395 lines
67 KiB
XML
395 lines
67 KiB
XML
<document id="734DB2AC0B16985EE21675D3D3BBA05A" ID-DOI="10.11646/zootaxa.4786.2.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="3874770" ID-ZooBank="82E64B92-EAA4-43FA-A33C-E7D1FA955C59" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591189189041" checkinUser="plazi" docAuthor="Molineri, Carlos & Derka, Tomáš" docDate="2020" docId="03CCAB054507FFAC818F3D07D7DDFE24" docLanguage="en" docName="zootaxa.4786.2.7.pdf" docOrigin="Zootaxa 4786 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptohyphes kukenan Molineri & Derka 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="267" masterDocId="FFF5D37D4506FFAA81183F6AD704FFAF" masterDocTitle="A rare new species of Leptohyphes Eaton (Ephemeroptera: Leptohyphidae) from the Lost World" masterLastPageNumber="268" masterPageNumber="261" pageNumber="262" updateTime="1700276350103" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="7DE45293C7CDB0C0423496615DDBD3DA">
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<mods:title id="35D98CE818867B96420630261D0E7E4C">A rare new species of Leptohyphes Eaton (Ephemeroptera: Leptohyphidae) from the Lost World</mods:title>
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<mods:name id="9FBA2B7C0228265EC8165EAD96E56502" type="personal">
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<mods:roleTerm id="3C14FE5B3C37C71CAA8B54B21651BFC7">Author</mods:roleTerm>
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<mods:namePart id="EC639573B99401D184A3C19EBE25DCC6">Molineri, Carlos</mods:namePart>
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<mods:affiliation id="50505518D483980BA8A596D8202CE099">Instituto de Biodiversidad Neotropical, CONICET (National Council of Scientific Research), Universidad Nacional de Tucumán, Facultad de Ciencias Naturales e Instituto M. Lillo, Tucumán, Argentina.</mods:affiliation>
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<mods:roleTerm id="0D54F9BDA0ABC6F68F2AF519F77A0286">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E0573A8843CCCB8E5799A3AD25638F3E">Derka, Tomáš</mods:namePart>
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<mods:affiliation id="EF7037161E5BBDC89BAE5E3847A5AB07">Department of Ecology, Faculty of Natural Sciences, Comenius University, Mlynská dolina, Ilkovičova 6, SK- 84215 Bratislava, Slo- vakia. derka @ fns. uniba. sk; https: // orcid. org / 0000 - 0002 - 9304 - 9668</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="288DCFD25C976EA1C7D6F1D0F5D0AC6C">text</mods:typeOfResource>
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<mods:title id="03A1FE43080D6600D7E9F33FE4BB31A3">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="DDDDDC43C14360DE8B0FFE028178C7B9">
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<mods:date id="0B7DC60A2C5AD54700ADDF97E1273A3B">2020</mods:date>
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<mods:number id="ABABB065789E79D3A6D8FA325138A52C">2020-06-02</mods:number>
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<mods:number id="F628BF609ED5ED12A04EDF1DE87E71D3">4786</mods:number>
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<mods:classification id="46611789BE36568C3AD0F657C997B6FD">journal article</mods:classification>
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<mods:identifier id="44B59BDB27724B9FD39287A3DBC692CF" type="DOI">10.11646/zootaxa.4786.2.7</mods:identifier>
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<mods:identifier id="341E6FD901030CCBE86E8E7147F8A81A" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="D7E6BC048445C9EAEAD1141B6CE3CBED" type="Zenodo-Dep">3874770</mods:identifier>
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<treatment id="03CCAB054507FFAC818F3D07D7DDFE24" LSID="urn:lsid:plazi:treatment:03CCAB054507FFAC818F3D07D7DDFE24" httpUri="http://treatment.plazi.org/id/03CCAB054507FFAC818F3D07D7DDFE24" lastPageId="6" lastPageNumber="267" pageId="1" pageNumber="262">
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<subSubSection id="C37F49984507FFAB818F3D07D503FD28" box="[151,519,621,648]" pageId="1" pageNumber="262" type="nomenclature">
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<paragraph id="8BDA1A134507FFAB818F3D07D503FD28" blockId="1.[151,519,621,683]" box="[151,519,621,648]" pageId="1" pageNumber="262">
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<heading id="D092AD7F4507FFAB818F3D07D503FD28" bold="true" box="[151,519,621,648]" fontSize="11" level="1" pageId="1" pageNumber="262" reason="1">
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<emphasis id="B911C6014507FFAB818F3D07D503FD28" bold="true" box="[151,519,621,648]" pageId="1" pageNumber="262">
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<taxonomicName id="4C6561904507FFAB818F3D07D6A6FD27" authority="Molineri & Derka, 2020" authorityName="Molineri & Derka" authorityYear="2020" box="[151,418,621,648]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="kukenan" status="sp. nov.">
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<emphasis id="B911C6014507FFAB818F3D07D6A6FD27" bold="true" box="[151,418,621,648]" italics="true" pageId="1" pageNumber="262">Leptohyphes kukenan</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2227B7A4507FFAB80B13D07D503FD28" box="[425,519,621,647]" pageId="1" pageNumber="262" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="C37F49984507FFAB818F3DFBD624FD04" box="[151,288,657,683]" pageId="1" pageNumber="262" type="description">
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<paragraph id="8BDA1A134507FFAB818F3DFBD624FD04" blockId="1.[151,519,621,683]" box="[151,288,657,683]" pageId="1" pageNumber="262">
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(
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<figureCitation id="135E06964507FFAB81873DFBD61CFD04" box="[159,280,657,683]" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–9" captionStart-2="FIGURES 10–20" captionStartId-0="2.[151,264,1930,1955]" captionStartId-1="3.[151,264,1886,1911]" captionStartId-2="4.[151,264,1948,1973]" captionTargetBox-0="[156,1431,181,1904]" captionTargetBox-1="[169,1426,453,1857]" captionTargetBox-2="[154,1436,846,1923]" captionTargetId-0="figure@2.[156,1431,181,1904]" captionTargetId-1="figure@3.[151,1436,453,1861]" captionTargetId-2="figure@4.[151,1436,839,1923]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 1–4. Leptohyphes kukenan sp. nov. 1, female mature nymph, habitus; 2, female head, dorsal view; 3, male nearly mature nymph, habitus; 4, male head, dorsal view." captionText-1="FIGURES 5–9. Leptohyphes kukenan sp. nov., mouthparts. 5, labrum dorsal view (setae on right side omitted), with detail of bifid microsetae on fore margin; 6, left mandible with detail of prostheca, dorsal view;; 7, right mandible with detail of prostheca, dorsal view; 8, maxilla, ventral view of distal part; 9, labium, ventral view with detail of microsetae and pits." captionText-2="FIGURES 10–20. Leptohyphes kukenan sp. nov. 10, foreleg, dorsal view with detail of spine; 11, fore tarsal claw; 12, hind leg with detail of spine, dorsal view; 13, schematic representation of abdominal terga setation; 14–16, operculate gill in dorsal, lateral and ventral view; 17–20, abdominal gills III–VI, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.3874772" figureDoi-1="http://doi.org/10.5281/zenodo.3874774" figureDoi-2="http://doi.org/10.5281/zenodo.3874776" httpUri-0="https://zenodo.org/record/3874772/files/figure.png" httpUri-1="https://zenodo.org/record/3874774/files/figure.png" httpUri-2="https://zenodo.org/record/3874776/files/figure.png" pageId="1" pageNumber="262">Figs. 1–20</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37F49984507FFAB818F3DB3D6B4FC2C" pageId="1" pageNumber="262" type="materials_examined">
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<paragraph id="8BDA1A134507FFAB818F3DB3D6B4FC2C" blockId="1.[151,1437,728,2015]" pageId="1" pageNumber="262">
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<emphasis id="B911C6014507FFAB818F3DB3D64CFD5C" bold="true" box="[151,328,728,755]" pageId="1" pageNumber="262">Type material:</emphasis>
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<materialsCitation id="3B0D104E4507FFAB80573DB3D33CFCB8" collectingDate="2015-11-25" collectionCode="MLBV, T" collectorName="Derka" country="Venezuela" elevation="1631" latitude="5.15601" location="Kukenan Tepui" longLatPrecision="1" longitude="-60.82507" municipality="Kukenan river" pageId="1" pageNumber="262" specimenCount="1" stateProvince="Bolivar" typeStatus="holotype">
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<typeStatus id="54DEA4B14507FFAB80573DB3D6B3FD5C" box="[335,439,729,755]" pageId="1" pageNumber="262" type="holotype">Holotype</typeStatus>
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female mature nymph (
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<collectionCode id="ED7482D64507FFAB83DD3DB3D41DFD5C" box="[709,793,729,755]" pageId="1" pageNumber="262">MLBV</collectionCode>
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) from
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<collectingCountry id="F3725A834507FFAB827E3DB3D30BFD5C" box="[870,1039,729,755]" name="Venezuela" pageId="1" pageNumber="262">VENEZUELA</collectingCountry>
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, Est.
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<collectingRegion id="49A1D4F14507FFAB85553DB3D3A7FD5C" box="[1101,1187,729,755]" country="Venezuela" name="Bolivar" pageId="1" pageNumber="262">Bolívar</collectingRegion>
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,
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<collectingMunicipality id="6BBE80694507FFAB85B73DB3D254FD5C" box="[1199,1360,729,755]" pageId="1" pageNumber="262">Kukenán river</collectingMunicipality>
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below
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<location id="8EBA4CC84507FFAB818F3D97D641FCB8" LSID="urn:lsid:plazi:treatment:03CCAB054507FFAC818F3D07D7DDFE24:8EBA4CC84507FFAB818F3D97D641FCB8" box="[151,325,765,791]" country="Venezuela" latitude="5.15601" longLatPrecision="1" longitude="-60.82507" municipality="Kukenan river" name="Kukenan Tepui" pageId="1" pageNumber="262" stateProvince="Bolivar">Kukenán Tepui</location>
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,
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<geoCoordinate id="EE517CD44507FFAB80483D97D6DBFCB8" box="[336,479,764,791]" degrees="05.15601" direction="north" orientation="latitude" pageId="1" pageNumber="262" precision="1" value="5.15601">N 05.15601º</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE517CD44507FFAB80F23D97D585FCB8" box="[490,641,764,791]" degrees="60.82507" direction="west" orientation="longitude" pageId="1" pageNumber="262" precision="1" value="-60.82507">W 60.82507º</geoCoordinate>
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||
,
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<elevation id="0048FD204507FFAB839D3D96D410FCB8" box="[645,788,764,791]" metricMagnitude="3" metricUnit="m" metricValue="1.6309999999999998" pageId="1" pageNumber="262" unit="m" value="1631.0">
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<quantity id="4C9DB7F64507FFAB839D3D96D5DCFCB8" box="[645,728,764,791]" metricMagnitude="3" metricUnit="m" metricValue="1.6309999999999998" pageId="1" pageNumber="262" unit="m" value="1631.0">1631 m</quantity>
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||
a.s.l.
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||
</elevation>
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||
,
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||
<date id="FFDB3CD34507FFAB82383D96D498FCB8" box="[800,924,764,791]" pageId="1" pageNumber="262" value="2015-11-25">
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||
<collectingDate id="EF9FC53B4507FFAB82383D96D498FCB8" box="[800,924,764,791]" pageId="1" pageNumber="262" value="2015-11-25">25/xi/2015</collectingDate>
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</date>
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||
,
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||
<collectionCode id="ED7482D64507FFAB82BF3D97D4BDFCB8" box="[935,953,765,791]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="262">T</collectionCode>
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||
.
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<collectorName id="26907FC54507FFAB82DC3D97D30CFCB8" box="[964,1032,765,791]" pageId="1" pageNumber="262">Derka</collectorName>
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||
col.
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||
</materialsCitation>
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<materialsCitation id="3B0D104E4507FFAB85273D97D47FFC94" collectingDate="2015-11-25" collectionCode="MLBV, IBN" collectorName="Derka" country="Venezuela" elevation="1631" latitude="5.15601" location="Kukenan Tepui" longLatPrecision="1" longitude="-60.82507" municipality="Kukenan river" pageId="1" pageNumber="262" specimenCount="8" specimenCount-female="7" specimenCount-male="1" stateProvince="Bolivar" typeStatus="paratype">
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<typeStatus id="54DEA4B14507FFAB85273D97D3B4FCB8" box="[1087,1200,765,791]" pageId="1" pageNumber="262" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="9D63D19A4507FFAB85A23D97D21FFCB8" box="[1210,1307,765,791]" pageId="1" pageNumber="262" type="female">6 female</specimenCount>
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and
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<specimenCount id="9D63D19A4507FFAB84493D97D298FCB8" box="[1361,1436,765,791]" pageId="1" pageNumber="262" type="male">1 male</specimenCount>
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nymphs, same data as holotype (
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<specimenCount id="9D63D19A4507FFAB83143C4BD569FC94" box="[524,621,801,827]" pageId="1" pageNumber="262" type="female">1 female</specimenCount>
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in
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<collectionCode id="ED7482D64507FFAB838B3C4BD5E7FC94" box="[659,739,801,827]" pageId="1" pageNumber="262">MLBV</collectionCode>
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, rest in
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<collectionCode id="ED7482D64507FFAB82263C4BD477FC94" box="[830,883,801,827]" pageId="1" pageNumber="262">IBN</collectionCode>
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)
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</materialsCitation>
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;
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<materialsCitation id="3B0D104E4507FFAB82903C4BD6B4FC2C" collectingDate="2015-11-25" collectionCode="IBN, CM, T" collectorName="Derka" country="Venezuela" elevation="1875" latitude="5.16636" location="Kukenan Tepui" longLatPrecision="1" longitude="-60.82384" pageId="1" pageNumber="262" specimenCode="IBN833" specimenCount="1" specimenCount-female="1" stateProvince="Bolivar" typeStatus="paratype">
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<specimenCount id="9D63D19A4507FFAB82903C4BD4EEFC94" box="[904,1002,801,827]" pageId="1" pageNumber="262" type="female">1 female</specimenCount>
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||
nymph (
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||
<collectionCode id="ED7482D64507FFAB85493C4BD381FC94" box="[1105,1157,801,827]" pageId="1" pageNumber="262">IBN</collectionCode>
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, slide
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||
<specimenCode id="DBC3B2684507FFAB85D63C4BD22DFC94" box="[1230,1321,801,827]" collectionCode="IBN" pageId="1" pageNumber="262">IBN833</specimenCode>
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||
<collectionCode id="ED7482D64507FFAB84313C4BD252FC94" box="[1321,1366,801,827]" country="China" name="Chongqing Museum" pageId="1" pageNumber="262">CM</collectionCode>
|
||
) from
|
||
<collectingCountry id="F3725A834507FFAB818F3C2FD644FCF0" box="[151,320,837,863]" name="Venezuela" pageId="1" pageNumber="262">VENEZUELA</collectingCountry>
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||
, Est.
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||
<collectingRegion id="49A1D4F14507FFAB80983C2FD6D2FCF0" box="[384,470,837,863]" country="Venezuela" name="Bolivar" pageId="1" pageNumber="262">Bolívar</collectingRegion>
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||
, small creek in
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<location id="8EBA4CC84507FFAB83913C2FD431FCF0" LSID="urn:lsid:plazi:treatment:03CCAB054507FFAC818F3D07D7DDFE24:8EBA4CC84507FFAB83913C2FD431FCF0" box="[649,821,837,863]" country="Venezuela" latitude="5.16636" longLatPrecision="1" longitude="-60.82384" name="Kukenan Tepui" pageId="1" pageNumber="262" stateProvince="Bolivar">Kukenán Tepui</location>
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||
base camp,
|
||
<geoCoordinate id="EE517CD44507FFAB82DB3C2FD351FCF0" box="[963,1109,836,863]" degrees="05.16636" direction="north" orientation="latitude" pageId="1" pageNumber="262" precision="1" value="5.16636">N 05.16636º</geoCoordinate>
|
||
,
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||
<geoCoordinate id="EE517CD44507FFAB85783C2FD3F3FCF0" box="[1120,1271,837,863]" degrees="60.82384" direction="west" orientation="longitude" pageId="1" pageNumber="262" precision="1" value="-60.82384">W 60.82384º</geoCoordinate>
|
||
,
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||
<elevation id="0048FD204507FFAB841B3C2ED293FCF0" box="[1283,1431,836,863]" metricMagnitude="3" metricUnit="m" metricValue="1.875" pageId="1" pageNumber="262" unit="m" value="1875.0">
|
||
<quantity id="4C9DB7F64507FFAB841B3C2ED25DFCF0" box="[1283,1369,836,863]" metricMagnitude="3" metricUnit="m" metricValue="1.875" pageId="1" pageNumber="262" unit="m" value="1875.0">1875 m</quantity>
|
||
a.s.l.
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||
</elevation>
|
||
,
|
||
<date id="FFDB3CD34507FFAB818F3C02D617FC2C" box="[151,275,872,899]" pageId="1" pageNumber="262" value="2015-11-25">
|
||
<collectingDate id="EF9FC53B4507FFAB818F3C02D617FC2C" box="[151,275,872,899]" pageId="1" pageNumber="262" value="2015-11-25">25/xi/2015</collectingDate>
|
||
</date>
|
||
,
|
||
<collectionCode id="ED7482D64507FFAB80063C03D634FC2C" box="[286,304,873,899]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="262">T</collectionCode>
|
||
.
|
||
<collectorName id="26907FC54507FFAB80233C03D67BFC2C" box="[315,383,873,899]" pageId="1" pageNumber="262">Derka</collectorName>
|
||
col.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F49984507FFA981DF3CE6D693FE00" lastPageId="3" lastPageNumber="264" pageId="1" pageNumber="262" type="diagnosis">
|
||
<paragraph id="8BDA1A134507FFAB81DF3CE6D438FBD0" blockId="1.[151,1437,728,2015]" pageId="1" pageNumber="262">
|
||
<emphasis id="B911C6014507FFAB81DF3CE6D640FC08" bold="true" box="[199,324,908,935]" pageId="1" pageNumber="262">Diagnosis:</emphasis>
|
||
1) male compound eyes enlarged and divided (
|
||
<figureCitation id="135E06964507FFAB82413CE7D4C3FC08" box="[857,967,909,935]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1930,1955]" captionTargetBox="[156,1431,181,1904]" captionTargetId="figure@2.[156,1431,181,1904]" captionTargetPageId="2" captionText="FIGURES 1–4. Leptohyphes kukenan sp. nov. 1, female mature nymph, habitus; 2, female head, dorsal view; 3, male nearly mature nymph, habitus; 4, male head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3874772" httpUri="https://zenodo.org/record/3874772/files/figure.png" pageId="1" pageNumber="262">Figs. 3–4</figureCitation>
|
||
); 2) tubercles on body absent (
|
||
<figureCitation id="135E06964507FFAB84393CE7D288FC08" box="[1313,1420,909,935]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1930,1955]" captionTargetBox="[156,1431,181,1904]" captionTargetId="figure@2.[156,1431,181,1904]" captionTargetPageId="2" captionText="FIGURES 1–4. Leptohyphes kukenan sp. nov. 1, female mature nymph, habitus; 2, female head, dorsal view; 3, male nearly mature nymph, habitus; 4, male head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3874772" httpUri="https://zenodo.org/record/3874772/files/figure.png" pageId="1" pageNumber="262">Figs. 1, 3</figureCitation>
|
||
); 3) forefemur length/maximum width 2.1 (
|
||
<figureCitation id="135E06964507FFAB83763CDBD5C6FC64" box="[622,706,945,971]" captionStart="FIGURES 10–20" captionStartId="4.[151,264,1948,1973]" captionTargetBox="[154,1436,846,1923]" captionTargetId="figure@4.[151,1436,839,1923]" captionTargetPageId="4" captionText="FIGURES 10–20. Leptohyphes kukenan sp. nov. 10, foreleg, dorsal view with detail of spine; 11, fore tarsal claw; 12, hind leg with detail of spine, dorsal view; 13, schematic representation of abdominal terga setation; 14–16, operculate gill in dorsal, lateral and ventral view; 17–20, abdominal gills III–VI, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3874776" httpUri="https://zenodo.org/record/3874776/files/figure.png" pageId="1" pageNumber="262">Fig. 10</figureCitation>
|
||
); 4) fore and hind margin of middle and hind femora with spines on strong elevated sockets (
|
||
<figureCitation id="135E06964507FFAB80D73CBFD527FC40" box="[463,547,981,1007]" captionStart="FIGURES 10–20" captionStartId="4.[151,264,1948,1973]" captionTargetBox="[154,1436,846,1923]" captionTargetId="figure@4.[151,1436,839,1923]" captionTargetPageId="4" captionText="FIGURES 10–20. Leptohyphes kukenan sp. nov. 10, foreleg, dorsal view with detail of spine; 11, fore tarsal claw; 12, hind leg with detail of spine, dorsal view; 13, schematic representation of abdominal terga setation; 14–16, operculate gill in dorsal, lateral and ventral view; 17–20, abdominal gills III–VI, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3874776" httpUri="https://zenodo.org/record/3874776/files/figure.png" pageId="1" pageNumber="262">Fig. 12</figureCitation>
|
||
); 5) small pale spots (”pale pits”) cover the surface of head, thorax and femora (dorsally) and submentum (ventrally); 6) dorsum of hind femora with a mediolongitudinal ridge and a row of 16–20 minute spines (
|
||
<figureCitation id="135E06964507FFAB80583B77D696FB98" box="[320,402,1053,1079]" captionStart="FIGURES 10–20" captionStartId="4.[151,264,1948,1973]" captionTargetBox="[154,1436,846,1923]" captionTargetId="figure@4.[151,1436,839,1923]" captionTargetPageId="4" captionText="FIGURES 10–20. Leptohyphes kukenan sp. nov. 10, foreleg, dorsal view with detail of spine; 11, fore tarsal claw; 12, hind leg with detail of spine, dorsal view; 13, schematic representation of abdominal terga setation; 14–16, operculate gill in dorsal, lateral and ventral view; 17–20, abdominal gills III–VI, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3874776" httpUri="https://zenodo.org/record/3874776/files/figure.png" pageId="1" pageNumber="262">Fig. 12</figureCitation>
|
||
); 7) tarsal claws with 4–5 marginal denticles and 2 subapical denticles (
|
||
<figureCitation id="135E06964507FFAB85B43B77D3F9FB98" box="[1196,1277,1053,1079]" captionStart="FIGURES 10–20" captionStartId="4.[151,264,1948,1973]" captionTargetBox="[154,1436,846,1923]" captionTargetId="figure@4.[151,1436,839,1923]" captionTargetPageId="4" captionText="FIGURES 10–20. Leptohyphes kukenan sp. nov. 10, foreleg, dorsal view with detail of spine; 11, fore tarsal claw; 12, hind leg with detail of spine, dorsal view; 13, schematic representation of abdominal terga setation; 14–16, operculate gill in dorsal, lateral and ventral view; 17–20, abdominal gills III–VI, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3874776" httpUri="https://zenodo.org/record/3874776/files/figure.png" pageId="1" pageNumber="262">Fig. 11</figureCitation>
|
||
); 8) pronotum with large rounded lateral projection (
|
||
<figureCitation id="135E06964507FFAB83243B2BD5A3FBF4" box="[572,679,1089,1115]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1930,1955]" captionTargetBox="[156,1431,181,1904]" captionTargetId="figure@2.[156,1431,181,1904]" captionTargetPageId="2" captionText="FIGURES 1–4. Leptohyphes kukenan sp. nov. 1, female mature nymph, habitus; 2, female head, dorsal view; 3, male nearly mature nymph, habitus; 4, male head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3874772" httpUri="https://zenodo.org/record/3874772/files/figure.png" pageId="1" pageNumber="262">Figs. 1–2</figureCitation>
|
||
); 9) hind wing pads absent in female; 10) gill formula 8/7/7/4/1, gill V without ventral extension on dorsal lamella (
|
||
<figureCitation id="135E06964507FFAB83BE3B0FD42BFBD0" box="[678,815,1125,1151]" captionStart="FIGURES 10–20" captionStartId="4.[151,264,1948,1973]" captionTargetBox="[154,1436,846,1923]" captionTargetId="figure@4.[151,1436,839,1923]" captionTargetPageId="4" captionText="FIGURES 10–20. Leptohyphes kukenan sp. nov. 10, foreleg, dorsal view with detail of spine; 11, fore tarsal claw; 12, hind leg with detail of spine, dorsal view; 13, schematic representation of abdominal terga setation; 14–16, operculate gill in dorsal, lateral and ventral view; 17–20, abdominal gills III–VI, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3874776" httpUri="https://zenodo.org/record/3874776/files/figure.png" pageId="1" pageNumber="262">Figs. 13–20</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDA1A134507FFA981DF3BE3D7DAFE23" blockId="1.[151,1437,728,2015]" lastBlockId="3.[151,1437,153,431]" lastPageId="3" lastPageNumber="264" pageId="1" pageNumber="262">
|
||
<emphasis id="B911C6014507FFAB81DF3BE3D680FB0C" bold="true" box="[199,388,1161,1187]" pageId="1" pageNumber="262">Mature nymph:</emphasis>
|
||
Length of female (mm) (n=3): body, 8.9–9.1; caudal filaments, 7.0–8.0. Length of male (mm) (n=1): body, 8.5; caudal filaments (broken at base). General coloration whitish to yellowish light brown, with small pale spots on head, thorax and legs (
|
||
<figureCitation id="135E06964507FFAB832B3BBBD5A4FB44" box="[563,672,1233,1259]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1930,1955]" captionTargetBox="[156,1431,181,1904]" captionTargetId="figure@2.[156,1431,181,1904]" captionTargetPageId="2" captionText="FIGURES 1–4. Leptohyphes kukenan sp. nov. 1, female mature nymph, habitus; 2, female head, dorsal view; 3, male nearly mature nymph, habitus; 4, male head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3874772" httpUri="https://zenodo.org/record/3874772/files/figure.png" pageId="1" pageNumber="262">Figs. 1, 3</figureCitation>
|
||
). Head: shaded with black widely except anteriorly to each ocellus, on occiput forming a net-like pattern (
|
||
<figureCitation id="135E06964507FFAB835F3B9FD5AEFAA0" box="[583,682,1269,1295]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1930,1955]" captionTargetBox="[156,1431,181,1904]" captionTargetId="figure@2.[156,1431,181,1904]" captionTargetPageId="2" captionText="FIGURES 1–4. Leptohyphes kukenan sp. nov. 1, female mature nymph, habitus; 2, female head, dorsal view; 3, male nearly mature nymph, habitus; 4, male head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3874772" httpUri="https://zenodo.org/record/3874772/files/figure.png" pageId="1" pageNumber="262">Fig. 2, 4</figureCitation>
|
||
); and with small pale spots. Antennae yellowish white. Compound eyes of female rounded and small as usual, male eyes enlarged and divided in two portions (
|
||
<figureCitation id="135E06964507FFAB85883A73D3FFFA9C" box="[1168,1275,1305,1331]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1930,1955]" captionTargetBox="[156,1431,181,1904]" captionTargetId="figure@2.[156,1431,181,1904]" captionTargetPageId="2" captionText="FIGURES 1–4. Leptohyphes kukenan sp. nov. 1, female mature nymph, habitus; 2, female head, dorsal view; 3, male nearly mature nymph, habitus; 4, male head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3874772" httpUri="https://zenodo.org/record/3874772/files/figure.png" pageId="1" pageNumber="262">Figs. 3–4</figureCitation>
|
||
), line between both parts hardly visible. Mouthparts (
|
||
<figureCitation id="135E06964507FFAB83493A57D5C5FAF8" box="[593,705,1340,1367]" captionStart="FIGURES 5–9" captionStartId="3.[151,264,1886,1911]" captionTargetBox="[169,1426,453,1857]" captionTargetId="figure@3.[151,1436,453,1861]" captionTargetPageId="3" captionText="FIGURES 5–9. Leptohyphes kukenan sp. nov., mouthparts. 5, labrum dorsal view (setae on right side omitted), with detail of bifid microsetae on fore margin; 6, left mandible with detail of prostheca, dorsal view;; 7, right mandible with detail of prostheca, dorsal view; 8, maxilla, ventral view of distal part; 9, labium, ventral view with detail of microsetae and pits." figureDoi="http://doi.org/10.5281/zenodo.3874774" httpUri="https://zenodo.org/record/3874774/files/figure.png" pageId="1" pageNumber="262">Figs. 5–9</figureCitation>
|
||
): labrum with single long setae dorsally, some of them forming a transverse submedian row (
|
||
<figureCitation id="135E06964507FFAB80F03A0BD52BFAD4" box="[488,559,1376,1403]" captionStart="FIGURES 5–9" captionStartId="3.[151,264,1886,1911]" captionTargetBox="[169,1426,453,1857]" captionTargetId="figure@3.[151,1436,453,1861]" captionTargetPageId="3" captionText="FIGURES 5–9. Leptohyphes kukenan sp. nov., mouthparts. 5, labrum dorsal view (setae on right side omitted), with detail of bifid microsetae on fore margin; 6, left mandible with detail of prostheca, dorsal view;; 7, right mandible with detail of prostheca, dorsal view; 8, maxilla, ventral view of distal part; 9, labium, ventral view with detail of microsetae and pits." figureDoi="http://doi.org/10.5281/zenodo.3874774" httpUri="https://zenodo.org/record/3874774/files/figure.png" pageId="1" pageNumber="262">Fig. 5</figureCitation>
|
||
); anteromedian emargination shallow, with a row of bifid micro-setae along ventral edge of fore margin (
|
||
<figureCitation id="135E06964507FFAB80CF3AEFD51FFA30" box="[471,539,1412,1439]" captionStart="FIGURES 5–9" captionStartId="3.[151,264,1886,1911]" captionTargetBox="[169,1426,453,1857]" captionTargetId="figure@3.[151,1436,453,1861]" captionTargetPageId="3" captionText="FIGURES 5–9. Leptohyphes kukenan sp. nov., mouthparts. 5, labrum dorsal view (setae on right side omitted), with detail of bifid microsetae on fore margin; 6, left mandible with detail of prostheca, dorsal view;; 7, right mandible with detail of prostheca, dorsal view; 8, maxilla, ventral view of distal part; 9, labium, ventral view with detail of microsetae and pits." figureDoi="http://doi.org/10.5281/zenodo.3874774" httpUri="https://zenodo.org/record/3874774/files/figure.png" pageId="1" pageNumber="262">Fig. 5</figureCitation>
|
||
, detail); hypopharynx with rounded small superlinguae and lingua; maxilla with well-developed 3-segmented palpus (
|
||
<figureCitation id="135E06964507FFAB832D3AC3D57CFA6C" box="[565,632,1449,1475]" captionStart="FIGURES 5–9" captionStartId="3.[151,264,1886,1911]" captionTargetBox="[169,1426,453,1857]" captionTargetId="figure@3.[151,1436,453,1861]" captionTargetPageId="3" captionText="FIGURES 5–9. Leptohyphes kukenan sp. nov., mouthparts. 5, labrum dorsal view (setae on right side omitted), with detail of bifid microsetae on fore margin; 6, left mandible with detail of prostheca, dorsal view;; 7, right mandible with detail of prostheca, dorsal view; 8, maxilla, ventral view of distal part; 9, labium, ventral view with detail of microsetae and pits." figureDoi="http://doi.org/10.5281/zenodo.3874774" httpUri="https://zenodo.org/record/3874774/files/figure.png" pageId="1" pageNumber="262">Fig. 8</figureCitation>
|
||
); mandibles relatively stout (
|
||
<figureCitation id="135E06964507FFAB82A23AC3D322FA6B" box="[954,1062,1449,1476]" captionStart="FIGURES 5–9" captionStartId="3.[151,264,1886,1911]" captionTargetBox="[169,1426,453,1857]" captionTargetId="figure@3.[151,1436,453,1861]" captionTargetPageId="3" captionText="FIGURES 5–9. Leptohyphes kukenan sp. nov., mouthparts. 5, labrum dorsal view (setae on right side omitted), with detail of bifid microsetae on fore margin; 6, left mandible with detail of prostheca, dorsal view;; 7, right mandible with detail of prostheca, dorsal view; 8, maxilla, ventral view of distal part; 9, labium, ventral view with detail of microsetae and pits." figureDoi="http://doi.org/10.5281/zenodo.3874774" httpUri="https://zenodo.org/record/3874774/files/figure.png" pageId="1" pageNumber="262">Figs. 6–7</figureCitation>
|
||
); labium with large and elongated submentum, completely covered with pale spots ventrally (
|
||
<figureCitation id="135E06964507FFAB823C3AA7D463FA48" box="[804,871,1485,1511]" captionStart="FIGURES 5–9" captionStartId="3.[151,264,1886,1911]" captionTargetBox="[169,1426,453,1857]" captionTargetId="figure@3.[151,1436,453,1861]" captionTargetPageId="3" captionText="FIGURES 5–9. Leptohyphes kukenan sp. nov., mouthparts. 5, labrum dorsal view (setae on right side omitted), with detail of bifid microsetae on fore margin; 6, left mandible with detail of prostheca, dorsal view;; 7, right mandible with detail of prostheca, dorsal view; 8, maxilla, ventral view of distal part; 9, labium, ventral view with detail of microsetae and pits." figureDoi="http://doi.org/10.5281/zenodo.3874774" httpUri="https://zenodo.org/record/3874774/files/figure.png" pageId="1" pageNumber="262">Fig. 9</figureCitation>
|
||
). Thorax. Nota brownish shaded with black widely except median area paler, wingbuds grayish on costal margin; sterna paler. Pronotum with rounded lateral projection (
|
||
<figureCitation id="135E06964507FFAB81CD397FD641F980" box="[213,325,1557,1583]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1930,1955]" captionTargetBox="[156,1431,181,1904]" captionTargetId="figure@2.[156,1431,181,1904]" captionTargetPageId="2" captionText="FIGURES 1–4. Leptohyphes kukenan sp. nov. 1, female mature nymph, habitus; 2, female head, dorsal view; 3, male nearly mature nymph, habitus; 4, male head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3874772" httpUri="https://zenodo.org/record/3874772/files/figure.png" pageId="1" pageNumber="262">Figs. 1, 3</figureCitation>
|
||
). Mesonotum with subtriangular blunt anterolateral projection (
|
||
<figureCitation id="135E06964507FFAB8539397FD395F980" box="[1057,1169,1557,1583]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1930,1955]" captionTargetBox="[156,1431,181,1904]" captionTargetId="figure@2.[156,1431,181,1904]" captionTargetPageId="2" captionText="FIGURES 1–4. Leptohyphes kukenan sp. nov. 1, female mature nymph, habitus; 2, female head, dorsal view; 3, male nearly mature nymph, habitus; 4, male head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3874772" httpUri="https://zenodo.org/record/3874772/files/figure.png" pageId="1" pageNumber="262">Figs. 1, 3</figureCitation>
|
||
). Hind wingpad absent in female. Legs yellowish slightly shaded gray dorsally except mediolongitudinal pale line on middle of femora, dorsum of all femora completely covered by small pale spots. Ratio length fore femur/hind femur = 0.55. Foreleg (
|
||
<figureCitation id="135E06964507FFAB81B839EBD7F0F934" box="[160,244,1665,1691]" captionStart="FIGURES 10–20" captionStartId="4.[151,264,1948,1973]" captionTargetBox="[154,1436,846,1923]" captionTargetId="figure@4.[151,1436,839,1923]" captionTargetPageId="4" captionText="FIGURES 10–20. Leptohyphes kukenan sp. nov. 10, foreleg, dorsal view with detail of spine; 11, fore tarsal claw; 12, hind leg with detail of spine, dorsal view; 13, schematic representation of abdominal terga setation; 14–16, operculate gill in dorsal, lateral and ventral view; 17–20, abdominal gills III–VI, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3874776" httpUri="https://zenodo.org/record/3874776/files/figure.png" pageId="1" pageNumber="262">Fig. 10</figureCitation>
|
||
): ratio femur length/maximum width 2.1; transversal row at 0.6 from base to apex; fore margin with small spines, hind margin with fringed rounded spines (
|
||
<figureCitation id="135E06964507FFAB83DE39CFD41DF910" box="[710,793,1701,1727]" captionStart="FIGURES 10–20" captionStartId="4.[151,264,1948,1973]" captionTargetBox="[154,1436,846,1923]" captionTargetId="figure@4.[151,1436,839,1923]" captionTargetPageId="4" captionText="FIGURES 10–20. Leptohyphes kukenan sp. nov. 10, foreleg, dorsal view with detail of spine; 11, fore tarsal claw; 12, hind leg with detail of spine, dorsal view; 13, schematic representation of abdominal terga setation; 14–16, operculate gill in dorsal, lateral and ventral view; 17–20, abdominal gills III–VI, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3874776" httpUri="https://zenodo.org/record/3874776/files/figure.png" pageId="1" pageNumber="262">Fig. 10</figureCitation>
|
||
, detail) distally to transversal row, all spines mounted on elevated sockets. Foretibia slightly longer than femur, with tiny spines along inner margin and a longitudinal ridge along outer margin. Tarsus 1/2 the length of tibia with row of setae in inner margin; tarsal claw with five marginal denticles, and 2 subapical submarginal denticles (
|
||
<figureCitation id="135E06964507FFAB83DB387BD410F884" box="[707,788,1809,1835]" captionStart="FIGURES 10–20" captionStartId="4.[151,264,1948,1973]" captionTargetBox="[154,1436,846,1923]" captionTargetId="figure@4.[151,1436,839,1923]" captionTargetPageId="4" captionText="FIGURES 10–20. Leptohyphes kukenan sp. nov. 10, foreleg, dorsal view with detail of spine; 11, fore tarsal claw; 12, hind leg with detail of spine, dorsal view; 13, schematic representation of abdominal terga setation; 14–16, operculate gill in dorsal, lateral and ventral view; 17–20, abdominal gills III–VI, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3874776" httpUri="https://zenodo.org/record/3874776/files/figure.png" pageId="1" pageNumber="262">Fig. 11</figureCitation>
|
||
). Middle and hind legs (
|
||
<figureCitation id="135E06964507FFAB853F387BD37DF884" box="[1063,1145,1809,1835]" captionStart="FIGURES 10–20" captionStartId="4.[151,264,1948,1973]" captionTargetBox="[154,1436,846,1923]" captionTargetId="figure@4.[151,1436,839,1923]" captionTargetPageId="4" captionText="FIGURES 10–20. Leptohyphes kukenan sp. nov. 10, foreleg, dorsal view with detail of spine; 11, fore tarsal claw; 12, hind leg with detail of spine, dorsal view; 13, schematic representation of abdominal terga setation; 14–16, operculate gill in dorsal, lateral and ventral view; 17–20, abdominal gills III–VI, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3874776" httpUri="https://zenodo.org/record/3874776/files/figure.png" pageId="1" pageNumber="262">Fig. 12</figureCitation>
|
||
) similar in form and setation to each other (middle leg smaller). Ratio hind femur length/maximum width 2.0; fore margin with row of tiny rounded spines mounted on large sockets (on basal half this row is double), hind margin with 38–40 rounded small spines on very elevated sockets (see detail in
|
||
<figureCitation id="135E06964507FFAB83B13817D5FAF838" box="[681,766,1917,1943]" captionStart="FIGURES 10–20" captionStartId="4.[151,264,1948,1973]" captionTargetBox="[154,1436,846,1923]" captionTargetId="figure@4.[151,1436,839,1923]" captionTargetPageId="4" captionText="FIGURES 10–20. Leptohyphes kukenan sp. nov. 10, foreleg, dorsal view with detail of spine; 11, fore tarsal claw; 12, hind leg with detail of spine, dorsal view; 13, schematic representation of abdominal terga setation; 14–16, operculate gill in dorsal, lateral and ventral view; 17–20, abdominal gills III–VI, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3874776" httpUri="https://zenodo.org/record/3874776/files/figure.png" pageId="1" pageNumber="262">Fig. 12</figureCitation>
|
||
), dorsal surface with longitudinal ridge covered by 18–20 small spines. Tibia slightly longer than femur, with dorsal ridge, inner and outer margins with spines, those on hind margin stronger and mounted on elevated sockets. Tarsus 0.25 the length of tibia, inner margin with setae, tarsal claw as in foreleg. Abdomen (
|
||
<figureCitation id="135E06964505FFA980EB3FF3D565FF1C" box="[499,609,153,179]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1930,1955]" captionTargetBox="[156,1431,181,1904]" captionTargetId="figure@2.[156,1431,181,1904]" captionTargetPageId="2" captionText="FIGURES 1–4. Leptohyphes kukenan sp. nov. 1, female mature nymph, habitus; 2, female head, dorsal view; 3, male nearly mature nymph, habitus; 4, male head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3874772" httpUri="https://zenodo.org/record/3874772/files/figure.png" pageId="3" pageNumber="264">Figs. 2, 3</figureCitation>
|
||
,
|
||
<figureCitation id="135E06964505FFA983683FF3D588FF1C" box="[624,652,153,179]" captionStart="FIGURES 10–20" captionStartId="4.[151,264,1948,1973]" captionTargetBox="[154,1436,846,1923]" captionTargetId="figure@4.[151,1436,839,1923]" captionTargetPageId="4" captionText="FIGURES 10–20. Leptohyphes kukenan sp. nov. 10, foreleg, dorsal view with detail of spine; 11, fore tarsal claw; 12, hind leg with detail of spine, dorsal view; 13, schematic representation of abdominal terga setation; 14–16, operculate gill in dorsal, lateral and ventral view; 17–20, abdominal gills III–VI, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3874776" httpUri="https://zenodo.org/record/3874776/files/figure.png" pageId="3" pageNumber="264">13</figureCitation>
|
||
) yellowish diffusely shaded with gray on terga as in
|
||
<figureCitation id="135E06964505FFA985EB3FF3D253FF1C" box="[1267,1367,153,179]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1930,1955]" captionTargetBox="[156,1431,181,1904]" captionTargetId="figure@2.[156,1431,181,1904]" captionTargetPageId="2" captionText="FIGURES 1–4. Leptohyphes kukenan sp. nov. 1, female mature nymph, habitus; 2, female head, dorsal view; 3, male nearly mature nymph, habitus; 4, male head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3874772" httpUri="https://zenodo.org/record/3874772/files/figure.png" pageId="3" pageNumber="264">Figure 2</figureCitation>
|
||
; terga glabrous, except rounded small paired spines submedially on terga VI–IX (
|
||
<figureCitation id="135E06964505FFA982CC3FD7D322FF78" box="[980,1062,189,215]" captionStart="FIGURES 10–20" captionStartId="4.[151,264,1948,1973]" captionTargetBox="[154,1436,846,1923]" captionTargetId="figure@4.[151,1436,839,1923]" captionTargetPageId="4" captionText="FIGURES 10–20. Leptohyphes kukenan sp. nov. 10, foreleg, dorsal view with detail of spine; 11, fore tarsal claw; 12, hind leg with detail of spine, dorsal view; 13, schematic representation of abdominal terga setation; 14–16, operculate gill in dorsal, lateral and ventral view; 17–20, abdominal gills III–VI, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3874776" httpUri="https://zenodo.org/record/3874776/files/figure.png" pageId="3" pageNumber="264">Fig. 13</figureCitation>
|
||
); segments III–VI forming lateral flanges; small posterolateral projections present on VII–IX. Gills (
|
||
<figureCitation id="135E06964505FFA982633F8BD301FF54" box="[891,1029,225,251]" captionStart="FIGURES 10–20" captionStartId="4.[151,264,1948,1973]" captionTargetBox="[154,1436,846,1923]" captionTargetId="figure@4.[151,1436,839,1923]" captionTargetPageId="4" captionText="FIGURES 10–20. Leptohyphes kukenan sp. nov. 10, foreleg, dorsal view with detail of spine; 11, fore tarsal claw; 12, hind leg with detail of spine, dorsal view; 13, schematic representation of abdominal terga setation; 14–16, operculate gill in dorsal, lateral and ventral view; 17–20, abdominal gills III–VI, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3874776" httpUri="https://zenodo.org/record/3874776/files/figure.png" pageId="3" pageNumber="264">Figs. 14–20</figureCitation>
|
||
): operculate gill light brownish-gray with whitish margins, ventrally with membranous and relatively short ”basal spine” and two fringed lamellae (
|
||
<figureCitation id="135E06964505FFA9847C3E6FD7E6FEEB" captionStart="FIGURES 10–20" captionStartId="4.[151,264,1948,1973]" captionTargetBox="[154,1436,846,1923]" captionTargetId="figure@4.[151,1436,839,1923]" captionTargetPageId="4" captionText="FIGURES 10–20. Leptohyphes kukenan sp. nov. 10, foreleg, dorsal view with detail of spine; 11, fore tarsal claw; 12, hind leg with detail of spine, dorsal view; 13, schematic representation of abdominal terga setation; 14–16, operculate gill in dorsal, lateral and ventral view; 17–20, abdominal gills III–VI, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3874776" httpUri="https://zenodo.org/record/3874776/files/figure.png" pageId="3" pageNumber="264">Figs. 14–16</figureCitation>
|
||
); other gills grayish to hyaline, gills III–IV also with fringed lamellae (
|
||
<figureCitation id="135E06964505FFA985183E43D38DFEEC" box="[1024,1161,297,323]" captionStart="FIGURES 10–20" captionStartId="4.[151,264,1948,1973]" captionTargetBox="[154,1436,846,1923]" captionTargetId="figure@4.[151,1436,839,1923]" captionTargetPageId="4" captionText="FIGURES 10–20. Leptohyphes kukenan sp. nov. 10, foreleg, dorsal view with detail of spine; 11, fore tarsal claw; 12, hind leg with detail of spine, dorsal view; 13, schematic representation of abdominal terga setation; 14–16, operculate gill in dorsal, lateral and ventral view; 17–20, abdominal gills III–VI, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3874776" httpUri="https://zenodo.org/record/3874776/files/figure.png" pageId="3" pageNumber="264">Figs. 17–18</figureCitation>
|
||
). Gill formula 8/7/7/4/1. Caudal filaments with whorl of short setae at joinings, with darker annuli at base on females (broken off and lost in male).
|
||
</paragraph>
|
||
<caption id="DF1A4A9B4504FFA8818F38E0D592F868" ID-DOI="http://doi.org/10.5281/zenodo.3874772" ID-Zenodo-Dep="3874772" httpUri="https://zenodo.org/record/3874772/files/figure.png" pageId="2" pageNumber="263" startId="2.[151,264,1930,1955]" targetBox="[156,1431,181,1904]" targetPageId="2">
|
||
<paragraph id="8BDA1A134504FFA8818F38E0D592F868" blockId="2.[151,1436,1930,1991]" pageId="2" pageNumber="263">
|
||
<emphasis id="B911C6014504FFA8818F38E0D638F80C" bold="true" box="[151,316,1930,1955]" pageId="2" pageNumber="263">FIGURES 1–4.</emphasis>
|
||
<taxonomicName id="4C6561904504FFA8805B38E1D51BF80C" authority="Molineri & Derka, 2020" authorityName="Molineri & Derka" authorityYear="2020" box="[323,543,1931,1955]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="kukenan" status="sp. nov.">
|
||
<emphasis id="B911C6014504FFA8805B38E1D51BF80C" box="[323,543,1931,1955]" italics="true" pageId="2" pageNumber="263">Leptohyphes kukenan</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911C6014504FFA8833E38E1D57CF80C" bold="true" box="[550,632,1931,1955]" pageId="2" pageNumber="263">
|
||
<taxonomicNameLabel id="A2227B7A4504FFA8833E38E1D57CF80C" box="[550,632,1931,1955]" pageId="2" pageNumber="263" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
1, female mature nymph, habitus; 2, female head, dorsal view; 3, male nearly mature nymph, habitus; 4, male head, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDA1A134505FFA981DF3EFED693FE00" blockId="3.[151,1437,153,431]" box="[199,407,404,431]" pageId="3" pageNumber="264">
|
||
<emphasis id="B911C6014505FFA981DF3EFED624FE00" bold="true" box="[199,288,404,431]" pageId="3" pageNumber="264">Adults:</emphasis>
|
||
unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF1A4A9B4505FFA9818F3834D3E3F810" ID-DOI="http://doi.org/10.5281/zenodo.3874774" ID-Zenodo-Dep="3874774" httpUri="https://zenodo.org/record/3874774/files/figure.png" pageId="3" pageNumber="264" startId="3.[151,264,1886,1911]" targetBox="[169,1426,453,1857]" targetPageId="3">
|
||
<paragraph id="8BDA1A134505FFA9818F3834D3E3F810" blockId="3.[151,1436,1886,1983]" pageId="3" pageNumber="264">
|
||
<emphasis id="B911C6014505FFA9818F3834D63DF8D8" bold="true" box="[151,313,1886,1911]" pageId="3" pageNumber="264">FIGURES 5–9.</emphasis>
|
||
<taxonomicName id="4C6561904505FFA980263835D513F8D8" authority="Molineri & Derka, 2020" authorityName="Molineri & Derka" authorityYear="2020" box="[318,535,1887,1911]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="kukenan" status="sp. nov.">
|
||
<emphasis id="B911C6014505FFA980263835D513F8D8" box="[318,535,1887,1911]" italics="true" pageId="3" pageNumber="264">Leptohyphes kukenan</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911C6014505FFA983053835D568F8D8" bold="true" box="[541,620,1887,1911]" pageId="3" pageNumber="264">
|
||
<taxonomicNameLabel id="A2227B7A4505FFA983053835D568F8D8" box="[541,620,1887,1911]" pageId="3" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, mouthparts. 5, labrum dorsal view (setae on right side omitted), with detail of bifid microsetae on fore margin; 6, left mandible with detail of prostheca, dorsal view;; 7, right mandible with detail of prostheca, dorsal view; 8, maxilla, ventral view of distal part; 9, labium, ventral view with detail of microsetae and pits.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37F49984502FFAE81DF3FF3D661FF78" pageId="4" pageNumber="265" type="etymology">
|
||
<paragraph id="8BDA1A134502FFAE81DF3FF3D661FF78" blockId="4.[151,1437,153,821]" pageId="4" pageNumber="265">
|
||
<emphasis id="B911C6014502FFAE81DF3FF3D64CFF1C" bold="true" box="[199,328,153,179]" pageId="4" pageNumber="265">Etymology</emphasis>
|
||
: The species is named after Kukenán tepui, a table mountain in southeastern
|
||
<collectingCountry id="F3725A834502FFAE85AA3FF3D22CFF1C" box="[1202,1320,153,179]" name="Venezuela" pageId="4" pageNumber="265">Venezuela</collectingCountry>
|
||
, on which talus it was found.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F49984502FFAE81DF3F8AD3C4FF54" box="[199,1216,224,251]" pageId="4" pageNumber="265" type="distribution">
|
||
<paragraph id="8BDA1A134502FFAE81DF3F8AD3C4FF54" blockId="4.[151,1437,153,821]" box="[199,1216,224,251]" pageId="4" pageNumber="265">
|
||
<emphasis id="B911C6014502FFAE81DF3F8AD665FF54" bold="true" box="[199,353,224,251]" pageId="4" pageNumber="265">Distribution.</emphasis>
|
||
Only known from Kukenán tepui (
|
||
<figureCitation id="135E06964502FFAE83F23F8BD477FF54" box="[746,883,225,251]" captionStart="FIGURES 21–22" captionStartId="5.[151,264,1969,1994]" captionTargetBox="[206,1381,181,1943]" captionTargetId="figure@5.[206,1381,181,1943]" captionTargetPageId="5" captionText="FIGURES 21–22. 21, Landscape of Kukenán tepui foothills where sampling localities are situated; 22, type locality. Photographs by Daniel Grul’a." figureDoi="http://doi.org/10.5281/zenodo.3874778" httpUri="https://zenodo.org/record/3874778/files/figure.png" pageId="4" pageNumber="265">Figs. 21–22</figureCitation>
|
||
), Pantepui region,
|
||
<collectingCountry id="F3725A834502FFAE855D3F8BD3B8FF54" box="[1093,1212,225,251]" name="Venezuela" pageId="4" pageNumber="265">Venezuela</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F49984502FFAC81DF3E6ED7DDFE24" lastPageId="6" lastPageNumber="267" pageId="4" pageNumber="265" type="discussion">
|
||
<paragraph id="8BDA1A134502FFAE81DF3E6ED26CFC9B" blockId="4.[151,1437,153,821]" pageId="4" pageNumber="265">
|
||
<emphasis id="B911C6014502FFAE81DF3E6ED64BFEB0" bold="true" box="[199,335,260,287]" pageId="4" pageNumber="265">Discussion:</emphasis>
|
||
<taxonomicName id="4C6561904502FFAE804E3E6ED543FEB1" authority="Molineri & Derka, 2020" authorityName="Molineri & Derka" authorityYear="2020" box="[342,583,260,286]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="kukenan" status="sp. nov.">
|
||
<emphasis id="B911C6014502FFAE804E3E6ED543FEB1" box="[342,583,260,286]" italics="true" pageId="4" pageNumber="265">Leptohyphes kukenan</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911C6014502FFAE83573E6FD5ACFEB0" bold="true" box="[591,680,261,287]" pageId="4" pageNumber="265">
|
||
<taxonomicNameLabel id="A2227B7A4502FFAE83573E6FD5ACFEB0" box="[591,680,261,287]" pageId="4" pageNumber="265" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from all its congeners by the list of characters given in the diagnosis. It is similar to
|
||
<taxonomicName id="4C6561904502FFAE83263E43D44DFEED" authority="Allen (1973)" authorityName="Allen" authorityYear="1973" box="[574,841,296,322]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="populus">
|
||
<emphasis id="B911C6014502FFAE83263E43D5B1FEED" box="[574,693,296,322]" italics="true" pageId="4" pageNumber="265">L. populus</emphasis>
|
||
<bibRefCitation id="EFF467E24502FFAE83A23E42D44DFEED" author="Allen, R. K." box="[698,841,296,322]" pageId="4" pageNumber="265" pagination="363 - 372" refId="ref2760" refString="Allen, R. K. (1973) New species of Leptohyphes Eaton (Ephemeroptera: Tricorythidae). Pan- Pacific Entomologist, 49, 363 - 372." type="journal article" year="1973">Allen (1973)</bibRefCitation>
|
||
</taxonomicName>
|
||
because of the uncommon division of male eyes in a dorsal and a ventral portion. Other characters common to both species may also appear in other species of the genus, for example the large elevated sockets on femoral spines. The new species can be separated from
|
||
<taxonomicName id="4C6561904502FFAE85E03E1AD274FE26" authorityName="Allen" authorityYear="1973" box="[1272,1392,367,393]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="populus">
|
||
<emphasis id="B911C6014502FFAE85E03E1AD274FE26" box="[1272,1392,367,393]" italics="true" pageId="4" pageNumber="265">L. populus</emphasis>
|
||
</taxonomicName>
|
||
because the spines on femora and tibia of the new species are relatively smaller and fewer; the form of the submentum (longer in the new species), and the paired subapical denticles on tarsal claws (absent in
|
||
<taxonomicName id="4C6561904502FFAE85993EDDD3FDFE7F" authorityName="Allen" authorityYear="1973" box="[1153,1273,438,464]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="populus">
|
||
<emphasis id="B911C6014502FFAE85993EDDD3FDFE7F" box="[1153,1273,438,464]" italics="true" pageId="4" pageNumber="265">L. populus</emphasis>
|
||
</taxonomicName>
|
||
). Gills are not described for
|
||
<taxonomicName id="4C6561904502FFAE80373EB0D6A1FE5C" authorityName="Allen" authorityYear="1973" box="[303,421,473,499]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="populus">
|
||
<emphasis id="B911C6014502FFAE80373EB0D6A1FE5C" box="[303,421,473,499]" italics="true" pageId="4" pageNumber="265">L. populus</emphasis>
|
||
</taxonomicName>
|
||
, but in the new species, although preserving the general shape and aspect of the genus, they are different in their structure from other
|
||
<taxonomicName id="4C6561904502FFAE83743E97D5FEFDB8" authorityName="Eaton" authorityYear="1882" box="[620,762,509,535]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B911C6014502FFAE83743E97D5FEFDB8" box="[620,762,509,535]" italics="true" pageId="4" pageNumber="265">Leptohyphes</emphasis>
|
||
</taxonomicName>
|
||
: smaller ventral lobes arise consecutively from larger ones, so they are imbricated. This pattern (as well as the divided eyes) is plesiomorphic in
|
||
<taxonomicName id="4C6561904502FFAE85483D4AD3FFFD95" box="[1104,1275,544,570]" class="Insecta" family="Leptohyphidae" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="family">Leptohyphidae</taxonomicName>
|
||
, being present only in
|
||
<taxonomicName id="4C6561904502FFAE81E83D2ED5B2FDF1" authority="Salles & Molineri (2006)" authorityName="Salles & Molineri" authorityYear="2006" box="[240,694,579,606]" class="Insecta" family="Leptohyphidae" genus="Amanahyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B911C6014502FFAE81E83D2ED689FDF1" box="[240,397,580,606]" italics="true" pageId="4" pageNumber="265">Amanahyphes</emphasis>
|
||
<bibRefCitation id="EFF467E24502FFAE808D3D29D5B2FDF1" author="Salles, F. F. & Molineri, C." box="[405,694,579,606]" pageId="4" pageNumber="265" pagination="1 - 12" refId="ref3594" refString="Salles, F. F. & Molineri, C. (2006) Amanahyphes saguassu, a new genus and species of Leptohyphidae (Ephemeroptera: Ephemerelloidea) from northern Brazil. Aquatic Insects, 28 (1), 1 - 12. https: // doi. org / 10.1080 / 13682820500343180" type="journal article" year="2006">Salles & Molineri (2006)</bibRefCitation>
|
||
</taxonomicName>
|
||
, and also in related families
|
||
<taxonomicName id="4C6561904502FFAE82E73D29D3CDFDF2" box="[1023,1225,579,605]" class="Insecta" family="Melanemerellidae" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="family">Melanemerellidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6561904502FFAE841A3D29D654FD2D" authority="(Molineri 2006)" baseAuthorityName="Molineri" baseAuthorityYear="2006" class="Insecta" family="Coryphoridae" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="family">
|
||
Coryphoridae (
|
||
<bibRefCitation id="EFF467E24502FFAE81873D0DD64CFD2D" author="Molineri, C. & Zuniga, M. C." box="[159,328,615,642]" pageId="4" pageNumber="265" pagination="139 - 151" refId="ref3401" refString="Molineri, C. & Zuniga, M. C. (2006) New species of Leptophyphidae (Insecta: Ephemeroptera) from Colombia with evidence of reproductive time segregation. Studies on Neotropical Fauna and Environment, 41, 139 - 151. https: // doi. org / 10.1080 / 0165020500210891" type="journal article" year="2006">Molineri 2006</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Another character that can be interpreted as plesiomorphic in the new species is the presence of paired subapical denticles on the tarsal claws, which is widespread in the family but in
|
||
<taxonomicName id="4C6561904502FFAE859E3DE0D210FD0B" authorityName="Eaton" authorityYear="1882" box="[1158,1300,650,676]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B911C6014502FFAE859E3DE0D210FD0B" box="[1158,1300,650,676]" italics="true" pageId="4" pageNumber="265">Leptohyphes</emphasis>
|
||
</taxonomicName>
|
||
it is almost always single and asymmetrical (
|
||
<bibRefCitation id="EFF467E24502FFAE830A3DC4D5BDFD66" author="Molineri, C." box="[530,697,686,713]" pageId="4" pageNumber="265" pagination="47 - 70" refId="ref3304" refString="Molineri, C. (2003) Revision of the South American species of Leptohyphes Eaton (Ephemeroptera: Leptohyphidae) with a key to the nymphs. Studies on Neotropical Fauna and Environment, 38 (1), 47 - 70. https: // doi. org / 10.1076 / snfe. 38.1.47.14031" type="journal article" year="2003">Molineri 2003</bibRefCitation>
|
||
, 2006). Small pale spots have been described in other species of
|
||
<taxonomicName id="4C6561904502FFAE818F3DB8D621FD43" authorityName="Eaton" authorityYear="1882" box="[151,293,722,748]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B911C6014502FFAE818F3DB8D621FD43" box="[151,293,722,748]" italics="true" pageId="4" pageNumber="265">Leptohyphes</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<taxonomicName id="4C6561904502FFAE802A3DB9D55EFD42" authority="Wang et al. (1998)" authorityName="Wang" authorityYear="1998" box="[306,602,722,749]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="liniti">
|
||
<emphasis id="B911C6014502FFAE802A3DB9D680FD43" box="[306,388,722,748]" italics="true" pageId="4" pageNumber="265">L. liniti</emphasis>
|
||
<bibRefCitation id="EFF467E24502FFAE80913DB9D55EFD42" author="Wang, T. Q. & Sites, R. W. & McCafferty, W. P." box="[393,602,722,749]" pageId="4" pageNumber="265" pagination="68 - 75" refId="ref3676" refString="Wang, T. Q., Sites, R. W. & McCafferty, W. P. (1998) Two new species of Leptohyphes (Ephemeroptera: Leptohyphidae) from Ecuador. Florida Entomologist, 81, 68 - 75. https: // doi. org / 10.2307 / 3495997" type="journal article" year="1998">
|
||
Wang
|
||
<emphasis id="B911C6014502FFAE80C83DB9D504FD43" box="[464,512,722,748]" italics="true" pageId="4" pageNumber="265">et al</emphasis>
|
||
. (1998)
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6561904502FFAE837D3DB9D467FD42" authority="Allen (1967)" authorityName="Allen" authorityYear="1967" box="[613,867,722,749]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="petersi">
|
||
<emphasis id="B911C6014502FFAE837D3DB9D5C9FD43" box="[613,717,723,748]" italics="true" pageId="4" pageNumber="265">L. petersi</emphasis>
|
||
<bibRefCitation id="EFF467E24502FFAE83CA3DB8D467FD42" author="Allen, R. K." box="[722,867,722,749]" pageId="4" pageNumber="265" pagination="350 - 375" refId="ref2713" refString="Allen, R. K. (1967) New species of new world Leptohyphinae (Ephemeroptera: Tricorythidae). The Canadian Entomologist, 99 (4), 350 - 375. https: // doi. org / 10.4039 / Ent 99350 - 4" type="journal article" year="1967">Allen (1967)</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6561904502FFAE82763DB9D38EFD42" authority="Allen (1967)" authorityName="Allen" authorityYear="1967" box="[878,1162,722,749]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="murdochi">
|
||
<emphasis id="B911C6014502FFAE82763DB9D4F0FD43" box="[878,1012,722,748]" italics="true" pageId="4" pageNumber="265">L. murdochi</emphasis>
