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<mods:title id="24898BAFFD0E613A5DB991EFAF2E138F">Taxonomic changes in Ichneumonoidea (Hymenoptera), and notes on certain type specimens</mods:title>
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<treatment id="03D35267FF9C8B65D8F3742CADF7C3B6" ID-DOI="http://doi.org/10.5281/zenodo.4595652" ID-GBIF-Taxon="180596298" ID-Zenodo-Dep="4595652" LSID="urn:lsid:plazi:treatment:03D35267FF9C8B65D8F3742CADF7C3B6" httpUri="http://treatment.plazi.org/id/03D35267FF9C8B65D8F3742CADF7C3B6" lastPageId="18" lastPageNumber="529" pageId="17" pageNumber="528">
<subSubSection id="C360B0FAFF9C8B66D8F3742CACACC756" box="[151,649,1583,1610]" pageId="17" pageNumber="528" type="nomenclature">
<paragraph id="8BC5E371FF9C8B66D8F3742CACACC756" blockId="17.[151,649,1583,1645]" box="[151,649,1583,1610]" pageId="17" pageNumber="528">
<heading id="D08D541DFF9C8B66D8F3742CACACC756" bold="true" box="[151,649,1583,1610]" fontSize="11" level="1" pageId="17" pageNumber="528" reason="1">
<emphasis id="B90E3F63FF9C8B66D8F3742CACACC756" bold="true" box="[151,649,1583,1610]" pageId="17" pageNumber="528">
15.
<taxonomicName id="4C7A98F2FF9C8B66D8A7742CACACC756" ID-CoL="3KT5G" authority="Morley 1912" authorityName="Morley" authorityYear="1912" box="[195,649,1583,1610]" class="Insecta" family="Ichneumonidae" genus="Hemiteles" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="528" phylum="Arthropoda" rank="species" species="cingulatorius">
<emphasis id="B90E3F63FF9C8B66D8A7742CAFC1C755" bold="true" box="[195,484,1583,1610]" italics="true" pageId="17" pageNumber="528">Hemiteles cingulatorius</emphasis>
<bibRefCitation id="EFEB9E80FF9C8B66D988742CACACC756" author="Morley, C." box="[492,649,1583,1609]" pageId="17" pageNumber="528" pagination="169 - 179" refId="ref10891" refString="Morley, C. (1912) No. XII. - Hymenoptera, Ichneumonidae. Transactions of the Linnean Society of London, 15, 169 - 179. https: // doi. org / 10.1111 / j. 1096 - 3642.1912. tb 00097. x" type="journal article" year="1912">Morley 1912</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C360B0FAFF9C8B66D8F37450AB44C7A9" pageId="17" pageNumber="528" type="description">
<paragraph id="8BC5E371FF9C8B66D8F37450AEDFC772" blockId="17.[151,649,1583,1645]" box="[151,250,1619,1645]" pageId="17" pageNumber="528">
(
<figureCitation id="1341FFF4FF9C8B66D8C47450AED4C772" box="[160,241,1619,1645]" captionStart="FIGURE 14" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[175,1411,181,1919]" captionTargetId="figure-15@19.[175,1411,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Hemiteles cingulatorius Morley 1912 lectotype female (NHMUK010634939), dorsal habitus (above) and head to second metasomal segment, dorsal (below), card mount and labels inset; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4595629" httpUri="https://zenodo.org/record/4595629/files/figure.png" pageId="17" pageNumber="528">Fig. 14</figureCitation>
)
</paragraph>
<paragraph id="8BC5E371FF9C8B66D8F37498AB44C7A9" blockId="17.[151,1437,1691,2042]" box="[151,1377,1691,1718]" pageId="17" pageNumber="528">
<emphasis id="B90E3F63FF9C8B66D8F37498AFB9C7AA" bold="true" box="[151,412,1691,1717]" pageId="17" pageNumber="528">Current combination.</emphasis>
<taxonomicName id="4C7A98F2FF9C8B66D9C77498AD67C7AA" authority="(Morley 1912)" authorityName="Broad" authorityYear="2021" baseAuthorityName="Morley" baseAuthorityYear="1912" box="[419,834,1691,1717]" class="Insecta" family="Ichneumonidae" genus="Tolonus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="528" phylum="Arthropoda" rank="species" species="cingulatorius" status="comb. nov.">
<emphasis id="B90E3F63FF9C8B66D9C77498ACB2C7AA" box="[419,663,1691,1717]" italics="true" pageId="17" pageNumber="528">Tolonus cingulatorius</emphasis>
(
<bibRefCitation id="EFEB9E80FF9C8B66DAC27498AD1FC7AA" author="Morley, C." box="[678,826,1691,1717]" pageId="17" pageNumber="528" pagination="169 - 179" refId="ref10891" refString="Morley, C. (1912) No. XII. - Hymenoptera, Ichneumonidae. Transactions of the Linnean Society of London, 15, 169 - 179. https: // doi. org / 10.1111 / j. 1096 - 3642.1912. tb 00097. x" type="journal article" year="1912">Morley 1912</bibRefCitation>
)
</taxonomicName>
<emphasis id="B90E3F63FF9C8B66DB2D7498AA30C7AA" bold="true" box="[841,1045,1691,1717]" pageId="17" pageNumber="528">
<taxonomicNameLabel id="A23D8218FF9C8B66DB2D7498AA30C7AA" box="[841,1045,1691,1717]" pageId="17" pageNumber="528" rank="species">new combination</taxonomicNameLabel>
</emphasis>
(
<taxonomicName id="4C7A98F2FF9C8B66DC407498AAF2C7AA" authorityName="Latreille" authorityYear="1802" box="[1060,1239,1691,1717]" class="Insecta" family="Ichneumonidae" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="528" phylum="Arthropoda" rank="family">Ichneumonidae</taxonomicName>
:
<taxonomicName id="4C7A98F2FF9C8B66DC857498AB70C7A9" baseAuthorityName="sensu Santos" baseAuthorityYear="2017" box="[1249,1365,1691,1718]" class="Insecta" family="Ichneumonidae" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="528" phylum="Arthropoda" rank="subFamily" subFamily="Cryptinae">Cryptinae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C360B0FAFF9C8B66D8A374BCAD50C6AE" pageId="17" pageNumber="528" type="materials_examined">
<paragraph id="8BC5E371FF9C8B66D8A374BCAD50C6AE" blockId="17.[151,1437,1691,2042]" pageId="17" pageNumber="528">
<emphasis id="B90E3F63FF9C8B66D8A374BCAF8AC7C6" bold="true" box="[199,431,1727,1753]" pageId="17" pageNumber="528">Material examined.</emphasis>
<taxonomicName id="4C7A98F2FF9C8B66D9DE74BCAD7FC7C6" authority="Morley 1912" authorityName="Morley" authorityYear="1912" box="[442,858,1727,1753]" class="Insecta" family="Ichneumonidae" genus="Hemiteles" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="528" phylum="Arthropoda" rank="species" species="cingulatorius">
<emphasis id="B90E3F63FF9C8B66D9DE74BCACE1C7C6" box="[442,708,1727,1753]" italics="true" pageId="17" pageNumber="528">Hemiteles cingulatorius</emphasis>
<bibRefCitation id="EFEB9E80FF9C8B66DAAE74BCAD7FC7C6" author="Morley, C." box="[714,858,1727,1753]" pageId="17" pageNumber="528" pagination="169 - 179" refId="ref10891" refString="Morley, C. (1912) No. XII. - Hymenoptera, Ichneumonidae. Transactions of the Linnean Society of London, 15, 169 - 179. https: // doi. org / 10.1111 / j. 1096 - 3642.1912. tb 00097. x" type="journal article" year="1912">Morley 1912</bibRefCitation>
</taxonomicName>
<specimenCount id="9D7C28F8FF9C8B66DB0474C3AD50C7C5" box="[864,885,1728,1754]" pageId="17" pageNumber="528" type="female"></specimenCount>
<typeStatus id="54C15DD3FF9C8B66DB1E74BCADC1C7C6" box="[890,996,1727,1753]" pageId="17" pageNumber="528" type="lectotype">lectotype</typeStatus>
, here designated,
<collectionCode id="ED6B7BB4FF9C8B66DCC374C3AB31C7C5" box="[1191,1300,1728,1754]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="17" pageNumber="528" type="Museum">NHMUK</collectionCode>
,
<collectingCountry id="F36DA3E1FF9C8B66DD7974BCABBDC7C6" box="[1309,1432,1727,1753]" name="Seychelles" pageId="17" pageNumber="528">Seychelles</collectingCountry>
, Praslin, according to
<bibRefCitation id="EFEB9E80FF9C8B66D9E974E0AC13C7E2" author="Morley, C." box="[397,566,1763,1789]" pageId="17" pageNumber="528" pagination="169 - 179" refId="ref10891" refString="Morley, C. (1912) No. XII. - Hymenoptera, Ichneumonidae. Transactions of the Linnean Society of London, 15, 169 - 179. https: // doi. org / 10.1111 / j. 1096 - 3642.1912. tb 00097. x" type="journal article" year="1912">Morley (1912)</bibRefCitation>
, Côtes dOr Estate, end of
<date id="FFC4C5B1FF9C8B66DB1274E7ADD4C7E2" box="[886,1009,1763,1790]" pageId="17" pageNumber="528" value="1908-11">Nov. 1908</date>
: 48; Praslin 08.
<collectingCountry id="F36DA3E1FF9C8B66DCB274E0AB6BC7E2" box="[1238,1358,1763,1789]" name="Seychelles" pageId="17" pageNumber="528">Seychelles</collectingCountry>
Exp.; Type C.M.;
<collectingCountry id="F36DA3E1FF9C8B66D9597504AF9FC63E" box="[317,442,1799,1825]" name="Seychelles" pageId="17" pageNumber="528">Seychelles</collectingCountry>
Islands. Pres. by Committee of the Percy Sladen Trust Fund. 1911-22.; Named by Claude Morley
<taxonomicName id="4C7A98F2FF9C8B66D9357528AC88C65A" authority="Morl." authorityName="Morley" authorityYear="1912" box="[337,685,1835,1861]" class="Insecta" family="Ichneumonidae" genus="Hemiteles" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="528" phylum="Arthropoda" rank="species" species="cingulatorius">Hemiteles cingulatorius Morl.</taxonomicName>
<date id="FFC4C5B1FF9C8B66DADC7528AD05C65A" box="[696,800,1835,1861]" pageId="17" pageNumber="528" value="1911-07">vii 1911</date>
; B.M.Type Hym. 3b391;
<collectionCode id="ED6B7BB4FF9C8B66DC0B7528AAFAC65A" box="[1135,1247,1835,1861]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="17" pageNumber="528" type="Museum">NHMUK</collectionCode>
010634939;
<specimenCount id="9D7C28F8FF9C8B66DD1D7528ABB9C65A" box="[1401,1436,1835,1861]" pageId="17" pageNumber="528" type="female">1♀</specimenCount>
(
<collectionCode id="ED6B7BB4FF9C8B66D8FB754CAF29C675" box="[159,268,1871,1898]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="17" pageNumber="528" type="Museum">NHMUK</collectionCode>
010634940), 2♂ (
<collectionCode id="ED6B7BB4FF9C8B66D987754CAC75C676" box="[483,592,1871,1897]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="17" pageNumber="528" type="Museum">NHMUK</collectionCode>
010634941,
<collectionCode id="ED6B7BB4FF9C8B66DA87754CAD6BC675" box="[739,846,1871,1898]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="17" pageNumber="528" type="Museum">NHMUK</collectionCode>
010634942)
<typeStatus id="54C15DD3FF9C8B66DB80754CAAACC676" box="[996,1161,1871,1897]" pageId="17" pageNumber="528" type="paralectotype">paralectotypes</typeStatus>
,
<collectingCountry id="F36DA3E1FF9C8B66DCF8754CAB32C676" box="[1180,1303,1871,1897]" name="Seychelles" pageId="17" pageNumber="528">Seychelles</collectingCountry>
, Silhouette (low coconut-planted country near the coast at Pointe Etienne,
<date id="FFC4C5B1FF9C8B66DB0B7570AA03C692" box="[879,1062,1907,1933]" pageId="17" pageNumber="528" value="1908-09-17">17th Sept. 1908</date>
, etc.; Mare aux Cochons marshy plateau and adjacent jungle,
<date id="FFC4C5B1FF9C8B66D9B1759BACE6C6AE" box="[469,707,1943,1970]" pageId="17" pageNumber="528" value="1908-08" valueMax="1908-09-31" valueMin="1908-08-00">Aug. and Sept. 1908</date>
:
<bibRefCitation id="EFEB9E80FF9C8B66DAB77594AD4DC6AE" author="Morley, C." box="[723,872,1943,1969]" pageId="17" pageNumber="528" pagination="169 - 179" refId="ref10891" refString="Morley, C. (1912) No. XII. - Hymenoptera, Ichneumonidae. Transactions of the Linnean Society of London, 15, 169 - 179. https: // doi. org / 10.1111 / j. 1096 - 3642.1912. tb 00097. x" type="journal article" year="1912">Morley 1912</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C360B0FAFF9C8B65D8A375B8ADF7C3B6" lastPageId="18" lastPageNumber="529" pageId="17" pageNumber="528" type="discussion">
<paragraph id="8BC5E371FF9C8B65D8A375B8AA96C05E" blockId="17.[151,1437,1691,2042]" lastBlockId="18.[151,1437,151,681]" lastPageId="18" lastPageNumber="529" pageId="17" pageNumber="528">
<emphasis id="B90E3F63FF9C8B66D8A375B8AF6AC6CA" bold="true" box="[199,335,1979,2005]" pageId="17" pageNumber="528">Comments.</emphasis>
<bibRefCitation id="EFEB9E80FF9C8B66D93F75B8AC26C6CA" author="Morley, C." box="[347,515,1979,2005]" pageId="17" pageNumber="528" pagination="169 - 179" refId="ref10891" refString="Morley, C. (1912) No. XII. - Hymenoptera, Ichneumonidae. Transactions of the Linnean Society of London, 15, 169 - 179. https: // doi. org / 10.1111 / j. 1096 - 3642.1912. tb 00097. x" type="journal article" year="1912">Morley (1912)</bibRefCitation>
described
<taxonomicName id="4C7A98F2FF9C8B66DAE175B8ADB3C6CA" authorityName="Morley" authorityYear="1912" box="[645,918,1979,2005]" class="Insecta" family="Ichneumonidae" genus="Hemiteles" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="528" phylum="Arthropoda" rank="species" species="cingulatorius">
<emphasis id="B90E3F63FF9C8B66DAE175B8ADB3C6CA" box="[645,918,1979,2005]" italics="true" pageId="17" pageNumber="528">Hemiteles cingulatorius</emphasis>
</taxonomicName>
from the
<collectingCountry id="F36DA3E1FF9C8B66DC7575B8AAACC6CA" box="[1041,1161,1979,2005]" name="Seychelles" pageId="17" pageNumber="528">Seychelles</collectingCountry>
(Praslin and Silhouette Islands), and the species has been ignored since the original description.
<taxonomicName id="4C7A98F2FF9C8B66DBAC75DCAAECC6E6" authority="Gravenhorst" authorityName="Gravenhorst" authorityYear="1829" box="[968,1225,2015,2041]" class="Insecta" family="Ichneumonidae" genus="Hemiteles" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="528" phylum="Arthropoda" rank="genus">
<emphasis id="B90E3F63FF9C8B66DBAC75DCAA1DC6E6" box="[968,1080,2015,2041]" italics="true" pageId="17" pageNumber="528">Hemiteles</emphasis>
Gravenhorst
</taxonomicName>
was a genus name used for many cryptines and phygadeuontines (in the modern sense;
<bibRefCitation id="EFEB9E80FF9F8B65DBFE7294AA03C1AE" author="Santos, B. F." box="[922,1062,151,177]" pageId="18" pageNumber="529" pagination="650 - 676" refId="ref11153" refString="Santos, B. F. (2017) Phylogeny and reclassification of Cryptini (Hymenoptera, Ichneumonidae, Cryptinae), with implications for ichneumonid higher-level classification. Systematic Entomology, 42, 650 - 676. https: // doi. org / 10.1111 / syen. 12238" type="journal article" year="2017">Santos 2017</bibRefCitation>
) with the fore wing areolet open. Now,
<taxonomicName id="4C7A98F2FF9F8B65D8BF72B8AF6EC1CA" authorityName="Gravenhorst" authorityYear="1829" box="[219,331,187,213]" class="Insecta" family="Ichneumonidae" genus="Hemiteles" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="B90E3F63FF9F8B65D8BF72B8AF6EC1CA" box="[219,331,187,213]" italics="true" pageId="18" pageNumber="529">Hemiteles</emphasis>
</taxonomicName>
is restricted to a small genus of
<taxonomicName id="4C7A98F2FF9F8B65DADF72B8AD5AC1CA" baseAuthorityName="Förster" baseAuthorityYear="1869" box="[699,895,187,213]" class="Insecta" family="Ichneumonidae" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="529" phylum="Arthropoda" rank="subFamily" subFamily="Phygadeuontinae">Phygadeuontinae</taxonomicName>
, parasitoids of spider egg sacs (e.g.,
<bibRefCitation id="EFEB9E80FF9F8B65DD7A72B8AF3FC1E6" author="Schwarz, M. &amp; Shaw, M. R." pageId="18" pageNumber="529" pagination="147 - 186" refId="ref11203" refString="Schwarz, M. &amp; Shaw, M. R. (2000) Western Palaearctic Cryptinae (Hymenoptera: Ichneumonidae) in the National Museums of Scotland, with nomenclatural changes, taxonomic notes, rearing records and special reference to the British check list. Part 3. Tribe Phygadeuontini, subtribes Chiroticina, Acrolytina, Hemitelina and Gelina (excluding Gelis), with descriptions of new species. Entomologist's Gazette, 51, 147 - 186." type="journal article" year="2000">Schwarz &amp; Shaw 2000</bibRefCitation>
).
<bibRefCitation id="EFEB9E80FF9F8B65D94972DCAFF7C1E6" author="Morley, C." box="[301,466,223,249]" pageId="18" pageNumber="529" pagination="169 - 179" refId="ref10891" refString="Morley, C. (1912) No. XII. - Hymenoptera, Ichneumonidae. Transactions of the Linnean Society of London, 15, 169 - 179. https: // doi. org / 10.1111 / j. 1096 - 3642.1912. tb 00097. x" type="journal article" year="1912">Morley (1912)</bibRefCitation>
described a type series of
<specimenCount id="9D7C28F8FF9F8B65DB6472DCAD5EC1E6" box="[768,891,223,249]" pageId="18" pageNumber="529" type="female">12 females</specimenCount>
and nine males, but only four of this apparently conspecific
<typeStatus id="54C15DD3FF9F8B65D97A7300AF50C002" box="[286,373,259,285]" pageId="18" pageNumber="529" type="syntype">syntype</typeStatus>
series seem to be in NHMUK now; I have designated the female that Morley labelled as type (NHMUK010634939) as
<typeStatus id="54C15DD3FF9F8B65D9D37324AC3BC05E" box="[439,542,295,321]" pageId="18" pageNumber="529" type="lectotype">lectotype</typeStatus>
(
<figureCitation id="1341FFF4FF9F8B65DA497324AC5BC05E" box="[557,638,295,321]" captionStart="FIGURE 14" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[175,1411,181,1919]" captionTargetId="figure-15@19.[175,1411,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Hemiteles cingulatorius Morley 1912 lectotype female (NHMUK010634939), dorsal habitus (above) and head to second metasomal segment, dorsal (below), card mount and labels inset; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4595629" httpUri="https://zenodo.org/record/4595629/files/figure.png" pageId="18" pageNumber="529">Fig. 14</figureCitation>
) in case the series are found not to be conspecific.
</paragraph>
<paragraph id="8BC5E371FF9F8B65D8A37348ADF7C3B6" blockId="18.[151,1437,151,681]" pageId="18" pageNumber="529">
<taxonomicName id="4C7A98F2FF9F8B65D8A37348AFF4C07A" authorityName="Morley" authorityYear="1912" box="[199,465,331,357]" class="Insecta" family="Ichneumonidae" genus="Hemiteles" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="529" phylum="Arthropoda" rank="species" species="cingulatorius">
<emphasis id="B90E3F63FF9F8B65D8A37348AFF4C07A" box="[199,465,331,357]" italics="true" pageId="18" pageNumber="529">Hemiteles cingulatorius</emphasis>
</taxonomicName>
clearly belongs to the subfamily
<taxonomicName id="4C7A98F2FF9F8B65DB5C7348AAAEC07A" authority="(sensu Santos 2017)" baseAuthorityName="sensu Santos" baseAuthorityYear="2017" box="[824,1163,331,357]" class="Insecta" family="Ichneumonidae" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="529" phylum="Arthropoda" rank="subFamily" subFamily="Cryptinae">
Cryptinae (
<emphasis id="B90E3F63FF9F8B65DBD17348ADD6C07B" box="[949,1011,331,356]" italics="true" pageId="18" pageNumber="529">sensu</emphasis>
<bibRefCitation id="EFEB9E80FF9F8B65DB9C7348AAA6C07A" author="Santos, B. F." box="[1016,1155,331,357]" pageId="18" pageNumber="529" pagination="650 - 676" refId="ref11153" refString="Santos, B. F. (2017) Phylogeny and reclassification of Cryptini (Hymenoptera, Ichneumonidae, Cryptinae), with implications for ichneumonid higher-level classification. Systematic Entomology, 42, 650 - 676. https: // doi. org / 10.1111 / syen. 12238" type="journal article" year="2017">Santos 2017</bibRefCitation>
)
</taxonomicName>
, and the
<taxonomicName id="4C7A98F2FF9F8B65DC8C7348AB73C07A" authorityName="Holmgren" authorityYear="1868" box="[1256,1366,331,357]" class="Insecta" family="Ichneumonidae" genus="Goryphus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="B90E3F63FF9F8B65DC8C7348AB73C07A" box="[1256,1366,331,357]" italics="true" pageId="18" pageNumber="529">Goryphus</emphasis>
</taxonomicName>
genus group. The nearest fit is the genus
<taxonomicName id="4C7A98F2FF9F8B65DA7A736CACCEC096" authority="Seyrig. In" authorityName="Seyrig. In" authorityYear="1952" box="[542,747,367,393]" class="Insecta" family="Ichneumonidae" genus="Tolonus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="B90E3F63FF9F8B65DA7A736CAC53C096" box="[542,630,367,393]" italics="true" pageId="18" pageNumber="529">Tolonus</emphasis>
Seyrig. In
</taxonomicName>
Towness (1970) key to genera of
<taxonomicName id="4C7A98F2FF9F8B65DC10736CAAC7C096" baseAuthorityName="sensu Santos" baseAuthorityYear="2017" box="[1140,1250,367,393]" class="Insecta" family="Ichneumonidae" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="529" phylum="Arthropoda" rank="subFamily" subFamily="Cryptinae">Cryptinae</taxonomicName>
(as Gelinae),
<taxonomicName id="4C7A98F2FF9F8B65DDE5736CAF09C0B2" authorityName="Morley" authorityYear="1912" class="Insecta" family="Ichneumonidae" genus="Hemiteles" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="529" phylum="Arthropoda" rank="species" species="cingulatorius">
<emphasis id="B90E3F63FF9F8B65DDE5736CAF09C0B2" italics="true" pageId="18" pageNumber="529">H. cingulatorius</emphasis>
</taxonomicName>
runs to
<taxonomicName id="4C7A98F2FF9F8B65D9E37390AFFAC0B2" authorityName="Seyrig. In" authorityYear="1952" box="[391,479,403,429]" class="Insecta" family="Ichneumonidae" genus="Tolonus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="B90E3F63FF9F8B65D9E37390AFFAC0B2" box="[391,479,403,429]" italics="true" pageId="18" pageNumber="529">Tolonus</emphasis>
</taxonomicName>
as the first tergite has anterior lateral teeth; the epomia is weak and diverges abruptly from the collar; and the scutellum has lateral carinae only very anteriorly. Additional characters supporting its placement in
<taxonomicName id="4C7A98F2FF9F8B65D94A73D8AFA3C0EA" authorityName="Seyrig. In" authorityYear="1952" box="[302,390,475,501]" class="Insecta" family="Ichneumonidae" genus="Tolonus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="B90E3F63FF9F8B65D94A73D8AFA3C0EA" box="[302,390,475,501]" italics="true" pageId="18" pageNumber="529">Tolonus</emphasis>
</taxonomicName>
are the fine longitudinal carina on the frons and the fore wing postnervulus intercepted above the middle (
<emphasis id="B90E3F63FF9F8B65D94473FCAF62C306" box="[288,327,511,537]" italics="true" pageId="18" pageNumber="529">CU</emphasis>
shorter than 2
<emphasis id="B90E3F63FF9F8B65D98E73FCAC39C307" box="[490,540,511,536]" italics="true" pageId="18" pageNumber="529">cu-a</emphasis>
). Five of the described species of
<taxonomicName id="4C7A98F2FF9F8B65DBC473FCADDDC306" authorityName="Seyrig. In" authorityYear="1952" box="[928,1016,511,537]" class="Insecta" family="Ichneumonidae" genus="Tolonus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="B90E3F63FF9F8B65DBC473FCADDDC306" box="[928,1016,511,537]" italics="true" pageId="18" pageNumber="529">Tolonus</emphasis>
</taxonomicName>
are known from
<collectingCountry id="F36DA3E1FF9F8B65DCDF73FCAB67C306" box="[1211,1346,511,537]" name="Madagascar" pageId="18" pageNumber="529">Madagascar</collectingCountry>
and the sixth from
<collectingCountry id="F36DA3E1FF9F8B65D9767020AF4EC322" box="[274,363,547,573]" name="Senegal" pageId="18" pageNumber="529">Senegal</collectingCountry>
or
<collectingCountry id="F36DA3E1FF9F8B65D9F57020AFCFC322" box="[401,490,547,573]" name="Gambia" pageId="18" pageNumber="529">Gambia</collectingCountry>
(
<bibRefCitation id="EFEB9E80FF9F8B65D99C7020ACA0C322" author="Seyrig, A." box="[504,645,546,573]" pageId="18" pageNumber="529" pagination="1 - 213" refId="ref11284" refString="Seyrig, A. (1952) Les Ichneumonides de Madagascar. IV Ichneumonidae Cryptinae. Memoires de l'Academie Malgache, Fascicule XIX, 1 - 213." type="book chapter" year="1952">Seyrig 1952</bibRefCitation>
;
<bibRefCitation id="EFEB9E80FF9F8B65DAF47020AD80C322" author="Townes, H. K. &amp; Townes, M." box="[656,933,547,573]" pageId="18" pageNumber="529" pagination="1 - 416" refId="ref11536" refString="Townes, H. K. &amp; Townes, M. (1973) A catalogue and reclassification of the Ethiopian Ichneumonidae. Memoirs of the American Entomological Institute, 19, 1 - 416." type="journal article" year="1973">Townes &amp; Townes 1973</bibRefCitation>
);
<bibRefCitation id="EFEB9E80FF9F8B65DBDC7020AA44C322" author="Townes, H. K." box="[952,1121,547,573]" pageId="18" pageNumber="529" pagination="1 - 537" refId="ref11506" refString="Townes, H. K. (1970) The genera of Ichneumonidae, Part 2. Memoirs of the American Entomological Institute, 12, 1 - 537." type="journal article" year="1970">Townes (1970)</bibRefCitation>
mentions three undescribed species from Africa.
<taxonomicName id="4C7A98F2FF9F8B65D91B7044AC57C37E" authorityName="Broad" authorityYear="2021" baseAuthorityName="Morley" baseAuthorityYear="1912" box="[383,626,583,609]" class="Insecta" family="Ichneumonidae" genus="Tolonus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="529" phylum="Arthropoda" rank="species" species="cingulatorius" status="comb. nov.">
<emphasis id="B90E3F63FF9F8B65D91B7044AC57C37E" box="[383,626,583,609]" italics="true" pageId="18" pageNumber="529">Tolonus cingulatorius</emphasis>
</taxonomicName>
<emphasis id="B90E3F63FF9F8B65DA1D7044ACDCC37E" bold="true" box="[633,761,583,609]" pageId="18" pageNumber="529">
<taxonomicNameLabel id="A23D8218FF9F8B65DA1D7044ACDCC37E" box="[633,761,583,609]" pageId="18" pageNumber="529" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
is a little smaller than the other described species (fore wing length
<quantity id="4C824E94FF9F8B65D8837068AF09C39A" box="[231,300,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="18" pageNumber="529" unit="mm" value="4.0">4 mm</quantity>
, in other species
<quantity id="4C824E94FF9F8B65D9927069AC70C39A" box="[502,597,618,645]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="18" pageNumber="529" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5-7 mm</quantity>
), although there is quite some variation between
<taxonomicName id="4C7A98F2FF9F8B65DCE97068AAC0C39A" authorityName="Seyrig. In" authorityYear="1952" box="[1165,1253,619,645]" class="Insecta" family="Ichneumonidae" genus="Tolonus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="B90E3F63FF9F8B65DCE97068AAC0C39A" box="[1165,1253,619,645]" italics="true" pageId="18" pageNumber="529">Tolonus</emphasis>
</taxonomicName>
species and the goryphine fauna of the Afrotropical and Malagasy regions needs revision.
</paragraph>
</subSubSection>
</treatment>
</document>