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<mods:title id="0F6EE778CFC4097932EC71CF79E5CAC6">Systematics of the genus Rhacophorus (Amphibia, Anura): identity of red-webbed forms and description of a new species from Assam</mods:title>
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<treatment id="03DC831FFFB9AD7D9F929C1EFE84F8EE" ID-DOI="http://doi.org/10.5281/zenodo.5629733" ID-GBIF-Taxon="119380396" ID-Zenodo-Dep="5629733" LSID="urn:lsid:plazi:treatment:03DC831FFFB9AD7D9F929C1EFE84F8EE" httpUri="http://treatment.plazi.org/id/03DC831FFFB9AD7D9F929C1EFE84F8EE" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
<subSubSection id="C36F6182FFB9AD7F9F929C1EFF0FFCED" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BCA3209FFB9AD7F9F929C1EFF0FFCED" blockId="4.[151,1423,791,852]" pageId="4" pageNumber="5">
<heading id="D0828565FFB9AD7F9F929C1EFF0FFCED" bold="true" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<emphasis id="B901EE1BFFB9AD7F9F929C1EFF0FFCED" bold="true" pageId="4" pageNumber="5">
Identity of
<taxonomicName id="4C75498AFFB9AD7F9E249C1EFCDAFC88" authority="Liu &amp; Hu, 1960" authorityName="Liu &amp; Hu" authorityYear="1960" box="[289,775,791,817]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="rhodopus">
<emphasis id="B901EE1BFFB9AD7F9E249C1EFDE0FC88" bold="true" box="[289,573,791,817]" italics="true" pageId="4" pageNumber="5">Rhacophorus rhodopus</emphasis>
<bibRefCitation id="EFE44FF8FFB9AD7F9D419C1EFCDAFC88" author="Liu" box="[580,775,791,817]" pageId="4" pageNumber="17" refString="Liu C. - C. &amp; Hu S. - Q. (1960) Preliminary report of amphibian from southern Yunnan. Acta Zoologica Sinica, &quot; 1959 &quot; 11 (4), 509 - 533." type="journal article" year="1960">Liu &amp; Hu, 1960</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C75498AFFB9AD7F9C439C1EFF0FFCED" authority="Sarkar &amp; Sanyal, 1985" authorityName="Sarkar &amp; Sanyal" authorityYear="1985" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="namdaphaensis">
<emphasis id="B901EE1BFFB9AD7F9C439C1EFB70FC88" bold="true" box="[838,1197,791,817]" italics="true" pageId="4" pageNumber="5">Rhacophorus namdaphaensis</emphasis>
<bibRefCitation id="EFE44FF8FFB9AD7F9BB19C1EFF0FFCED" author="Sarkar" pageId="4" pageNumber="17" refString="Sarkar, A. K. &amp; Sanyal, D. P. (1985) Amphibia. Records of the zoological Survey of India, 82, 285 - 295, pl. 1." type="journal article" year="1985">Sarkar &amp; Sanyal, 1985</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36F6182FFB9AD7D9F929C8CFE84F8EE" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="description">
<paragraph id="8BCA3209FFB9AD7F9F929C8CFCA7FB36" blockId="4.[151,1437,901,2007]" pageId="4" pageNumber="5">
<taxonomicName id="4C75498AFFB9AD7F9F929C8CFE74FC27" box="[151,425,901,926]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="rhodopus">
<emphasis id="B901EE1BFFB9AD7F9F929C8CFE74FC27" box="[151,425,901,926]" italics="true" pageId="4" pageNumber="5">Rhacophorus rhodopus</emphasis>
</taxonomicName>
was described by Liu &amp; Hu in 1960 on a holophoront (CIB 571171) and
<specimenCount id="9D73F980FFB9AD7F9A099C8CFAB7FC26" box="[1292,1386,901,927]" pageId="4" pageNumber="5" type="male">20 male</specimenCount>
and
<specimenCount id="9D73F980FFB9AD7F9F929CA4FEDDFC7E" box="[151,256,941,967]" pageId="4" pageNumber="5" type="female">2 female</specimenCount>
specimens from Meng-yang in Yunnan (
<collectingCountry id="F3627299FFB9AD7F9DF59CA4FCE4FC7E" box="[752,825,941,967]" name="China" pageId="4" pageNumber="5">China</collectingCountry>
) (copy of original description in appendix). It is a rather small frog (males
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, females
<quantity id="4C8D9FECFFB9AD7F9DD39CDCFC59FC56" box="[726,900,981,1007]" metricMagnitude="-2" metricUnit="m" metricValue="5.13" metricValueMax="5.34" metricValueMin="4.92" pageId="4" pageNumber="5" unit="mm" value="51.3" valueMax="53.4" valueMin="49.2">49.253.4 mm</quantity>
) with pointed snout, with a red brown or yellow dorsum, indistinct dark markings, usually with dark brown X-shaped marking, posterior back with dark transverse stripes; flanks brightly yellow; legs with dark crossbars, and red web. In the original description it was compared to the species of Anderson and was distinguished by the dorsal colour (green, violet after death, in
<taxonomicName id="4C75498AFFB9AD7F9FB09B7CFE93FB37" box="[181,334,1141,1166]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B901EE1BFFB9AD7F9FB09B7CFE93FB37" box="[181,334,1141,1166]" italics="true" pageId="4" pageNumber="5">R. maculatus</emphasis>
</taxonomicName>
) and shape of snout (rounded in
<taxonomicName id="4C75498AFFB9AD7F9DD59B7CFCB4FB37" box="[720,873,1141,1166]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B901EE1BFFB9AD7F9DD59B7CFCB4FB37" box="[720,873,1141,1166]" italics="true" pageId="4" pageNumber="5">R. maculatus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BCA3209FFB9AD7F9FC39B94FEDFFABE" blockId="4.[151,1437,901,2007]" pageId="4" pageNumber="5">
Specimens mentioned by
<bibRefCitation id="EFE44FF8FFB9AD7F9EF79B94FD4FFB0E" author="Taylor" box="[498,658,1181,1207]" pageId="4" pageNumber="17" refString="Taylor, E. H. (1962) The amphibian fauna of Thailand. University of Kansas Science Bulletin, 63, 265 - 599." type="journal article" year="1962">Taylor (1962)</bibRefCitation>
from Chiang Mai (
<collectingCountry id="F3627299FFB9AD7F9C779B94FC03FB0E" box="[882,990,1181,1207]" name="Thailand" pageId="4" pageNumber="5">Thailand</collectingCountry>
) as
<taxonomicName id="4C75498AFFB9AD7F9B0E9B94FAEAFB0F" box="[1035,1335,1181,1206]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="B901EE1BFFB9AD7F9B0E9B94FAEAFB0F" box="[1035,1335,1181,1206]" italics="true" pageId="4" pageNumber="5">Rhacophorus bipunctatus</emphasis>
</taxonomicName>
are similar to the specimens of
<taxonomicName id="4C75498AFFB9AD7F9EBA9BCCFD8CFB67" box="[447,593,1221,1246]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="rhodopus">
<emphasis id="B901EE1BFFB9AD7F9EBA9BCCFD8CFB67" box="[447,593,1221,1246]" italics="true" pageId="4" pageNumber="5">R. rhodopus</emphasis>
</taxonomicName>
by the presence of a dark line between eyes, faint x-shaped mark on shoulder.
</paragraph>
<paragraph id="8BCA3209FFB9AD7F9FC39A1CFD25F9A6" blockId="4.[151,1437,901,2007]" pageId="4" pageNumber="5">
Inger
<emphasis id="B901EE1BFFB9AD7F9E0E9A1CFE98FA97" box="[267,325,1301,1326]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
(1999) reported specimens of
<taxonomicName id="4C75498AFFB9AD7F9DAE9A1CFC0AFA97" box="[683,983,1301,1326]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="B901EE1BFFB9AD7F9DAE9A1CFC0AFA97" box="[683,983,1301,1326]" italics="true" pageId="4" pageNumber="5">Rhacophorus bipunctatus</emphasis>
</taxonomicName>
from Boun Luoi (
<collectingCountry id="F3627299FFB9AD7F9BAE9A1CFAC8FA96" box="[1195,1301,1301,1327]" name="Vietnam" pageId="4" pageNumber="5">Vietnam</collectingCountry>
), compared them to the description of
<taxonomicName id="4C75498AFFB9AD7F9ED29A34FD36FAEF" box="[471,747,1341,1366]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="rhodopus">
<emphasis id="B901EE1BFFB9AD7F9ED29A34FD36FAEF" box="[471,747,1341,1366]" italics="true" pageId="4" pageNumber="5">Rhacophorus rhodopus</emphasis>
</taxonomicName>
and concluded that they belonged in the same biological species. Inger
<emphasis id="B901EE1BFFB9AD7F9E3A9A6CFEA7FAC7" box="[319,378,1381,1406]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
(1999) did not consider
<bibRefCitation id="EFE44FF8FFB9AD7F9D9F9A6CFCBCFAC6" author="Anderson" box="[666,865,1381,1407]" pageId="4" pageNumber="16" refString="Anderson, J. (1871) A list of the reptilian accession to the Indian Museum, Calcutta, from 1865 to 1870, with a description of some new species. Journal of the Asiatic Society of Bengal, 40, 12 - 39." type="journal article" year="1871">Anderson (1871)</bibRefCitation>
as the original author of the taxon with the valid name
<taxonomicName id="4C75498AFFB9AD7F9FE59A84FDCDFA1F" box="[224,528,1421,1446]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="B901EE1BFFB9AD7F9FE59A84FDCDFA1F" box="[224,528,1421,1446]" italics="true" pageId="4" pageNumber="5">Rhacophorus bipunctatus</emphasis>
</taxonomicName>
. They did not compare their specimens with the onomatophores of Andersons taxon. Though they mentioned specimens from Assam,
<collectingCountry id="F3627299FFB9AD7F9C8B9ABCFBDFFA76" box="[910,1026,1461,1487]" name="Myanmar" pageId="4" pageNumber="5">Myanmar</collectingCountry>
,
<collectingCountry id="F3627299FFB9AD7F9B179ABCFBA7FA76" box="[1042,1146,1461,1487]" name="Thailand" pageId="4" pageNumber="5">Thailand</collectingCountry>
and
<collectingCountry id="F3627299FFB9AD7F9BB99ABCFAFFFA76" box="[1212,1314,1461,1487]" name="Vietnam" pageId="4" pageNumber="5">Vietnam</collectingCountry>
, they discussed only differences between
<collectingCountry id="F3627299FFB9AD7F9D259AD4FD8BFA4E" box="[544,598,1501,1527]" name="Thailand" pageId="4" pageNumber="5">Thai</collectingCountry>
and Vietnamese specimens, thus no specimens of the same biological species as
<taxonomicName id="4C75498AFFB9AD7F9E16990CFD29F9A6" authority="Anderson, 1871" authorityName="Anderson" authorityYear="1871" box="[275,756,1541,1567]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B901EE1BFFB9AD7F9E16990CFDF2F9A7" box="[275,559,1541,1566]" italics="true" pageId="4" pageNumber="5">Rhacophorus maculatus</emphasis>
<bibRefCitation id="EFE44FF8FFB9AD7F9D33990CFD29F9A6" author="Anderson" box="[566,756,1541,1567]" pageId="4" pageNumber="16" refString="Anderson, J. (1871) A list of the reptilian accession to the Indian Museum, Calcutta, from 1865 to 1870, with a description of some new species. Journal of the Asiatic Society of Bengal, 40, 12 - 39." type="journal article" year="1871">Anderson, 1871</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BCA3209FFB9AD7F9FC39924FA84F92E" blockId="4.[151,1437,901,2007]" pageId="4" pageNumber="5">
In conclusion, specimens of
<collectingCountry id="F3627299FFB9AD7F9D119924FDAAF9FE" box="[532,631,1581,1607]" name="Vietnam" pageId="4" pageNumber="5">Vietnam</collectingCountry>
studied by Inger
<emphasis id="B901EE1BFFB9AD7F9C419924FCA2F9FF" box="[836,895,1581,1606]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
(1999) are to be assigned to the same biological species as
<taxonomicName id="4C75498AFFB9AD7F9E3B995CFE13F9D7" box="[318,462,1621,1646]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="rhodopus">
<emphasis id="B901EE1BFFB9AD7F9E3B995CFE13F9D7" box="[318,462,1621,1646]" italics="true" pageId="4" pageNumber="5">R. rhodopus</emphasis>
</taxonomicName>
from Yunnan. In contrast these authors did not provide evidence of the identity of this species with
<taxonomicName id="4C75498AFFB9AD7F9E659974FCE2F92E" authority="Anderson, 1871" authorityName="Anderson" authorityYear="1871" box="[352,831,1661,1687]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B901EE1BFFB9AD7F9E659974FDA6F92F" box="[352,635,1661,1686]" italics="true" pageId="4" pageNumber="5">Rhacophorus maculatus</emphasis>
<bibRefCitation id="EFE44FF8FFB9AD7F9D869974FCE2F92E" author="Anderson" box="[643,831,1661,1687]" pageId="4" pageNumber="16" refString="Anderson, J. (1871) A list of the reptilian accession to the Indian Museum, Calcutta, from 1865 to 1870, with a description of some new species. Journal of the Asiatic Society of Bengal, 40, 12 - 39." type="journal article" year="1871">Anderson, 1871</bibRefCitation>
</taxonomicName>
and their synonymy therefore has no support.
</paragraph>
<paragraph id="8BCA3209FFB9AD7F9FC399ACFA40F8B6" blockId="4.[151,1437,901,2007]" pageId="4" pageNumber="5">
It is true that both
<taxonomicName id="4C75498AFFB9AD7F9E9E99ACFD99F907" box="[411,580,1701,1726]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="B901EE1BFFB9AD7F9E9E99ACFD99F907" box="[411,580,1701,1726]" italics="true" pageId="4" pageNumber="5">R. bipunctatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C75498AFFB9AD7F9D7999ACFCD7F907" box="[636,778,1701,1726]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="rhodopus">
<emphasis id="B901EE1BFFB9AD7F9D7999ACFCD7F907" box="[636,778,1701,1726]" italics="true" pageId="4" pageNumber="5">R. rhodopus</emphasis>
</taxonomicName>
have pointed snout, but the dorsal coloration of all symphoronts of
<taxonomicName id="4C75498AFFB9AD7F9E2F99C4FE1AF95F" box="[298,455,1741,1766]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B901EE1BFFB9AD7F9E2F99C4FE1AF95F" box="[298,455,1741,1766]" italics="true" pageId="4" pageNumber="5">R. maculatus</emphasis>
</taxonomicName>
from Assam is bluish, thus green in life, whereas
<taxonomicName id="4C75498AFFB9AD7F9B2A99C4FB1DF95F" box="[1071,1216,1741,1766]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="rhodopus">
<emphasis id="B901EE1BFFB9AD7F9B2A99C4FB1DF95F" box="[1071,1216,1741,1766]" italics="true" pageId="4" pageNumber="5">R. rhodopus</emphasis>
</taxonomicName>
had brown dorsal colour. The two species show several other morphological differences which confirm their specific difference.
</paragraph>
<paragraph id="8BCA3209FFB9AD7E9FC39814FA48FE73" blockId="4.[151,1437,901,2007]" lastBlockId="5.[151,1437,152,458]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
In 1985, Sarkar &amp; Sanyal described a new species,
<taxonomicName id="4C75498AFFB9AD7F9C6D9814FB17F88F" box="[872,1226,1821,1846]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="namdaphaensis">
<emphasis id="B901EE1BFFB9AD7F9C6D9814FB17F88F" box="[872,1226,1821,1846]" italics="true" pageId="4" pageNumber="5">Rhacophorus namdaphaensis</emphasis>
</taxonomicName>
from Arunachal Pradesh,
<collectingCountry id="F3627299FFB9AD7F9E06984CFEE2F8E6" box="[259,319,1861,1887]" name="India" pageId="4" pageNumber="5">India</collectingCountry>
(holophoront, ZSI A.7180, ♂;
<figureCitation id="134E2E8CFFB9AD7F9DAE984CFD2DF8E6" box="[683,752,1861,1887]" captionStart="FIGURE 2" captionStartId="5.[151,255,1836,1860]" captionTargetBox="[150,1436,498,1758]" captionTargetId="figure@5.[147,1441,494,1759]" captionTargetPageId="5" captionText="FIGURE 2. Rhacophorus rhodopus Liu &amp; Hu, 1960. A, B, C: Rhacophorus namdaphaensis Sarkar &amp; Sanyal, 1985. ZSI A 7180, holophoront, ♂, SVL 36.5 mm. Dorsal, ventral and lateral view. D: MNHN 2006.2518, ♂ from Kengkoung, Luang Prabang Province, Laos." httpUri="https://zenodo.org/record/179815/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
A, B, C), which they compared to
<taxonomicName id="4C75498AFFB9AD7F9B82984CFEB1F83E" authority="Boulenger, 1892" authorityName="Boulenger" authorityYear="1892" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dulitensis">
<emphasis id="B901EE1BFFB9AD7F9B82984CFA41F8E7" box="[1159,1436,1861,1886]" italics="true" pageId="4" pageNumber="5">Rhacophorus dulitensis</emphasis>
(Boulenger, 1892)
</taxonomicName>
from Sumatra. It is a small sized
<taxonomicName id="4C75498AFFB9AD7F9DF39864FC4FF83F" box="[758,914,1901,1926]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B901EE1BFFB9AD7F9DF39864FC4FF83F" box="[758,914,1901,1926]" italics="true" pageId="4" pageNumber="5">Rhacophorus</emphasis>
</taxonomicName>
(SVL ♂
<quantity id="4C8D9FECFFB9AD7F9CFE9864FB7BF83E" box="[1019,1190,1901,1927]" metricMagnitude="-2" metricUnit="m" metricValue="3.405" metricValueMax="3.65" metricValueMin="3.16" pageId="4" pageNumber="5" unit="mm" value="34.05" valueMax="36.5" valueMin="31.6">31.636.5 mm</quantity>
) with reddish-brown dorsal colour intermixed with variable black spots. Upper surface of limbs show faint cross bars and undersurface of hindlimbs is reddish brown. A large, more or less round black spot is present on side behind the armpit. Study of the hypodigm confirms this description (see detailed redescription of the holophront in appendix). It corresponds in size and coloration pattern to the specimens described as
<taxonomicName id="4C75498AFFB8AD7E9B8F9FC9FA41FF60" box="[1162,1436,192,217]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="rhodopus">
<emphasis id="B901EE1BFFB8AD7E9B8F9FC9FA41FF60" box="[1162,1436,192,217]" italics="true" pageId="5" pageNumber="6">Rhacophorus rhodopus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFE44FF8FFB8AD7E9FBB9FE1FE4EFEBB" author="Liu" box="[190,403,232,258]" pageId="5" pageNumber="17" refString="Liu C. - C. &amp; Hu S. - Q. (1960) Preliminary report of amphibian from southern Yunnan. Acta Zoologica Sinica, &quot; 1959 &quot; 11 (4), 509 - 533." type="journal article" year="1960">Liu &amp; Hu (1960)</bibRefCitation>
. These specimens match also the specimens mentioned by
