="true" pageId="43" pageNumber="44">Deretaphrus parviceps</emphasis>
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<bibRefCitation id="F30D76B77D65FA2EAD44861AFDA8F77E" author="Lea" box="[378,528,1581,1602]" pageId="43" pageNumber="44" pagination="521 - 645" refId="ref63142" refString="Lea, A. M. 1898. Descriptions of new species of Australian Coleoptera. Part V. The Proceedings of the Linnean Society of New South Wales 23: 521 - 645." type="journal article" year="1898">Lea, 1898: 548</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
.
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<treatmentCitation id="163D2D577D65FA2EAE25861AFEAEF760" author="Blackburn" page="120" pageId="43" pageNumber="44" year="1903">
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||
<bibRefCitation id="F30D76B77D65FA2EAE25861AFEAEF760" author="Blackburn" pageId="43" pageNumber="44" pagination="91 - 182" refId="ref61766" refString="Blackburn, T. 1903. Further notes on Australian Coleoptera, with descriptions of new genera and species. Transactions of the Royal Society of South Australia 27: 91 - 182." type="journal article" year="1903">Blackburn 1903: 120</bibRefCitation>
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</treatmentCitation>
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||
; Carter and Zeck 1937: 200;
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<treatmentCitation id="163D2D577D65FA2EAE058670FEADF74B" author="Heinze" page="119" pageId="43" pageNumber="44" year="1943">
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||
<bibRefCitation id="F30D76B77D65FA2EAE058670FEADF74B" author="Heinze" pageId="43" pageNumber="44" pagination="97 - 124" refId="ref62411" refString="Heinze, E. 1943. Studien zur Kenntnis der Tribus Deretaphrini und deren Stellung im System (Colydiidae). Entomologische Blatter fur Biologie und Systematik der Kafer 39: 85 - 93, 97 - 124." type="journal article" year="1943">Heinze 1943: 119</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="DF8658CD7D65FA2EAC9186A6FBC5F239" pageId="43" pageNumber="44" type="diagnosis">
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<paragraph id="97230B467D65FA2EAC9186A6FDC6F7FD" blockId="43.[151,638,1681,1729]" pageId="43" pageNumber="44">
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<emphasis id="A5E8D7547D65FA2EAC9186A6FEA0F79A" bold="true" box="[175,280,1681,1702]" pageId="43" pageNumber="44">Diagnosis.</emphasis>
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This species can be readily distinguished from congeners by the nearly glabrous
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</paragraph>
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<paragraph id="97230B467D65FA2EAE9380F5FBC5F239" blockId="43.[685,1173,194,1729]" pageId="43" pageNumber="44">
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antennae, the extremely convex eyes (nearing angulate), the weak pronotal median longitudinal depression, elytral interstitial intervals 3, 5, and 7 being moderately carinate, the carina on intervals 3 and 5 merging near apex, and a distribution restricted to
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<collectingRegion id="5558C5A47D65FA2EAF6E8170FC45F060" box="[848,1021,327,348]" country="Australia" name="Western Australia" pageId="43" pageNumber="44">Western Australia</collectingRegion>
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. It most closely resembles
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<taxonomicName id="509C70C57D65FA2EAF258155FC0CF04B" box="[795,948,354,375]" class="Insecta" family="Bothrideridae" genus="Deretaphrus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="bucculentus">
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<emphasis id="A5E8D7547D65FA2EAF258155FC0CF04B" box="[795,948,354,375]" italics="true" pageId="43" pageNumber="44">D. bucculentus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="509C70C57D65FA2EAFF98155FBE6F04B" authorityName="Lea. Carter and Zeck" authorityYear="1898" box="[967,1118,354,375]" class="Insecta" family="Bothrideridae" genus="Deretaphrus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="puncticollis">
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<emphasis id="A5E8D7547D65FA2EAFF98155FBE6F04B" box="[967,1118,354,375]" italics="true" pageId="43" pageNumber="44">D. puncticollis</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="509C70C57D65FA2EAE93814BFCA1F0AD" authorityName="Carter and Zeck" authorityYear="1937" box="[685,793,380,401]" class="Insecta" family="Bothrideridae" genus="Deretaphrus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="incultus">
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<emphasis id="A5E8D7547D65FA2EAE93814BFCA1F0AD" box="[685,793,380,401]" italics="true" pageId="43" pageNumber="44">D. incultus</emphasis>
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</taxonomicName>
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.
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<taxonomicName id="509C70C57D65FA2EAF16814BFBBCF0AD" box="[808,1028,380,401]" class="Insecta" family="Bothrideridae" genus="Deretaphrus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="parviceps">
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<emphasis id="A5E8D7547D65FA2EAF16814BFBBCF0AD" box="[808,1028,380,401]" italics="true" pageId="43" pageNumber="44">Deretaphrus parviceps</emphasis>
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</taxonomicName>
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can be distinguished from
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<taxonomicName id="509C70C57D65FA2EAF0681A0FC72F090" box="[824,970,407,428]" class="Insecta" family="Bothrideridae" genus="Deretaphrus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="bucculentus">
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||
<emphasis id="A5E8D7547D65FA2EAF0681A0FC72F090" box="[824,970,407,428]" italics="true" pageId="43" pageNumber="44">D. bucculentus</emphasis>
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||
</taxonomicName>
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by the elytral interstitial interval 5 curving inward, merging or narrowly merging with interstitial interval 3 near the apex and the head lacking laterally expanded supra-ocular ridges. It can be distinguished from
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<taxonomicName id="509C70C57D65FA2EAE93822BFC86F30D" authorityName="Lea. Carter and Zeck" authorityYear="1898" box="[685,830,540,561]" class="Insecta" family="Bothrideridae" genus="Deretaphrus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="puncticollis">
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<emphasis id="A5E8D7547D65FA2EAE93822BFC86F30D" box="[685,830,540,561]" italics="true" pageId="43" pageNumber="44">D. puncticollis</emphasis>
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</taxonomicName>
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by the less densely and coarsely punctate pronotal disc and the median longitudinal depression being more strongly punctate near the base, not bordered on either side by a small keel or carina. It can be distinguished from
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<taxonomicName id="509C70C57D65FA2EA82F82B1FBCEF3A7" authorityName="Carter and Zeck" authorityYear="1937" box="[1041,1142,646,667]" class="Insecta" family="Bothrideridae" genus="Deretaphrus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="incultus">
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<emphasis id="A5E8D7547D65FA2EA82F82B1FBCEF3A7" box="[1041,1142,646,667]" italics="true" pageId="43" pageNumber="44">D. incultus</emphasis>
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</taxonomicName>
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by the more weakly developed (but present) pronotal median longitudinal depression and elytral interstitial interval 5 curving inward and merging or narrowly merging with interstitial interval 3 near the apex.
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</paragraph>
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</subSubSection>
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<subSubSection id="DF8658CD7D65FA29AEFB833DFEEBF592" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44" type="description">
|
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<paragraph id="97230B467D65FA29AEFB833DFDE1F578" blockId="43.[685,1173,194,1729]" lastBlockId="44.[173,660,194,1729]" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">
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<emphasis id="A5E8D7547D65FA2EAEFB833DFCE1F223" bold="true" box="[709,857,778,799]" pageId="43" pageNumber="44">Redescription.</emphasis>
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Length
|
||
<quantity id="5064A6A37D65FA2EAF83833DFBFDF223" box="[957,1093,778,800]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" metricValueMax="9.3" metricValueMin="8.5" pageId="43" pageNumber="44" unit="mm" value="8.9" valueMax="9.3" valueMin="8.5">8.5–9.3 mm</quantity>
|
||
. Width
|
||
<quantity id="5064A6A37D65FA2EAE938312FC9FF206" box="[685,807,805,827]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="43" pageNumber="44" unit="mm" value="2.6" valueMax="2.7" valueMin="2.5">2.5–2.7 mm</quantity>
|
||
. Body elongate, parallel, dark red to black; dorsal surface glabrous, moderately shiny; ventral surface moderately shiny, glabrous except for minute setae that arise from punctures.
|
||
<emphasis id="A5E8D7547D65FA2EA8638342FB2BF2B6" bold="true" box="[1117,1171,885,906]" pageId="43" pageNumber="44">Head</emphasis>
|
||
(
|
||
<figureCitation id="0FA717C37D65FA2EAE8B83B8FCBDF298" box="[693,773,911,932]" captionStart="Figs" captionStartId="71.[175,215,1597,1616]" captionTargetBox="[151,1171,194,1569]" captionTargetId="figure-1@71.[151,1172,193,1619]" captionTargetPageId="71" captionText="Figs. 35–41. Deretaphrus species, head. 35) D. bucculentus, dorsal view; 36) D. carinatus, dorsal view; 37) D. incultus, dorsal view; 38) D. parviceps, dorsal view; 39) D. rodmani, dorsal view; 40) D. ocularis, dorsal view; 41) D. ocularis, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4907486" httpUri="https://zenodo.org/record/4907486/files/figure.png" pageId="43" pageNumber="44">Fig. 38</figureCitation>
|
||
): In dorsal view, narrowed anteriorly; without laterally expanded supra-ocular ridges; eyes strongly protuberant (nearing angulate), visible from above; punctures small and dense except sparse along posterior margin. Frontoclypeal suture not distinct, arcuate anteromedially. Clypeus distinctly narrower than head. Anterior margin of clypeus slightly arcuate medially. Submentum shape as in
|
||
<figureCitation id="0FA717C37D65FA2EAF188453FCD7F545" box="[806,879,1124,1145]" captionStart="Figs" captionStartId="74.[197,237,1496,1515]" captionTargetBox="[173,1193,236,1467]" captionTargetId="figure-1@74.[173,1194,183,1524]" captionTargetPageId="74" captionText="Figs. 48–53. Deretaphrus species, head capsule, ventral view, showing setae and shape of submentum. 48) D. ignarus;" figureDoi="http://doi.org/10.5281/zenodo.4907492" httpUri="https://zenodo.org/record/4907492/files/figure.png" pageId="43" pageNumber="44">Fig. 52</figureCitation>
|
||
; triangular, fused to subgenal braces, without paired setose pits; anterior margin arcuate, strongly produced anteroventrally over oral cavity, mentum completely concealed. Antennal groove well-developed, impunctate internally. Antenna sparsely setose, appearing glabrous except under high magnification. Antennal club distinctly asymmetrical, leading edge of club segments much more inflated than trailing edge. Dorsal surface (external face) of mandible without median setose groove.
|
||
<emphasis id="A5E8D7547D65FA2EAF3A8559FCE9F4BF" bold="true" box="[772,849,1390,1411]" pageId="43" pageNumber="44">Thorax</emphasis>
|
||
(
|
||
<figureCitation id="0FA717C37D65FA2EAF5B8559FC0DF4BF" box="[869,949,1390,1411]" captionStart="Figs" captionStartId="78.[197,237,1618,1637]" captionTargetBox="[173,1193,193,1590]" captionTargetId="figure-1@78.[173,1194,193,1623]" captionTargetPageId="78" captionText="Figs. 75–81. Deretaphrus species, pronotum. 75) D. parviceps; 76) D. piceus; 77) D. puncticollis; 78) D. rodmani;" figureDoi="http://doi.org/10.5281/zenodo.4907500" httpUri="https://zenodo.org/record/4907500/files/figure.png" pageId="43" pageNumber="44">Fig. 75</figureCitation>
|
||
): Pronotum quadrate, slightly tapering to base, widest near anterior margin; dorsal surface nearly flat, deflexed near anterior and posterior angles; anterior margin straight; anterior angles broadly rounded; base narrower than elytral bases; posterior margin weakly sinuate; posterior angles with small denticle; lateral margin with incomplete carina, forming a distinct, large raised border in dorsal view. Pronotal disc with large, deep, round to oval punctures. Pronotal median longitudinal depression present but very faint; punctures largest within depression, oval anteriorly, round posteriorly. Hypomeron sparsely punctate, punctures large and shallow, each bearing a seta; lateral walls strongly deflexed. Prosternum flat medially, strongly deflexed at lateral margin. Tergosternal suture more or less straight.
