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<mods:title id="F46745BCDBC77D2D9A7299B8E34309F0">Two new species and records of Olophrum laxum species group (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) from China and Nepal</mods:title>
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<emphasis id="B916EAED29001868DEB8F9B9781B0D7D" bold="true" box="[151,1115,1591,1618]" pageId="4" pageNumber="299">
Key to species of
<taxonomicName id="4C624D7C29001868DF42F9B67E7F0D7E" ID-CoL="4997R" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[365,575,1591,1617]" class="Insecta" family="Staphylinidae" genus="Olophrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="299" phylum="Arthropoda" rank="species" species="laxum">
<emphasis id="B916EAED29001868DF42F9B67E7F0D7E" bold="true" box="[365,575,1591,1617]" italics="true" pageId="4" pageNumber="299">Olophrum laxum</emphasis>
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group (modified Shavrin &amp; Smetana 2017)
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1 Head with slightly protruded eyes and extremely flattened postocular ridges. Aedeagus (
<figureCitation id="13592A7A29001868DA6EF9FC783C0DBC" box="[1089,1148,1661,1683]" captionStart="FIGURES 36" captionStartId="2.[151,264,1024,1047]" captionTargetBox="[197,1397,196,982]" captionTargetId="figure@2.[170,1417,193,1003]" captionTargetPageId="2" captionText="FIGURES 36. Aedeagus of Olophrum lama (34: holotype) and O. schuelkei (56: holotype): 3, 5—parameral view, 4, 6— lateral view. Scale bar:0.1 mm." httpUri="https://zenodo.org/record/1206574/files/figure.png" pageId="4" pageNumber="299">
Fig.
<bibRefCitation id="EFF34B0E29001868DA5FF9FC783C0DBC" box="[1136,1148,1661,1683]" pageId="4" pageNumber="299" refId="ref3309" refString="3 Punctation of pronotum small and sparse. Aedeagus with apices of parameres about reaching apex of median lobe (see Shavrin &amp; Smetana 2017: Fig. 6) Body length: 3.79 - 5.02 mm. Habitus as in Fig. 3 (see Shavrin &amp; Smetana 2017). Hubei, Shaanxi, Sichuan.................................................................. O. pacei Shavrin &amp; Smetana, 2017" type="journal article">3</bibRefCitation>
</figureCitation>
) with very wide internal sac. Body length:
<quantity id="4C9A9B1A29001868DF4CF91A7DFB0D9E" box="[355,443,1691,1713]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" pageId="4" pageNumber="299" unit="mm" value="4.65">4.65 mm</quantity>
. Habitus as in
<figureCitation id="13592A7A29001868DC67F91A7EC00D9E" box="[584,640,1691,1713]" captionStart="FIGURES 12" captionStartId="1.[151,264,1925,1948]" captionTargetBox="[259,1400,1031,1900]" captionTargetId="figure@1.[151,1436,1020,1903]" captionTargetPageId="1" captionText="FIGURES 12. Habitus of Olophrum lama (1: holotype) and O. schuelkei (2: holotype).Scale bar:1.0 mm." httpUri="https://zenodo.org/record/1206572/files/figure.png" pageId="4" pageNumber="299">Fig. 1</figureCitation>
. Nepal.........................................
<taxonomicName id="4C624D7C29001868DAEDF91D79220D9E" ID-CoL="4997N" authority="Shavrin" authorityName="Shavrin" authorityYear="2018" box="[1218,1378,1691,1713]" class="Insecta" family="Staphylinidae" genus="Olophrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="299" phylum="Arthropoda" rank="species" species="lama" status="sp. nov.">
<emphasis id="B916EAED29001868DAEDF91D794F0D9E" box="[1218,1295,1692,1713]" italics="true" pageId="4" pageNumber="299">O. lama</emphasis>
Shavrin
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,
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<emphasis id="B916EAED29001868DB43F91A79DC0D9E" bold="true" box="[1388,1436,1691,1713]" pageId="4" pageNumber="299">sp.n.</emphasis>
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<keyLead id="309395CA29001868DEB8F93879DB0DC4" pageId="4" pageNumber="299">- Head with distinctly protruded eyes, with obtuse or sharp well-defined postocular ridges. Aedeagus with moderately narrow internal sac. Chinese species............................................................................. 2</keyLead>
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<keyLead id="309395CA29001868DEB8F97279DB0C26" box="[151,1435,1779,1801]" pageId="4" pageNumber="299">2 Head without or with very short, fine grooves in front of ocelli................................................. 3</keyLead>
</paragraph>
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<keyLead id="309395CA29001868DEB8F89079DC0C09" box="[151,1436,1808,1830]" pageId="4" pageNumber="299">- Head with moderately long, deeper grooves in front of ocelli, reaching (at least almost) midlength of eye............... 4</keyLead>
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<keyLead id="309395CA29001868DEB8F8AC79DC0C51" pageId="4" pageNumber="299">
3
<title id="F966DDC929001868DECEF8AC7E2A0C6C" box="[225,618,1837,1859]" jopScore="0.07034105" pageId="4" pageNumber="299">Punctation of pronotum small and sparse</title>
. Aedeagus with apices of parameres about reaching apex of median lobe (see Shavrin &amp; Smetana
<year id="FFCA77AC29001868DF7AF8CA7DC90C4E" box="[341,393,1867,1889]" pageId="4" pageNumber="299">2017</year>
:
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Fig.
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) Body length:
<quantity id="4C9A9B1A29001868DC4DF8CA7EB20C4E" box="[610,754,1867,1889]" metricMagnitude="-3" metricUnit="m" metricValue="4.404999999999999" metricValueMax="5.02" metricValueMin="3.79" pageId="4" pageNumber="299" unit="mm" value="4.404999999999999" valueMax="5.02" valueMin="3.79">3.795.02 mm</quantity>
. Habitus as in
<figureCitation id="13592A7A29001868DDAAF8CA7FFE0C4E" box="[901,958,1867,1889]" captionStart="FIGURES 36" captionStartId="2.[151,264,1024,1047]" captionTargetBox="[197,1397,196,982]" captionTargetId="figure@2.[170,1417,193,1003]" captionTargetPageId="2" captionText="FIGURES 36. Aedeagus of Olophrum lama (34: holotype) and O. schuelkei (56: holotype): 3, 5—parameral view, 4, 6— lateral view. Scale bar:0.1 mm." httpUri="https://zenodo.org/record/1206574/files/figure.png" pageId="4" pageNumber="299">
Fig.
<pagination id="B2ABC80B29001868DD9DF8CA7FFE0C4E" box="[946,958,1867,1889]" pageId="4" pageNumber="299" type="pagination">3</pagination>
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(see Shavrin &amp; Smetana 2017).
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<author id="53DFDE1A29001868DB2FF8CA79D80C4E" box="[1280,1432,1867,1889]" fns="N" kam="kam" npo="LF" pageId="4" pageNumber="299">Hubei, Shaanxi</author>
, Sichuan
</collectingRegion>
..................................................................
<emphasis id="B916EAED29001868DA62F8E8781E0C51" box="[1101,1118,1897,1918]" italics="true" pageId="4" pageNumber="299">O</emphasis>
.
<emphasis id="B916EAED29001868DA45F8E878DE0C51" box="[1130,1182,1897,1918]" italics="true" pageId="4" pageNumber="299">pacei</emphasis>
Shavrin &amp; Smetana, 2017
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<materialsCitation id="3B0A3CA229001868DEB8F80779DC0C96" ID-GBIF-Occurrence="1836204409" location="Shaanxi, Gansu" pageId="4" pageNumber="299" specimenCount="1" stateProvince="Shaanxi, Gansu">
<keyLead id="309395CA29001868DEB8F80779DC0C96" pageId="4" pageNumber="299">
- Punctation of pronotum moderately large, coarse and dense. Aedeagus with long parameres significantly exceeding apex of median lobe (
<figureCitation id="13592A7A29001868DF4BF8227DDF0C96" box="[356,415,1955,1977]" captionStart="FIGURES 36" captionStartId="2.[151,264,1024,1047]" captionTargetBox="[197,1397,196,982]" captionTargetId="figure@2.[170,1417,193,1003]" captionTargetPageId="2" captionText="FIGURES 36. Aedeagus of Olophrum lama (34: holotype) and O. schuelkei (56: holotype): 3, 5—parameral view, 4, 6— lateral view. Scale bar:0.1 mm." httpUri="https://zenodo.org/record/1206574/files/figure.png" pageId="4" pageNumber="299">Fig. 5</figureCitation>
). Body length:
<quantity id="4C9A9B1A29001868DC1CF8227E810C96" box="[563,705,1955,1977]" metricMagnitude="-3" metricUnit="m" metricValue="4.17" metricValueMax="4.64" metricValueMin="3.7" pageId="4" pageNumber="299" unit="mm" value="4.17" valueMax="4.64" valueMin="3.7">3.704.64 mm</quantity>
. Habitus as in
<figureCitation id="13592A7A29001868DD62F8227FC70C96" box="[845,903,1955,1977]" captionStart="FIGURES 12" captionStartId="1.[151,264,1925,1948]" captionTargetBox="[259,1400,1031,1900]" captionTargetId="figure@1.[151,1436,1020,1903]" captionTargetPageId="1" captionText="FIGURES 12. Habitus of Olophrum lama (1: holotype) and O. schuelkei (2: holotype).Scale bar:1.0 mm." httpUri="https://zenodo.org/record/1206572/files/figure.png" pageId="4" pageNumber="299">Fig. 2</figureCitation>
.
<collectingRegion id="49A6F81D29001868DDBEF82278670C96" box="[913,1063,1955,1977]" isEnumeration="true" pageId="4" pageNumber="299">Shaanxi, Gansu</collectingRegion>
.........
<taxonomicName id="4C624D7C29001868DAB6F82579220C96" ID-CoL="4998Q" authority="Shavrin" authorityName="Shavrin" authorityYear="2018" box="[1177,1378,1955,1977]" class="Insecta" family="Staphylinidae" genus="Olophrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="299" phylum="Arthropoda" rank="species" species="schuelkei" status="sp. nov.">
<emphasis id="B916EAED29001868DAB6F825794F0C96" box="[1177,1295,1956,1977]" italics="true" pageId="4" pageNumber="299">O. schuelkei</emphasis>
Shavrin
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,
<taxonomicNameLabel id="A225579629001868DB43F82279DC0C96" box="[1388,1436,1955,1977]" pageId="4" pageNumber="299" rank="species">
<emphasis id="B916EAED29001868DB43F82279DC0C96" bold="true" box="[1388,1436,1955,1977]" pageId="4" pageNumber="299">sp.n.</emphasis>
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<keyLead id="309395CA29001868DEB8F84179DB0CDC" pageId="4" pageNumber="299">4 Head behind sharp postocular ridges markedly narrowed toward neck. Lateral margins of pronotum evenly narrowed anteriad and posteriad, disc with vague transverse mediobasal impression. Apices of parameres slightly exceeding apex of median lobe</keyLead>
</paragraph>
<paragraph id="8BDD36FF29011869DECEFF1679DC0A10" blockId="5.[151,1436,151,319]" pageId="5" pageNumber="300">
(see Shavrin &amp; Smetana 2017:
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). Body larger:
<quantity id="4C9A9B1A29011869DCF9FF167F0B0B83" box="[726,843,151,173]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.5" metricValueMin="4.2" pageId="5" pageNumber="300" unit="mm" value="4.35" valueMax="4.5" valueMin="4.2">4.24.5 mm</quantity>
. Habitus as in
<figureCitation id="13592A7A29011869DDF7FF1678500B82" box="[984,1040,151,173]" captionStart="FIGURES 12" captionStartId="1.[151,264,1925,1948]" captionTargetBox="[259,1400,1031,1900]" captionTargetId="figure@1.[151,1436,1020,1903]" captionTargetPageId="1" captionText="FIGURES 12. Habitus of Olophrum lama (1: holotype) and O. schuelkei (2: holotype).Scale bar:1.0 mm." httpUri="https://zenodo.org/record/1206572/files/figure.png" pageId="5" pageNumber="300">Fig. 2</figureCitation>
(see Shavrin &amp; Smetana 2017). Yunnan.....................................................................
<taxonomicName id="4C624D7C29011869DA39FF3479DC0BE5" ID-CoL="4997F" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[1046,1436,180,202]" class="Insecta" family="Staphylinidae" genus="Olophrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="300" phylum="Arthropoda" rank="species" species="hromadkai">
<emphasis id="B916EAED29011869DA39FF34786B0BE5" box="[1046,1067,181,202]" italics="true" pageId="5" pageNumber="300">O</emphasis>
.
<emphasis id="B916EAED29011869DA18FF3478DE0BE5" box="[1079,1182,181,202]" italics="true" pageId="5" pageNumber="300">hromadkai</emphasis>
Shavrin &amp; Smetana, 2017
</taxonomicName>
<keyLead id="309395CA29011869DEB8FF5379DC0A10" pageId="5" pageNumber="300">
- Head behind somewhat obtuse postocular ridges moderately narrowed toward neck. Lateral margins of pronotum markedly more narrowed anteriad than posteriad, disc with indistinct to distinct, deep transverse impression. Apices of parameres distinctly exceeding apex of median lobe (see Shavrin &amp; Smetana 2017:
<figureCitation id="13592A7A29011869DD55FE8D7FF30A0D" box="[890,947,268,290]" captionStart="FIGURES 36" captionStartId="2.[151,264,1024,1047]" captionTargetBox="[197,1397,196,982]" captionTargetId="figure@2.[170,1417,193,1003]" captionTargetPageId="2" captionText="FIGURES 36. Aedeagus of Olophrum lama (34: holotype) and O. schuelkei (56: holotype): 3, 5—parameral view, 4, 6— lateral view. Scale bar:0.1 mm." httpUri="https://zenodo.org/record/1206574/files/figure.png" pageId="5" pageNumber="300">Fig. 4</figureCitation>
) Body smaller:
<quantity id="4C9A9B1A29011869DA63FE8D78990A0D" box="[1100,1241,268,290]" metricMagnitude="-3" metricUnit="m" metricValue="3.825" metricValueMax="4.25" metricValueMin="3.4" pageId="5" pageNumber="300" unit="mm" value="3.825" valueMax="4.25" valueMin="3.4">3.404.25 mm</quantity>
. Habitus as in
<figureCitation id="13592A7A29011869DB4BFE8D79DC0A0D" box="[1380,1436,268,290]" captionStart="FIGURES 12" captionStartId="1.[151,264,1925,1948]" captionTargetBox="[259,1400,1031,1900]" captionTargetId="figure@1.[151,1436,1020,1903]" captionTargetPageId="1" captionText="FIGURES 12. Habitus of Olophrum lama (1: holotype) and O. schuelkei (2: holotype).Scale bar:1.0 mm." httpUri="https://zenodo.org/record/1206572/files/figure.png" pageId="5" pageNumber="300">Fig. 1</figureCitation>
(see see Shavrin &amp; Smetana 2017). Sichuan, Shaanxi, Yunnan......................
<taxonomicName id="4C624D7C29011869DA69FEAB79DC0A10" ID-CoL="4997R" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[1094,1436,297,319]" class="Insecta" family="Staphylinidae" genus="Olophrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="300" phylum="Arthropoda" rank="species" species="laxum">
<emphasis id="B916EAED29011869DA69FEAB78170A10" box="[1094,1111,298,319]" italics="true" pageId="5" pageNumber="300">O</emphasis>
.
<emphasis id="B916EAED29011869DA4BFEAB78DF0A10" box="[1124,1183,298,319]" italics="true" pageId="5" pageNumber="300">laxum</emphasis>
Shavrin &amp; Smetana, 2017
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</paragraph>
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