<bibRefCitation id="EFE44FF8FFB8AD7E9B659FE1FADAFEBB" author="Taylor" box="[1120,1287,232,258]" pageId="5" pageNumber="17" refString="Taylor, E. H. (1962) The amphibian fauna of Thailand. University of Kansas Science Bulletin, 63, 265 - 599." type="journal article" year="1962">Taylor (1962)</bibRefCitation>
, Inger
<emphasis id="B901EE1BFFB8AD7E9A5B9FE1FA41FEB8" box="[1374,1436,232,257]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
(1999) and
<bibRefCitation id="EFE44FF8FFB8AD7E9E1B9E19FDF0FE93" box="[286,557,272,298]" pageId="5" pageNumber="17" refString="Wilkinson, J. A., Thin, T., Lwin K. S. &amp; Shein, A. K. (2005) A new species of Rhacophorus (Anura: Rhacophoridae) from Myanmar (Burma). Proceedings of the California Academy of Sciences, 56 (4), 42 - 52." type="journal article">
Wilkinson
<emphasis id="B901EE1BFFB8AD7E9E9B9E19FE05FE90" box="[414,472,272,297]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
(2005)
</bibRefCitation>
as
<taxonomicName id="4C75498AFFB8AD7E9D519E19FC5CFE90" box="[596,897,272,297]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="B901EE1BFFB8AD7E9D519E19FC5CFE90" box="[596,897,272,297]" italics="true" pageId="5" pageNumber="6">Rhacophorus bipunctatus</emphasis>
</taxonomicName>
(or
<taxonomicName id="4C75498AFFB8AD7E9CB79E19FBBCFE90" box="[946,1121,272,297]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B901EE1BFFB8AD7E9CB79E19FBBCFE90" box="[946,1121,272,297]" italics="true" pageId="5" pageNumber="6">R. bimaculatus</emphasis>
</taxonomicName>
). The dorsal pattern corresponds to the orange to tan coloration with a darker brown pattern (blotching or an X mark) on the dorsum described by
<bibRefCitation id="EFE44FF8FFB8AD7E9E319E69FD9BFEC3" box="[308,582,352,378]" pageId="5" pageNumber="17" refString="Wilkinson, J. A., Thin, T., Lwin K. S. &amp; Shein, A. K. (2005) A new species of Rhacophorus (Anura: Rhacophoridae) from Myanmar (Burma). Proceedings of the California Academy of Sciences, 56 (4), 42 - 52." type="journal article">
Wilkinson
<emphasis id="B901EE1BFFB8AD7E9EB09E69FE2DFEC0" box="[437,496,352,377]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
(2005)
</bibRefCitation>
as
<taxonomicName id="4C75498AFFB8AD7E9D6B9E69FCC4FEC0" box="[622,793,352,377]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="B901EE1BFFB8AD7E9D6B9E69FCC4FEC0" box="[622,793,352,377]" italics="true" pageId="5" pageNumber="6">R. bipunctatus</emphasis>
</taxonomicName>
. The specimens included in the original description of
<taxonomicName id="4C75498AFFB8AD7E9F929E81FEACFE18" box="[151,369,392,417]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="namdaphaensis">
<emphasis id="B901EE1BFFB8AD7E9F929E81FEACFE18" box="[151,369,392,417]" italics="true" pageId="5" pageNumber="6">R. namdaphaensis</emphasis>
</taxonomicName>
have the distinct crossbands on the limbs and reduced webbing between fingers. These specimens have a single round spot behind armpit as mentioned by
<bibRefCitation id="EFE44FF8FFB8AD7E9CA29EB9FB6BFE73" box="[935,1206,432,458]" pageId="5" pageNumber="17" refString="Wilkinson, J. A., Thin, T., Lwin K. S. &amp; Shein, A. K. (2005) A new species of Rhacophorus (Anura: Rhacophoridae) from Myanmar (Burma). Proceedings of the California Academy of Sciences, 56 (4), 42 - 52." type="journal article">
Wilkinson
<emphasis id="B901EE1BFFB8AD7E9B229EB9FBBCFE70" box="[1063,1121,432,457]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
(2005)
</bibRefCitation>
for
<taxonomicName id="4C75498AFFB8AD7E9BE09EB9FA53FE70" box="[1253,1422,432,457]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="B901EE1BFFB8AD7E9BE09EB9FA53FE70" box="[1253,1422,432,457]" italics="true" pageId="5" pageNumber="6">R. bipunctatus</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF0A6281FFB8AD7E9F929825FE3AF829" httpUri="https://zenodo.org/record/179815/files/figure.png" pageId="5" pageNumber="6" targetBox="[150,1436,498,1758]" targetPageId="5">
<paragraph id="8BCA3209FFB8AD7E9F929825FE3AF829" blockId="5.[151,1436,1836,1936]" pageId="5" pageNumber="6">
<emphasis id="B901EE1BFFB8AD7E9F929825FEC4F8FD" bold="true" box="[151,281,1836,1860]" pageId="5" pageNumber="6">FIGURE 2.</emphasis>
<taxonomicName id="4C75498AFFB8AD7E9E1A9825FD16F8FA" authority="Liu &amp; Hu, 1960" authorityName="Liu &amp; Hu" authorityYear="1960" box="[287,715,1836,1859]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="rhodopus">
<emphasis id="B901EE1BFFB8AD7E9E1A9825FDC5F8FA" box="[287,536,1836,1859]" italics="true" pageId="5" pageNumber="6">Rhacophorus rhodopus</emphasis>
Liu &amp; Hu, 1960
</taxonomicName>
. A, B, C:
<taxonomicName id="4C75498AFFB8AD7E9C319825FAB7F8FA" authority="Sarkar &amp; Sanyal, 1985" authorityName="Sarkar &amp; Sanyal" authorityYear="1985" box="[820,1386,1836,1859]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="namdaphaensis">
<emphasis id="B901EE1BFFB8AD7E9C319825FBB3F8FA" box="[820,1134,1836,1859]" italics="true" pageId="5" pageNumber="6">Rhacophorus namdaphaensis</emphasis>
Sarkar &amp; Sanyal, 1985
</taxonomicName>
. ZSI A7180, holophoront, ♂, SVL 36.5 mm. Dorsal, ventral and lateral view. D: MNHN 2006.2518, ♂ from Kengkoung, Luang Prabang Province, Laos.
</paragraph>
</caption>
<paragraph id="8BCA3209FFBBAD7D9FC39F91FC2CFE1B" blockId="6.[151,1436,152,418]" pageId="6" pageNumber="7">
Specimens of
<taxonomicName id="4C75498AFFBBAD7D9E6E9F91FE24FF08" box="[363,505,152,177]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="rhodopus">
<emphasis id="B901EE1BFFBBAD7D9E6E9F91FE24FF08" box="[363,505,152,177]" italics="true" pageId="6" pageNumber="7">R. rhodopus</emphasis>
</taxonomicName>
have been collected in Phongsaly province and Luang Prabang province (
<collectingCountry id="F3627299FFBBAD7D9A5D9F91FA49FF0B" box="[1368,1428,152,178]" name="Laos" pageId="6" pageNumber="7">Laos</collectingCountry>
) (MNHN 2004.04090410, MNHN 2006.25182519) and in Doi Chiang
<collectingCountry id="F3627299FFBBAD7D9CEC9FC9FBC7FF63" box="[1001,1050,192,218]" name="Iceland" pageId="6" pageNumber="7">Dao</collectingCountry>
(
<collectingCountry id="F3627299FFBBAD7D9B2C9FC9FB45FF63" box="[1065,1176,192,218]" name="Thailand" pageId="6" pageNumber="7">Thailand</collectingCountry>
). These specimens are morphologically very similar and show the colour pattern described above. Thus they are distinct from the specimens described by
<bibRefCitation id="EFE44FF8FFBBAD7D9EB09E19FD5DFE93" author="Anderson" box="[437,640,272,298]" pageId="6" pageNumber="16" refString="Anderson, J. (1871) A list of the reptilian accession to the Indian Museum, Calcutta, from 1865 to 1870, with a description of some new species. Journal of the Asiatic Society of Bengal, 40, 12 - 39." type="journal article" year="1871">Anderson (1871)</bibRefCitation>
. Morphometrical comparison of specimens including the hypodigm of
<taxonomicName id="4C75498AFFBBAD7D9FBD9E31FE0BFEE8" box="[184,470,312,337]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B901EE1BFFBBAD7D9FBD9E31FE0BFEE8" box="[184,470,312,337]" italics="true" pageId="6" pageNumber="7">Rhacophorus maculatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C75498AFFBBAD7D9D169E31FD36FEE8" box="[531,747,312,337]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="namdaphaensis">
<emphasis id="B901EE1BFFBBAD7D9D169E31FD36FEE8" box="[531,747,312,337]" italics="true" pageId="6" pageNumber="7">R. namdaphaensis</emphasis>
</taxonomicName>
showed statistically significant differences for snout-vent length, head width, head length and tibia length and for ratio of head width/snout-vent length (
<tableCitation id="C6F707B2FFBBAD7D9BE59E69FAE5FEC3" box="[1248,1336,352,378]" captionStart="TABLE 1" captionStartId="6.[151,239,511,535]" captionTargetBox="[156,1425,647,1431]" captionTargetPageId="6" captionText="TABLE 1. Measurements of males and females of Rhacophorus bipunctatus and R. rhodopus. Mean value, standard deviation, minimum maximum values are given. U Mann-Whitney U; p Probability; *** p = 0.001; ** 0.001 &lt;p = 0.01; 0.1 &lt;p = 0.05; ns result not significant." httpUri="http://table.plazi.org/id/DF0A6281FFBBAD7D9F929EF6FD07FDE5" pageId="6" pageNumber="7" tableUuid="DF0A6281FFBBAD7D9F929EF6FD07FDE5">Table 1</tableCitation>
;
<figureCitation id="134E2E8CFFBBAD7D9A429E69FA51FEC3" box="[1351,1420,352,378]" captionStart="FIGURE 3" captionStartId="7.[151,255,1104,1128]" captionTargetBox="[271,1375,191,1084]" captionTargetId="figure@7.[223,1383,176,1098]" captionTargetPageId="7" captionText="FIGURE 3. Plot of size (SVL) versus relative head width (HW / SVL) for adult male specimens of Rhacophorus bipunctatus (including measurements of symphoronts of Rhacophorus maculatus) and R. rhodopus (including measurements of hypodigm of R. namdaphaensis)." httpUri="https://zenodo.org/record/179816/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
).
<taxonomicName id="4C75498AFFBBAD7D9F929E81FE9DFE18" box="[151,320,392,417]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="B901EE1BFFBBAD7D9F929E81FE9DFE18" box="[151,320,392,417]" italics="true" pageId="6" pageNumber="7">R. bipunctatus</emphasis>
</taxonomicName>
is larger and has a relatively larger head than
<taxonomicName id="4C75498AFFBBAD7D9C5E9E81FC37FE18" box="[859,1002,392,417]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="rhodopus">
<emphasis id="B901EE1BFFBBAD7D9C5E9E81FC37FE18" box="[859,1002,392,417]" italics="true" pageId="6" pageNumber="7">R. rhodopus</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF0A6281FFBBAD7D9F929EF6FD07FDE5" ID-Table-UUID="DF0A6281FFBBAD7D9F929EF6FD07FDE5" httpUri="http://table.plazi.org/id/DF0A6281FFBBAD7D9F929EF6FD07FDE5" pageId="6" pageNumber="7" targetBox="[156,1425,647,1431]" targetIsTable="true" targetPageId="6">
<paragraph id="8BCA3209FFBBAD7D9F929EF6FD07FDE5" blockId="6.[151,1436,511,604]" pageId="6" pageNumber="7">
<emphasis id="B901EE1BFFBBAD7D9F929EF6FED1FDAE" bold="true" box="[151,268,511,535]" pageId="6" pageNumber="7">TABLE 1.</emphasis>
Measurements of males and females of
<taxonomicName id="4C75498AFFBBAD7D9DCA9D09FC3FFDAE" box="[719,994,512,535]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="B901EE1BFFBBAD7D9DCA9D09FC3FFDAE" box="[719,994,512,535]" italics="true" pageId="6" pageNumber="7">Rhacophorus bipunctatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C75498AFFBBAD7D9B199D09FB7CFDAE" box="[1052,1185,512,535]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="rhodopus">
<emphasis id="B901EE1BFFBBAD7D9B199D09FB7CFDAE" box="[1052,1185,512,535]" italics="true" pageId="6" pageNumber="7">R. rhodopus</emphasis>
</taxonomicName>
. Mean value, standard deviation, minimum maximum values are given. U Mann-Whitney U; p Probability; *** p=0.001; ** 0.001&lt;p=0.01; 0.1&lt;p=0.05; ns result not significant.
</paragraph>
</caption>
<paragraph id="8BCA3209FFBBAD7D9F999DB9FA4CFA2E" pageId="6" pageNumber="7">
<table id="F975C0A9FFBB52849F999D8EFA4CFA2E" box="[156,1425,647,1431]" gridcols="4" gridrows="9" pageId="6" pageNumber="7">
<tr id="3545304BFFBB52849F999D8EFA4CFD7C" box="[156,1425,647,709]" gridrow="0" pageId="6" pageNumber="7">
<th id="76945937FFBB52849F999D8EFEF5FD7C" box="[156,296,647,709]" gridcol="0" gridrow="0" pageId="6" pageNumber="7">Measurements</th>
<th id="76945937FFBB52849E669D8EFD7FFD7C" box="[355,674,647,709]" gridcol="1" gridrow="0" pageId="6" pageNumber="7">
<taxonomicName id="4C75498AFFBBAD7D9E909D81FDA8FD24" box="[405,629,648,669]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="rhodopus">
<emphasis id="B901EE1BFFBBAD7D9E909D81FDA8FD24" box="[405,629,648,669]" italics="true" pageId="6" pageNumber="7">Rhacophorus rhodopus</emphasis>
</taxonomicName>
</th>
<th id="76945937FFBB52849DF59D8EFB9AFD7C" box="[752,1095,647,709]" gridcol="2" gridrow="0" pageId="6" pageNumber="7">
<taxonomicName id="4C75498AFFBBAD7D9C249D81FBCAFD24" box="[801,1047,648,669]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="B901EE1BFFBBAD7D9C249D81FBCAFD24" box="[801,1047,648,669]" italics="true" pageId="6" pageNumber="7">Rhacophorus bipunctatus</emphasis>
</taxonomicName>
</th>
<th id="76945937FFBB52849B7E9D8EFA4CFD7C" box="[1147,1425,647,709]" gridcol="3" gridrow="0" pageId="6" pageNumber="7">Mann-Whitney U test</th>
</tr>
<tr id="3545304BFFBB52849F999DEFFA4CFCA3" box="[156,1425,742,794]" gridrow="1" pageId="6" pageNumber="7">
<th id="76945937FFBB52849F999DEFFEF5FCA3" box="[156,296,742,794]" gridcol="0" gridrow="1" pageId="6" pageNumber="7">/ ratios</th>
<td id="76945937FFBB52849E669DEFFD7FFCA3" box="[355,674,742,794]" gridcol="1" gridrow="1" pageId="6" pageNumber="7">Males Females N=10 N=2</td>
<td id="76945937FFBB52849DF59DEFFB9AFCA3" box="[752,1095,742,794]" gridcol="2" gridrow="1" pageId="6" pageNumber="7">Males Females N=10 N=5</td>
<td id="76945937FFBB52849B7E9DEFFA4CFCA3" box="[1147,1425,742,794]" gridcol="3" gridrow="1" pageId="6" pageNumber="7">Males Females</td>
</tr>
<tr id="3545304BFFBB52849F999C32FA4CFCCC" box="[156,1425,827,885]" gridrow="2" pageId="6" pageNumber="7">
<th id="76945937FFBB52849F999C32FEF5FCCC" box="[156,296,827,885]" gridcol="0" gridrow="2" pageId="6" pageNumber="7">SVL</th>
<td id="76945937FFBB52849E669C32FD7FFCCC" box="[355,674,827,885]" gridcol="1" gridrow="2" pageId="6" pageNumber="7">34.6 ± 2.07 50.3 ± 0.35 (31.637.2) (50.050.5)</td>
<td id="76945937FFBB52849DF59C32FB9AFCCC" box="[752,1095,827,885]" gridcol="2" gridrow="2" pageId="6" pageNumber="7">38.9 ± 2.89 53.4 ± 4.64 (31.541.7) (49.259.1)</td>
<td id="76945937FFBB52849B7E9C32FA4CFCCC" box="[1147,1425,827,885]" gridcol="3" gridrow="2" pageId="6" pageNumber="7">U=10 U=4 p=0.002 ** p=0.857 ns</td>
</tr>
<tr id="3545304BFFBB52849F999C9FFA4CFC69" box="[156,1425,918,976]" gridrow="3" pageId="6" pageNumber="7">
<th id="76945937FFBB52849F999C9FFEF5FC69" box="[156,296,918,976]" gridcol="0" gridrow="3" pageId="6" pageNumber="7">HW</th>
<td id="76945937FFBB52849E669C9FFD7FFC69" box="[355,674,918,976]" gridcol="1" gridrow="3" pageId="6" pageNumber="7">11.7 ± 0.84 16.9 ± 0.07 (10.312.9) (16.816.9)</td>
<td id="76945937FFBB52849DF59C9FFB9AFC69" box="[752,1095,918,976]" gridcol="2" gridrow="3" pageId="6" pageNumber="7">13.9 ± 0.88 19.0 ± 1.73 (12.315.3) (17.420.9)</td>
<td id="76945937FFBB52849B7E9C9FFA4CFC69" box="[1147,1425,918,976]" gridcol="3" gridrow="3" pageId="6" pageNumber="7">U=3 U=0 p=0.000 *** p=0.95 ns</td>
</tr>
<tr id="3545304BFFBB52849F999CF8FA4CFB92" box="[156,1425,1009,1067]" gridrow="4" pageId="6" pageNumber="7">
<th id="76945937FFBB52849F999CF8FEF5FB92" box="[156,296,1009,1067]" gridcol="0" gridrow="4" pageId="6" pageNumber="7">HL</th>
<td id="76945937FFBB52849E669CF8FD7FFB92" box="[355,674,1009,1067]" gridcol="1" gridrow="4" pageId="6" pageNumber="7">12.3 ± 0.79 17.7 ± 1.06 (10.913.2) (16.918.4)</td>
<td id="76945937FFBB52849DF59CF8FB9AFB92" box="[752,1095,1009,1067]" gridcol="2" gridrow="4" pageId="6" pageNumber="7">13.7 ± 0.78 18.0 ± 1.06 (12.515.2) (16.819.7)</td>
<td id="76945937FFBB52849B7E9CF8FA4CFB92" box="[1147,1425,1009,1067]" gridcol="3" gridrow="4" pageId="6" pageNumber="7">U=8 U=5 p=0.001 *** p=1.000 ns</td>
</tr>
<tr id="3545304BFFBB52849F999B45FA4CFB3F" box="[156,1425,1100,1158]" gridrow="5" pageId="6" pageNumber="7">
<th id="76945937FFBB52849F999B45FEF5FB3F" box="[156,296,1100,1158]" gridcol="0" gridrow="5" pageId="6" pageNumber="7">TL</th>
<td id="76945937FFBB52849E669B45FD7FFB3F" box="[355,674,1100,1158]" gridcol="1" gridrow="5" pageId="6" pageNumber="7">17.4 ± 0.66 24.4 ± 1.20 (16.518.3) (23.525.2)</td>
<td id="76945937FFBB52849DF59B45FB9AFB3F" box="[752,1095,1100,1158]" gridcol="2" gridrow="5" pageId="6" pageNumber="7">18.8 ± 1.30 26.3 ± 2.19 (16.420.9) (24.228.8)</td>
<td id="76945937FFBB52849B7E9B45FA4CFB3F" box="[1147,1425,1100,1158]" gridcol="3" gridrow="5" pageId="6" pageNumber="7">U=15 U=2 p=0.008 ** p=0.381 ns</td>
</tr>
<tr id="3545304BFFBB52849F999BAEFA4CFB58" box="[156,1425,1191,1249]" gridrow="6" pageId="6" pageNumber="7">
<th id="76945937FFBB52849F999BAEFEF5FB58" box="[156,296,1191,1249]" gridcol="0" gridrow="6" pageId="6" pageNumber="7">HW/SVL</th>
<td id="76945937FFBB52849E669BAEFD7FFB58" box="[355,674,1191,1249]" gridcol="1" gridrow="6" pageId="6" pageNumber="7">337 ± 9.80 335 ± 3.77 (326353) (333338)</td>
<td id="76945937FFBB52849DF59BAEFB9AFB58" box="[752,1095,1191,1249]" gridcol="2" gridrow="6" pageId="6" pageNumber="7">358 ± 16.83 357 ± 12.83 (338390) (335366)</td>
<td id="76945937FFBB52849B7E9BAEFA4CFB58" box="[1147,1425,1191,1249]" gridcol="3" gridrow="6" pageId="6" pageNumber="7">U=13 U=1 p=0.005 ** p=0.190 ns</td>
</tr>
<tr id="3545304BFFBB52849F999A0BFA4CFA85" box="[156,1425,1282,1340]" gridrow="7" pageId="6" pageNumber="7">
<th id="76945937FFBB52849F999A0BFEF5FA85" box="[156,296,1282,1340]" gridcol="0" gridrow="7" pageId="6" pageNumber="7">HL/SVL</th>
<td id="76945937FFBB52849E669A0BFD7FFA85" box="[355,674,1282,1340]" gridcol="1" gridrow="7" pageId="6" pageNumber="7">355 ± 6.51 351 ± 18.64 (345365) (338364)</td>
<td id="76945937FFBB52849DF59A0BFB9AFA85" box="[752,1095,1282,1340]" gridcol="2" gridrow="7" pageId="6" pageNumber="7">354 ± 19.28 339 ± 15.67 (328397) (315358)</td>
<td id="76945937FFBB52849B7E9A0BFA4CFA85" box="[1147,1425,1282,1340]" gridcol="3" gridrow="7" pageId="6" pageNumber="7">U=43 U=3 p=0.597 ns p=0.571 ns</td>
</tr>
<tr id="3545304BFFBB52849F999A54FA4CFA2E" box="[156,1425,1373,1431]" gridrow="8" pageId="6" pageNumber="7">
<th id="76945937FFBB52849F999A54FEF5FA2E" box="[156,296,1373,1431]" gridcol="0" gridrow="8" pageId="6" pageNumber="7">TL/SVL</th>
<td id="76945937FFBB52849E669A54FD7FFA2E" box="[355,674,1373,1431]" gridcol="1" gridrow="8" pageId="6" pageNumber="7">503 ± 29.82 485 ± 20.51 (473572) (470499)</td>
<td id="76945937FFBB52849DF59A54FB9AFA2E" box="[752,1095,1373,1431]" gridcol="2" gridrow="8" pageId="6" pageNumber="7">484 ± 18.98 492 ± 4.23 (463521) (487498)</td>
<td id="76945937FFBB52849B7E9A54FA4CFA2E" box="[1147,1425,1373,1431]" gridcol="3" gridrow="8" pageId="6" pageNumber="7">U=31 U=5 p=0.151 ns p=1.000 ns</td>
</tr>
</table>
</paragraph>
<paragraph id="8BCA3209FFBBAD7D9FC3992CFE84F8EE" blockId="6.[151,1437,1573,1879]" pageId="6" pageNumber="7">
Thus the specimens referred to of
<taxonomicName id="4C75498AFFBBAD7D9D5F992CFCD8F987" box="[602,773,1573,1598]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="B901EE1BFFBBAD7D9D5F992CFCD8F987" box="[602,773,1573,1598]" italics="true" pageId="6" pageNumber="7">R. bipunctatus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFE44FF8FFBBAD7D9C34992CFB99F986" box="[817,1092,1573,1599]" pageId="6" pageNumber="17" refString="Wilkinson, J. A., Thin, T., Lwin K. S. &amp; Shein, A. K. (2005) A new species of Rhacophorus (Anura: Rhacophoridae) from Myanmar (Burma). Proceedings of the California Academy of Sciences, 56 (4), 42 - 52." type="journal article">
Wilkinson
<emphasis id="B901EE1BFFBBAD7D9CB7992CFC30F987" box="[946,1005,1573,1598]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
(2005)
</bibRefCitation>
are to be considered conspecific with those described as
<taxonomicName id="4C75498AFFBBAD7D9EE39944FD60F9DF" box="[486,701,1613,1638]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="namdaphaensis">
<emphasis id="B901EE1BFFBBAD7D9EE39944FD60F9DF" box="[486,701,1613,1638]" italics="true" pageId="6" pageNumber="7">R. namdaphaensis</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFE44FF8FFBBAD7D9DE29944FC20F9DE" author="Sarkar" box="[743,1021,1613,1639]" pageId="6" pageNumber="17" refString="Sarkar, A. K. &amp; Sanyal, D. P. (1985) Amphibia. Records of the zoological Survey of India, 82, 285 - 295, pl. 1." type="journal article" year="1985">Sarkar &amp; Sanyal (1985)</bibRefCitation>
and those described as
<taxonomicName id="4C75498AFFBBAD7D9A0B9944FA41F9DF" box="[1294,1436,1613,1638]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="rhodopus">
<emphasis id="B901EE1BFFBBAD7D9A0B9944FA41F9DF" box="[1294,1436,1613,1638]" italics="true" pageId="6" pageNumber="7">R. rhodopus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFE44FF8FFBBAD7D9FBE997CFE5CF936" author="Liu" box="[187,385,1653,1679]" pageId="6" pageNumber="17" refString="Liu C. - C. &amp; Hu S. - Q. (1960) Preliminary report of amphibian from southern Yunnan. Acta Zoologica Sinica, &quot; 1959 &quot; 11 (4), 509 - 533." type="journal article" year="1960">Liu &amp; Hu (1960)</bibRefCitation>
and this also applies to the specimens mentioned as
<taxonomicName id="4C75498AFFBBAD7D9CE3997CFB52F937" box="[998,1167,1653,1678]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="B901EE1BFFBBAD7D9CE3997CFB52F937" box="[998,1167,1653,1678]" italics="true" pageId="6" pageNumber="7">R. bipunctatus</emphasis>
</taxonomicName>
(or
<taxonomicName id="4C75498AFFBBAD7D9BBB997CFAB3F937" box="[1214,1390,1653,1678]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B901EE1BFFBBAD7D9BBB997CFAB3F937" box="[1214,1390,1653,1678]" italics="true" pageId="6" pageNumber="7">R. bimaculatus</emphasis>
</taxonomicName>
) by
<bibRefCitation id="EFE44FF8FFBBAD7D9F929994FEE4F90E" author="Taylor" box="[151,313,1693,1719]" pageId="6" pageNumber="17" refString="Taylor, E. H. (1962) The amphibian fauna of Thailand. University of Kansas Science Bulletin, 63, 265 - 599." type="journal article" year="1962">Taylor (1962)</bibRefCitation>
from Chiang Mai (
<collectingCountry id="F3627299FFBBAD7D9D249994FD52F90E" box="[545,655,1693,1719]" name="Thailand" pageId="6" pageNumber="7">Thailand</collectingCountry>
), Inger
<emphasis id="B901EE1BFFBBAD7D9DEE9994FCFAF90F" box="[747,807,1693,1718]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
(1999) from Buon Loi (
<collectingCountry id="F3627299FFBBAD7D9B4C9994FB6EF90E" box="[1097,1203,1693,1719]" name="Vietnam" pageId="6" pageNumber="7">Vietnam</collectingCountry>
). Its valid nomen is
<taxonomicName id="4C75498AFFBBAD7D9F9299CCFE75F967" box="[151,424,1733,1758]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="rhodopus">
<emphasis id="B901EE1BFFBBAD7D9F9299CCFE75F967" box="[151,424,1733,1758]" italics="true" pageId="6" pageNumber="7">Rhacophorus rhodopus</emphasis>
</taxonomicName>
. This species has a distribution area which extends from NE
<collectingCountry id="F3627299FFBBAD7D9B7199CCFB6DF966" box="[1140,1200,1733,1759]" name="India" pageId="6" pageNumber="7">India</collectingCountry>
to
<collectingCountry id="F3627299FFBBAD7D9BD099CCFA94F966" box="[1237,1353,1733,1759]" name="Myanmar" pageId="6" pageNumber="7">Myanmar</collectingCountry>
, northern
<collectingCountry id="F3627299FFBBAD7D9FC099E4FEEDF8BE" box="[197,304,1773,1799]" name="Thailand" pageId="6" pageNumber="7">Thailand</collectingCountry>
,
<collectingCountry id="F3627299FFBBAD7D9E3899E4FEA4F8BE" box="[317,377,1773,1799]" name="Laos" pageId="6" pageNumber="7">Laos</collectingCountry>
,
<collectingCountry id="F3627299FFBBAD7D9E8399E4FE34F8BE" box="[390,489,1773,1799]" name="Vietnam" pageId="6" pageNumber="7">Vietnam</collectingCountry>
and southern
<collectingCountry id="F3627299FFBBAD7D9D9799E4FD01F8BE" box="[658,732,1773,1799]" name="China" pageId="6" pageNumber="7">China</collectingCountry>
. Thus, it is sympatric in part of its range (NE
<collectingCountry id="F3627299FFBBAD7D9A0799E4FA9CF8BE" box="[1282,1345,1773,1799]" name="India" pageId="6" pageNumber="7">India</collectingCountry>
,
<collectingCountry id="F3627299FFBBAD7D9A4B99E4FF1BF896" name="Myanmar" pageId="6" pageNumber="7">Myanmar</collectingCountry>
) with
<taxonomicName id="4C75498AFFBBAD7D9E11981CFD1FF896" authority="Ahl, 1927" authorityName="Ahl" authorityYear="1927" box="[276,706,1813,1839]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="B901EE1BFFBBAD7D9E11981CFD9FF897" box="[276,578,1813,1838]" italics="true" pageId="6" pageNumber="7">Rhacophorus bipunctatus</emphasis>
<bibRefCitation id="EFE44FF8FFBBAD7D9D4E981CFD1FF896" author="Ahl" box="[587,706,1813,1839]" pageId="6" pageNumber="16" refString="Ahl, E. (1927) Zur Systematik der asiatischen Arten der Froschgattung Rhacophorus. Sitzungsberichte der Gesellschaft naturforschender Freunde Berlin, 1927, 35 - 47." type="journal article" year="1927">Ahl, 1927</bibRefCitation>
</taxonomicName>
as defined by the onomatophores of
<taxonomicName id="4C75498AFFBBAD7D9B7B981CFE88F8EE" authority="Anderson, 1871" authorityName="Anderson" authorityYear="1871" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B901EE1BFFBBAD7D9B7B981CFA41F897" box="[1150,1436,1813,1838]" italics="true" pageId="6" pageNumber="7">Rhacophorus maculatus</emphasis>
<bibRefCitation id="EFE44FF8FFBBAD7D9F929834FE88F8EE" author="Anderson" box="[151,341,1853,1879]" pageId="6" pageNumber="16" refString="Anderson, J. (1871) A list of the reptilian accession to the Indian Museum, Calcutta, from 1865 to 1870, with a description of some new species. Journal of the Asiatic Society of Bengal, 40, 12 - 39." type="journal article" year="1871">Anderson, 1871</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>