|
||
<emphasis id="A5E8D7547D62FA29AD3D811BFEFBF07D" bold="true" box="[259,323,300,321]" pageId="44" pageNumber="45">Elytra</emphasis>
|
||
(
|
||
<figureCitation id="0FA717C37D62FA29AD6D811BFE59F07D" box="[339,481,300,322]" captionStart="Figs" captionStartId="85.[175,215,1605,1624]" captionTargetBox="[151,1171,193,1577]" captionTargetId="figure-1@85.[151,1172,193,1612]" captionTargetPageId="85" captionText="Figs. 118–123. Deretaphrus species, elytra, dorsal and lateral views. 118–119) D. parviceps; 120–121) D. piceus;" figureDoi="http://doi.org/10.5281/zenodo.4907516" httpUri="https://zenodo.org/record/4907516/files/figure.png" pageId="44" pageNumber="45">Figs. 118–119</figureCitation>
|
||
): Elytron flat, not evenly convex; interstitial intervals 3, 5, 7, and 9 weakly raised, more pronounced near apex; interval 5 curves inward and merges with interval 3 near the apex. Crest of interstitial intervals with micropunctures bearing minute setae. Basal elytral margin with intervals 3 and 5 each ending in a knob-like protuberance; protuberance of interval 5 merges with finger-like callosity on the elytral shoulder.
|
||
<emphasis id="A5E8D7547D62FA29AD34822CFE81F30C" bold="true" box="[266,313,539,560]" pageId="44" pageNumber="45">Legs</emphasis>
|
||
: Femur and tibia with short, sparse setae; inner face of tibia with small fringe of setae. Tarsus sparsely setose except for 2 pockets of dense setae on the ventral surface at the apex of tarsomeres 1–3.
|
||
<emphasis id="A5E8D7547D62FA29AD5282B1FE77F3A7" bold="true" box="[364,463,646,667]" pageId="44" pageNumber="45">Abdomen</emphasis>
|
||
: Ventrite V simple.