|
||
<bibRefCitation id="EFF467E24502FFAE82E13DB8D38EFD42" author="Allen, R. K." box="[1017,1162,722,749]" pageId="4" pageNumber="265" pagination="350 - 375" refId="ref2713" refString="Allen, R. K. (1967) New species of new world Leptohyphinae (Ephemeroptera: Tricorythidae). The Canadian Entomologist, 99 (4), 350 - 375. https: // doi. org / 10.4039 / Ent 99350 - 4" type="journal article" year="1967">Allen (1967)</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6561904502FFAE858D3DB9D29CFD42" authority="Brusca (1971)" authorityName="Brusca" authorityYear="1971" box="[1173,1432,722,749]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="alleni">
|
||
<emphasis id="B911C6014502FFAE858D3DB9D3F5FD43" box="[1173,1265,722,748]" italics="true" pageId="4" pageNumber="265">L. alleni</emphasis>
|
||
<bibRefCitation id="EFF467E24502FFAE85EF3DB9D29CFD42" author="Brusca, R. C." box="[1271,1432,723,749]" pageId="4" pageNumber="265" pagination="146 - 148" refId="ref2846" refString="Brusca, R. C. (1971) A new species of Leptohyphes (Epemeroptera: Tricorythidae) from Mexico Pan-Pacific Entomologist, 47, 146 - 148." type="journal article" year="1971">Brusca (1971)</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C6561904502FFAE81D03D9DD531FCBE" authority="Molineri et al. (2016)" authorityName="Molineri" authorityYear="2016" box="[200,565,758,785]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="andina">
|
||
<emphasis id="B911C6014502FFAE81D03D9DD630FCBF" box="[200,308,758,784]" italics="true" pageId="4" pageNumber="265">L. andina</emphasis>
|
||
<bibRefCitation id="EFF467E24502FFAE80243D9CD531FCBE" author="Molineri, C. & Zuniga, M. C. & Ramos, B. C. & Giraldo, L. P. & Cardona, W." box="[316,565,758,785]" pageId="4" pageNumber="265" pagination="1 - 12" refId="ref3456" refString="Molineri, C., Zuniga, M. C., Ramos, B. C., Giraldo, L. P. & Cardona, W. (2016) Three new species of Leptohyphes Eaton (Ephemeroptera: Leptohyphidae) from Colombia. Iheringia, Serie Zoologia, 106, 1 - 12. https: // doi. org / 10.1590 / 1678 - 4766 e 2016024" type="journal article" year="2016">
|
||
Molineri
|
||
<emphasis id="B911C6014502FFAE80BE3D9DD6DDFCBF" box="[422,473,758,784]" italics="true" pageId="4" pageNumber="265">et al</emphasis>
|
||
. (2016)
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
, but their function is unknown (
|
||
<bibRefCitation id="EFF467E24502FFAE82B83D9CD22CFCBE" author="Baumgardner, D. E. & McCafferty, W. P." box="[928,1320,758,785]" pageId="4" pageNumber="265" pagination="1 - 33" refId="ref2790" refString="Baumgardner, D. E. & McCafferty, W. P. (2010) Revision of the genus Leptohyphes Eaton (Ephemeroptera: Leptohyphidae) in North and Central America. Zootaxa, 2360 (1), 1 - 33. https: // doi. org / 10.11646 / zootaxa. 2360.1.1" type="journal article" year="2010">Baumgardner & McCafferty 2010</bibRefCitation>
|
||
). We have observed in other species of the genus some intraspecific variation on the number and extent of such pale pits.
|
||
</paragraph>
|
||
<caption id="DF1A4A9B4502FFAE818F38F6D457F852" ID-DOI="http://doi.org/10.5281/zenodo.3874776" ID-Zenodo-Dep="3874776" httpUri="https://zenodo.org/record/3874776/files/figure.png" pageId="4" pageNumber="265" startId="4.[151,264,1948,1973]" targetBox="[154,1436,846,1923]" targetPageId="4">
|
||
<paragraph id="8BDA1A134502FFAE818F38F6D457F852" blockId="4.[151,1436,1948,2045]" pageId="4" pageNumber="265">
|
||
<emphasis id="B911C6014502FFAE818F38F6D652F81A" bold="true" box="[151,342,1948,1973]" pageId="4" pageNumber="265">FIGURES 10–20.</emphasis>
|
||
<taxonomicName id="4C6561904502FFAE804638F7D53EF81A" authority="Molineri & Derka, 2020" authorityName="Molineri & Derka" authorityYear="2020" box="[350,570,1949,1973]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="kukenan" status="sp. nov.">
|
||
<emphasis id="B911C6014502FFAE804638F7D53EF81A" box="[350,570,1949,1973]" italics="true" pageId="4" pageNumber="265">Leptohyphes kukenan</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911C6014502FFAE835A38F7D591F81A" bold="true" box="[578,661,1949,1973]" pageId="4" pageNumber="265">
|
||
<taxonomicNameLabel id="A2227B7A4502FFAE835A38F7D591F81A" box="[578,661,1949,1973]" pageId="4" pageNumber="265" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
10, foreleg, dorsal view with detail of spine; 11, fore tarsal claw; 12, hind leg with detail of spine, dorsal view; 13, schematic representation of abdominal terga setation; 14–16, operculate gill in dorsal, lateral and ventral view; 17–20, abdominal gills III–VI, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1A4A9B4503FFAF818F38DBD690F841" ID-DOI="http://doi.org/10.5281/zenodo.3874778" ID-Zenodo-Dep="3874778" httpUri="https://zenodo.org/record/3874778/files/figure.png" pageId="5" pageNumber="266" startId="5.[151,264,1969,1994]" targetBox="[206,1381,181,1943]" targetPageId="5">
|
||
<paragraph id="8BDA1A134503FFAF818F38DBD690F841" blockId="5.[151,1437,1969,2030]" pageId="5" pageNumber="266">
|
||
<emphasis id="B911C6014503FFAF818F38DBD652F865" bold="true" box="[151,342,1969,1994]" pageId="5" pageNumber="266">FIGURES 21–22.</emphasis>
|
||
21, Landscape of Kukenán tepui foothills where sampling localities are situated; 22, type locality. Photographs by Daniel Grul’a.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDA1A134500FFAC81DF3FF2D7DDFE24" blockId="6.[151,1437,152,395]" pageId="6" pageNumber="267">
|
||
<emphasis id="B911C6014500FFAC81DF3FF2D695FF1C" bold="true" box="[199,401,152,179]" pageId="6" pageNumber="267">Ecological notes:</emphasis>
|
||
Both sites were located on mountain watercourses with relatively cool water (
|
||
<figureCitation id="135E06964500FFAC841E3FF3D294FF1C" box="[1286,1424,153,179]" captionStart="FIGURES 21–22" captionStartId="5.[151,264,1969,1994]" captionTargetBox="[206,1381,181,1943]" captionTargetId="figure@5.[206,1381,181,1943]" captionTargetPageId="5" captionText="FIGURES 21–22. 21, Landscape of Kukenán tepui foothills where sampling localities are situated; 22, type locality. Photographs by Daniel Grul’a." figureDoi="http://doi.org/10.5281/zenodo.3874778" httpUri="https://zenodo.org/record/3874778/files/figure.png" pageId="6" pageNumber="267">Figs. 21–22</figureCitation>
|
||
). The
|
||
<typeStatus id="54DEA4B14500FFAC81D13FD7D7FDFF78" box="[201,249,189,215]" pageId="6" pageNumber="267">type</typeStatus>
|
||
locality is situated on a cascading,
|
||
<quantity id="4C9DB7F64500FFAC83993FD6D5C3FF78" box="[641,711,188,215]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="6" pageNumber="267" unit="m" value="5.5" valueMax="6.0" valueMin="5.0">5–6 m</quantity>
|
||
wide river with bottom formed by boulders and stones, the water temperature oscillated around 17 ºC. The surrounding area was a secondary savanna, trees were preserved on the banks. The stream originates on the plateau of Kukenán tepui at an altitude about
|
||
<quantity id="4C9DB7F64500FFAC853A3E6FD379FEB0" box="[1058,1149,261,287]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="6" pageNumber="267" unit="m" value="2600.0">2,600 m</quantity>
|
||
a.s.l. The second site was a stream
|
||
<quantity id="4C9DB7F64500FFAC81E53E43D639FEEC" box="[253,317,297,323]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="267" unit="m" value="0.4">0.4 m</quantity>
|
||
wide, with gravel bottom with many tree roots and debris. The water temperature 15.4 ºC indicates that the sampling site was situated near the spring. The surrounding area was a dense young forest regenerating after a fire.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |