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<document id="B11B4E050FB74665FF9DDA12F94D6150" ID-DOI="10.5281/zenodo.190974" ID-GBIF-Dataset="4a7a3f4f-958b-4396-b234-bca1e1a100ab" ID-ISSN="1175-5326" ID-Zenodo-Dep="190974" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461122289397" checkinUser="plazi" docAuthor="Craig, Matthew T. &amp; Randall, John E." docDate="2009" docId="1967E85CFF814F44DCF3309BFCADFAB4" docLanguage="en" docName="zt02271p068.pdf" docOrigin="Zootaxa 2271" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Briggsia hastingsi Craig &amp; Randall, 2009, new species" docType="treatment" docUuid="29FC6306-D6F8-49CF-AD2D-6FB39255D283" docUuidSource="ZooBank" docVersion="9" lastPageNumber="67" masterDocId="E55E9024FF804F47DC643322FF82FFAB" masterDocTitle="Briggsia hastingsi, a new genus and species of clingfish from Oman" masterLastPageNumber="68" masterPageNumber="64" pageNumber="65" updateTime="1698596131515" updateUser="plazi">
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<mods:title id="DDE9DD811466B137371C12F5B2A29AF2">Briggsia hastingsi, a new genus and species of clingfish from Oman</mods:title>
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<mods:namePart id="7717F567E8F0CCD8E73B8BFD73F8DBA1">Craig, Matthew T.</mods:namePart>
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<mods:namePart id="8E6929F440BE607606B15824C733EA2D">Randall, John E.</mods:namePart>
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<mods:date id="A40F3740F9B77CAB0EB2A754B57F8564">2009</mods:date>
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<treatment id="1967E85CFF814F44DCF3309BFCADFAB4" ID-DOI="http://doi.org/10.5281/zenodo.6216283" ID-GBIF-Taxon="119625898" ID-Zenodo-Dep="6216283" LSID="urn:lsid:zoobank.org:act:29FC6306-D6F8-49CF-AD2D-6FB39255D283" httpUri="http://treatment.plazi.org/id/1967E85CFF814F44DCF3309BFCADFAB4" lastPageId="3" lastPageNumber="67" pageId="1" pageNumber="65">
<subSubSection id="D9D40AC1FF814F46DCF3309BFE0AFC5D" pageId="1" pageNumber="65" type="nomenclature">
<paragraph id="9171594AFF814F46DCF3309BFD92FC79" blockId="1.[151,528,952,1014]" box="[151,528,952,979]" pageId="1" pageNumber="65">
<heading id="CA39EE26FF814F46DCF3309BFD92FC79" bold="true" box="[151,528,952,979]" fontSize="11" level="1" pageId="1" pageNumber="65" reason="1">
<emphasis id="A3BA8558FF814F46DCF3309BFD92FC79" bold="true" box="[151,528,952,979]" pageId="1" pageNumber="65">
<taxonomicName id="56CE22C9FF814F46DCF3309BFEF0FC78" ID-CoL="NB4N" box="[151,370,953,979]" class="Actinopterygii" family="Gobiesocidae" genus="Briggsia" kingdom="Animalia" order="Gobiesociformes" pageId="1" pageNumber="65" phylum="Chordata" rank="species" species="hastingsi" status="sp. nov.">
<emphasis id="A3BA8558FF814F46DCF3309BFEF0FC78" bold="true" box="[151,370,953,979]" italics="true" pageId="1" pageNumber="65">Briggsia hastingsi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="B8893823FF814F46DDE4309AFD92FC79" box="[384,528,952,978]" pageId="1" pageNumber="65" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9171594AFF814F46DCF330FEFE0AFC5D" blockId="1.[151,528,952,1014]" box="[151,392,988,1014]" pageId="1" pageNumber="65">
<figureCitation id="09F545CFFF814F46DCF330FEFEA6FC5D" box="[151,292,988,1014]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,255,796,820]" captionStartId-1="2.[151,255,1528,1552]" captionStartId-2="3.[151,255,1783,1807]" captionTargetBox-0="[472,1115,506,771]" captionTargetBox-2="[296,1284,1392,1764]" captionTargetId-0="figure@2.[472,1115,506,771]" captionTargetId-2="figure@3.[291,1289,1391,1773]" captionTargetPageId-0="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Holotype of Briggsia hastingsi, BPBM 36539, 22 mm SL, Oman (Photo: J. E. Randall)." captionText-1="FIGURE 2. Disc of holotype of Briggsia hastingsi (M. T. Craig)." captionText-2="FIGURE 3. Cephalic sensory pores of holotype of Briggsia hastingsi (M. T Craig, following Briggs, 1993). PO—Post orbital canals, LC—Lachrymal canals, NC—Nasal canals." httpUri-0="https://zenodo.org/record/190975/files/figure.png" httpUri-2="https://zenodo.org/record/190976/files/figure.png" pageId="1" pageNumber="65">Figures 13</figureCitation>
;
<tableCitation id="DC4C6CF1FF814F46DD5530FEFE0AFC5D" box="[305,392,988,1014]" captionStart="TABLE 1" captionStartId="3.[151,239,383,407]" captionText="TABLE 1. Proportional Measurements of the holotype of Briggsia hastingsi as percentages of the standard length. Standard Length (22 mm) -" pageId="1" pageNumber="65">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="D9D40AC1FF814F46DCF3370AFD28FBC1" pageId="1" pageNumber="65" type="materials_examined">
<paragraph id="9171594AFF814F46DCF3370AFD28FBC1" blockId="1.[151,1437,1064,2019]" pageId="1" pageNumber="65">
<emphasis id="A3BA8558FF814F46DCF3370AFE9EFBE9" bold="true" box="[151,284,1064,1090]" pageId="1" pageNumber="65">
<typeStatus id="4E75E7E8FF814F46DCF3370AFE94FBE9" box="[151,278,1064,1090]" pageId="1" pageNumber="65" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="F7DFC18FFF814F46DD40370BFEF8FBE8" LSID="urn:lsid:biocol.org:col:34697" box="[292,378,1065,1091]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="1" pageNumber="65">BPBM</collectionCode>
36539, 22 mm SL,
<collectingCountry id="E9D919DAFF814F46DE1D370BFD4AFBE8" box="[633,712,1065,1091]" name="Oman" pageId="1" pageNumber="65">Oman</collectingCountry>
, southeastern coast, Rahah Bay, 16°57°N,
<geoCoordinate id="F4FA3F8DFF814F46D88D370BFAD7FBE8" box="[1257,1365,1065,1091]" direction="east" orientation="longitude" pageId="1" pageNumber="65" precision="925" value="54.816666">54°49E</geoCoordinate>
,
<quantity id="5636F4AFFF814F46D900370BFA15FBE8" box="[1380,1431,1065,1091]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="65" unit="m" value="2.0">2 m</quantity>
, quinaldine, John P. Hoover,
<date id="E5707F8AFF814F46DD853772FD24FBC1" box="[481,678,1104,1130]" pageId="1" pageNumber="65" value="1993-02-08">8 February 1993</date>
.
</paragraph>
</subSubSection>
<subSubSection id="D9D40AC1FF814F46DCA23754FEE8FAD3" pageId="1" pageNumber="65" type="diagnosis">
<paragraph id="9171594AFF814F46DCA23754FEE8FAD3" blockId="1.[151,1437,1064,2019]" pageId="1" pageNumber="65">
<emphasis id="A3BA8558FF814F46DCA23754FECAFB3B" bold="true" box="[198,328,1142,1168]" pageId="1" pageNumber="65">Diagnosis:</emphasis>
A species of aspasimine gobiesocid with dorsal and anal fins posterior on body, each with 4 rays; pectoral rays 23; caudal fin broad and rounded, with 10 principal rays; gill rakers on first gill arch 9, on third arch 10; head large, its length
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SL; body depth
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SL; disc double and well-separated; disc length
<quantity id="5636F4AFFF814F46DC8C37C8FEAEFAAF" box="[232,300,1258,1284]" metricMagnitude="-2" metricUnit="m" metricValue="9.906" pageId="1" pageNumber="65" unit="in" value="3.9">3.9 in</quantity>
SL; disc width
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SL; color in alcohol uniform tan; color when fresh dark reddish gray with numerous blue dots on side of body except posteriorly, and a few on postorbital head; a dark purplish brown stripe from front of snout, continuing posterior to eye; head ventral to stripe yellowish white; a distinct orange ring around pupil.
</paragraph>
</subSubSection>
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<emphasis id="A3BA8558FF814F46DCA236A7FEE3FA34" bold="true" box="[198,353,1413,1439]" pageId="1" pageNumber="65">Description:</emphasis>
Dorsal rays 4; anal rays 4; pectoral rays 23; principal caudal rays 10; upper procurrent caudal rays 6; lower procurrent caudal rays 5; gill rakers on second gill arch 9; gill rakers on third gill arch 10; vertebrae 29.
</paragraph>
<paragraph id="9171594AFF814F46DCA136DBFEABF9CB" blockId="1.[151,1437,1064,2019]" pageId="1" pageNumber="65">
Body depth
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SL; body rounded anteriorly, progressively more compressed posteriorly; head length
<quantity id="5636F4AFFF814F46DCF33502FF58F991" box="[151,218,1568,1594]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" pageId="1" pageNumber="65" unit="in" value="2.5">2.5 in</quantity>
SL; head width
<quantity id="5636F4AFFF814F46DDFA3502FE60F991" box="[414,482,1568,1594]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" pageId="1" pageNumber="65" unit="in" value="2.5">2.5 in</quantity>
SL; snout length
<quantity id="5636F4AFFF814F46DED03502FD7AF991" box="[692,760,1568,1594]" metricMagnitude="-2" metricUnit="m" metricValue="6.604" pageId="1" pageNumber="65" unit="in" value="2.6">2.6 in</quantity>
HL; eye diameter
<quantity id="5636F4AFFF814F46DFB13502FBAAF991" box="[981,1064,1568,1594]" metricMagnitude="-1" metricUnit="m" metricValue="1.6128999999999998" pageId="1" pageNumber="65" unit="in" value="6.35">6.35 in</quantity>
HL; caudal-peduncle depth
<quantity id="5636F4AFFF814F46D9133502FF2CF9CB" metricMagnitude="-2" metricUnit="m" metricValue="2.794" pageId="1" pageNumber="65" unit="in" value="1.1">1.1 in</quantity>
its length.
</paragraph>
<paragraph id="9171594AFF814F46DCA1354FFDC3F889" blockId="1.[151,1437,1064,2019]" pageId="1" pageNumber="65">
Disc double, well-separated, and of moderate size; disc length
<quantity id="5636F4AFFF814F46DFC0354FFC65F92C" box="[932,999,1645,1671]" metricMagnitude="-2" metricUnit="m" metricValue="9.906" pageId="1" pageNumber="65" unit="in" value="3.9">3.9 in</quantity>
SL; disc width
<quantity id="5636F4AFFF814F46D8C5354FFB66F92C" box="[1185,1252,1645,1671]" metricMagnitude="-1" metricUnit="m" metricValue="1.1176" pageId="1" pageNumber="65" unit="in" value="4.4">4.4 in</quantity>
SL; anus about half way between origin of anal fin and posterior edge of disc. Disc region “A” (
<emphasis id="A3BA8558FF814F46D80935B6FB32F906" box="[1133,1200,1684,1709]" italics="true" pageId="1" pageNumber="65">sensu</emphasis>
<bibRefCitation id="F55F24BBFF814F46D8DC35B6FAD7F905" author="Briggs" box="[1208,1365,1684,1710]" pageId="1" pageNumber="68" refString="Briggs, J. C. (1955) A monograph of the clingfishes (Order Xenopterygii). Stanford Ichthyological Bulletin 6, iv + 224 pp." type="book" year="1955">Briggs, 1955</bibRefCitation>
) with several rows of flattened papillae; disc region “B” with three rows of flattened papillae; disc region “C” with four rows of flattened papillae; a small, oval region slightly posterior to disc region “A” with two circular pads formed by flattened papillae (
<figureCitation id="09F545CFFF814F46DD96342AFDBBF889" box="[498,569,1800,1826]" captionStart="FIGURE 2" captionStartId="2.[151,255,1528,1552]" captionText="FIGURE 2. Disc of holotype of Briggsia hastingsi (M. T. Craig)." pageId="1" pageNumber="65">Fig. 2</figureCitation>
)
</paragraph>
<paragraph id="9171594AFF814F46DCA1340CFA17F8C4" blockId="1.[151,1437,1064,2019]" pageId="1" pageNumber="65">
Mouth slightly inferior, oblique, and moderately large, the upper-jaw length
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SL; front of jaws with a sandpaper-like pad of small teeth, followed by two or three well-developed canines and two rows of incisors.
</paragraph>
<paragraph id="9171594AFF814F46DCA1345EFD4EF817" blockId="1.[151,1437,1064,2019]" pageId="1" pageNumber="65">
Nostrils well separated and distinctly tubular; five pairs of cephalic sensory pores; nasal sensory canals 2, postorbital canals 1, lachrymal canals 2 (
<figureCitation id="09F545CFFF814F46DE113480FD3EF817" box="[629,700,1954,1980]" captionStart="FIGURE 3" captionStartId="3.[151,255,1783,1807]" captionTargetBox="[296,1284,1392,1764]" captionTargetId="figure@3.[291,1289,1391,1773]" captionTargetPageId="3" captionText="FIGURE 3. Cephalic sensory pores of holotype of Briggsia hastingsi (M. T Craig, following Briggs, 1993). PO—Post orbital canals, LC—Lachrymal canals, NC—Nasal canals." httpUri="https://zenodo.org/record/190976/files/figure.png" pageId="1" pageNumber="65">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="9171594AFF814F45DCA134EBFCA6FF19" blockId="1.[151,1437,1064,2019]" lastBlockId="2.[151,1436,152,448]" lastPageId="2" lastPageNumber="66" pageId="1" pageNumber="65">
Origins of dorsal and anal fins in vertical alignment; postdorsal-caudal distance
<quantity id="5636F4AFFF814F46D81A34EBFB40F848" box="[1150,1218,1993,2019]" metricMagnitude="-1" metricUnit="m" metricValue="2.1082" pageId="1" pageNumber="65" unit="in" value="8.3">8.3 in</quantity>
SL; dorsal-caudal distance
<quantity id="5636F4AFFF824F45DC9A33BAFEC3FF19" box="[254,321,152,178]" metricMagnitude="-1" metricUnit="m" metricValue="1.3462" pageId="2" pageNumber="66" unit="in" value="5.3">5.3 in</quantity>
SL; no fleshy pad at base of pectoral fin.
</paragraph>
<paragraph id="9171594AFF824F45DCA9339CFE1DFE6B" blockId="2.[151,1436,152,448]" pageId="2" pageNumber="66">
Color of
<typeStatus id="4E75E7E8FF824F45DD5D339CFE23FF73" box="[313,417,190,216]" pageId="2" pageNumber="66" type="holotype">holotype</typeStatus>
in alcohol uniform tan. Color when fresh: body and dorsal part of head reddish gray; numerous blue dots on side of body except posteriorly; a few blue dots on postorbital head; a dark purplish brown stripe from upper lip to eye, continuing obliquely downward a short distance behind eye; head below stripe abruptly yellowish white; a short blue line at ventral edge of dark stripe posterior to eye; eye colored like adjacent head except for a bright orange ring around pupil; dorsal and anal fins translucent with vague pale spots; caudal fin with reddish rays and clear membranes, the upper and lower rays with a row of pale spots; paired fins pale.
</paragraph>
</subSubSection>
<caption id="C5B109C2FF824F45DCF3303EFB46FC98" box="[151,1220,796,820]" httpUri="https://zenodo.org/record/190975/files/figure.png" pageId="2" pageNumber="66" targetBox="[472,1115,506,771]" targetPageId="2">
<paragraph id="9171594AFF824F45DCF3303EFB46FC98" blockId="2.[151,1220,796,820]" box="[151,1220,796,820]" pageId="2" pageNumber="66">
<emphasis id="A3BA8558FF824F45DCF3303EFE9BFC9F" bold="true" box="[151,281,796,820]" pageId="2" pageNumber="66">FIGURE 1.</emphasis>
Holotype of
<taxonomicName id="56CE22C9FF824F45DDC2303FFDEBFC9F" box="[422,617,797,820]" class="Actinopterygii" family="Gobiesocidae" genus="Briggsia" kingdom="Animalia" order="Gobiesociformes" pageId="2" pageNumber="66" phylum="Chordata" rank="species" species="hastingsi">
<emphasis id="A3BA8558FF824F45DDC2303FFDEBFC9F" box="[422,617,797,820]" italics="true" pageId="2" pageNumber="66">Briggsia hastingsi</emphasis>
</taxonomicName>
, BPBM 36539, 22 mm SL, Oman (Photo: J.E. Randall).
</paragraph>
</caption>
<caption id="C5B109C2FF824F45DCF336DAFCCFF9A4" box="[151,845,1528,1552]" pageId="2" pageNumber="66">
<paragraph id="9171594AFF824F45DCF336DAFCCFF9A4" blockId="2.[151,845,1528,1552]" box="[151,845,1528,1552]" pageId="2" pageNumber="66">
<emphasis id="A3BA8558FF824F45DCF336DAFE9BF9BB" bold="true" box="[151,281,1528,1552]" pageId="2" pageNumber="66">FIGURE 2.</emphasis>
Disc of holotype of
<taxonomicName id="56CE22C9FF824F45DD9136DBFCCBF9A4" authority="M.T. Craig" authorityName="M.T. Craig" box="[501,841,1528,1552]" class="Actinopterygii" family="Gobiesocidae" genus="Briggsia" kingdom="Animalia" order="Gobiesociformes" pageId="2" pageNumber="66" phylum="Chordata" rank="species" species="hastingsi">
<emphasis id="A3BA8558FF824F45DD9136DBFD35F9BB" box="[501,695,1529,1552]" italics="true" pageId="2" pageNumber="66">Briggsia hastingsi</emphasis>
(M.T. Craig)
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="D9D40AC1FF824F45DCA1351CFBA9F9D5" pageId="2" pageNumber="66" type="etymology">
<paragraph id="9171594AFF824F45DCA1351CFBA9F9D5" blockId="2.[151,1437,1598,2011]" pageId="2" pageNumber="66">
<emphasis id="A3BA8558FF824F45DCA1351CFED4F9F3" bold="true" box="[197,342,1598,1624]" pageId="2" pageNumber="66">Etymology:</emphasis>
This species name
<taxonomicName id="56CE22C9FF824F45DE5D351CFD21F9FC" box="[569,675,1598,1623]" class="Actinopterygii" family="Gobiesocidae" genus="Briggsia" kingdom="Animalia" order="Gobiesociformes" pageId="2" pageNumber="66" phylum="Chordata" rank="species" species="hastingsi">
<emphasis id="A3BA8558FF824F45DE5D351CFD21F9FC" box="[569,675,1598,1623]" italics="true" pageId="2" pageNumber="66">hastingsi</emphasis>
</taxonomicName>
is named in honor of Dr. Philip A. Hastings, the thesis advisor of the first author, who initially aroused the first authors curiosity in clingfishes.
</paragraph>
</subSubSection>
<subSubSection id="D9D40AC1FF824F44DCA135A9FCADFAB4" lastPageId="3" lastPageNumber="67" pageId="2" pageNumber="66" type="discussion">
<paragraph id="9171594AFF824F45DCA135A9FD2DF8B2" blockId="2.[151,1437,1598,2011]" pageId="2" pageNumber="66">
<emphasis id="A3BA8558FF824F45DCA135A9FEC1F90E" bold="true" box="[197,323,1675,1701]" pageId="2" pageNumber="66">Remarks:</emphasis>
<bibRefCitation id="F55F24BBFF824F45DD2835A9FD81F90E" author="Randall" box="[332,515,1675,1701]" pageId="2" pageNumber="68" refString="Randall, J. E. (1996). Coastal Fishes of Oman. University of Hawaii Press, Honolulu. xiii + 432 pp." type="book" year="1996">Randall (1996)</bibRefCitation>
published a book entitled
<emphasis id="A3BA8558FF824F45DF2235A9FBEEF90F" box="[838,1132,1675,1700]" italics="true" pageId="2" pageNumber="66">
Coastal Fishes of
<collectingCountry id="E9D919DAFF824F45D84735A9FBEEF90F" box="[1059,1132,1675,1700]" name="Oman" pageId="2" pageNumber="66">Oman</collectingCountry>
</emphasis>
. Fish collections for this volume include a single specimen of a clingfish taken in 1993 that he was unable to identify to genus or species. We are aware of no additional material of this species, which is so distinctive that we no longer hesitate to describe it from a single specimen.
</paragraph>
<paragraph id="9171594AFF824F44DCA13404FBE8FEE7" blockId="2.[151,1437,1598,2011]" lastBlockId="3.[151,1437,152,332]" lastPageId="3" lastPageNumber="67" pageId="2" pageNumber="66">
<bibRefCitation id="F55F24BBFF824F45DCA13404FE75F8EB" author="Shiogaki" box="[197,503,1830,1856]" pageId="2" pageNumber="68" refString="Shiogaki, M. &amp; Y. Dotsu (1983) Two new genera and two new species of clingfishes from Japan with comments on head sensory canals of the Gobiesocidae. Japanese Journal of Ichthyology, 30, 111 - 121" type="journal article" year="1983">Shiogaki &amp; Dotsu (1983)</bibRefCitation>
reported on the head sensory canal pores of the Gobieosocidae and proposed that the orientation and number of these pores may be useful to differentiate clingfish genera. The pattern observed in
<taxonomicName id="56CE22C9FF824F45DD4A3451FE42F827" box="[302,448,1907,1932]" class="Actinopterygii" family="Gobiesocidae" genus="Briggsia" kingdom="Animalia" order="Gobiesociformes" pageId="2" pageNumber="66" phylum="Chordata" rank="species" species="hastingsi">
<emphasis id="A3BA8558FF824F45DD4A3451FE42F827" box="[302,448,1907,1932]" italics="true" pageId="2" pageNumber="66">B. hastingsi</emphasis>
</taxonomicName>
(nasal canals 2, postorbital canals 1, lachrymal canals 2) would indicate a close relationship with species of the genera
<taxonomicName id="56CE22C9FF824F45DE0234B8FD74F818" box="[614,758,1946,1971]" class="Actinopterygii" family="Gobiesocidae" genus="Pherallodus" kingdom="Animalia" order="Gobiesociformes" pageId="2" pageNumber="66" phylum="Chordata" rank="genus">
<emphasis id="A3BA8558FF824F45DE0234B8FD74F818" box="[614,758,1946,1971]" italics="true" pageId="2" pageNumber="66">Pherallodus</emphasis>
</taxonomicName>
and
<taxonomicName id="56CE22C9FF824F45DF5534B8FC38F818" box="[817,954,1946,1971]" class="Actinopterygii" family="Gobiesocidae" genus="Parvicrepis" kingdom="Animalia" order="Gobiesociformes" pageId="2" pageNumber="66" phylum="Chordata" rank="genus">
<emphasis id="A3BA8558FF824F45DF5534B8FC38F818" box="[817,954,1946,1971]" italics="true" pageId="2" pageNumber="66">Parvicrepis</emphasis>
</taxonomicName>
of the subfamily Diplocrepinae. Species of the Diplocrepinae, however, have gill membranes that are free from the isthmus, thus
<taxonomicName id="56CE22C9FF824F45D8D434E3FABEF871" box="[1200,1340,1985,2010]" class="Actinopterygii" family="Gobiesocidae" genus="Briggsia" kingdom="Animalia" order="Gobiesociformes" pageId="2" pageNumber="66" phylum="Chordata" rank="species" species="hastingsi">
<emphasis id="A3BA8558FF824F45D8D434E3FABEF871" box="[1200,1340,1985,2010]" italics="true" pageId="2" pageNumber="66">B. hastingsi</emphasis>
</taxonomicName>
with its attached gill membranes is placed in the Aspaminae, following
<bibRefCitation id="F55F24BBFF834F44DFE533BAFBAAFF19" author="Briggs" box="[897,1064,152,178]" pageId="3" pageNumber="68" refString="Briggs, J. C. (1955) A monograph of the clingfishes (Order Xenopterygii). Stanford Ichthyological Bulletin 6, iv + 224 pp." type="book" year="1955">Briggs (1955)</bibRefCitation>
.
<bibRefCitation id="F55F24BBFF834F44D85033BAFB5AFF19" author="Briggs" box="[1076,1240,152,178]" pageId="3" pageNumber="68" refString="Briggs, J. C. (1993) New genus and species of clingfish (Gobiesocidae) from southern Australia. Copeia, 1993 (1), 196 - 199." type="journal article" year="1993">Briggs (1993)</bibRefCitation>
described a new genus and species,
<taxonomicName id="56CE22C9FF834F44DDE8339DFD48FF73" box="[396,714,191,216]" class="Actinopterygii" family="Gobiesocidae" genus="Posidonichthys" kingdom="Animalia" order="Gobiesociformes" pageId="3" pageNumber="67" phylum="Chordata" rank="species" species="hutchinsi">
<emphasis id="A3BA8558FF834F44DDE8339DFD48FF73" box="[396,714,191,216]" italics="true" pageId="3" pageNumber="67">Posidonichthys hutchinsi</emphasis>
</taxonomicName>
, which also possesses three gills, a double disc, and gill membranes that are attached to the isthmus, indicating its placement within the Aspasminae. However, Briggs noted several characters in the general osteology of
<taxonomicName id="56CE22C9FF834F44DF47322EFC37FE8E" box="[803,949,268,293]" class="Actinopterygii" family="Gobiesocidae" genus="Posidonichthys" kingdom="Animalia" order="Gobiesociformes" pageId="3" pageNumber="67" phylum="Chordata" rank="species" species="hutchinsi">
<emphasis id="A3BA8558FF834F44DF47322EFC37FE8E" box="[803,949,268,293]" italics="true" pageId="3" pageNumber="67">P. hutchinsi</emphasis>
</taxonomicName>
that suggest a close relationship to the Diplocrepinae. The relationship of these two subfamilies is in need of further study.
</paragraph>
<caption id="C5B109C2FF834F44DCF3325DFC95FE62" pageId="3" pageNumber="67">
<paragraph id="9171594AFF834F44DCF3325DFC95FE62" blockId="3.[151,1373,383,1311]" pageId="3" pageNumber="67">
<emphasis id="A3BA8558FF834F44DCF3325DFE8AFE3C" bold="true" box="[151,264,383,407]" pageId="3" pageNumber="67">TABLE 1.</emphasis>
Proportional Measurements of the holotype of
<taxonomicName id="56CE22C9FF834F44DF6032A2FC44FE3C" box="[772,966,384,407]" class="Actinopterygii" family="Gobiesocidae" genus="Briggsia" kingdom="Animalia" order="Gobiesociformes" pageId="3" pageNumber="67" phylum="Chordata" rank="species" species="hastingsi">
<emphasis id="A3BA8558FF834F44DF6032A2FC44FE3C" box="[772,966,384,407]" italics="true" pageId="3" pageNumber="67">Briggsia hastingsi</emphasis>
</taxonomicName>
as percentages of the standard length. Standard Length (22 mm) -
</paragraph>
</caption>
<paragraph id="9171594AFF834F44DCFB32FFFCADFE5F" blockId="3.[151,1373,383,1311]" box="[159,815,477,500]" pageId="3" pageNumber="67">Body Depth 6.1</paragraph>
<paragraph id="9171594AFF834F44DCFB312AFCADFDB4" blockId="3.[151,1373,383,1311]" box="[159,815,520,543]" pageId="3" pageNumber="67">Body Width 4.6</paragraph>
<paragraph id="9171594AFF834F44DCFB3110FCADFDE2" blockId="3.[151,1373,383,1311]" box="[159,815,562,585]" pageId="3" pageNumber="67">Head Length 2.5</paragraph>
<paragraph id="9171594AFF834F44DCFB317FFCADFDDF" blockId="3.[151,1373,383,1311]" box="[159,815,605,628]" pageId="3" pageNumber="67">Head Width 3.9</paragraph>
<paragraph id="9171594AFF834F44DCFB31AAFCADFD34" blockId="3.[151,1373,383,1311]" box="[159,815,648,671]" pageId="3" pageNumber="67">Snout Length 6.5</paragraph>
<paragraph id="9171594AFF834F44DCFB3190FCBEFD62" blockId="3.[151,1373,383,1311]" box="[159,828,690,713]" pageId="3" pageNumber="67">Orbit Diameter 16.0</paragraph>
<paragraph id="9171594AFF834F44DCFB31FFFCBEFD5F" blockId="3.[151,1373,383,1311]" box="[159,828,733,756]" pageId="3" pageNumber="67">Interorbital Width 13.9</paragraph>
<paragraph id="9171594AFF834F44DCFB302AFCADFCB4" blockId="3.[151,1373,383,1311]" box="[159,815,776,799]" pageId="3" pageNumber="67">Upper-jaw Length 6.3</paragraph>
<paragraph id="9171594AFF834F44DCFB3010FCBEFCE2" blockId="3.[151,1373,383,1311]" box="[159,828,818,841]" pageId="3" pageNumber="67">Caudal-peduncle Depth 16.0</paragraph>
<paragraph id="9171594AFF834F44DCFB307FFCBEFCDF" blockId="3.[151,1373,383,1311]" box="[159,828,861,884]" pageId="3" pageNumber="67">Caudal-peduncle Length 13.9</paragraph>
<paragraph id="9171594AFF834F44DCFB30AAFCADFC34" blockId="3.[151,1373,383,1311]" box="[159,815,904,927]" pageId="3" pageNumber="67">Predorsal Length 1.2</paragraph>
<paragraph id="9171594AFF834F44DCFB3090FCADFC62" blockId="3.[151,1373,383,1311]" box="[159,815,946,969]" pageId="3" pageNumber="67">Preanal Length 1.3</paragraph>
<paragraph id="9171594AFF834F44DCFB30FFFCBEFC5F" blockId="3.[151,1373,383,1311]" box="[159,828,989,1012]" pageId="3" pageNumber="67">Longest Dorsal-fin Ray 18.9</paragraph>
<paragraph id="9171594AFF834F44DCFB372AFCBEFBB4" blockId="3.[151,1373,383,1311]" box="[159,828,1032,1055]" pageId="3" pageNumber="67">Longest Anal-fin Ray 20.8</paragraph>
<paragraph id="9171594AFF834F44DCFB3710FCADFBE2" blockId="3.[151,1373,383,1311]" box="[159,815,1074,1097]" pageId="3" pageNumber="67">Caudal-fin Length 7.7</paragraph>
<paragraph id="9171594AFF834F44DCFB377FFCBEFBDF" blockId="3.[151,1373,383,1311]" box="[159,828,1117,1140]" pageId="3" pageNumber="67">Pectoral-fin Length 10.4</paragraph>
<paragraph id="9171594AFF834F44DCFB37AAFCADFB34" blockId="3.[151,1373,383,1311]" box="[159,815,1160,1183]" pageId="3" pageNumber="67">Dorsal-caudal Distance 5.3</paragraph>
<paragraph id="9171594AFF834F44DCFB3790FCADFB62" blockId="3.[151,1373,383,1311]" box="[159,815,1202,1225]" pageId="3" pageNumber="67">Postdorsal-caudal Distance 8.3</paragraph>
<paragraph id="9171594AFF834F44DCFB37FFFCADFB5F" blockId="3.[151,1373,383,1311]" box="[159,815,1245,1268]" pageId="3" pageNumber="67">Disc Length 3.9</paragraph>
<paragraph id="9171594AFF834F44DCFB362AFCADFAB4" blockId="3.[151,1373,383,1311]" box="[159,815,1288,1311]" pageId="3" pageNumber="67">Disc Width 4.4</paragraph>
</subSubSection>
</treatment>
</document>