|
||
<emphasis id="A5E8D7547D62FA29AC938297FEB6F389" bold="true" box="[173,270,672,693]" pageId="44" pageNumber="45">Aedeagus</emphasis>
|
||
: Phallobase as in
|
||
<figureCitation id="0FA717C37D62FA29AD858297FDB5F389" box="[443,525,672,693]" captionStart="Figs" captionStartId="91.[175,215,1597,1616]" captionTargetBox="[170,1163,195,1569]" captionTargetId="figure-1@91.[151,1173,183,1616]" captionTargetPageId="91" captionText="Figs. 158–164. Deretaphrus species, phallobase. 158) D. parviceps; 159) D. piceus; 160) D. puncticollis;" figureDoi="http://doi.org/10.5281/zenodo.4907528" httpUri="https://zenodo.org/record/4907528/files/figure.png" pageId="44" pageNumber="45">Fig. 158</figureCitation>
|
||
. Tegmen consisting of paired, transverse plates at middle; plates narrowly touching medially. Basal piece with short anterolateral struts; struts slightly curved anteriorly. Parameres moderately short, somewhat elongate, rounded at apices, widely separated and individually articulated to phallobase, internal lateral margins arcuate; paramere sparsely setose with short setae at apical margin; dorsal surface of paramere with short subtriangular process; process thickest at base, narrowing apically, about as long as 1/3 the length of the paramere, apex slightly swollen; processes widely separated. Penis as in
|
||
<figureCitation id="0FA717C37D62FA29AE3D83E8FDEBF2C8" box="[515,595,991,1012]" captionStart="Figs" captionStartId="92.[197,237,1321,1340]" captionTargetBox="[186,1185,196,1294]" captionTargetId="figure-1@92.[173,1195,183,1358]" captionTargetPageId="92" captionText="Figs. 165–172. Deretaphrus species, penis (left: dorsal view; right: lateral view; phallobase, to scale with penis). 165) D. boliviensis; 166) D. bucculentus; 167) D. gracilis; 168) D. hoplites; 169) D. ignarus; 170) D. oregonensis; 171) D. parviceps; 172) D. puncticollis." figureDoi="http://doi.org/10.5281/zenodo.4907530" httpUri="https://zenodo.org/record/4907530/files/figure.png" pageId="44" pageNumber="45">Fig. 171</figureCitation>
|
||
; short, moderately thick, more or less straight; apex narrowed and pointed; base slightly expanded, bearing short, paired anterior struts. Penis unicolored.
|
||
</paragraph>
|
||
<paragraph id="97230B467D62FA29ACFB847EFEEBF592" blockId="44.[173,660,194,1729]" pageId="44" pageNumber="45">
|
||
<emphasis id="A5E8D7547D62FA29ACFB847EFE91F562" bold="true" box="[197,297,1097,1118]" pageId="44" pageNumber="45">Variation.</emphasis>
|
||
The punctures in the center of the pronotal disc are sometimes less dense, and the puncture shape on the elytra varies from round to slightly irregular.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DF8658CD7D62FA29ACFB8483FEAAF5DF" pageId="44" pageNumber="45" type="distribution">
|
||
<paragraph id="97230B467D62FA29ACFB8483FEAAF5DF" blockId="44.[173,660,194,1729]" pageId="44" pageNumber="45">
|
||
<emphasis id="A5E8D7547D62FA29ACFB8483FEF6F5F5" bold="true" box="[197,334,1204,1225]" pageId="44" pageNumber="45">Distribution.</emphasis>
|
||
<collectingCountry id="EF8B4BD67D62FA29AD638483FE7BF5F5" box="[349,451,1204,1225]" name="Australia" pageId="44" pageNumber="45">Australia</collectingCountry>
|
||
:
|
||
<collectingRegion id="5558C5A47D62FA29ADEF8483FD2BF5F5" box="[465,659,1204,1225]" country="Australia" name="Western Australia" pageId="44" pageNumber="45">Western Australia</collectingRegion>
|
||
(
|
||
<figureCitation id="0FA717C37D62FA29AC8A84F9FEBFF5DF" box="[180,263,1230,1251]" captionStart="Figs" captionStartId="93.[175,215,1480,1499]" captionTargetBox="[252,1071,238,1451]" captionTargetId="figure-1@93.[151,1173,183,1504]" captionTargetPageId="93" captionText="Figs. 173–174. Distribution of Deretaphrus species. 173) D. alveolatus, D. antennatus, D. parviceps, and D. rodmani;" figureDoi="http://doi.org/10.5281/zenodo.4907532" httpUri="https://zenodo.org/record/4907532/files/figure.png" pageId="44" pageNumber="45">Fig. 173</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DF8658CD7D62FA29ACFB84DEFE2FF5C2" box="[197,407,1257,1278]" pageId="44" pageNumber="45" type="biology_ecology">
|
||
<paragraph id="97230B467D62FA29ACFB84DEFE2FF5C2" blockId="44.[173,660,194,1729]" box="[197,407,1257,1278]" pageId="44" pageNumber="45">
|
||
<emphasis id="A5E8D7547D62FA29ACFB84DEFEAEF5C2" bold="true" box="[197,278,1257,1278]" pageId="44" pageNumber="45">Biology.</emphasis>
|
||
Not known.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DF8658CD7D62FA29ACFB8534FD2CF454" pageId="44" pageNumber="45" type="discussion">
|
||
<paragraph id="97230B467D62FA29ACFB8534FD2CF454" blockId="44.[173,660,194,1729]" pageId="44" pageNumber="45">
|
||
<emphasis id="A5E8D7547D62FA29ACFB8534FE90F424" bold="true" box="[197,296,1283,1304]" pageId="44" pageNumber="45">Remarks.</emphasis>
|
||
<bibRefCitation id="F30D76B77D62FA29AD0B8534FE16F424" author="Lea" box="[309,430,1283,1304]" pageId="44" pageNumber="45" pagination="521 - 645" refId="ref63142" refString="Lea, A. M. 1898. Descriptions of new species of Australian Coleoptera. Part V. The Proceedings of the Linnean Society of New South Wales 23: 521 - 645." type="journal article" year="1898">Lea (1898)</bibRefCitation>
|
||
did not mention the number of specimens he examined. In order to stabilize this name, a
|
||
<typeStatus id="4827B5E47D62FA29AD4E850FFE73F471" box="[368,459,1336,1357]" pageId="44" pageNumber="45" type="lectotype">lectotype</typeStatus>
|
||
is
|
||
<emphasis id="A5E8D7547D62FA29ADD3850FFD2BF471" bold="true" box="[493,659,1336,1357]" pageId="44" pageNumber="45">here designated</emphasis>
|
||
from the only known type material of
|
||
<emphasis id="A5E8D7547D62FA29AE288564FD2CF454" box="[534,660,1363,1384]" italics="true" pageId="44" pageNumber="45">
|
||
<taxonomicName id="509C70C57D62FA29AE288564FD28F454" box="[534,656,1363,1384]" class="Insecta" family="Bothrideridae" genus="Deretaphrus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="parviceps">D. parviceps</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DF8658CD7D62FA29ACFB8559FCFDF0AD" pageId="44" pageNumber="45" type="materials_examined">
|
||
<paragraph id="97230B467D62FA29ACFB8559FD2BF4BF" blockId="44.[173,660,194,1729]" box="[197,659,1390,1411]" pageId="44" pageNumber="45">
|
||
<materialsCitation id="27F4011B7D62FA29ACFB8559FD36F4BF" box="[197,654,1390,1411]" location="Swan River" pageId="44" pageNumber="45" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
|
||
<emphasis id="A5E8D7547D62FA29ACFB8559FEECF4BF" bold="true" box="[197,340,1390,1411]" pageId="44" pageNumber="45">
|
||
<typeStatus id="4827B5E47D62FA29ACFB8559FF4DF4BF" box="[197,245,1390,1411]" pageId="44" pageNumber="45">Type</typeStatus>
|
||
Locality.
|
||
</emphasis>
|
||
<location id="92435D9D7D62FA29AD5C8559FE6DF4BF" LSID="urn:lsid:plazi:treatment:1F35BA507D65FA29ACDA85D0FCFDF0AD:92435D9D7D62FA29AD5C8559FE6DF4BF" box="[354,469,1390,1411]" name="Swan River" pageId="44" pageNumber="45" stateProvince="Western Australia">Swan River</location>
|
||
,
|
||
<collectingRegion id="5558C5A47D62FA29ADDE8559FD36F4BF" box="[480,654,1390,1411]" country="Australia" name="Western Australia" pageId="44" pageNumber="45">Western Australia</collectingRegion>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="97230B467D62FA29ACFB85BFFD2CF7B0" blockId="44.[173,660,194,1729]" pageId="44" pageNumber="45">
|
||
<materialsCitation id="27F4011B7D62FA29ACFB85BFFED0F4EE" collectionCode="SAMA" collectorName="Examined. L E C T O T Y P E" pageId="44" pageNumber="45" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<emphasis id="A5E8D7547D62FA29ACFB85BFFE64F4A1" bold="true" box="[197,476,1416,1437]" pageId="44" pageNumber="45">
|
||
<typeStatus id="4827B5E47D62FA29ACFB85BFFF4EF4A1" box="[197,246,1416,1437]" pageId="44" pageNumber="45">Type</typeStatus>
|
||
Material
|
||
<collectorName id="3A696E907D62FA29AD5485BFFE64F4A1" box="[362,476,1416,1437]" pageId="44" pageNumber="45">Examined.</collectorName>
|
||
</emphasis>
|
||
L E C T O T Y P E (
|
||
<figureCitation id="0FA717C37D62FA29AC8A8594FE87F484" box="[180,319,1442,1464]" captionStart="Figs" captionStartId="102.[197,237,1618,1637]" captionTargetBox="[173,1193,193,1590]" captionTargetId="figure-1@102.[173,1194,193,1623]" captionTargetPageId="102" captionText="Figs. 219–227. Primary type specimens and specimen labels of Deretaphrus species. 219) D. iridescens, lectotype labels; 220–221) D. lateropunctatus, holotype, BMNH; 222–223) D. ocularis, holotype, SAMA; 224–225) D. oregonensis, lectotype, MCZ; 226–227) D. parviceps, lectotype, SAMA." figureDoi="http://doi.org/10.5281/zenodo.4907552" httpUri="https://zenodo.org/record/4907552/files/figure.png" pageId="44" pageNumber="45">Figs. 226–227</figureCitation>
|
||
) (
|
||
<specimenCount id="819AC0CF7D62FA29AD698595FED1F484" box="[343,361,1442,1464]" pageId="44" pageNumber="45" type="male">♂</specimenCount>
|
||
,
|
||
<collectionCode id="F18D93837D62FA29AD468594FE06F484" box="[376,446,1443,1464]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="44" pageNumber="45" type="Museum">SAMA</collectionCode>
|
||
, point-mounted) label data: “
|
||
<taxonomicName id="509C70C57D62FA29ACD6858AFED0F4EE" authority="Lea" authorityName="Lea" box="[232,360,1469,1490]" class="Insecta" family="Bothrideridae" genus="Deretaphrus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="parviceps">parviceps Lea</taxonomicName>
|
||
</materialsCitation>
|
||
<materialsCitation id="27F4011B7D62FA29AD53858AFD2BF4D1" collectorName="Swan R" country="Australia" location="Australia" pageId="44" pageNumber="45" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="4827B5E47D62FA29AD53858AFE10F4EE" box="[365,424,1469,1490]" pageId="44" pageNumber="45">TYPE</typeStatus>
|
||
<collectorName id="3A696E907D62FA29AD95858AFE4CF4EE" box="[427,500,1469,1490]" pageId="44" pageNumber="45">Swan R</collectorName>
|
||
[in Lea’ s hand] // 9560
|
||
<taxonomicName id="509C70C57D62FA29ACD485EFFD93F4D1" authority="Lea W." authorityName="Lea W." box="[234,555,1496,1517]" class="Insecta" family="Bothrideridae" genus="Deretaphrus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="parviceps">Deretaphrus parviceps Lea W.</taxonomicName>
|
||
<collectingCountry id="EF8B4BD67D62FA29AE0A85EFFD2BF4D1" box="[564,659,1496,1517]" name="Australia" pageId="44" pageNumber="45">Australia</collectingCountry>
|
||
</materialsCitation>
|
||
<materialsCitation id="27F4011B7D62FA29AC9385C5FD2CF7B0" collectionCode="NL" collectorName="S. Aust. Museum" country="Australia" pageId="44" pageNumber="45" specimenCode="SAMA 278" specimenCount="2" typeStatus="holotype">
|
||
<typeStatus id="4827B5E47D62FA29AC9385C5FF51F73B" box="[173,233,1522,1543]" pageId="44" pageNumber="45">TYPE</typeStatus>
|
||
[handwritten in black ink, “
|
||
<typeStatus id="4827B5E47D62FA29AE3F85C5FD85F73B" box="[513,573,1522,1543]" pageId="44" pageNumber="45">TYPE</typeStatus>
|
||
” written in red ink on right side of label] //
|
||
<collectorName id="3A696E907D62FA29ADCE863AFD2BF71E" box="[496,659,1549,1570]" pageId="44" pageNumber="45">S. Aust. Museum</collectorName>
|
||
specimen [orange label]” [
|
||
<specimenCode id="C73AA33D7D62FA29AD8D8610FD90F700" box="[435,552,1575,1596]" collectionCode="SAMA" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="44" pageNumber="45" type="Museum">SAMA 278</specimenCode>
|
||
<collectionCode id="F18D93837D62FA29AE168610FDF2F700" box="[552,586,1575,1596]" pageId="44" pageNumber="45">NL</collectionCode>
|
||
, examined]. The abdomen of this specimen was dissected [glued to point-mount] and the genitalia is in glycerin in a genitalia vial pinned beneath the specimen.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="97230B467D62FA29ACFB86A5FCFDF0AD" blockId="44.[173,660,194,1729]" lastBlockId="44.[707,1194,193,401]" pageId="44" pageNumber="45">
|
||
<materialsCitation id="27F4011B7D62FA29ACFB86A5FCF9F0AD" collectionCode="NL" collectorName="R. P. McMillan" location="Western Australian Museum Entomology Reg." pageId="44" pageNumber="45" specimenCode="WAMP 21" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
|
||
<emphasis id="A5E8D7547D62FA29ACFB86A5FDB0F79B" bold="true" box="[197,520,1682,1703]" pageId="44" pageNumber="45">Additional Material Examined.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="4827B5E47D62FA29AD36869BFEE7F7FD" box="[264,351,1708,1729]" pageId="44" pageNumber="45" type="lectotype">lectotype</typeStatus>
|
||
and one other specimen. Label data: “
|
||
<collectorName id="3A696E907D62FA29AF3680F5FC25F1EB" box="[776,925,194,215]" pageId="44" pageNumber="45">R.P. McMillan</collectorName>
|
||
DATE 16.6.51 Culham //
|
||
<location id="92435D9D7D62FA29AEFD80EAFB12F1CE" LSID="urn:lsid:plazi:treatment:1F35BA507D65FA29ACDA85D0FCFDF0AD:92435D9D7D62FA29AEFD80EAFB12F1CE" box="[707,1194,221,242]" name="Western Australian Museum Entomology Reg." pageId="44" pageNumber="45">Western Australian Museum Entomology Reg.</location>
|
||
no. 65202” [
|
||
<specimenCode id="C73AA33D7D62FA29AF7C80C0FC08F030" box="[834,944,247,268]" collectionCode="WAMP" pageId="44" pageNumber="45">WAMP 21</specimenCode>
|
||
<collectionCode id="F18D93837D62FA29AF8E80C0FC6DF030" box="[944,981,247,268]" pageId="44" pageNumber="45">NL</collectionCode>
|
||
]. This specimen was removed from the original card-mount and pointmounted. The original card-mount, abdomen (glued to a card, bearing a
|
||
<specimenCount id="819AC0CF7D62FA29AF858171FC75F060" box="[955,973,326,348]" pageId="44" pageNumber="45" type="female">♀</specimenCount>
|
||
symbol)], and genitalia (in glycerin in genitalia vial) are pinned beneath the specimen
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |