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<document id="CD024AF468FB8F61AF1776CD0051910E" ID-CLB-Dataset="24720" ID-DOI="10.5852/ejt.2019.583" ID-GBIF-Dataset="104a1ea7-b7d6-45cd-9402-bc84ab16b143" ID-Zenodo-Dep="3564532" ID-ZooBank="4F834ADD-57E7-4CE3-B16A-127F1BE4AE14" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1575557973180" checkinUser="plazi" docAuthor="Yoshitomi, Hiroyuki" docDate="2019" docId="03F9EF199F51FFCAFDB4FEEB5A162CAC" docLanguage="en" docName="ejt-583_yoshitomi.pdf.imf" docOrigin="European Journal of Taxonomy 583" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Acontosceles borneensis Yoshitomi 2019, sp. nov." docType="treatment" docUuid="2582F411-8790-4937-B6A0-248899FD9208" docUuidSource="ZooBank" docVersion="13" lastPageNumber="9" masterDocId="FFC097619F56FFC2FFE2FFE05B3C2E73" masterDocTitle="Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera" masterLastPageNumber="45" masterPageNumber="1" pageNumber="8" updateTime="1698772818425" updateUser="ExternalLinkService">
<mods:mods id="DD5FA5E55CB1F3D832041E6624163EAD" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="E81ADA1C3090CE9790E9CC7D5BF2483C">Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera</mods:title>
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<mods:name id="125B24D0518435FFE301021D0A648FE5" type="personal">
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<mods:namePart id="502DEE67557B03A5C646228CEF72E5D8">Yoshitomi, Hiroyuki</mods:namePart>
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<mods:title id="F3A37583AFB9F70C5CC69B74B40BDB25">European Journal of Taxonomy</mods:title>
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<mods:part id="826CF48F45045F37E8F6C907D9D52CD1">
<mods:date id="EA621FE77FAC831F8AC441BFB6ECA9D6">2019</mods:date>
<mods:detail id="8E58A942BD10BB5C158E39F00FE28B99" type="pubDate">
<mods:number id="D42F17D5C2E13B1C673B72DDDB465C3F">2019-09-04</mods:number>
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<mods:number id="791C1EFEFC78BC251B6257A45A45D5F6">583</mods:number>
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<treatment id="03F9EF199F51FFCAFDB4FEEB5A162CAC" ID-DOI="http://doi.org/10.5281/zenodo.5386691" ID-GBIF-Taxon="160630991" ID-Zenodo-Dep="5386691" LSID="urn:lsid:zoobank.org:act:2582F411-8790-4937-B6A0-248899FD9208" httpUri="http://treatment.plazi.org/id/03F9EF199F51FFCAFDB4FEEB5A162CAC" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<subSubSection id="C34A0D849F51FFC5FDB4FEEB5F892F39" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8BEF5E0F9F51FFC5FDB4FEEB58E22F55" blockId="7.[383,1205,267,366]" box="[598,990,267,294]" pageId="7" pageNumber="8">
<heading id="D0A7E9639F51FFC5FDB4FEEB58E22F55" box="[598,990,267,294]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="8" reason="2">
<taxonomicName id="4C50258C9F51FFC5FDB4FEEB58402F55" authority="Yoshitomi, 2019" authorityName="Yoshitomi" authorityYear="2019" box="[598,892,267,294]" class="Insecta" family="Limnichidae" genus="Acontosceles" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="borneensis" status="sp. nov.">
<emphasis id="B924821D9F51FFC5FDB4FEEB58402F55" bold="true" box="[598,892,267,294]" italics="true" pageId="7" pageNumber="8">Acontosceles borneensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2173F669F51FFC5FC61FEEC58E22F55" box="[899,990,268,294]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BEF5E0F9F51FFC5FE9DFECF5F892F39" blockId="7.[383,1205,267,366]" box="[383,1205,303,330]" pageId="7" pageNumber="8">
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urn:lsid:zoobank.org:act:
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<date id="FFEE78CF9F51FFC5FD79FECF59E52F39" box="[667,729,303,330]" pageId="7" pageNumber="8" value="2582">2582</date>
F411-8790-4937-B6A0-248899FD9208
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C34A0D849F51FFC5FD3EFEB358642F1E" box="[732,856,339,366]" pageId="7" pageNumber="8" type="description">
<paragraph id="8BEF5E0F9F51FFC5FD3EFEB358642F1E" blockId="7.[383,1205,267,366]" box="[732,856,339,366]" pageId="7" pageNumber="8">
<figureCitation id="136B428A9F51FFC5FD3EFEB358002F1D" box="[732,828,339,366]" captionStart="Fig" captionStartId="8.[189,232,1930,1956]" captionTargetBox="[189,1384,804,1875]" captionTargetId="figure@8.[189,1399,797,1888]" captionTargetPageId="8" captionText="Fig. 5. Acontosceles spp. A. A. borneensis sp. nov., holotype (BPBM). B. A. yorioi Satô, 1966 (EUMJ). C. A. hydroporoides Champion, 1924 (NME). D. A. chujoi Yoshitomi &amp; Satô, 2005 (EUMJ). E. A. siwalikensis Pütz, 2008 (NME). F. A. zetteli Pütz, 2008 (BPBM). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3564542" httpUri="https://zenodo.org/record/3564542/files/figure.png" pageId="7" pageNumber="8">Figs 5A</figureCitation>
,
<figureCitation id="136B428A9F51FFC5FCABFEB358642F1E" box="[841,856,339,365]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[196,1392,354,1877]" captionTargetId="figure@9.[189,1398,348,1883]" captionTargetPageId="9" captionText="Fig. 6. Acontosceles borneensis sp. nov., male (AE) and female (FH) genitalia (holotype and paratype, BPBM). A, F. Sternite VII. B. Sternite VIII. C. Sternite IX. D. Aedeagus, left lateral view. E. Aedeagus, ventral view. G. Urosternite. H. Ovipositor." figureDoi="http://doi.org/10.5281/zenodo.3564544" httpUri="https://zenodo.org/record/3564544/files/figure.png" pageId="7" pageNumber="8">6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34A0D849F51FFC5FF5FFE7659812FAC" box="[189,701,406,479]" pageId="7" pageNumber="8" type="etymology">
<paragraph id="8BEF5E0F9F51FFC5FF5FFE765A782FC3" blockId="7.[189,324,406,432]" box="[189,324,406,432]" pageId="7" pageNumber="8">
<heading id="D0A7E9639F51FFC5FF5FFE765A782FC3" bold="true" box="[189,324,406,432]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B924821D9F51FFC5FF5FFE765A782FC3" bold="true" box="[189,324,406,432]" pageId="7" pageNumber="8">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF5E0F9F51FFC5FF5FFE2559812FAC" blockId="7.[189,701,453,479]" box="[189,701,453,479]" pageId="7" pageNumber="8">
This species is named after its
<typeStatus id="54EBE0AD9F51FFC5FDC7FE2559642FAC" box="[549,600,453,479]" pageId="7" pageNumber="8">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C34A0D849F51FFC5FF5FFDE759CC2D18" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="8BEF5E0F9F51FFC5FF5FFDE75A962C52" blockId="7.[189,426,519,545]" box="[189,426,519,545]" pageId="7" pageNumber="8">
<heading id="D0A7E9639F51FFC5FF5FFDE75A962C52" bold="true" box="[189,426,519,545]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B924821D9F51FFC5FF5FFDE75A962C52" bold="true" box="[189,426,519,545]" pageId="7" pageNumber="8">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF5E0F9F51FFC5FF08FDD65A602C23" blockId="7.[189,1398,566,665]" box="[234,348,566,592]" pageId="7" pageNumber="8">
<typeStatus id="54EBE0AD9F51FFC5FF08FDD65A602C23" box="[234,348,566,592]" pageId="7" pageNumber="8" type="holotype">
<emphasis id="B924821D9F51FFC5FF08FDD65A602C23" bold="true" box="[234,348,566,592]" pageId="7" pageNumber="8">Holotype</emphasis>
</typeStatus>
</paragraph>
<paragraph id="8BEF5E0F9F51FFC5FF5FFDBA5A2A2CEA" blockId="7.[189,1398,566,665]" pageId="7" pageNumber="8">
<materialsCitation id="3B3854529F51FFC5FF5FFDBA5A2A2CEA" ID-GBIF-Occurrence="2464648267" collectionCode="BPBM" collectorName="T. C. Maa Collector" country="Malaysia" location="MALAYSIA" pageId="7" pageNumber="8" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="F3471E9F9F51FFC5FF5FFDBA5A6F2C07" box="[189,339,602,628]" name="Malaysia" pageId="7" pageNumber="8">MALAYSIA</collectingCountry>
<specimenCount id="9D5695869F51FFC5FE94FDBB5AB32C06" box="[374,399,603,629]" pageId="7" pageNumber="8" type="male"></specimenCount>
; “SARAWAK: Nanga Pelagus
<date id="FFEE78CF9F51FFC5FCF3FDBA58A82C06" box="[785,916,602,629]" pageId="7" pageNumber="8">7-14. VIII.</date>
58”; “
<collectorName id="26A53BD99F51FFC5FC11FDBA5FD02C07" box="[1011,1260,602,629]" pageId="7" pageNumber="8">T. C. Maa Collector</collectorName>
BISHOP”;
<collectionCode id="ED41C6CA9F51FFC5FF5FFD9F5A2E2CEA" box="[189,274,639,665]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="7" pageNumber="8" type="Museum">BPBM</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BEF5E0F9F51FFC5FF08FD215A5B2CA8" blockId="7.[189,1399,705,875]" box="[234,359,705,731]" pageId="7" pageNumber="8">
<typeStatus id="54EBE0AD9F51FFC5FF08FD215A5B2CA8" box="[234,359,705,731]" pageId="7" pageNumber="8" type="paratype">
<emphasis id="B924821D9F51FFC5FF08FD215A5B2CA8" bold="true" box="[234,359,705,731]" pageId="7" pageNumber="8">Paratypes</emphasis>
</typeStatus>
</paragraph>
<paragraph id="8BEF5E0F9F51FFC5FF5FFD0459CC2D18" blockId="7.[189,1399,705,875]" pageId="7" pageNumber="8">
<collectingCountry id="F3471E9F9F51FFC5FF5FFD045A6F2C8D" box="[189,339,740,766]" name="Malaysia" pageId="7" pageNumber="8">MALAYSIA</collectingCountry>
<specimenCount id="9D5695869F51FFC5FE88FD045AAB2C8C" box="[362,407,740,767]" pageId="7" pageNumber="8" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D5695869F51FFC5FE40FD045AD82C8C" box="[418,484,740,767]" pageId="7" pageNumber="8" type="female">2 ♀♀</specimenCount>
; same collection data as for
<typeStatus id="54EBE0AD9F51FFC5FCCEFD0458AA2C8D" box="[812,918,740,766]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
;
<collectionCode id="ED41C6CA9F51FFC5FC43FD0558CF2C8C" box="[929,1011,741,767]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="7" pageNumber="8" type="Museum">BPBM</collectionCode>
<materialsCitation id="3B3854529F51FFC5FBEBFD045AB02D50" ID-GBIF-Occurrence="2464648307" collectionCode="EUMJ" pageId="7" pageNumber="8" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D5695869F51FFC5FBEBFD045F092C8C" box="[1033,1077,740,767]" pageId="7" pageNumber="8" type="female">1 ♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED41C6CA9F51FFC5FEDDFCE95AB02D50" box="[319,396,777,803]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="7" pageNumber="8">EUMJ</collectionCode>
</materialsCitation>
<materialsCitation id="3B3854529F51FFC5FE41FCE859952D34" ID-GBIF-Occurrence="2464648302" collectingDate="1958-07-01" collectingDateMax="1958-07-06" collectingDateMin="1958-07-01" collectionCode="BPBM" collectorName="T. C. Maa Collector" country="Malaysia" elevation="165" location="Merirai Valley" municipality="Kapit" pageId="7" pageNumber="8" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="9D5695869F51FFC5FE41FCE85AD82D50" box="[419,484,776,803]" pageId="7" pageNumber="8" type="female">2 ♀♀</specimenCount>
; “BORNEO:
<collectingCountry id="F3471E9F9F51FFC5FD64FCE858292D51" box="[646,789,776,802]" name="Malaysia" pageId="7" pageNumber="8">SARAWAK</collectingCountry>
<location id="8E8F08D49F51FFC5FCF9FCE858F82D51" LSID="urn:lsid:plazi:treatment:03F9EF199F51FFCAFDB4FEEB5A162CAC:8E8F08D49F51FFC5FCF9FCE858F82D51" box="[795,964,776,802]" country="Malaysia" municipality="Kapit" name="Merirai Valley" pageId="7" pageNumber="8">Merirai Valley</location>
nr
<collectingMunicipality id="6B8BC4759F51FFC5FC0BFCE85F112D51" box="[1001,1069,776,802]" pageId="7" pageNumber="8">Kapit</collectingMunicipality>
,
<quantity id="4CA8F3EA9F51FFC5FBD5FCE85F982D51" box="[1079,1188,776,802]" metricMagnitude="2" metricUnit="m" metricValue="1.65" metricValueMax="3.0" metricValueMin="0.3" pageId="7" pageNumber="8" unit="m" value="165.0" valueMax="300.0" valueMin="30.0">
<elevation id="007DB93C9F51FFC5FBD5FCE85F982D51" box="[1079,1188,776,802]" metricMagnitude="2" metricUnit="m" metricValue="1.65" metricValueMax="3.0" metricValueMin="0.3" pageId="7" pageNumber="8" unit="m" value="165.0" valueMax="300.0" valueMin="30.0">30-300m</elevation>
</quantity>
,
<date id="FFEE78CF9F51FFC5FB4DFCE85E002D50" box="[1199,1340,776,803]" pageId="7" pageNumber="8" value="1958-07-01" valueMax="1958-07-06" valueMin="1958-07-01">
<collectingDate id="EFAA81279F51FFC5FB4DFCE85E002D50" box="[1199,1340,776,803]" pageId="7" pageNumber="8" value="1958-07-01" valueMax="1958-07-06" valueMin="1958-07-01">1-6. VII. 58</collectingDate>
</date>
”; “
<collectorName id="26A53BD99F51FFC5FABCFCE85AB42D35" pageId="7" pageNumber="8">T. C. Maa Collector</collectorName>
BISHOP Mus.”;
<collectionCode id="ED41C6CA9F51FFC5FDB5FCCD59952D34" box="[599,681,813,839]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="7" pageNumber="8" type="Museum">BPBM</collectionCode>
</materialsCitation>
<materialsCitation id="3B3854529F51FFC5FD20FCCC59D02D18" ID-GBIF-Occurrence="2464648271" collectingDate="1958-08-29" collectingDateMax="1958-08-30" collectingDateMin="1958-08-29" collectionCode="BPBM" collectorName="T. C. Maa Collector" country="Malaysia" location="Bau" municipality="Lake Area" pageId="7" pageNumber="8" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D5695869F51FFC5FD20FCCC59CC2D34" box="[706,752,812,839]" pageId="7" pageNumber="8" type="female">1 ♀</specimenCount>
; “BORNEO:
<collectingCountry id="F3471E9F9F51FFC5FC76FCCC5F1F2D35" box="[916,1059,812,838]" name="Malaysia" pageId="7" pageNumber="8">SARAWAK</collectingCountry>
<location id="8E8F08D49F51FFC5FBC8FCCD5F602D34" LSID="urn:lsid:plazi:treatment:03F9EF199F51FFCAFDB4FEEB5A162CAC:8E8F08D49F51FFC5FBC8FCCD5F602D34" box="[1066,1116,813,839]" country="Malaysia" municipality="Lake Area" name="Bau" pageId="7" pageNumber="8">Bau</location>
,
<collectingMunicipality id="6B8BC4759F51FFC5FB8AFCCC5FDE2D34" box="[1128,1250,812,839]" pageId="7" pageNumber="8">Lake Area</collectingMunicipality>
<date id="FFEE78CF9F51FFC5FB08FCCC5BC12D18" pageId="7" pageNumber="8" value="1958-08-29" valueMax="1958-08-30" valueMin="1958-08-29">
<collectingDate id="EFAA81279F51FFC5FB08FCCC5BC12D18" pageId="7" pageNumber="8" value="1958-08-29" valueMax="1958-08-30" valueMin="1958-08-29">
29-30. VIII.
<date id="FFEE78CF9F51FFC5FF5FFCB05BC12D18" box="[189,253,848,875]" pageId="7" pageNumber="8" value="1958" year="1958">1958</date>
</collectingDate>
</date>
”; “
<collectorName id="26A53BD99F51FFC5FEC2FCB059362D19" box="[288,522,848,875]" pageId="7" pageNumber="8">T. C. Maa Collector</collectorName>
BISHOP”;
<collectionCode id="ED41C6CA9F51FFC5FD75FCB159D02D18" box="[663,748,849,875]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="7" pageNumber="8" type="Museum">BPBM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A0D849F51FFCAFF5FFC7259C62FAF" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
<paragraph id="8BEF5E0F9F51FFC5FF5FFC725A722DDF" blockId="7.[189,334,914,940]" box="[189,334,914,940]" pageId="7" pageNumber="8">
<heading id="D0A7E9639F51FFC5FF5FFC725A722DDF" bold="true" box="[189,334,914,940]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B924821D9F51FFC5FF5FFC725A722DDF" bold="true" box="[189,334,914,940]" pageId="7" pageNumber="8">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF5E0F9F51FFC5FF08FC225A162DAF" blockId="7.[189,1399,962,1096]" box="[234,298,962,988]" pageId="7" pageNumber="8">
<emphasis id="B924821D9F51FFC5FF08FC225A162DAF" bold="true" box="[234,298,962,988]" pageId="7" pageNumber="8">Male</emphasis>
</paragraph>
<paragraph id="8BEF5E0F9F51FFC5FF5FFC0659F02A3B" blockId="7.[189,1399,962,1096]" pageId="7" pageNumber="8">BODY. Oblong, subparallel-sided, slightly convex dorsally, weakly shiny, closely covered with suberect silver setae. Coloration of body blackish brown, but mouthparts and legs yellowish brown; elytra with obscure silver spots consisting of erect setae.</paragraph>
<paragraph id="8BEF5E0F9F51FFC5FF5FFB9058A62ADD" blockId="7.[189,1398,1136,1198]" pageId="7" pageNumber="8">HEAD. Moderate in size; frons and clypeus closely covered with erect, long silver setae; clypeus with straight anterior margin. Eyes large, prominent dorso-laterally.</paragraph>
<paragraph id="8BEF5E0F9F51FFC5FF5FFB36587C2A82" blockId="7.[189,832,1238,1265]" box="[189,832,1238,1265]" pageId="7" pageNumber="8">ANTENNAE. Reaching about proximal half of pronotum.</paragraph>
<paragraph id="8BEF5E0F9F51FFC5FF5FFAF9585C2B25" blockId="7.[189,1399,1304,1367]" pageId="7" pageNumber="8">PRONOTUM. Widest at basal third, densely bearing short yellowish setae and minute scale-like setae; PW/ PL 2.252.50 (2.34). Scutellum small, subtriangular.</paragraph>
<paragraph id="8BEF5E0F9F51FFC5FF5FFA9F58FC2B93" blockId="7.[189,1399,1406,1505]" pageId="7" pageNumber="8">ELYTRA. Oblong, subparallel-sided near base to apical fifth, densely covered with short yellowish setae and minute scale-like setae, projecting ventrally in apical part; EL/ EW 1.591.80 (1.72); EL/ PL 4.17 4.63 (4.39); EW/ PW 1.021.16 (1.10); TL/ EW 1.972.22 (2.11).</paragraph>
<paragraph id="8BEF5E0F9F51FFC5FF5FF9E95AE528FD" blockId="7.[189,1399,1544,1678]" pageId="7" pageNumber="8">
ABDOMEN. Sternite VII (
<figureCitation id="136B428A9F51FFC5FE06F9E859792850" box="[484,581,1544,1571]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[196,1392,354,1877]" captionTargetId="figure@9.[189,1398,348,1883]" captionTargetPageId="9" captionText="Fig. 6. Acontosceles borneensis sp. nov., male (AE) and female (FH) genitalia (holotype and paratype, BPBM). A, F. Sternite VII. B. Sternite VIII. C. Sternite IX. D. Aedeagus, left lateral view. E. Aedeagus, ventral view. G. Urosternite. H. Ovipositor." figureDoi="http://doi.org/10.5281/zenodo.3564544" httpUri="https://zenodo.org/record/3564544/files/figure.png" pageId="7" pageNumber="8">Fig. 6A</figureCitation>
) broad, gently arcuate along caudal margin, shallowly concave in median part, bearing about 20 long setae and three pairs of long and stout setae. Sternite VIII (
<figureCitation id="136B428A9F51FFC5FAF1F9CC5E522834" box="[1299,1390,1580,1607]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[196,1392,354,1877]" captionTargetId="figure@9.[189,1398,348,1883]" captionTargetPageId="9" captionText="Fig. 6. Acontosceles borneensis sp. nov., male (AE) and female (FH) genitalia (holotype and paratype, BPBM). A, F. Sternite VII. B. Sternite VIII. C. Sternite IX. D. Aedeagus, left lateral view. E. Aedeagus, ventral view. G. Urosternite. H. Ovipositor." figureDoi="http://doi.org/10.5281/zenodo.3564544" httpUri="https://zenodo.org/record/3564544/files/figure.png" pageId="7" pageNumber="8">Fig. 6B</figureCitation>
) membranous, bearing minute spines in apical parts. Sternite IX (
<figureCitation id="136B428A9F51FFC5FC53F9B05F302818" box="[945,1036,1616,1643]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[196,1392,354,1877]" captionTargetId="figure@9.[189,1398,348,1883]" captionTargetPageId="9" captionText="Fig. 6. Acontosceles borneensis sp. nov., male (AE) and female (FH) genitalia (holotype and paratype, BPBM). A, F. Sternite VII. B. Sternite VIII. C. Sternite IX. D. Aedeagus, left lateral view. E. Aedeagus, ventral view. G. Urosternite. H. Ovipositor." figureDoi="http://doi.org/10.5281/zenodo.3564544" httpUri="https://zenodo.org/record/3564544/files/figure.png" pageId="7" pageNumber="8">Fig. 6C</figureCitation>
) moderately sclerotized, large, with small apical plates.
</paragraph>
<paragraph id="8BEF5E0F9F51FFC5FF5FF9575939294F" blockId="7.[189,1399,1719,1853]" pageId="7" pageNumber="8">
AEDEAGUS (
<figureCitation id="136B428A9F51FFC5FEA1F9575ABA28A1" box="[323,390,1719,1746]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[196,1392,354,1877]" captionTargetId="figure@9.[189,1398,348,1883]" captionTargetPageId="9" captionText="Fig. 6. Acontosceles borneensis sp. nov., male (AE) and female (FH) genitalia (holotype and paratype, BPBM). A, F. Sternite VII. B. Sternite VIII. C. Sternite IX. D. Aedeagus, left lateral view. E. Aedeagus, ventral view. G. Urosternite. H. Ovipositor." figureDoi="http://doi.org/10.5281/zenodo.3564544" httpUri="https://zenodo.org/record/3564544/files/figure.png" pageId="7" pageNumber="8">Fig. 6</figureCitation>
DE). Long, well sclerotized, strongly curved ventrally; basal piece oblong, expanded basally, evenly curved ventrally; lateral lobes long and slender, serrate on dorso-lateral margins, curved ventrally; median lobe long and slender, about 0.6 times as long as lateral lobe, gently curved ventrally; ML/BL 0.36; ML/ LL 0.68.
</paragraph>
<paragraph id="8BEF5E0F9F51FFC5FF08F8855A79290C" blockId="7.[189,1399,1893,2028]" box="[234,325,1893,1919]" pageId="7" pageNumber="8">
<emphasis id="B924821D9F51FFC5FF08F8855A79290C" bold="true" box="[234,325,1893,1919]" pageId="7" pageNumber="8">Female</emphasis>
</paragraph>
<paragraph id="8BEF5E0F9F51FFCAFF5FF8695E4A2F3A" blockId="7.[189,1399,1893,2028]" lastBlockId="8.[189,1398,267,329]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
Sexual dimorphism distinct in the following characteristics: 1) frons and clypeus covered with short yellowish setae; 2) apical part of elytra projecting ventro-apically; 3) sternite VII (
<figureCitation id="136B428A9F51FFC5FB92F84D5FFA29BB" box="[1136,1222,1965,1992]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[196,1392,354,1877]" captionTargetId="figure@9.[189,1398,348,1883]" captionTargetPageId="9" captionText="Fig. 6. Acontosceles borneensis sp. nov., male (AE) and female (FH) genitalia (holotype and paratype, BPBM). A, F. Sternite VII. B. Sternite VIII. C. Sternite IX. D. Aedeagus, left lateral view. E. Aedeagus, ventral view. G. Urosternite. H. Ovipositor." figureDoi="http://doi.org/10.5281/zenodo.3564544" httpUri="https://zenodo.org/record/3564544/files/figure.png" pageId="7" pageNumber="8">Fig. 6F</figureCitation>
) rather pointed on caudal margin, bearing about 40 long setae. Urosternite (
<figureCitation id="136B428A9F51FFC5FC71F83158CD299F" box="[915,1009,2001,2028]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[196,1392,354,1877]" captionTargetId="figure@9.[189,1398,348,1883]" captionTargetPageId="9" captionText="Fig. 6. Acontosceles borneensis sp. nov., male (AE) and female (FH) genitalia (holotype and paratype, BPBM). A, F. Sternite VII. B. Sternite VIII. C. Sternite IX. D. Aedeagus, left lateral view. E. Aedeagus, ventral view. G. Urosternite. H. Ovipositor." figureDoi="http://doi.org/10.5281/zenodo.3564544" httpUri="https://zenodo.org/record/3564544/files/figure.png" pageId="7" pageNumber="8">Fig. 6G</figureCitation>
) well sclerotized, with long and slender apodeme; lateral projections stout, bifid in apical parts. Ovipositor (
<figureCitation id="136B428A9F5EFFCAFBA6FEEB5F9E2F55" box="[1092,1186,267,294]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[196,1392,354,1877]" captionTargetId="figure@9.[189,1398,348,1883]" captionTargetPageId="9" captionText="Fig. 6. Acontosceles borneensis sp. nov., male (AE) and female (FH) genitalia (holotype and paratype, BPBM). A, F. Sternite VII. B. Sternite VIII. C. Sternite IX. D. Aedeagus, left lateral view. E. Aedeagus, ventral view. G. Urosternite. H. Ovipositor." figureDoi="http://doi.org/10.5281/zenodo.3564544" httpUri="https://zenodo.org/record/3564544/files/figure.png" pageId="8" pageNumber="9">Fig. 6H</figureCitation>
) well sclerotized; coxite sparsely punctuate, pointed at apices; approximate ratio of coxite and baculus (n = 1) of 1.0:1.7.
</paragraph>
<paragraph id="8BEF5E0F9F5EFFCAFF5FFE905A492FF9" blockId="8.[189,373,368,394]" box="[189,373,368,394]" pageId="8" pageNumber="9">
<heading id="D0A7E9639F5EFFCAFF5FFE905A492FF9" bold="true" box="[189,373,368,394]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B924821D9F5EFFCAFF5FFE905A492FF9" bold="true" box="[189,373,368,394]" pageId="8" pageNumber="9">Measurements</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF5E0F9F5EFFCAFF5FFE7E59C62FAF" blockId="8.[189,1399,414,476]" pageId="8" pageNumber="9">Unsexed (n = 4): TL 2.172.25 (2.21) mm; PW 0.901.00 (0.96) mm; PL 0.400.42 (0.41) mm; EL 1.751.85 (1.80) mm; EW 1.001.10 (1.05) mm.</paragraph>
</subSubSection>
<subSubSection id="C34A0D849F5EFFCAFF5FFDE35A8C2C3E" box="[189,432,515,589]" pageId="8" pageNumber="9" type="distribution">
<paragraph id="8BEF5E0F9F5EFFCAFF5FFDE35A6A2C6E" blockId="8.[189,342,515,541]" box="[189,342,515,541]" pageId="8" pageNumber="9">
<heading id="D0A7E9639F5EFFCAFF5FFDE35A6A2C6E" bold="true" box="[189,342,515,541]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B924821D9F5EFFCAFF5FFDE35A6A2C6E" bold="true" box="[189,342,515,541]" pageId="8" pageNumber="9">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF5E0F9F5EFFCAFF5FFDD25A8C2C3E" blockId="8.[189,432,562,589]" box="[189,432,562,589]" pageId="8" pageNumber="9">
<collectingCountry id="F3471E9F9F5EFFCAFF5FFDD25A152C3F" box="[189,297,562,588]" name="Malaysia" pageId="8" pageNumber="9">Malaysia</collectingCountry>
(
<collectingCountry id="F3471E9F9F5EFFCAFEDAFDD25A982C3E" box="[312,420,562,589]" name="Malaysia" pageId="8" pageNumber="9">Sarawak</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C34A0D849F5EFFCAFF5FFD935A162CAC" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8BEF5E0F9F5EFFCAFF5FFD935A132CFE" blockId="8.[189,303,627,653]" box="[189,303,627,653]" pageId="8" pageNumber="9">
<heading id="D0A7E9639F5EFFCAFF5FFD935A132CFE" bold="true" box="[189,303,627,653]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B924821D9F5EFFCAFF5FFD935A132CFE" bold="true" box="[189,303,627,653]" pageId="8" pageNumber="9">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF5E0F9F5EFFCAFF5FFD425A162CAC" blockId="8.[189,1398,674,735]" pageId="8" pageNumber="9">
This species belongs to the
<taxonomicName id="4C50258C9F5EFFCAFDFFFD4259FA2CCF" authorityName="Champion" authorityYear="1924" box="[541,710,674,700]" class="Insecta" family="Limnichidae" genus="Acontosceles" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="hydroporoides">
<emphasis id="B924821D9F5EFFCAFDFFFD4259FA2CCF" box="[541,710,674,700]" italics="true" pageId="8" pageNumber="9">hydroporoides</emphasis>
</taxonomicName>
species group judging from characteristics of the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>